171 lines
19 KiB
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171 lines
19 KiB
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<document id="31550AC47B0C406CA9C4A0DA3A3E9969" ID-DOI="10.11646/phytotaxa.614.1.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="8389307" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1695979127173" checkinUser="plazi" docAuthor="Henderson, Andrew" docDate="2023" docId="03B387DAFFC81F58FF50FD2FFC138F4F" docLanguage="en" docName="phytotaxa.614.1.1.pdf" docOrigin="Phytotaxa 614 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.614.1.1" docStyle="DocumentStyle:F08184CE06D8A97EA3E6DE35D99648B0.3:Phytotaxa.2014-.monograph" docStyleId="F08184CE06D8A97EA3E6DE35D99648B0" docStyleName="Phytotaxa.2014-.monograph" docStyleVersion="3" docTitle="Coccothrinax jimenezii Mejia & Garcia 2013" docType="treatment" docVersion="2" lastPageNumber="56" masterDocId="FF8AFFA2FFFF1F6FFFD8FFB9FFF9881B" masterDocTitle="A revision of Coccothrinax, Hemithrinax, Leucothrinax, Thrinax, and Zombia (Arecaceae)" masterLastPageNumber="115" masterPageNumber="1" pageNumber="56" updateTime="1695980191673" updateUser="ExternalLinkService">
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<mods:mods id="5D2A7148DB862ACC735B9750B55F86B7" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="DB81550B43987FA0729C5ACF67FBE54F">
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<mods:title id="654A614C901473312AE9CD40C6C6A966">A revision of Coccothrinax, Hemithrinax, Leucothrinax, Thrinax, and Zombia (Arecaceae)</mods:title>
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</mods:titleInfo>
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<mods:name id="67497785D0A4F09F9EA99D74625AA103" type="personal">
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<mods:roleTerm id="8D7CA92CF904F3011F9E0ED549146E21">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="5554F7170A0E5B2E941C0A7F3BB42CDD">Henderson, Andrew</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="EB8B97DFAE6BA75C482B4A761C10B927">text</mods:typeOfResource>
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<mods:titleInfo id="FDCF90C4031B3CB6D2F7DDDD863DCF57">
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<mods:title id="794AD34ED8E632EC3B3EB509DA3C63E3">Phytotaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="6320C03C318EB1A5E541C1F546A76F8B">
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<mods:date id="C354FAA6E1159E65BA5BD7DE2E5D5BB8">2023</mods:date>
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<mods:detail id="8AE396692F305E1F3E2A2E4550D37B00" type="pubDate">
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<mods:number id="943D54FC0E44987A8D5653492C833296">2023-09-19</mods:number>
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</mods:detail>
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<mods:number id="EDAFCCD727D4461CD03E0B558B43C4FB">614</mods:number>
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</mods:detail>
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<mods:detail id="36D04D76D9A81C69DDDD46A3E3FB70E9" type="issue">
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<mods:number id="BC7210AA4B9665951991FD7DCDF115D9">1</mods:number>
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<mods:start id="3AA869C5D9EFC034A39D869F2742CE95">1</mods:start>
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<mods:end id="AAC4CEC525266CD991559C1CBCF1E58F">115</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location id="A8FD6A53ED7D9B2AD7095D529BC5FD7C">
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<mods:url id="2CE6641389B041BA1FC14207B853DA5C">http://dx.doi.org/10.11646/phytotaxa.614.1.1</mods:url>
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</mods:location>
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<mods:classification id="DFAB5F7447405C9AC77FFD87DD0C97AB">journal article</mods:classification>
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<mods:identifier id="93601A66B2C2CE191AFA55C06461A602" type="DOI">10.11646/phytotaxa.614.1.1</mods:identifier>
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<mods:identifier id="786CB462D846D9E37380827D96A88368" type="ISSN">1179-3163</mods:identifier>
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<mods:identifier id="84B1659B3D4603EDFABA2253D26CC67F" type="Zenodo-Dep">8389307</mods:identifier>
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</mods:mods>
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<treatment id="03B387DAFFC81F58FF50FD2FFC138F4F" ID-DOI="http://doi.org/10.5281/zenodo.8389313" ID-Zenodo-Dep="8389313" LSID="urn:lsid:plazi:treatment:03B387DAFFC81F58FF50FD2FFC138F4F" httpUri="http://treatment.plazi.org/id/03B387DAFFC81F58FF50FD2FFC138F4F" lastPageNumber="56" pageId="55" pageNumber="56">
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<subSubSection id="C3006547FFC81F58FF50FD2FFBA98AE2" pageId="55" pageNumber="56" type="nomenclature">
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<paragraph id="8BA536CCFFC81F58FF50FD2FFBA98AE2" blockId="55.[136,1452,661,761]" pageId="55" pageNumber="56">
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<emphasis id="B96EEADEFFC81F58FF50FD2FFD078AAB" bold="true" box="[136,766,661,689]" pageId="55" pageNumber="56">
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1.20.
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<taxonomicName id="4C1A4D4FFFC81F58FF1DFD2FFD018AAB" ID-CoL="WLYP" authority="Mejia & Garcia (2013: 9)" authorityName="Mejia & Garcia" authorityPageNumber="9" authorityYear="2013" box="[197,760,661,689]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="55" pageNumber="56" phylum="Tracheophyta" rank="species" species="jimenezii">
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<emphasis id="B96EEADEFFC81F58FF1DFD2FFE368AAB" bold="true" box="[197,463,662,689]" italics="true" pageId="55" pageNumber="56">Coccothrinax jimenezii</emphasis>
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<bibRefCitation id="EF8B4B3DFFC81F58FE0BFD2CFD018AAB" author="Mejia, M. & Garcia, R." box="[467,760,661,688]" pageId="55" pageNumber="56" pagination="9 - 13" refId="ref62950" refString="Mejia, M. & Garcia, R. (2013) Una nueva especie de Coccothrinax (Arecaceae) para la isla Espanola. Moscosoa 18: 9 - 13." type="journal article" year="2013">Mejía & García (2013: 9)</bibRefCitation>
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</taxonomicName>
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.
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</emphasis>
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<materialsCitation id="3B723C91FFC81F58FCDAFD2FFBA98AE2" collectingDate="2010-07-24" collectionCode="W, R, M, JBSD" collectorName="de Neiba & de Postrer Rio & La Descubierta" country="Dominican Republic" elevation="50" location="Sierra de Neiba" pageId="55" pageNumber="56" specimenCode="N 71" specimenCount="1" stateProvince="Independencia" typeStatus="holotype">
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Type:—
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<collectingCountry id="F30D765CFFC81F58FC83FD2FFB688AAB" box="[859,1169,662,688]" name="Dominican Republic" pageId="55" pageNumber="56">DOMINICAN REPUBLIC</collectingCountry>
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. Province
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<collectingRegion id="49DEF82EFFC81F58FAD9FD2FFA5E8AAB" box="[1281,1447,662,688]" country="Dominican Republic" name="Independencia" pageId="55" pageNumber="56">Independencia</collectingRegion>
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,
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<location id="8EC56017FFC81F58FF50FD03FEC18ACF" LSID="urn:lsid:plazi:treatment:03B387DAFFC81F58FF50FD2FFC138F4F:8EC56017FFC81F58FF50FD03FEC18ACF" box="[136,312,698,724]" country="Dominican Republic" name="Sierra de Neiba" pageId="55" pageNumber="56" stateProvince="Independencia">
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Sierra
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<collectorName id="26EF531AFFC81F58FF09FD03FEC18ACF" box="[209,312,698,724]" pageId="55" pageNumber="56">de Neiba</collectorName>
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</location>
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,
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<quantity id="4CE29B29FFC81F58FE9AFD03FE838ACF" box="[322,378,698,724]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="55" pageNumber="56" unit="km" value="9.0">9 km</quantity>
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al oeste
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<collectorName id="26EF531AFFC81F58FE02FD03FD828ACF" box="[474,635,698,725]" pageId="55" pageNumber="56">de Postrer Río</collectorName>
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, carrertera hacia
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<collectorName id="26EF531AFFC81F58FCE1FD02FC1C8ACF" box="[825,997,698,725]" pageId="55" pageNumber="56">La Descubierta</collectorName>
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, lado norte del Lago Enriquillo, 18˚34’
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<specimenCode id="DBBC9EB7FFC81F58FA4CFD03FF5C8AE2" collectionCode="N" country="China" lsid="urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="55" pageNumber="56" type="Herbarium">N 71</specimenCode>
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˚42’
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<collectionCode id="ED0BAE09FFC81F58FF0FFD66FF0B8AE2" box="[215,242,735,761]" country="Austria" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="55" pageNumber="56" type="Herbarium">W</collectionCode>
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,
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<quantity id="4CE29B29FFC81F58FF25FD67FEC18AE2" box="[253,312,734,761]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="55" pageNumber="56" unit="m" value="50.0">
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<elevation id="0037D1FFFFC81F58FF25FD67FEC18AE2" box="[253,312,734,761]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="55" pageNumber="56" unit="m" value="50.0">50 m</elevation>
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</quantity>
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,
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<date id="FFA4100CFFC81F58FE9CFD67FE2E8AE3" box="[324,471,734,760]" pageId="55" pageNumber="56" value="2010-07-24">
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<collectingDate id="EFE0E9E4FFC81F58FE9CFD67FE2E8AE3" box="[324,471,734,760]" pageId="55" pageNumber="56" value="2010-07-24">24 July 2010</collectingDate>
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</date>
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,
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<emphasis id="B96EEADEFFC81F58FE3AFD66FCE08AE3" box="[482,793,734,761]" italics="true" pageId="55" pageNumber="56">
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<collectionCode id="ED0BAE09FFC81F58FE3AFD66FE0F8AE3" box="[482,502,735,760]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="55" pageNumber="56">R</collectionCode>
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. García &
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<collectionCode id="ED0BAE09FFC81F58FDACFD66FD768AE3" box="[628,655,735,760]" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="55" pageNumber="56" type="Herbarium">M</collectionCode>
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. Mejía 7640
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</emphasis>
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(
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<typeStatus id="54A1886EFFC81F58FCF0FD67FC758AE3" box="[808,908,734,760]" pageId="55" pageNumber="56">holotype</typeStatus>
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<collectionCode id="ED0BAE09FFC81F58FC4BFD67FC278AE3" box="[915,990,734,760]" country="Dominican Republic" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15541" lsid="urn:lsid:biocol.org:col:15541" name="Jardín Botánico Nacional Dr. Rafael M. Moscoso" pageId="55" pageNumber="56" type="Herbarium">JBSD</collectionCode>
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!).
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<figureCitation id="13212A49FFC81F58FC2DFD67FBA98AE2" box="[1013,1104,734,761]" captionStart="PLATE 15" captionStartId="56.[136,214,1917,1938]" captionTargetBox="[151,1433,194,1891]" captionTargetId="figure-27@56.[150,1434,193,1893]" captionTargetPageId="56" captionText="PLATE 15. Coccothrinax jimenezii, Dominican Republic, with clustered stems and wedge-shaped leaf blades. (Image by Eladio Fernández)." figureDoi="http://doi.org/10.5281/zenodo.8389389" httpUri="https://zenodo.org/record/8389389/files/figure.png" pageId="55" pageNumber="56">Plate 15</figureCitation>
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3006547FFC81F58FF50FC9FFDF38D46" pageId="55" pageNumber="56" type="description">
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<paragraph id="8BA536CCFFC81F58FF50FC9FFDF38D46" blockId="55.[136,1452,806,1877]" pageId="55" pageNumber="56">
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<emphasis id="B96EEADEFFC81F58FF50FC9FFF298B5B" bold="true" box="[136,208,806,832]" pageId="55" pageNumber="56">Stems</emphasis>
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4.0(2.5–5.5) m long, diameter not recorded, solitary or clustered.
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<emphasis id="B96EEADEFFC81F58FC18FC9FFBEB8B5B" bold="true" box="[960,1042,806,832]" pageId="55" pageNumber="56">Leaves</emphasis>
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more or less deciduous or only leaf bases persisting on stem; leaf sheath fibers 0.4(0.2–0.7) mm diameter, stout, loosely woven and forming a loose, hexagonal mesh, initially forming ligules at the apices; petioles 7.6(6.2–10.0) mm diameter just below the apex; palmans 4.4(1.5–8.0) cm long, relatively short, with the adaxial veins prominent and terminating in a slight raised ridge and distinct pulvinus; leaf blades wedge-shaped; segments 23(16–29) per leaf, the middle ones 36.1(30.0–45.0) cm long and 1.9(1.7–2.2) cm wide; segments not pendulous at the apices, giving the leaf a flat appearance; middle leaf segments tapering from base to apex, often folded, stiff and leathery, with or without scarcely developed shoulders, the apices sharply pointed and briefly splitting; middle leaf segment apices attenuate; leaf segments not waxy or sometimes with a deciduous, thin layer of wax adaxially, without indumentum, scales, or wax abaxially, without or with poorly developed transverse veinlets.
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<emphasis id="B96EEADEFFC81F58FDADFBD3FCE78C9F" bold="true" box="[629,798,1130,1156]" pageId="55" pageNumber="56">Inflorescences</emphasis>
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curving, arching, or pendulous amongst the leaves, with few partial inflorescences; rachis bracts narrow, closely sheathing, sparsely tomentose, usually without hairs at the apex; partial inflorescences 4; proximalmost rachillae straight, 5.2(3.0–7.7) cm long and 0.8(0.5–1.0) mm diameter in fruit; rachillae uneven at or near anthesis with lines of warty outgrowths, these often becoming more pronounced as fruits develop; stamens not recorded; fruit pedicels 0.6(0.3–1.1) mm long;
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<emphasis id="B96EEADEFFC81F58FC0CFB43FBEC8D0F" bold="true" box="[980,1045,1274,1300]" pageId="55" pageNumber="56">fruits</emphasis>
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<quantity id="4CE29B29FFC81F58FBC5FB43FB8D8D0E" box="[1053,1140,1274,1301]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="55" pageNumber="56" unit="mm" value="5.1">5.1 mm</quantity>
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long and
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<quantity id="4CE29B29FFC81F58FB3DFB43FAC58D0E" box="[1253,1340,1274,1301]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="55" pageNumber="56" unit="mm" value="5.6">5.6 mm</quantity>
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diameter, yellowish; fruit surfaces smooth or sometimes with projecting fibers; seed surfaces lobed, the lobes running from base of seeds approximately to equator.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3006547FFC81F58FF65FADFFC138F4F" pageId="55" pageNumber="56" type="distribution">
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<paragraph id="8BA536CCFFC81F58FF65FADFFC738DBF" blockId="55.[136,1452,806,1877]" pageId="55" pageNumber="56">
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<emphasis id="B96EEADEFFC81F58FF65FADFFDFC8D9B" bold="true" box="[189,517,1382,1408]" pageId="55" pageNumber="56">Distribution and habitat:—</emphasis>
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<collectingCountry id="F30D765CFFC81F58FDDDFADFFD178D9B" box="[517,750,1382,1408]" name="Dominican Republic" pageId="55" pageNumber="56">Dominican Republic</collectingCountry>
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(
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<collectingRegion id="49DEF82EFFC81F58FD24FADFFC5A8D9B" box="[764,931,1382,1408]" country="Dominican Republic" name="Independencia" pageId="55" pageNumber="56">Independencia</collectingRegion>
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) (
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<figureCitation id="13212A49FFC81F58FC61FADFFBF38D9B" box="[953,1034,1382,1408]" captionStart="FIGURE 13" captionStartId="52.[136,229,1506,1528]" captionTargetBox="[155,1429,198,1477]" captionTargetId="figure-27@52.[150,1434,193,1482]" captionTargetPageId="52" captionText="FIGURE 13. Distribution of Coccothrinax gracilis, C. gundlachii, C. hioramii, and C. jimenezii" figureDoi="http://doi.org/10.5281/zenodo.8389385" httpUri="https://zenodo.org/record/8389385/files/figure.png" pageId="55" pageNumber="56">Fig. 13</figureCitation>
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) in dry forest on limestone rocks and coral sediments, or dry, broad leaf forest, at 161(0–500) m elevation.
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</paragraph>
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<paragraph id="8BA536CCFFC81F58FF65FA17FB788E42" blockId="55.[136,1452,806,1877]" pageId="55" pageNumber="56">
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<emphasis id="B96EEADEFFC81F58FF65FA17FE568DD3" bold="true" box="[189,431,1454,1480]" pageId="55" pageNumber="56">Taxonomic notes:—</emphasis>
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As a preliminary species,
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<taxonomicName id="4C1A4D4FFFC81F58FD06FA17FC1F8DD3" authorityName="Mejia & Garcia" authorityYear="2013" box="[734,998,1454,1480]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="55" pageNumber="56" phylum="Tracheophyta" rank="species" species="jimenezii">
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<emphasis id="B96EEADEFFC81F58FD06FA17FC1F8DD3" box="[734,998,1454,1480]" italics="true" pageId="55" pageNumber="56">Coccothrinax jimenezii</emphasis>
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</taxonomicName>
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has a unique combination of character states and is recognized as a phylogenetic species. It is a member of a group of nine Hispaniolan species (see notes under
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<taxonomicName id="4C1A4D4FFFC81F58FF0AFA4EFE938E0B" authorityName="Mejia & Garcia" authorityYear="1997" box="[210,362,1526,1553]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="55" pageNumber="56" phylum="Tracheophyta" rank="species" species="boschiana">
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<emphasis id="B96EEADEFFC81F58FF0AFA4EFE938E0B" box="[210,362,1526,1553]" italics="true" pageId="55" pageNumber="56">C. boschiana</emphasis>
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</taxonomicName>
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). An unidentified specimen (
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<emphasis id="B96EEADEFFC81F58FD18FA4EFC998E0B" box="[704,864,1526,1552]" italics="true" pageId="55" pageNumber="56">Zanoni 33375</emphasis>
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) from near Pestel in southwestern
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<collectingCountry id="F30D765CFFC81F58FB24FA4FFACD8E0B" box="[1276,1332,1526,1552]" name="Haiti" pageId="55" pageNumber="56">Haiti</collectingCountry>
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shares all character states (although lacking fruits) with
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<taxonomicName id="4C1A4D4FFFC81F58FD50F9A2FCF48E2F" authorityName="Mejia & Garcia" authorityYear="2013" box="[648,781,1563,1589]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="55" pageNumber="56" phylum="Tracheophyta" rank="species" species="jimenezii">
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<emphasis id="B96EEADEFFC81F58FD50F9A2FCF48E2F" box="[648,781,1563,1589]" italics="true" pageId="55" pageNumber="56">C. jimenezii</emphasis>
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</taxonomicName>
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. However, it occurs almost
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<quantity id="4CE29B29FFC81F58FB9DF9A3FB638E2F" box="[1093,1178,1562,1589]" metricMagnitude="5" metricUnit="m" metricValue="2.5" pageId="55" pageNumber="56" unit="km" value="250.0">250 km</quantity>
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west of the
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|||
|
<typeStatus id="54A1886EFFC81F58FAF9F9A2FAA88E2E" box="[1313,1361,1563,1589]" pageId="55" pageNumber="56">type</typeStatus>
|
|||
|
locality of
|
|||
|
<taxonomicName id="4C1A4D4FFFC81F58FF7EF986FED58E43" authorityName="Mejia & Garcia" authorityYear="2013" box="[166,300,1599,1625]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="55" pageNumber="56" phylum="Tracheophyta" rank="species" species="jimenezii">
|
|||
|
<emphasis id="B96EEADEFFC81F58FF7EF986FED58E43" box="[166,300,1599,1625]" italics="true" pageId="55" pageNumber="56">C. jimenezii</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
and on biogeographic grounds it seems likely to represent a different taxon.
|
|||
|
</paragraph>
|
|||
|
<paragraph id="8BA536CCFFC81F58FF65F9DBFC138F4F" blockId="55.[136,1452,806,1877]" pageId="55" pageNumber="56">
|
|||
|
<emphasis id="B96EEADEFFC81F58FF65F9DBFE418E67" bold="true" box="[189,440,1634,1660]" pageId="55" pageNumber="56">Subspecific variation</emphasis>
|
|||
|
:—There is some local variation and there appear to be three populations. Five of the seven known specimens are from
|
|||
|
<quantity id="4CE29B29FFC81F58FE1FF93FFDE78EBA" box="[455,542,1670,1697]" metricMagnitude="1" metricUnit="m" metricValue="2.5" metricValueMax="5.0" metricValueMin="0.0" pageId="55" pageNumber="56" unit="m" value="25.0" valueMax="50.0" valueMin="0.0">0–50 m</quantity>
|
|||
|
elevation on the margins of Lago Enriquillo, the
|
|||
|
<typeStatus id="54A1886EFFC81F58FB84F93EFB758EBA" box="[1116,1164,1671,1697]" pageId="55" pageNumber="56">type</typeStatus>
|
|||
|
locality, in dry forest on limestone. Here the population is very small, and plants appear to have clustered stems. Another specimen (
|
|||
|
<emphasis id="B96EEADEFFC81F58FA84F912FF378EF3" italics="true" pageId="55" pageNumber="56">Zanoni 39724</emphasis>
|
|||
|
) occurs in a nearby locality at
|
|||
|
<quantity id="4CE29B29FFC81F58FDEFF977FD798EF2" box="[567,640,1742,1769]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="55" pageNumber="56" unit="m" value="500.0">500 m</quantity>
|
|||
|
elevation and is said to grow in “bosque latifoliado medio seco”. This specimen has a longer palman, more, longer segments, and longer pedicels than the specimens from the
|
|||
|
<typeStatus id="54A1886EFFC81F58FAC3F94AFAB28F16" box="[1307,1355,1779,1805]" pageId="55" pageNumber="56">type</typeStatus>
|
|||
|
locality. One other specimen (
|
|||
|
<emphasis id="B96EEADEFFC81F58FEA3F8AEFDF08F2B" box="[379,521,1815,1840]" italics="true" pageId="55" pageNumber="56">Montero s.n.</emphasis>
|
|||
|
) has been collected from a large population approximately
|
|||
|
<quantity id="4CE29B29FFC81F58FB76F8AFFB0F8F2B" box="[1198,1270,1814,1840]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="55" pageNumber="56" unit="km" value="20.0">20 km</quantity>
|
|||
|
due west of the
|
|||
|
<typeStatus id="54A1886EFFC81F58FF50F882FF418F4E" box="[136,184,1851,1877]" pageId="55" pageNumber="56">type</typeStatus>
|
|||
|
locality. This specimen has a solitary stem and thicker leaf sheath fibers.
|
|||
|
</paragraph>
|
|||
|
</subSubSection>
|
|||
|
</treatment>
|
|||
|
</document>
|