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<mods:title id="558A41842040364CD22DB9E7E26E4D1B">The land snail genus Pincerna Preston, 1907 (Gastropoda: Alycaeidae) from Vietnam and Laos, with description of a new species</mods:title>
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<mods:namePart id="7B075EEE51D256556F5D06C39A3E3485">Sang, Do Duc</mods:namePart>
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<treatment id="03A68780FFC1FFA98C97028A7CEF1459" ID-DOI="http://doi.org/10.5281/zenodo.7171076" ID-Zenodo-Dep="7171076" LSID="urn:lsid:plazi:treatment:03A68780FFC1FFA98C97028A7CEF1459" httpUri="http://treatment.plazi.org/id/03A68780FFC1FFA98C97028A7CEF1459" lastPageNumber="369" pageId="5" pageNumber="369">
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<paragraph id="8BB03696FFC1FFA98C97028A7A9E1579" blockId="5.[247,662,764,821]" box="[247,662,764,788]" pageId="5" pageNumber="369">
<heading id="D0F881FAFFC1FFA98C97028A7A9E1579" bold="true" box="[247,662,764,788]" centered="true" fontSize="10" level="2" pageId="5" pageNumber="369" reason="2">
<taxonomicName id="4C0F4D15FFC1FFA98C97028A7A9E1579" ID-CoL="7SQ3X" authority="(L. Pfeiffer, 1862)" baseAuthorityName="L. Pfeiffer" baseAuthorityYear="1862" box="[247,662,764,788]" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="5" pageNumber="369" phylum="Mollusca" rank="species" species="mouhoti">
<emphasis id="B97BEA84FFC1FFA98C97028A7A9E1579" bold="true" box="[247,662,764,788]" pageId="5" pageNumber="369">
<emphasis id="B97BEA84FFC1FFA98C97028A79C91579" bold="true" box="[247,449,764,788]" italics="true" pageId="5" pageNumber="369">Pincerna mouhoti</emphasis>
(L. Pfeiffer, 1862)
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</heading>
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<paragraph id="8BB03696FFC1FFA98D75036B7A711558" blockId="5.[247,662,764,821]" box="[277,633,796,821]" pageId="5" pageNumber="369">
(
<figureCitation id="13342A13FFC1FFA98D7D036B79601558" box="[285,360,797,821]" captionStart="Fig" captionStartId="2.[113,148,1941,1963]" captionTargetBox="[113,1452,311,1926]" captionTargetId="figure-11@2.[113,1452,311,1926]" captionTargetPageId="2" captionText="Fig. 1. Distribution of Pincerna species in Vietnam and Laos. Triangle: Pincerna anceyi; Grey circle: Pincerna costulosus; Black circles: Pincerna mouhoti; Squares: Pincerna vanbuensis; Star: Pincerna clausa, new species. The localities are detailed in Table 1." figureDoi="http://doi.org/10.5281/zenodo.7174166" httpUri="https://zenodo.org/record/7174166/files/figure.png" pageId="5" pageNumber="369">Figs. 1</figureCitation>
,
<figureCitation id="13342A13FFC1FFA98D18036B79B21558" box="[376,442,797,821]" captionStart="Fig" captionStartId="3.[136,171,2006,2028]" captionTargetBox="[168,1442,189,1991]" captionTargetId="figure-8@3.[168,1442,189,1991]" captionTargetPageId="3" captionText="Fig. 2. Shells (apertural view, dorsal view, ventral view, and side view) of Pincerna species. A, Pincerna anceyi (Mabille, 1887), syntype MNHN-IM-2000-31797; B, C, Pincerna mouhoti (L. Pfeiffer, 1862): B, specimen from MƯờng La, SƠn La, Vietnam, ZMHU; C, syntype NHMUK 20170120; D, E, Pincerna costulosus Bavay &amp; Dautzenberg, 1912: D, syntype MNHN-IM-2000-31786; E, specimen from Phong Thổ, Lai Châu, Vietnam, ZMHU." figureDoi="http://doi.org/10.5281/zenodo.7174168" httpUri="https://zenodo.org/record/7174168/files/figure.png" pageId="5" pageNumber="369">2B, C</figureCitation>
,
<figureCitation id="13342A13FFC1FFA98DA7036A79E11559" box="[455,489,796,820]" captionStart="Fig" captionStartId="8.[113,149,1150,1172]" captionTargetBox="[113,1451,188,1135]" captionTargetId="figure-34@8.[113,1451,188,1135]" captionTargetPageId="8" captionText="Fig. 6. Synoptic view of Vietnamese and Laos Pincerna species. A, MNHN-IM-2000-31797; B, NHMUK 20170120; C, MNHN- IM-2000-31786; D, VNMN-IZ 002.304; E, MNHN-IM-2000-31798." figureDoi="http://doi.org/10.5281/zenodo.7174176" httpUri="https://zenodo.org/record/7174176/files/figure.png" pageId="5" pageNumber="369">6B</figureCitation>
;
<tableCitation id="C68D032DFFC1FFA98D97036B7A5B1558" box="[503,595,797,821]" captionStart="Table 1" captionStartId="9.[136,191,189,211]" captionText="Table 1. Shell size variation of Pincerna spp. specimen collections in this study" pageId="5" pageNumber="369">Tables 1</tableCitation>
,
<tableCitation id="C68D032DFFC1FFA98E03036B7A791558" box="[611,625,797,821]" captionStart="Table 2" captionStartId="9.[136,191,1135,1157]" captionText="Table 2. Comparison of diagnostic conchological characters among Pincerna species in this study." pageId="5" pageNumber="369">2</tableCitation>
)
</paragraph>
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<subSubSection id="C315651DFFC1FFA98CE8032D7A741219" pageId="5" pageNumber="369" type="reference_group">
<paragraph id="8BB03696FFC1FFA98CE8032D7A3E15E0" blockId="5.[136,775,859,1140]" pageId="5" pageNumber="369">
<taxonomicName id="4C0F4D15FFC1FFA98CE8032D7A33151C" ID-CoL="7PYJ7" authority="L. Pfeiffer, 1863 a" authorityName="L. Pfeiffer" authorityYear="1863" box="[136,571,859,881]" class="Gastropoda" family="Alycaeidae" genus="Alycaeus" kingdom="Animalia" order="Architaenioglossa" pageId="5" pageNumber="369" phylum="Mollusca" rank="species" species="mouhoti">
<emphasis id="B97BEA84FFC1FFA98CE8032D793E151D" box="[136,310,859,880]" italics="true" pageId="5" pageNumber="369">Alycaeus mouhoti</emphasis>
<materialsCitation id="3B673CCBFFC1FFA98D5F032A79E7151C" box="[319,495,859,881]" country="Laos" location="Lao Mountains" pageId="5" pageNumber="369" specimenCount="1">
L.
<bibRefCitation id="EF9E4B67FFC1FFA98D3C032D79E7151C" author="Pfeiffer L" box="[348,495,859,881]" pageId="5" pageNumber="369" pagination="268 - 278" refId="ref7228" refString="Pfeiffer L (1863 a) Descriptions of thirty-six new land shells from the collection of H. Cuming, Esq. Proceedings of the Zoological Society of London, 30 (1862): 268 - 278." type="journal article" year="1863">Pfeiffer, 1863a</bibRefCitation>
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(1862)
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: 275, pl. 36, figs. 1, 2. (
<typeStatus id="54B48834FFC1FFA98CAE030E78F115E0" box="[206,249,888,909]" pageId="5" pageNumber="369">type</typeStatus>
locality:
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,
<collectingCountry id="F3187606FFC1FFA98D98030E7A2315E0" box="[504,555,888,909]" name="Laos" pageId="5" pageNumber="369">Laos</collectingCountry>
).
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<treatmentCitationGroup id="AB1F11B8FFC1FFA98CE803E37AB3126C" pageId="5" pageNumber="369">
<taxonomicName id="4C0F4D15FFC1FFA98CE803E3793B15C7" ID-CoL="7PYJ7" authorityName="L. Pfeiffer" authorityYear="1863" box="[136,307,917,938]" class="Gastropoda" family="Alycaeidae" genus="Alycaeus" kingdom="Animalia" order="Architaenioglossa" pageId="5" pageNumber="369" phylum="Mollusca" rank="species" species="mouhoti">
<emphasis id="B97BEA84FFC1FFA98CE803E3793B15C7" box="[136,307,917,938]" italics="true" pageId="5" pageNumber="369">Alycaeus mouhoti</emphasis>
</taxonomicName>
– L.
<treatmentCitation id="0AAE1087FFC1FFA98D0403E37A2F15C7" author="Pfeiffer L" box="[356,551,917,939]" page="228" pageId="5" pageNumber="369" year="1863">
<bibRefCitation id="EF9E4B67FFC1FFA98D0403E37A2F15C7" author="Pfeiffer L" box="[356,551,917,939]" pageId="5" pageNumber="369" refId="ref7267" refString="Pfeiffer L (1863 b) Novitates Conchologicae. Series prima. Mollusca extramarina. Descriptions et figures de coquilles, estramarines nouvelles, ou peu connues. Beschreibung und Abbidung neuer order kritischer Land- und Susswasser Mollusken. Tome 2, Lief. 19, 20. Verlag von Theodore Fischer, Cassel, pp. 139 - 303, pls. 37 - 72." type="book" year="1863">Pfeiffer, 1863b: 228</bibRefCitation>
</treatmentCitation>
, 229, pl. 59, figs. 9–11;
<treatmentCitation id="0AAE1087FFC1FFA98CCD03C4798015A5" author="von Martens E" box="[173,392,946,968]" page="67" pageId="5" pageNumber="369" year="1867">
<bibRefCitation id="EF9E4B67FFC1FFA98CCD03C4798015A5" author="von Martens E" box="[173,392,946,968]" pageId="5" pageNumber="369" refId="ref7748" refString="von Martens E (1867) Die Preussische Expedition nach Ost-Asien. Verlag der Koniglichen Geheimen Ober-Hofbuchdruckerei, Berlin, 477 pp." type="book" year="1867">von Martens, 1867: 67</bibRefCitation>
</treatmentCitation>
;
<bibRefCitation id="EF9E4B67FFC1FFA98DF103C47A0515AA" author="Reeve LA" box="[401,525,946,967]" pageId="5" pageNumber="369" refId="ref7474" refString="Reeve LA (1878) Monograph of the genus Alycaeus. In: Reeve LA (ed.) Conchologica Iconica: or, Illustrations of the Shells of Molluscous Animals. Volume 20. Reeve and Co., London, 13 pp., 6 pls." type="book" year="1878">Reeve, 1878</bibRefCitation>
: pl. 3, species 19;
<treatmentCitation id="0AAE1087FFC1FFA98EDA03C479191589" author="Fischer P" page="108" pageId="5" pageNumber="369" year="1891">
<bibRefCitation id="EF9E4B67FFC1FFA98EDA03C479191589" author="Fischer P" pageId="5" pageNumber="369" refId="ref6347" refString="Fischer P (1891) Catalogue et distribution geographique des Mollusques terrestres, fluviatilies &amp; marins d'une partie de l'Indo-Chine (Siam, Laos, Cambodge, Cochinchine, Annam, Tonkin). Imprimerie Dejussieu Pere et Fils, Autun, 192 pp." type="book" year="1891">Fischer, 1891: 108</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AAE1087FFC1FFA98D7B03B87A631589" author="Fischer H &amp; Dautzenberg P" box="[283,619,974,996]" page="432" pageId="5" pageNumber="369" year="1904">
<bibRefCitation id="EF9E4B67FFC1FFA98D7B03B87A631589" author="Fischer H &amp; Dautzenberg P" box="[283,619,974,996]" pageId="5" pageNumber="369" pagination="390 - 450" refId="ref6307" refString="Fischer H &amp; Dautzenberg P (1904) Catalogue des mollusques terrestres et fluviatiles de l'IndoChine orientale cites jusqu' a ce jour. In: Mission Pavie, Etudes diverses. Volume 3, pp. 390 - 450." type="journal article" year="1904">Fischer &amp; Dautzenberg, 1904: 432</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AAE1087FFC1FFA98E1403B87923126D" author="Pall-Gergely B &amp; Hunyadi A &amp; Do SD &amp; Naggs F &amp; Asami T" page="10" pageId="5" pageNumber="369" year="2017">
<bibRefCitation id="EF9E4B67FFC1FFA98E1403B87923126D" author="Pall-Gergely B &amp; Hunyadi A &amp; Do SD &amp; Naggs F &amp; Asami T" pageId="5" pageNumber="369" pagination="1 - 124" refId="ref6902" refString="Pall-Gergely B, Hunyadi A, Do SD, Naggs F &amp; Asami T (2017) Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus. Zootaxa, 4331: 1 - 124." type="journal article" year="2017">Páll-Gergely et al., 2017: 10</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AAE1087FFC1FFA98D58039D7A35126D" author="Inkhavilay K &amp; Sutcharit C &amp; Bantaowong U &amp; Chanabun R &amp; Siriwut W &amp; Srisonchai R &amp; Pholyotha A &amp; Jirapatrasilp P &amp; Panha S" box="[312,573,1003,1025]" httpUri="http://treatment.plazi.org/id/4284033A72BF3D735EF75D37A82D9F07" page="12" pageId="5" pageNumber="369" year="2019">
<bibRefCitation id="EF9E4B67FFC1FFA98D58039D7A35126D" author="Inkhavilay K &amp; Sutcharit C &amp; Bantaowong U &amp; Chanabun R &amp; Siriwut W &amp; Srisonchai R &amp; Pholyotha A &amp; Jirapatrasilp P &amp; Panha S" box="[312,573,1003,1025]" pageId="5" pageNumber="369" pagination="1 - 166" refId="ref6528" refString="Inkhavilay K, Sutcharit C, Bantaowong U, Chanabun R, Siriwut W, Srisonchai R, Pholyotha A, Jirapatrasilp P &amp; Panha S (2019) Annotated checklist of the terrestrial molluscs from Laos (Mollusca, Gastropoda). ZooKeys, 834: 1 - 166." type="journal article" year="2019">Inkhavilay et al., 2019: 12</bibRefCitation>
</treatmentCitation>
, figs. 4A, B.
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</paragraph>
<paragraph id="8BB03696FFC1FFA98CE8047E799E1257" blockId="5.[136,775,859,1140]" pageId="5" pageNumber="369">
<treatmentCitationGroup id="AB1F11B8FFC1FFA98CE8047E799E1257" pageId="5" pageNumber="369">
<taxonomicName id="4C0F4D15FFC1FFA98CE8047E79C61270" ID-CoL="7PYJ7" authorityName="L. Pfeiffer" authorityYear="1863" box="[136,462,1032,1053]" class="Gastropoda" family="Alycaeidae" genus="Alycaeus" kingdom="Animalia" order="Architaenioglossa" pageId="5" pageNumber="369" phylum="Mollusca" rank="species" species="mouhoti" subGenus="Orthalycaeus">
<emphasis id="B97BEA84FFC1FFA98CE8047E78D71270" box="[136,223,1032,1053]" italics="true" pageId="5" pageNumber="369">Alycaeus</emphasis>
(
<emphasis id="B97BEA84FFC1FFA98C8E047E79671270" box="[238,367,1032,1053]" italics="true" pageId="5" pageNumber="369">Orthalycaeus</emphasis>
)
<emphasis id="B97BEA84FFC1FFA98D1F047E79C61270" box="[383,462,1032,1053]" italics="true" pageId="5" pageNumber="369">mouhoti</emphasis>
</taxonomicName>
–
<treatmentCitation id="0AAE1087FFC1FFA98D89047E78DC1257" author="Kobelt W &amp; von Mollendorff O" page="147" pageId="5" pageNumber="369" year="1897">
<bibRefCitation id="EF9E4B67FFC1FFA98D89047E78DC1257" author="Kobelt W &amp; von Mollendorff O" pageId="5" pageNumber="369" pagination="137 - 152" refId="ref6703" refString="Kobelt W &amp; von Mollendorff O (1897) Catalog der gegenvartig lebend bekannten Pneumonopomen. Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft, jahrg 29, pp. 73 - 88, 105 - 120, 137 - 152." type="journal article" year="1897">Kobelt &amp; Möllendorff, 1897: 147</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AAE1087FFC1FFA98C800453799A1257" author="Ancey C-F" box="[224,402,1061,1082]" page="138" pageId="5" pageNumber="369" year="1898">
<bibRefCitation id="EF9E4B67FFC1FFA98C800453799A1257" author="Ancey C-F" box="[224,402,1061,1082]" pageId="5" pageNumber="369" pagination="125 - 150" refId="ref5783" refString="Ancey C-F (1898) Notes malacologiques. A - Observation sur les mollusques terrestres et fluviatiles recueillies dans l'IndoChine et particulierement au Laos par Henri Counillon. B - Description d'especes nouvelle du centre d'Afrique; C - Notes sur quelques coupes generiques ou sous-generiques de Mollusques. D - Description d'un mollusque mediterraneen nouveau. Bulletin du Musee d'Histoire Naturelle de Marseille, Serie II 1 (1): 125 - 150." type="journal article" year="1898">Ancey, 1898: 138</bibRefCitation>
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.
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</paragraph>
<paragraph id="8BB03696FFC1FFA98CE804347B0F123A" blockId="5.[136,775,859,1140]" box="[136,775,1089,1112]" pageId="5" pageNumber="369">
<treatmentCitationGroup id="AB1F11B8FFC1FFA98CE804347B0F123A" box="[136,775,1089,1112]" pageId="5" pageNumber="369">
<taxonomicName id="4C0F4D15FFC1FFA98CE80434799E123A" ID-CoL="7PYJ7" authorityName="L. Pfeiffer" authorityYear="1863" box="[136,406,1090,1111]" class="Gastropoda" family="Alycaeidae" genus="Alycaeus" kingdom="Animalia" order="Architaenioglossa" pageId="5" pageNumber="369" phylum="Mollusca" rank="species" species="mouhoti" subGenus="Alycaeus">
<emphasis id="B97BEA84FFC1FFA98CE8043478D5123A" box="[136,221,1090,1111]" italics="true" pageId="5" pageNumber="369">Alycaeus</emphasis>
(
<emphasis id="B97BEA84FFC1FFA98C8804347934123A" box="[232,316,1090,1111]" italics="true" pageId="5" pageNumber="369">Alycaeus</emphasis>
)
<emphasis id="B97BEA84FFC1FFA98D290434799E123A" box="[329,406,1090,1111]" italics="true" pageId="5" pageNumber="369">mouhoti</emphasis>
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–
<treatmentCitation id="0AAE1087FFC1FFA98DCA04347A5C123A" author="Kobelt W" box="[426,596,1090,1112]" page="347" pageId="5" pageNumber="369" year="1902">
<bibRefCitation id="EF9E4B67FFC1FFA98DCA04347A5C123A" author="Kobelt W" box="[426,596,1090,1112]" pageId="5" pageNumber="369" refId="ref6654" refString="Kobelt W (1902) Das Tierreich. Eine Zusammenstellung und Kennzeichnung der rezenten Tierformen. In Verbindung mit der Deutschen Zoologischen Gesellschaft herausgegeben von der Koniglich Preussischen Akademie der Wissenschaften zu Berlin. Mollusca: Cyclophoridae. R. Friedlander und Sohn, Berlin, 662 pp." type="book" year="1902">Kobelt, 1902: 347</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AAE1087FFC1FFA98E3C04347B0B123A" author="Saurin E" box="[604,771,1089,1112]" page="113" pageId="5" pageNumber="369" year="1953">
<bibRefCitation id="EF9E4B67FFC1FFA98E3C04347B0B123A" author="Saurin E" box="[604,771,1089,1112]" pageId="5" pageNumber="369" pagination="113 - 120" refId="ref7523" refString="Saurin E (1953) Coquilles nouvelles de l'Indochine. Journal de Conchyliologie, 93: 113 - 120." type="journal article" year="1953">Saurin, 1953: 113</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB03696FFC1FFA98CE804297A741219" blockId="5.[136,775,859,1140]" box="[136,636,1118,1140]" pageId="5" pageNumber="369">
<taxonomicName id="4C0F4D15FFC1FFA98CE80429793E121E" ID-CoL="7SQ3X" baseAuthorityName="L. Pfeiffer" baseAuthorityYear="1862" box="[136,310,1118,1140]" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="5" pageNumber="369" phylum="Mollusca" rank="species" species="mouhoti">
<emphasis id="B97BEA84FFC1FFA98CE80429793E121E" box="[136,310,1118,1140]" italics="true" pageId="5" pageNumber="369">Pincerna mouhoti</emphasis>
</taxonomicName>
– Páll-Gergely et al., 2020: 180.
</paragraph>
</subSubSection>
<subSubSection id="C315651DFFC1FFA98CE804EB7A1E12B9" pageId="5" pageNumber="369" type="description">
<paragraph id="8BB03696FFC1FFA98CE804EB7A1E12B9" blockId="5.[136,775,1180,1237]" pageId="5" pageNumber="369">
<emphasis id="B97BEA84FFC1FFA98CE804EB794312D9" bold="true" box="[136,331,1180,1205]" pageId="5" pageNumber="369">Types examined.</emphasis>
<materialsCitation id="3B673CCBFFC1FFA98D3404EB7A1B12B9" collectionCode="NHMUK" collectorName="Mouhot" location="Lao Mountains" pageId="5" pageNumber="369" specimenCode="NHMUK 20170120" specimenCount="1" typeStatus="syntype">
<location id="8ED0604DFFC1FFA98D3404EB79F512D8" LSID="urn:lsid:plazi:treatment:03A68780FFC1FFA98C97028A7CEF1459:8ED0604DFFC1FFA98D3404EB79F512D8" box="[340,509,1181,1205]" name="Lao Mountains" pageId="5" pageNumber="369">Lao Mountains</location>
, leg.
<collectorName id="26FA5340FFC1FFA98E5C04EB7A9B12D8" box="[572,659,1181,1205]" pageId="5" pageNumber="369">Mouhot</collectorName>
,
<specimenCode id="DBA99EEDFFC1FFA98EC204EB78FB12B8" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="5" pageNumber="369" type="Museum">NHMUK 20170120</specimenCode>
(
<typeStatus id="54B48834FFC1FFA98D6204CB796A12B8" box="[258,354,1213,1237]" pageId="5" pageNumber="369" type="syntype">syntypes</typeStatus>
) (
<figureCitation id="13342A13FFC1FFA98D1B04CB79D212B8" box="[379,474,1213,1237]" captionStart="Fig" captionStartId="3.[136,171,2006,2028]" captionTargetBox="[168,1442,189,1991]" captionTargetId="figure-8@3.[168,1442,189,1991]" captionTargetPageId="3" captionText="Fig. 2. Shells (apertural view, dorsal view, ventral view, and side view) of Pincerna species. A, Pincerna anceyi (Mabille, 1887), syntype MNHN-IM-2000-31797; B, C, Pincerna mouhoti (L. Pfeiffer, 1862): B, specimen from MƯờng La, SƠn La, Vietnam, ZMHU; C, syntype NHMUK 20170120; D, E, Pincerna costulosus Bavay &amp; Dautzenberg, 1912: D, syntype MNHN-IM-2000-31786; E, specimen from Phong Thổ, Lai Châu, Vietnam, ZMHU." figureDoi="http://doi.org/10.5281/zenodo.7174168" httpUri="https://zenodo.org/record/7174168/files/figure.png" pageId="5" pageNumber="369">Figs. 2C</figureCitation>
,
<figureCitation id="13342A13FFC1FFA98D8704CA7A0112B9" box="[487,521,1212,1236]" captionStart="Fig" captionStartId="8.[113,149,1150,1172]" captionTargetBox="[113,1451,188,1135]" captionTargetId="figure-34@8.[113,1451,188,1135]" captionTargetPageId="8" captionText="Fig. 6. Synoptic view of Vietnamese and Laos Pincerna species. A, MNHN-IM-2000-31797; B, NHMUK 20170120; C, MNHN- IM-2000-31786; D, VNMN-IZ 002.304; E, MNHN-IM-2000-31798." figureDoi="http://doi.org/10.5281/zenodo.7174176" httpUri="https://zenodo.org/record/7174176/files/figure.png" pageId="5" pageNumber="369">6B</figureCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C315651DFFC1FFA98CE8048A7AB91198" pageId="5" pageNumber="369" type="materials_examined">
<paragraph id="8BB03696FFC1FFA98CE8048A7AB91198" blockId="5.[135,775,1276,2037]" pageId="5" pageNumber="369">
<emphasis id="B97BEA84FFC1FFA98CE8048A79E11379" bold="true" box="[136,489,1276,1300]" pageId="5" pageNumber="369">Additional material examined.</emphasis>
<materialsCitation id="3B673CCBFFC1FFA98D92048B7AA813F8" collectingDate="2017-09-03" collectionCode="ZMHU" collectorName="D. S. Do" country="Vietnam" county="Mai Chau District" elevation="829" latitude="20.763056" location="No." longLatPrecision="21" longitude="104.87389" municipality="Hang Kia - Pa Co Nature Reserve" pageId="5" pageNumber="369" specimenCode="ZMHU/ 7" specimenCount="1" stateProvince="Hoa Binh">
<specimenCode id="DBA99EEDFFC1FFA98D92048B7A681379" box="[498,608,1277,1301]" collectionCode="ZMHU" country="Germany" httpUri="http://grbio.org/cool/99a5-d0z2" name="Zoologisches Museum der Humboldt Universitaet" pageId="5" pageNumber="369">ZMHU/ 7</specimenCode>
,
<collectingCountry id="F3187606FFC1FFA98E0E048B7AC51378" box="[622,717,1277,1301]" name="Vietnam" pageId="5" pageNumber="369">Vietnam</collectingCountry>
,
<collectingRegion id="49CBF874FFC1FFA98EBA048B792C1358" country="Vietnam" name="Hoa Binh" pageId="5" pageNumber="369">Hòa Bình Province</collectingRegion>
,
<collectingCounty id="62D14E1AFFC1FFA98D4E056B79F91358" box="[302,497,1309,1333]" pageId="5" pageNumber="369">Mai Châu District</collectingCounty>
,
<collectingMunicipality id="6BD4ACECFFC1FFA98D9B056B78E91339" pageId="5" pageNumber="369">Hang Kia - Pà Cò Nature Reserve</collectingMunicipality>
, near the road
<location id="8ED0604DFFC1FFA98DE8054B79B81339" LSID="urn:lsid:plazi:treatment:03A68780FFC1FFA98C97028A7CEF1459:8ED0604DFFC1FFA98DE8054B79B81339" box="[392,432,1341,1364]" country="Vietnam" county="Mai Chau District" latitude="20.763056" longLatPrecision="21" longitude="104.87389" municipality="Hang Kia - Pa Co Nature Reserve" name="No." pageId="5" pageNumber="369" stateProvince="Hoa Binh">No.</location>
6, between
<collectingRegion id="49CBF874FFC1FFA98E59054B7AAB1338" box="[569,675,1341,1365]" country="Vietnam" name="Hoa Binh" pageId="5" pageNumber="369">Hòa Bình</collectingRegion>
and
<collectingRegion id="49CBF874FFC1FFA98EBC054B78AF1319" country="Vietnam" name="Son La" pageId="5" pageNumber="369">SƠn La</collectingRegion>
, under rocks in primary forest,
<geoCoordinate id="EE3B5051FFC1FFA98D8D052D7A651319" box="[493,621,1371,1396]" degrees="20" direction="north" minutes="45" orientation="latitude" pageId="5" pageNumber="369" precision="15" seconds="47" value="20.763056">20°45′47″N</geoCoordinate>
,
<geoCoordinate id="EE3B5051FFC1FFA98E17052D7B0A1319" box="[631,770,1371,1396]" degrees="104" direction="east" minutes="52" orientation="longitude" pageId="5" pageNumber="369" precision="15" seconds="26" value="104.87389">104°52′26″E</geoCoordinate>
,
<elevation id="0022D1A5FFC1FFA98CE8050B791813F8" box="[136,272,1405,1429]" metricMagnitude="2" metricUnit="m" metricValue="8.29" pageId="5" pageNumber="369" unit="m" value="829.0">
<quantity id="4CF79B73FFC1FFA98CE8050B78D813F9" box="[136,208,1405,1429]" metricMagnitude="2" metricUnit="m" metricValue="8.29" pageId="5" pageNumber="369" unit="m" value="829.0">829 m</quantity>
a.s.l.
</elevation>
, col.
<collectorName id="26FA5340FFC1FFA98D33050B79BA13F9" box="[339,434,1405,1429]" pageId="5" pageNumber="369">D.S. Do</collectorName>
,
<date id="FFB11056FFC1FFA98DA1050B7AA813F8" box="[449,672,1405,1429]" pageId="5" pageNumber="369" value="2017-09-03">
<collectingDate id="EFF5E9BEFFC1FFA98DA1050B7AA813F8" box="[449,672,1405,1429]" pageId="5" pageNumber="369" value="2017-09-03">03 September 2017</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B673CCBFFC1FFA98ECF050B7B0F1399" collectionCode="ZMHU" collectorName="D. S. Do" country="Vietnam" county="Van" elevation="1013" latitude="21.765556" location="Van" longLatPrecision="21" longitude="104.74166" municipality="District" pageId="5" pageNumber="369" specimenCode="ZMHU/ 8" specimenCount="1" stateProvince="Son La">
<specimenCode id="DBA99EEDFFC1FFA98ECF050B789F13D8" collectionCode="ZMHU" country="Germany" httpUri="http://grbio.org/cool/99a5-d0z2" name="Zoologisches Museum der Humboldt Universitaet" pageId="5" pageNumber="369">ZMHU/ 8</specimenCode>
,
<collectingCountry id="F3187606FFC1FFA98CC705EB790013D8" box="[167,264,1437,1461]" name="Vietnam" pageId="5" pageNumber="369">Vietnam</collectingCountry>
,
<collectingRegion id="49CBF874FFC1FFA98D7705EB79D013D8" box="[279,472,1437,1463]" country="Vietnam" name="Son La" pageId="5" pageNumber="369">SƠn La Province</collectingRegion>
,
<collectingCounty id="62D14E1AFFC1FFA98D8605EB7A1B13D8" box="[486,531,1437,1461]" pageId="5" pageNumber="369">Vân</collectingCounty>
Hồ
<collectingMunicipality id="6BD4ACECFFC1FFA98E2A05EB7A9713D8" box="[586,671,1437,1461]" pageId="5" pageNumber="369">District</collectingMunicipality>
,
<location id="8ED0604DFFC1FFA98ECE05EB7AD313D8" LSID="urn:lsid:plazi:treatment:03A68780FFC1FFA98C97028A7CEF1459:8ED0604DFFC1FFA98ECE05EB7AD313D8" box="[686,731,1437,1461]" country="Vietnam" county="Van" latitude="21.765556" longLatPrecision="21" longitude="104.74166" municipality="District" name="Van" pageId="5" pageNumber="369" stateProvince="Son La">Vân</location>
Hồ
<location id="8ED0604DFFC1FFA98CE805CB78F413B8" LSID="urn:lsid:plazi:treatment:03A68780FFC1FFA98C97028A7CEF1459:8ED0604DFFC1FFA98CE805CB78F413B8" box="[136,252,1469,1493]" country="Vietnam" county="Van" latitude="21.765556" longLatPrecision="21" longitude="104.74166" municipality="District" name="Commune" pageId="5" pageNumber="369" stateProvince="Son La">Commune</location>
,
<location id="8ED0604DFFC1FFA98D6B05CB792E13B9" LSID="urn:lsid:plazi:treatment:03A68780FFC1FFA98C97028A7CEF1459:8ED0604DFFC1FFA98D6B05CB792E13B9" box="[267,294,1469,1492]" country="Vietnam" county="Van" latitude="21.765556" longLatPrecision="21" longitude="104.74166" municipality="District" name="Pa" pageId="5" pageNumber="369" stateProvince="Son La">Pa</location>
Cốp
<location id="8ED0604DFFC1FFA98D0805CB79B313B8" LSID="urn:lsid:plazi:treatment:03A68780FFC1FFA98C97028A7CEF1459:8ED0604DFFC1FFA98D0805CB79B313B8" box="[360,443,1469,1493]" country="Vietnam" county="Van" latitude="21.765556" longLatPrecision="21" longitude="104.74166" municipality="District" name="Village" pageId="5" pageNumber="369" stateProvince="Son La">Village</location>
, densely vegetated limestone hill,
<geoCoordinate id="EE3B5051FFC1FFA98CDB05AD79341399" box="[187,316,1499,1524]" degrees="21" direction="north" minutes="45" orientation="latitude" pageId="5" pageNumber="369" precision="15" seconds="56" value="21.765556">21°45′56″N</geoCoordinate>
,
<geoCoordinate id="EE3B5051FFC1FFA98D2905AD79DD1399" box="[329,469,1499,1524]" degrees="104" direction="east" minutes="44" orientation="longitude" pageId="5" pageNumber="369" precision="15" seconds="30" value="104.74166">104°44′30″E</geoCoordinate>
,
<elevation id="0022D1A5FFC1FFA98D8205AB7A781398" box="[482,624,1501,1525]" metricMagnitude="3" metricUnit="m" metricValue="1.013" pageId="5" pageNumber="369" unit="m" value="1013.0">
<quantity id="4CF79B73FFC1FFA98D8205AB7A3D1399" box="[482,565,1501,1525]" metricMagnitude="3" metricUnit="m" metricValue="1.013" pageId="5" pageNumber="369" unit="m" value="1013.0">1013 m</quantity>
a.s.l.
</elevation>
, col.
<collectorName id="26FA5340FFC1FFA98ECE05AB7B0F1399" box="[686,775,1501,1525]" pageId="5" pageNumber="369">D.S. Do</collectorName>
</materialsCitation>
&amp;
<materialsCitation id="3B673CCBFFC1FFA98CC2058B79B61078" box="[162,446,1532,1557]" collectingDate="2013-10-26" collectionCode="V, T" country="Vietnam" location="Sin" pageId="5" pageNumber="369" specimenCount="1" stateProvince="Son La">
<collectionCode id="ED1EAE53FFC1FFA98CC2058B78B11079" box="[162,185,1533,1556]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="369" type="Museum">V</collectionCode>
.
<collectionCode id="ED1EAE53FFC1FFA98CDC058B78C71079" box="[188,207,1533,1556]" name="Tavera, Department of Geology and Geophysics" pageId="5" pageNumber="369">T</collectionCode>
.
<location id="8ED0604DFFC1FFA98CB9058B78F71078" LSID="urn:lsid:plazi:treatment:03A68780FFC1FFA98C97028A7CEF1459:8ED0604DFFC1FFA98CB9058B78F71078" box="[217,255,1533,1557]" country="Vietnam" name="Sin" pageId="5" pageNumber="369" stateProvince="Son La">Sin</location>
,
<date id="FFB11056FFC1FFA98D69058A79B61078" box="[265,446,1532,1557]" pageId="5" pageNumber="369" value="2013-10-26">
<collectingDate id="EFF5E9BEFFC1FFA98D69058A79B61078" box="[265,446,1532,1557]" pageId="5" pageNumber="369" value="2013-10-26">26 October 2013</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B673CCBFFC1FFA98DA8058B79331058" collectionCode="ZMHU" country="Vietnam" location="SOn La Province" pageId="5" pageNumber="369" specimenCode="ZMHU/ 7" specimenCount="1" stateProvince="Son La">
<specimenCode id="DBA99EEDFFC1FFA98DA8058B7A381079" box="[456,560,1533,1557]" collectionCode="ZMHU" country="Germany" httpUri="http://grbio.org/cool/99a5-d0z2" name="Zoologisches Museum der Humboldt Universitaet" pageId="5" pageNumber="369">ZMHU/ 7</specimenCode>
(
<figureCitation id="13342A13FFC1FFA98E5E058B7A991078" box="[574,657,1533,1557]" captionStart="Fig" captionStartId="3.[136,171,2006,2028]" captionTargetBox="[168,1442,189,1991]" captionTargetId="figure-8@3.[168,1442,189,1991]" captionTargetPageId="3" captionText="Fig. 2. Shells (apertural view, dorsal view, ventral view, and side view) of Pincerna species. A, Pincerna anceyi (Mabille, 1887), syntype MNHN-IM-2000-31797; B, C, Pincerna mouhoti (L. Pfeiffer, 1862): B, specimen from MƯờng La, SƠn La, Vietnam, ZMHU; C, syntype NHMUK 20170120; D, E, Pincerna costulosus Bavay &amp; Dautzenberg, 1912: D, syntype MNHN-IM-2000-31786; E, specimen from Phong Thổ, Lai Châu, Vietnam, ZMHU." figureDoi="http://doi.org/10.5281/zenodo.7174168" httpUri="https://zenodo.org/record/7174168/files/figure.png" pageId="5" pageNumber="369">Fig. 2B</figureCitation>
),
<collectingCountry id="F3187606FFC1FFA98EC4058B7B0B1078" box="[676,771,1533,1557]" name="Vietnam" pageId="5" pageNumber="369">Vietnam</collectingCountry>
,
<collectingRegion id="49CBF874FFC1FFA98CE8066B79331058" box="[136,315,1565,1591]" country="Vietnam" name="Son La" pageId="5" pageNumber="369">SƠn La Province</collectingRegion>
</materialsCitation>
,
<materialsCitation id="3B673CCBFFC1FFA98D25066B7A061058" box="[325,526,1565,1591]" collectionCode="MU" country="Vietnam" location="La District" pageId="5" pageNumber="369" specimenCount="1" stateProvince="La District">
<collectionCode id="ED1EAE53FFC1FFA98D25066B7967105A" box="[325,367,1565,1591]" country="USA" name="Midwestern University" pageId="5" pageNumber="369">MƯ</collectionCode>
ờng
<collectingRegion id="49CBF874FFC1FFA98DF9066B7A061058" box="[409,526,1565,1589]" inferredBySuffix="true" pageId="5" pageNumber="369">La District</collectingRegion>
</materialsCitation>
,
<materialsCitation id="3B673CCBFFC1FFA98E77066B78A110F9" collectionCode="MU" collectorName="D. S. Do" country="Vietnam" elevation="715" latitude="21.401388" location="Th am Bo Cave" longLatPrecision="21" longitude="104.10361" municipality="Bu Commune" pageId="5" pageNumber="369" specimenCount="1" stateProvince="La District">
<collectionCode id="ED1EAE53FFC1FFA98E77066B7A49105A" box="[535,577,1565,1591]" country="USA" name="Midwestern University" pageId="5" pageNumber="369">MƯ</collectionCode>
ờng
<collectingMunicipality id="6BD4ACECFFC1FFA98E0C066B7B0B1058" box="[620,771,1565,1589]" pageId="5" pageNumber="369">Bú Commune</collectingMunicipality>
, entrance of the
<location id="8ED0604DFFC1FFA98D4C064B79D81038" LSID="urn:lsid:plazi:treatment:03A68780FFC1FFA98C97028A7CEF1459:8ED0604DFFC1FFA98D4C064B79D81038" box="[300,464,1597,1623]" country="Vietnam" latitude="21.401388" longLatPrecision="21" longitude="104.10361" municipality="Bu Commune" name="Th am Bo Cave" pageId="5" pageNumber="369" stateProvince="La District">Thẳm Bó Cave</location>
, limestone slope with woody regrowth,
<geoCoordinate id="EE3B5051FFC1FFA98C96062D79701019" box="[246,376,1627,1652]" degrees="21" direction="north" minutes="24" orientation="latitude" pageId="5" pageNumber="369" precision="15" seconds="05" value="21.401388">21°24′05″N</geoCoordinate>
,
<geoCoordinate id="EE3B5051FFC1FFA98DE4062D7A181019" box="[388,528,1627,1652]" degrees="104" direction="east" minutes="06" orientation="longitude" pageId="5" pageNumber="369" precision="15" seconds="13" value="104.10361">104°06′13″E</geoCoordinate>
,
<elevation id="0022D1A5FFC1FFA98E7B062A7A921018" box="[539,666,1628,1653]" metricMagnitude="2" metricUnit="m" metricValue="7.15" pageId="5" pageNumber="369" unit="m" value="715.0">
<quantity id="4CF79B73FFC1FFA98E7B062A7A681019" box="[539,608,1628,1652]" metricMagnitude="2" metricUnit="m" metricValue="7.15" pageId="5" pageNumber="369" unit="m" value="715.0">715 m</quantity>
a.s.l.
</elevation>
, col.
<collectorName id="26FA5340FFC1FFA98EB7062B78A110F9" pageId="5" pageNumber="369">D.S. Do</collectorName>
</materialsCitation>
&amp;
<materialsCitation id="3B673CCBFFC1FFA98CAD060B7A1710F8" box="[205,543,1661,1685]" collectingDate="2012-10-27" collectionCode="T" country="Vietnam" location="Nguyen" pageId="5" pageNumber="369" specimenCount="1" stateProvince="La District">
<collectionCode id="ED1EAE53FFC1FFA98CAD060B78E810F9" box="[205,224,1661,1684]" name="Tavera, Department of Geology and Geophysics" pageId="5" pageNumber="369">T</collectionCode>
.L.
<location id="8ED0604DFFC1FFA98D63060B795210F8" LSID="urn:lsid:plazi:treatment:03A68780FFC1FFA98C97028A7CEF1459:8ED0604DFFC1FFA98D63060B795210F8" box="[259,346,1661,1685]" country="Vietnam" name="Nguyen" pageId="5" pageNumber="369" stateProvince="La District">Nguyen</location>
,
<date id="FFB11056FFC1FFA98D05060B7A1710F8" box="[357,543,1661,1685]" pageId="5" pageNumber="369" value="2012-10-27">
<collectingDate id="EFF5E9BEFFC1FFA98D05060B7A1710F8" box="[357,543,1661,1685]" pageId="5" pageNumber="369" value="2012-10-27">27 October 2012</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B673CCBFFC1FFA98E4A060B7A0D10D8" collectionCode="ZMHU" country="Vietnam" location="SOn La Province" pageId="5" pageNumber="369" specimenCode="ZMHU/ 2" specimenCount="1" stateProvince="Son La">
<specimenCode id="DBA99EEDFFC1FFA98E4A060B7A9F10F8" box="[554,663,1661,1685]" collectionCode="ZMHU" country="Germany" httpUri="http://grbio.org/cool/99a5-d0z2" name="Zoologisches Museum der Humboldt Universitaet" pageId="5" pageNumber="369">ZMHU/ 2</specimenCode>
,
<collectingCountry id="F3187606FFC1FFA98EC4060B7B0B10F8" box="[676,771,1661,1685]" name="Vietnam" pageId="5" pageNumber="369">Vietnam</collectingCountry>
,
<collectingRegion id="49CBF874FFC1FFA98CE806EB794810D8" box="[136,320,1693,1719]" country="Vietnam" name="Son La" pageId="5" pageNumber="369">SƠn La Province</collectingRegion>
,
<collectingRegion id="49CBF874FFC1FFA98D2C06EB7A0D10D8" box="[332,517,1693,1717]" country="Vietnam" name="Phu Yen" pageId="5" pageNumber="369">Phù Yên District</collectingRegion>
</materialsCitation>
,
<materialsCitation id="3B673CCBFFC1FFA98E7106EB78C91178" collectingDate="2014-07-12" collectionCode="MU" collectorName="D. S. Do" country="Vietnam" elevation="674" latitude="21.19611" location="Village" longLatPrecision="21" longitude="104.784996" municipality="Do Commune" pageId="5" pageNumber="369" specimenCount="1" stateProvince="Son La">
<collectionCode id="ED1EAE53FFC1FFA98E7106EB7A3310DA" box="[529,571,1693,1719]" country="USA" name="Midwestern University" pageId="5" pageNumber="369">MƯ</collectionCode>
ờng
<collectingMunicipality id="6BD4ACECFFC1FFA98E0806EB7B0B10D8" box="[616,771,1693,1717]" pageId="5" pageNumber="369">Do Commune</collectingMunicipality>
, Lằn
<location id="8ED0604DFFC1FFA98CD906CB790210B8" LSID="urn:lsid:plazi:treatment:03A68780FFC1FFA98C97028A7CEF1459:8ED0604DFFC1FFA98CD906CB790210B8" box="[185,266,1725,1749]" country="Vietnam" latitude="21.19611" longLatPrecision="21" longitude="104.784996" municipality="Do Commune" name="Village" pageId="5" pageNumber="369" stateProvince="Son La">Village</location>
, steep limestone cliff, woody regrowth at foot,
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,
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,
<elevation id="0022D1A5FFC1FFA98DC706AA7A291098" box="[423,545,1756,1781]" metricMagnitude="2" metricUnit="m" metricValue="6.74" pageId="5" pageNumber="369" unit="m" value="674.0">
<quantity id="4CF79B73FFC1FFA98DC706AA79E21099" box="[423,490,1756,1780]" metricMagnitude="2" metricUnit="m" metricValue="6.74" pageId="5" pageNumber="369" unit="m" value="674.0">674 m</quantity>
a.s.l.
</elevation>
, col.
<collectorName id="26FA5340FFC1FFA98E3906AB7AB91099" box="[601,689,1757,1781]" pageId="5" pageNumber="369">D.S. Do</collectorName>
,
<date id="FFB11056FFC1FFA98EDA06AB78C91178" pageId="5" pageNumber="369" value="2014-07-12">
<collectingDate id="EFF5E9BEFFC1FFA98EDA06AB78C91178" pageId="5" pageNumber="369" value="2014-07-12">12 July 2014</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B673CCBFFC1FFA98CAE068B7A421119" collectionCode="ZMHU" collectorName="D. S. Do" country="Vietnam" county="Thu" elevation="1246" latitude="21.357224" location="Co" longLatPrecision="21" longitude="103.52167" municipality="Chau District" pageId="5" pageNumber="369" specimenCode="ZMHU/ 3" specimenCount="1" stateProvince="Son La">
<specimenCode id="DBA99EEDFFC1FFA98CAE068B79351178" box="[206,317,1789,1813]" collectionCode="ZMHU" country="Germany" httpUri="http://grbio.org/cool/99a5-d0z2" name="Zoologisches Museum der Humboldt Universitaet" pageId="5" pageNumber="369">ZMHU/ 3</specimenCode>
,
<collectingCountry id="F3187606FFC1FFA98D2B068B79A21178" box="[331,426,1789,1813]" name="Vietnam" pageId="5" pageNumber="369">Vietnam</collectingCountry>
,
<collectingRegion id="49CBF874FFC1FFA98DD8068B7A7B1178" box="[440,627,1789,1815]" country="Vietnam" name="Son La" pageId="5" pageNumber="369">SƠn La Province</collectingRegion>
,
<collectingCounty id="62D14E1AFFC1FFA98EE0068B7AA31178" box="[640,683,1789,1813]" pageId="5" pageNumber="369">Thu</collectingCounty>
ận
<collectingMunicipality id="6BD4ACECFFC1FFA98EAE068B78D51158" pageId="5" pageNumber="369">Châu District</collectingMunicipality>
,
<location id="8ED0604DFFC1FFA98C8B076B79021158" LSID="urn:lsid:plazi:treatment:03A68780FFC1FFA98C97028A7CEF1459:8ED0604DFFC1FFA98C8B076B79021158" box="[235,266,1821,1845]" country="Vietnam" county="Thu" latitude="21.357224" longLatPrecision="21" longitude="103.52167" municipality="Chau District" name="Co" pageId="5" pageNumber="369" stateProvince="Son La">Co</location>
Mạ
<location id="8ED0604DFFC1FFA98D23076B79BF1158" LSID="urn:lsid:plazi:treatment:03A68780FFC1FFA98C97028A7CEF1459:8ED0604DFFC1FFA98D23076B79BF1158" box="[323,439,1821,1845]" country="Vietnam" county="Thu" latitude="21.357224" longLatPrecision="21" longitude="103.52167" municipality="Chau District" name="Commune" pageId="5" pageNumber="369" stateProvince="Son La">Commune</location>
,
<location id="8ED0604DFFC1FFA98DA6076B7AB71159" LSID="urn:lsid:plazi:treatment:03A68780FFC1FFA98C97028A7CEF1459:8ED0604DFFC1FFA98DA6076B7AB71159" box="[454,703,1821,1845]" country="Vietnam" county="Thu" latitude="21.357224" longLatPrecision="21" longitude="103.52167" municipality="Chau District" name="Copia Nature Reserve" pageId="5" pageNumber="369" stateProvince="Son La">Copia Nature Reserve</location>
, steep limestone slope with dense disturbed vegetation,
<geoCoordinate id="EE3B5051FFC1FFA98EE3074D7B0B1139" box="[643,771,1851,1876]" degrees="21" direction="north" minutes="21" orientation="latitude" pageId="5" pageNumber="369" precision="15" seconds="26" value="21.357224">21°21′26″N</geoCoordinate>
,
<geoCoordinate id="EE3B5051FFC1FFA98CE7072D791B1119" box="[135,275,1883,1908]" degrees="103" direction="east" minutes="31" orientation="longitude" pageId="5" pageNumber="369" precision="15" seconds="18" value="103.52167">103°31′18″E</geoCoordinate>
,
<elevation id="0022D1A5FFC1FFA98D41072A79B81118" box="[289,432,1884,1909]" metricMagnitude="3" metricUnit="m" metricValue="1.246" pageId="5" pageNumber="369" unit="m" value="1246.0">
<quantity id="4CF79B73FFC1FFA98D41072A797D1119" box="[289,373,1884,1908]" metricMagnitude="3" metricUnit="m" metricValue="1.246" pageId="5" pageNumber="369" unit="m" value="1246.0">1246 m</quantity>
a.s.l.
</elevation>
, col.
<collectorName id="26FA5340FFC1FFA98D90072B7A421119" box="[496,586,1885,1909]" pageId="5" pageNumber="369">D.S. Do</collectorName>
</materialsCitation>
&amp;
<materialsCitation id="3B673CCBFFC1FFA98E11072B791611F8" collectingDate="2013-06-08" collectionCode="T" country="Vietnam" location="Nguyen" pageId="5" pageNumber="369" specimenCount="1" stateProvince="Son La">
<collectionCode id="ED1EAE53FFC1FFA98E11072B7A8C1119" box="[625,644,1885,1908]" name="Tavera, Department of Geology and Geophysics" pageId="5" pageNumber="369">T</collectionCode>
.H.
<location id="8ED0604DFFC1FFA98ECB072B7B0A1118" LSID="urn:lsid:plazi:treatment:03A68780FFC1FFA98C97028A7CEF1459:8ED0604DFFC1FFA98ECB072B7B0A1118" box="[683,770,1885,1909]" country="Vietnam" name="Nguyen" pageId="5" pageNumber="369" stateProvince="Son La">Nguyen</location>
,
<date id="FFB11056FFC1FFA98CE8070B791611F8" box="[136,286,1917,1941]" pageId="5" pageNumber="369" value="2013-06-08">
<collectingDate id="EFF5E9BEFFC1FFA98CE8070B791611F8" box="[136,286,1917,1941]" pageId="5" pageNumber="369" value="2013-06-08">08 June 2013</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B673CCBFFC1FFA98D4B070B7AA61198" collectingDate="2017-09-02" collectionCode="ZMHU" collectorName="D. S. Do &amp; X. H. Nguyen" country="Vietnam" county="Yen Chau District" elevation="815" latitude="21.141666" location="Commune" longLatPrecision="21" longitude="104.17472" municipality="Yen" pageId="5" pageNumber="369" specimenCode="ZMHU/ 2" specimenCount="1" stateProvince="Son La">
<specimenCode id="DBA99EEDFFC1FFA98D4B070B799111F8" box="[299,409,1917,1941]" collectionCode="ZMHU" country="Germany" httpUri="http://grbio.org/cool/99a5-d0z2" name="Zoologisches Museum der Humboldt Universitaet" pageId="5" pageNumber="369">ZMHU/ 2</specimenCode>
,
<collectingCountry id="F3187606FFC1FFA98DC7070B7A0E11F8" box="[423,518,1917,1941]" name="Vietnam" pageId="5" pageNumber="369">Vietnam</collectingCountry>
,
<collectingRegion id="49CBF874FFC1FFA98E73070B7AC511F8" box="[531,717,1917,1943]" country="Vietnam" name="Son La" pageId="5" pageNumber="369">SƠn La Province</collectingRegion>
,
<collectingCounty id="62D14E1AFFC1FFA98EBA070B791311D8" pageId="5" pageNumber="369">Yên Châu District</collectingCounty>
,
<collectingMunicipality id="6BD4ACECFFC1FFA98D4607EB795A11D8" box="[294,338,1949,1973]" pageId="5" pageNumber="369">Yên</collectingMunicipality>
SƠn
<location id="8ED0604DFFC1FFA98DED07EB79F711D8" LSID="urn:lsid:plazi:treatment:03A68780FFC1FFA98C97028A7CEF1459:8ED0604DFFC1FFA98DED07EB79F711D8" box="[397,511,1949,1973]" country="Vietnam" county="Yen Chau District" latitude="21.141666" longLatPrecision="21" longitude="104.17472" municipality="Yen" name="Commune" pageId="5" pageNumber="369" stateProvince="Son La">Commune</location>
,
<location id="8ED0604DFFC1FFA98E6A07EB7A3911D8" LSID="urn:lsid:plazi:treatment:03A68780FFC1FFA98C97028A7CEF1459:8ED0604DFFC1FFA98E6A07EB7A3911D8" box="[522,561,1949,1973]" country="Vietnam" county="Yen Chau District" latitude="21.141666" longLatPrecision="21" longitude="104.17472" municipality="Yen" name="Chi" pageId="5" pageNumber="369" stateProvince="Son La">Chi</location>
Đảy
<location id="8ED0604DFFC1FFA98E0D07EB7AAE11D8" LSID="urn:lsid:plazi:treatment:03A68780FFC1FFA98C97028A7CEF1459:8ED0604DFFC1FFA98E0D07EB7AAE11D8" box="[621,678,1949,1973]" country="Vietnam" county="Yen Chau District" latitude="21.141666" longLatPrecision="21" longitude="104.17472" municipality="Yen" name="Cave" pageId="5" pageNumber="369" stateProvince="Son La">Cave</location>
, primary forest on limestone,
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,
<geoCoordinate id="EE3B5051FFC1FFA98D8E07CD7A7211B9" box="[494,634,1979,2004]" degrees="104" direction="east" minutes="10" orientation="longitude" pageId="5" pageNumber="369" precision="15" seconds="29" value="104.17472">104°10′29″E</geoCoordinate>
,
<elevation id="0022D1A5FFC1FFA98EE407CA7B0811B8" box="[644,768,1980,2005]" metricMagnitude="2" metricUnit="m" metricValue="8.15" pageId="5" pageNumber="369" unit="m" value="815.0">
<quantity id="4CF79B73FFC1FFA98EE407CA7AC011B9" box="[644,712,1980,2004]" metricMagnitude="2" metricUnit="m" metricValue="8.15" pageId="5" pageNumber="369" unit="m" value="815.0">815 m</quantity>
a.s.l.
</elevation>
, col.
<collectorName id="26FA5340FFC1FFA98CD807AB79191199" box="[184,273,2013,2037]" pageId="5" pageNumber="369">D.S. Do</collectorName>
&amp;
<collectorName id="26FA5340FFC1FFA98D5607AB79C21198" box="[310,458,2013,2037]" pageId="5" pageNumber="369">X.H. Nguyen</collectorName>
,
<date id="FFB11056FFC1FFA98DB707AB7AA61198" box="[471,686,2013,2037]" pageId="5" pageNumber="369" value="2017-09-02">
<collectingDate id="EFF5E9BEFFC1FFA98DB707AB7AA61198" box="[471,686,2013,2037]" pageId="5" pageNumber="369" value="2017-09-02">02 September 2017</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C315651DFFC1FFA98F2300CA7DCB1778" pageId="5" pageNumber="369" type="diagnosis">
<paragraph id="8BB03696FFC1FFA98F2300CA7DCB1778" blockId="5.[835,1475,188,277]" pageId="5" pageNumber="369">
<emphasis id="B97BEA84FFC1FFA98F2300CA7BB016B9" bold="true" box="[835,952,188,212]" pageId="5" pageNumber="369">Diagnosis.</emphasis>
<taxonomicName id="4C0F4D15FFC1FFA98FDE00CB7C7516B8" baseAuthorityName="L. Pfeiffer" baseAuthorityYear="1862" box="[958,1149,189,213]" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="5" pageNumber="369" phylum="Mollusca" rank="species" species="mouhoti">
<emphasis id="B97BEA84FFC1FFA98FDE00CB7C7516B8" box="[958,1149,189,213]" italics="true" pageId="5" pageNumber="369">Pincerna mouhoti</emphasis>
</taxonomicName>
is characterised by a mediumsized, translucent shell with rather regular ribs on R1, R2 long, smooth, slender and equal-spaced ribs and open umbilicus.
</paragraph>
</subSubSection>
<subSubSection id="C315651DFFC1FFA98F23014A7C0A17D8" pageId="5" pageNumber="369" type="distribution">
<paragraph id="8BB03696FFC1FFA98F23014A7C0A17D8" blockId="5.[835,1474,316,437]" pageId="5" pageNumber="369">
<emphasis id="B97BEA84FFC1FFA98F23014A7BDE1739" bold="true" box="[835,982,316,340]" pageId="5" pageNumber="369">Distribution.</emphasis>
<taxonomicName id="4C0F4D15FFC1FFA98FBA014B7C901738" baseAuthorityName="L. Pfeiffer" baseAuthorityYear="1862" box="[986,1176,317,341]" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="5" pageNumber="369" phylum="Mollusca" rank="species" species="mouhoti">
<emphasis id="B97BEA84FFC1FFA98FBA014B7C901738" box="[986,1176,317,341]" italics="true" pageId="5" pageNumber="369">Pincerna mouhoti</emphasis>
</taxonomicName>
is known from
<collectingCountry id="F3187606FFC1FFA9895C014B7D781739" box="[1340,1392,317,340]" name="Laos" pageId="5" pageNumber="369">Laos</collectingCountry>
(Luang Phrabang Province) and
<collectingCountry id="F3187606FFC1FFA98839012B7CBE1718" box="[1113,1206,349,373]" name="Vietnam" pageId="5" pageNumber="369">Vietnam</collectingCountry>
(
<collectingRegion id="49CBF874FFC1FFA988A7012B7D121719" box="[1223,1306,349,375]" country="Vietnam" name="Son La" pageId="5" pageNumber="369">SƠn La</collectingRegion>
and
<collectingRegion id="49CBF874FFC1FFA98936012B7DCA1718" box="[1366,1474,349,373]" country="Vietnam" name="Hoa Binh" pageId="5" pageNumber="369">Hòa Bình</collectingRegion>
Provinces) (
<bibRefCitation id="EF9E4B67FFC1FFA98FA6010B7C5717F8" author="Fischer P" box="[966,1119,381,405]" pageId="5" pageNumber="369" refId="ref6347" refString="Fischer P (1891) Catalogue et distribution geographique des Mollusques terrestres, fluviatilies &amp; marins d'une partie de l'Indo-Chine (Siam, Laos, Cambodge, Cochinchine, Annam, Tonkin). Imprimerie Dejussieu Pere et Fils, Autun, 192 pp." type="book" year="1891">Fischer, 1891</bibRefCitation>
;
<bibRefCitation id="EF9E4B67FFC1FFA9880D010B7CF317F9" author="Saurin E" box="[1133,1275,380,405]" pageId="5" pageNumber="369" pagination="113 - 120" refId="ref7523" refString="Saurin E (1953) Coquilles nouvelles de l'Indochine. Journal de Conchyliologie, 93: 113 - 120." type="journal article" year="1953">Saurin, 1953</bibRefCitation>
;
<bibRefCitation id="EF9E4B67FFC1FFA98969010B7B7417D8" author="Inkhavilay K &amp; Sutcharit C &amp; Bantaowong U &amp; Chanabun R &amp; Siriwut W &amp; Srisonchai R &amp; Pholyotha A &amp; Jirapatrasilp P &amp; Panha S" pageId="5" pageNumber="369" pagination="1 - 166" refId="ref6528" refString="Inkhavilay K, Sutcharit C, Bantaowong U, Chanabun R, Siriwut W, Srisonchai R, Pholyotha A, Jirapatrasilp P &amp; Panha S (2019) Annotated checklist of the terrestrial molluscs from Laos (Mollusca, Gastropoda). ZooKeys, 834: 1 - 166." type="journal article" year="2019">Inkhavilay et al., 2019</bibRefCitation>
; this study).
</paragraph>
</subSubSection>
<subSubSection id="C315651DFFC1FFA98F2301AB7CEF1459" pageId="5" pageNumber="369" type="discussion">
<paragraph id="8BB03696FFC1FFA98F2301AB7CEF1459" blockId="5.[835,1475,477,565]" pageId="5" pageNumber="369">
<emphasis id="B97BEA84FFC1FFA98F2301AB7BBE1798" bold="true" box="[835,950,477,501]" pageId="5" pageNumber="369">Remarks.</emphasis>
<figureCitation id="13342A13FFC1FFA98FA101AB7C3B1798" box="[961,1075,477,501]" captionStart="Fig" captionStartId="6.[113,148,2063,2085]" captionTargetBox="[126,1440,188,2047]" captionTargetId="figure-11@6.[126,1440,188,2047]" captionTargetPageId="6" captionText="Fig. 4. Shell (apertural view, dorsal view, ventral view, and side view) of Pincerna clausa, new species. A, holotype VNMN-IZ 002.304; B, C, paratypes ZMHU. MOL 044." figureDoi="http://doi.org/10.5281/zenodo.7174172" httpUri="https://zenodo.org/record/7174172/files/figure.png" pageId="5" pageNumber="369">Figure 4E</figureCitation>
in
<bibRefCitation id="EF9E4B67FFC1FFA9883F01AB7D7A1798" author="Inkhavilay K &amp; Sutcharit C &amp; Bantaowong U &amp; Chanabun R &amp; Siriwut W &amp; Srisonchai R &amp; Pholyotha A &amp; Jirapatrasilp P &amp; Panha S" box="[1119,1394,477,501]" pageId="5" pageNumber="369" pagination="1 - 166" refId="ref6528" refString="Inkhavilay K, Sutcharit C, Bantaowong U, Chanabun R, Siriwut W, Srisonchai R, Pholyotha A, Jirapatrasilp P &amp; Panha S (2019) Annotated checklist of the terrestrial molluscs from Laos (Mollusca, Gastropoda). ZooKeys, 834: 1 - 166." type="journal article" year="2019">Inkhavilay et al. (2019)</bibRefCitation>
shows that the shell surface is smooth, R2 long and smooth, and therefore it may belong to
<taxonomicName id="4C0F4D15FFC1FFA98809026B7CE81458" baseAuthorityName="L. Pfeiffer" baseAuthorityYear="1862" box="[1129,1248,541,565]" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="5" pageNumber="369" phylum="Mollusca" rank="species" species="mouhoti">
<emphasis id="B97BEA84FFC1FFA98809026B7CE81458" box="[1129,1248,541,565]" italics="true" pageId="5" pageNumber="369">P. mouhoti</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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