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<document id="0DE8F2A242A96A520E7F5CC3FB447072" ID-CLB-Dataset="8975" ID-DOI="10.11646/zootaxa.4903.4.3" ID-GBIF-Dataset="0a98a7e5-af6f-40b6-821e-c56567584422" ID-ISSN="1175-5326" ID-Zenodo-Dep="4431309" ID-ZooBank="CF96683E-75B8-4AC4-90A2-20AD8CC2F14B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1610349665964" checkinUser="plazi" docAuthor="Carrejo, Nancy, Obando, Ranulfo González, Aldrete, Alfonso Neri García &amp; Mendivil, Julián" docDate="2021" docId="03C49710FFEFFFB1598DFE5EB791FA15" docLanguage="en" docName="zootaxa.4903.4.3.pdf" docOrigin="Zootaxa 4903 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Goja caldasensis Carrejo &amp; Obando &amp; Aldrete &amp; Mendivil 2021, n. sp." docType="treatment" docVersion="7" lastPageNumber="505" masterDocId="FFFDEF68FFEBFFB5591AFF94B216FFED" masterDocTitle="New species of Goja Navás (Psocodea: ‘ Psocoptera’: Epipsocidae) from Colombia Mexico and Peru" masterLastPageNumber="541" masterPageNumber="501" pageNumber="505" updateTime="1698924142089" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="D5710ECBECA3ABE75F1E03F59D8FF40F">New species of Goja Navás (Psocodea: ‘ Psocoptera’: Epipsocidae) from Colombia Mexico and Peru</mods:title>
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<mods:namePart id="CD875E01436A1F556814977066595700">Carrejo, Nancy</mods:namePart>
<mods:nameIdentifier id="C88B28D1D2BD4E5F019E71D7F43145BB" type="ORCID">0000-0002-2395-6887</mods:nameIdentifier>
<mods:affiliation id="5193034B8FD655C99515554E9FA5F997">Departamento de Biología, Facultad de Ciencias Naturales y Exactas, Universidad del Valle, Santiago de Cali, COLOMBIA. Departamento de Zoología, Instituto de Biología, Universidad Nacional Autónoma de México, Apdo. Postal 70 - 153, CdMx, MÉXICO. anga @ ib. unam. mx; https: // orcid. org / 0000 - 0001 - 7214 - 7966 &amp; nancy. carrejo @ correounivalle. edu. co; https: // orcid. org / 0000 - 0002 - 2395 - 6887</mods:affiliation>
<mods:nameIdentifier id="3105CE9A3C5870F176C6ACB728E98DB8" type="email">nancy.carrejo@correounivalle.edu.co</mods:nameIdentifier>
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<mods:namePart id="FA7CB208C062271789DFBB6BB8E26D9F">Obando, Ranulfo González</mods:namePart>
<mods:nameIdentifier id="584A4CA3C6CE84B8EA37270EFBF26B82" type="ORCID">0000-0003-4047-8677</mods:nameIdentifier>
<mods:affiliation id="8C27D4DE2128F02BA50387F3B2DD6356">Departamento de Biología, Facultad de Ciencias Naturales y Exactas, Universidad del Valle, Santiago de Cali, COLOMBIA. Departamento de Zoología, Instituto de Biología, Universidad Nacional Autónoma de México, Apdo. Postal 70 - 153, CdMx, MÉXICO. anga @ ib. unam. mx; https: // orcid. org / 0000 - 0001 - 7214 - 7966 &amp; ranulfo. gonzalez @ correounivalle. edu. co; https: // orcid. org / 0000 - 0003 - 4047 - 8677</mods:affiliation>
<mods:nameIdentifier id="AB68FA87A6E3497B98A42F04B4708A97" type="email">ranulfo.gonzalez@correounivalle.edu.co</mods:nameIdentifier>
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<mods:namePart id="E832238984C7D338BCA87735F7685FC3">Aldrete, Alfonso Neri García</mods:namePart>
<mods:nameIdentifier id="541CF5175CAF1A2ED09F8FBA5A32DA99" type="ORCID">0000-0001-7214-7966</mods:nameIdentifier>
<mods:nameIdentifier id="835CD9E64DAE90B15A21ABAFEF977B59" type="email">anga@ib.unam.mx</mods:nameIdentifier>
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<mods:namePart id="C202B87286D7E86A7BD02B4D662FAB32">Mendivil, Julián</mods:namePart>
<mods:nameIdentifier id="B86C0BE9BBC64DA31C845E205AA6D817" type="ORCID">0000-0002-6339-2000</mods:nameIdentifier>
<mods:affiliation id="E03013599A7BCDD193377C11B7F01BBB">Departamento de Biología, Facultad de Ciencias Naturales y Exactas, Universidad del Valle, Santiago de Cali, COLOMBIA. Departamento de Zoología, Instituto de Biología, Universidad Nacional Autónoma de México, Apdo. Postal 70 - 153, CdMx, MÉXICO. anga @ ib. unam. mx; https: // orcid. org / 0000 - 0001 - 7214 - 7966 &amp; chalcidoidea 88 @ gmail. com; https: // orcid. org / 0000 - 0002 - 6339 - 2000</mods:affiliation>
<mods:nameIdentifier id="0636D63FC8611B63FBC5A5BBFAA7D6DE" type="email">chalcidoidea88@gmail.com</mods:nameIdentifier>
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<mods:title id="FB881F273A9DA97390D1610FA927E9FB">Zootaxa</mods:title>
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<mods:date id="14E7EC3F90B1079DE1C8E9C6B272CFD1">2021</mods:date>
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<mods:number id="A45F32846A9B99038A4A879BEBD0E11A">2021-01-11</mods:number>
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<mods:number id="526C8696FCB9FC9A8E8587683BF36271">4903</mods:number>
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<mods:identifier id="82DE10D766D2A57F4D0B93A32EE49114" type="DOI">10.11646/zootaxa.4903.4.3</mods:identifier>
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<treatment id="03C49710FFEFFFB1598DFE5EB791FA15" ID-DOI="http://doi.org/10.5281/zenodo.4431311" ID-GBIF-Taxon="176570444" ID-Zenodo-Dep="4431311" LSID="urn:lsid:plazi:treatment:03C49710FFEFFFB1598DFE5EB791FA15" httpUri="http://treatment.plazi.org/id/03C49710FFEFFFB1598DFE5EB791FA15" lastPageNumber="505" pageId="4" pageNumber="505">
<subSubSection id="C377758DFFEFFFB1598DFE5EB3EFFE08" box="[151,505,458,485]" pageId="4" pageNumber="505" type="nomenclature">
<paragraph id="8BD22606FFEFFFB1598DFE5EB3EFFE08" blockId="4.[151,505,458,521]" box="[151,505,458,485]" pageId="4" pageNumber="505">
<heading id="D09A916AFFEFFFB1598DFE5EB3EFFE08" bold="true" box="[151,505,458,485]" fontSize="11" level="1" pageId="4" pageNumber="505" reason="1">
<emphasis id="B919FA14FFEFFFB1598DFE5EB3EFFE08" bold="true" box="[151,505,458,485]" pageId="4" pageNumber="505">
<taxonomicName id="4C6D5D85FFEFFFB1598DFE5EB374FE08" ID-CoL="9HZXR" authority="Carrejo &amp; Obando &amp; Aldrete &amp; Mendivil, 2021" authorityName="Carrejo &amp; Obando &amp; Aldrete &amp; Mendivil" authorityYear="2021" box="[151,354,458,485]" class="Insecta" family="Epipsocidae" genus="Goja" kingdom="Animalia" order="Psocodea" pageId="4" pageNumber="505" phylum="Arthropoda" rank="species" species="caldasensis" status="sp. nov.">
<emphasis id="B919FA14FFEFFFB1598DFE5EB374FE08" bold="true" box="[151,354,458,485]" italics="true" pageId="4" pageNumber="505">Goja caldasensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22A476FFFEFFFB15873FE5FB3BAFE08" box="[361,428,459,485]" pageId="4" pageNumber="505" rank="species">n. sp.</taxonomicNameLabel>
Male.
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C377758DFFEFFFB1598DFE7AB31DFDE4" box="[151,267,494,521]" pageId="4" pageNumber="505" type="description">
<paragraph id="8BD22606FFEFFFB1598DFE7AB31DFDE4" blockId="4.[151,505,458,521]" box="[151,267,494,521]" pageId="4" pageNumber="505">
(
<figureCitation id="13563A83FFEFFFB15985FE7AB314FDE4" box="[159,258,494,521]" captionStart="FIGURES 1‒6" captionStartId="5.[151,264,1918,1942]" captionTargetBox="[156,1432,197,1866]" captionTargetId="figure-17@5.[151,1435,181,1894]" captionTargetPageId="5" captionText="FIGURES 1‒6. G. caldasensis n. sp. Male. 1. Forewing. 2. Hindwing. 3. Front view of head. 4. Left paraproct and epiproct. 5. Hypandrium. 6. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.4431313" httpUri="https://zenodo.org/record/4431313/files/figure.png" pageId="4" pageNumber="505">Figs 1–6</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C377758DFFEFFFB1598DFDA2B001FAA9" pageId="4" pageNumber="505" type="diagnosis">
<paragraph id="8BD22606FFEFFFB1598DFDA2B779FD0D" blockId="4.[151,1437,566,1529]" pageId="4" pageNumber="505">
<emphasis id="B919FA14FFEFFFB1598DFDA2B304FDBD" bold="true" box="[151,274,566,592]" pageId="4" pageNumber="505">Diagnosis.</emphasis>
Forewing Rs dichotomously branched, R2+3 forked, R4+5 dichotomously branched, resulting in four veins. M with five primary branches, M2, M3 and M5 forked as illustrated, resulting in ten veins. Hindwing Rs forked, M three branched, with M2 forked.Anterior pair of endophallic sclerites directed posteriorly, curved, distally acuminate; mesal pair with 2–3 long, acuminate processes directed outward (
<figureCitation id="13563A83FFEFFFB15AF0FD36B63BFD50" box="[1002,1069,674,701]" captionStart="FIGURES 1‒6" captionStartId="5.[151,264,1918,1942]" captionTargetBox="[156,1432,197,1866]" captionTargetId="figure-17@5.[151,1435,181,1894]" captionTargetPageId="5" captionText="FIGURES 1‒6. G. caldasensis n. sp. Male. 1. Forewing. 2. Hindwing. 3. Front view of head. 4. Left paraproct and epiproct. 5. Hypandrium. 6. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.4431313" httpUri="https://zenodo.org/record/4431313/files/figure.png" pageId="4" pageNumber="505">Fig. 6</figureCitation>
). It is similar to
<taxonomicName id="4C6D5D85FFEFFFB15DFAFD37B78BFD51" authority="Carrejo &amp; Obando &amp; Aldrete &amp; Mendivil, 2021" authorityName="Carrejo &amp; Obando &amp; Aldrete &amp; Mendivil" authorityYear="2021" box="[1248,1437,674,701]" class="Insecta" family="Epipsocidae" genus="Goja" kingdom="Animalia" order="Psocodea" pageId="4" pageNumber="505" phylum="Arthropoda" rank="species" species="pillcopatensis" status="n. sp. and g.">
<emphasis id="B919FA14FFEFFFB15DFAFD37B78BFD51" box="[1248,1437,674,701]" italics="true" pageId="4" pageNumber="505">G. pillcopatensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22A476FFFEFFFB1598DFD53B33EFD0C" box="[151,296,710,737]" pageId="4" pageNumber="505" rank="genus">
<emphasis id="B919FA14FFEFFFB1598DFD53B2C0FD0C" bold="true" box="[151,214,711,737]" pageId="4" pageNumber="505">n. sp.</emphasis>
and
<taxonomicName id="4C6D5D85FFEFFFB15817FD53B33EFD0C" authority="Carrejo &amp; Obando &amp; Aldrete &amp; Mendivil, 2021" authorityName="Carrejo &amp; Obando &amp; Aldrete &amp; Mendivil" authorityYear="2021" box="[269,296,711,737]" class="Insecta" family="Epipsocidae" genus="Goja" kingdom="Animalia" order="Psocodea" pageId="4" pageNumber="505" phylum="Arthropoda" rank="species" species="yarumosensis" status="sp. nov.">
<emphasis id="B919FA14FFEFFFB15817FD53B33EFD0C" box="[269,296,711,737]" italics="true" pageId="4" pageNumber="505">G.</emphasis>
</taxonomicName>
</taxonomicNameLabel>
<emphasis id="B919FA14FFEFFFB15835FD53B3D1FD0D" box="[303,455,711,736]" italics="true" pageId="4" pageNumber="505">yarumosensis</emphasis>
<emphasis id="B919FA14FFEFFFB158D4FD53B01BFD0C" bold="true" box="[462,525,711,737]" pageId="4" pageNumber="505">
<taxonomicNameLabel id="A22A476FFFEFFFB158D4FD53B01BFD0C" box="[462,525,711,737]" pageId="4" pageNumber="505" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, differing from them in wing venation, phallosome and endophallic sclerites.
</paragraph>
<paragraph id="8BD22606FFEFFFB159DDFD7EB2F0FC31" blockId="4.[151,1437,566,1529]" pageId="4" pageNumber="505">
<emphasis id="B919FA14FFEFFFB159DDFD7EB31DFCE9" bold="true" box="[199,267,746,772]" pageId="4" pageNumber="505">Color</emphasis>
(in 80% ethanol). Body brown-yellowish. Head pale brown (
<figureCitation id="13563A83FFEFFFB15AA9FD7EB1E0FCE8" box="[947,1014,746,773]" captionStart="FIGURES 1‒6" captionStartId="5.[151,264,1918,1942]" captionTargetBox="[156,1432,197,1866]" captionTargetId="figure-17@5.[151,1435,181,1894]" captionTargetPageId="5" captionText="FIGURES 1‒6. G. caldasensis n. sp. Male. 1. Forewing. 2. Hindwing. 3. Front view of head. 4. Left paraproct and epiproct. 5. Hypandrium. 6. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.4431313" httpUri="https://zenodo.org/record/4431313/files/figure.png" pageId="4" pageNumber="505">Fig. 3</figureCitation>
), postclypeus with ochre striae. Compound eyes dark brown, ocelli hyaline, with ochre centripetal crescents. Maxillary palps pale brown. Antennae pale brown. Thorax pale brown, pronotum cream, with brown spots; tergal lobes of meso- and metathorax with brown spots, mesopleura pale brown. Legs pale brown. Forewings (
<figureCitation id="13563A83FFEFFFB15A5FFCC2B191FC9C" box="[837,903,854,881]" captionStart="FIGURES 1‒6" captionStartId="5.[151,264,1918,1942]" captionTargetBox="[156,1432,197,1866]" captionTargetId="figure-17@5.[151,1435,181,1894]" captionTargetPageId="5" captionText="FIGURES 1‒6. G. caldasensis n. sp. Male. 1. Forewing. 2. Hindwing. 3. Front view of head. 4. Left paraproct and epiproct. 5. Hypandrium. 6. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.4431313" httpUri="https://zenodo.org/record/4431313/files/figure.png" pageId="4" pageNumber="505">Fig. 1</figureCitation>
), pale yellowish, veins pale brown. Hindwings (
<figureCitation id="13563A83FFEFFFB159BAFCEEB2F5FC78" box="[160,227,890,917]" captionStart="FIGURES 1‒6" captionStartId="5.[151,264,1918,1942]" captionTargetBox="[156,1432,197,1866]" captionTargetId="figure-17@5.[151,1435,181,1894]" captionTargetPageId="5" captionText="FIGURES 1‒6. G. caldasensis n. sp. Male. 1. Forewing. 2. Hindwing. 3. Front view of head. 4. Left paraproct and epiproct. 5. Hypandrium. 6. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.4431313" httpUri="https://zenodo.org/record/4431313/files/figure.png" pageId="4" pageNumber="505">Fig. 2</figureCitation>
), hyaline, with a yellowish hue, veins pale brown. Abdomen pale brown. Hypandrium brown, darker distally on sides; epiproct and paraprocts pale brown. Phallosome brown, with endophallic sclerites and aedeagal arch pale brown.
</paragraph>
<paragraph id="8BD22606FFEFFFB159DDFC73B001FAA9" blockId="4.[151,1437,566,1529]" pageId="4" pageNumber="505">
<emphasis id="B919FA14FFEFFFB159DDFC73B34DFBEC" bold="true" box="[199,347,999,1025]" pageId="4" pageNumber="505">Morphology</emphasis>
. As in diagnosis, plus the following: Head: H/MxW: 1.41; compound eyes large, H/d: 4.22; IO/ MxW: 0.74. Outer cusp of lacinial tips broad, with nine denticles. Mx4/Mx2: 1.08. Forewings (
<figureCitation id="13563A83FFEFFFB15DA8FB9EB6E2FBC8" box="[1202,1268,1034,1061]" captionStart="FIGURES 1‒6" captionStartId="5.[151,264,1918,1942]" captionTargetBox="[156,1432,197,1866]" captionTargetId="figure-17@5.[151,1435,181,1894]" captionTargetPageId="5" captionText="FIGURES 1‒6. G. caldasensis n. sp. Male. 1. Forewing. 2. Hindwing. 3. Front view of head. 4. Left paraproct and epiproct. 5. Hypandrium. 6. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.4431313" httpUri="https://zenodo.org/record/4431313/files/figure.png" pageId="4" pageNumber="505">Fig. 1</figureCitation>
): FW/W: 2.51; M 5-branched, resulting in 10 veins at wing margin as illustrated. Pterostigma: lp/wp: 3.46, areola postica tall, triangular: al/ah: 2.54. R2+3 forked, R4+5 dichotomously branched, resulting in four veins at wing margin; Hindwings (
<figureCitation id="13563A83FFEFFFB159BAFBE2B2F0FB7C" box="[160,230,1142,1169]" captionStart="FIGURES 1‒6" captionStartId="5.[151,264,1918,1942]" captionTargetBox="[156,1432,197,1866]" captionTargetId="figure-17@5.[151,1435,181,1894]" captionTargetPageId="5" captionText="FIGURES 1‒6. G. caldasensis n. sp. Male. 1. Forewing. 2. Hindwing. 3. Front view of head. 4. Left paraproct and epiproct. 5. Hypandrium. 6. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.4431313" httpUri="https://zenodo.org/record/4431313/files/figure.png" pageId="4" pageNumber="505">Fig. 2</figureCitation>
): HW/w: 2.95. Rs two-branched, M three-branched, M2 forked. Hypandrium (
<figureCitation id="13563A83FFEFFFB15D79FBE2B6BEFB7C" box="[1123,1192,1142,1169]" captionStart="FIGURES 1‒6" captionStartId="5.[151,264,1918,1942]" captionTargetBox="[156,1432,197,1866]" captionTargetId="figure-17@5.[151,1435,181,1894]" captionTargetPageId="5" captionText="FIGURES 1‒6. G. caldasensis n. sp. Male. 1. Forewing. 2. Hindwing. 3. Front view of head. 4. Left paraproct and epiproct. 5. Hypandrium. 6. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.4431313" httpUri="https://zenodo.org/record/4431313/files/figure.png" pageId="4" pageNumber="505">Fig. 5</figureCitation>
), concave posteriorly in the middle, setose as illustrated; phallosome (
<figureCitation id="13563A83FFEFFFB15BA0FB0EB0E8FB58" box="[698,766,1178,1205]" captionStart="FIGURES 1‒6" captionStartId="5.[151,264,1918,1942]" captionTargetBox="[156,1432,197,1866]" captionTargetId="figure-17@5.[151,1435,181,1894]" captionTargetPageId="5" captionText="FIGURES 1‒6. G. caldasensis n. sp. Male. 1. Forewing. 2. Hindwing. 3. Front view of head. 4. Left paraproct and epiproct. 5. Hypandrium. 6. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.4431313" httpUri="https://zenodo.org/record/4431313/files/figure.png" pageId="4" pageNumber="505">Fig. 6</figureCitation>
) with external parameres dilated distally, apically truncate, bearing pores. Endophallic sclerites (
<figureCitation id="13563A83FFEFFFB15B2EFB2AB06EFB34" box="[564,632,1214,1241]" captionStart="FIGURES 1‒6" captionStartId="5.[151,264,1918,1942]" captionTargetBox="[156,1432,197,1866]" captionTargetId="figure-17@5.[151,1435,181,1894]" captionTargetPageId="5" captionText="FIGURES 1‒6. G. caldasensis n. sp. Male. 1. Forewing. 2. Hindwing. 3. Front view of head. 4. Left paraproct and epiproct. 5. Hypandrium. 6. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.4431313" httpUri="https://zenodo.org/record/4431313/files/figure.png" pageId="4" pageNumber="505">Fig. 6</figureCitation>
). Paraprocts (
<figureCitation id="13563A83FFEFFFB15A08FB2AB140FB34" box="[786,854,1214,1241]" captionStart="FIGURES 1‒6" captionStartId="5.[151,264,1918,1942]" captionTargetBox="[156,1432,197,1866]" captionTargetId="figure-17@5.[151,1435,181,1894]" captionTargetPageId="5" captionText="FIGURES 1‒6. G. caldasensis n. sp. Male. 1. Forewing. 2. Hindwing. 3. Front view of head. 4. Left paraproct and epiproct. 5. Hypandrium. 6. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.4431313" httpUri="https://zenodo.org/record/4431313/files/figure.png" pageId="4" pageNumber="505">Fig. 4</figureCitation>
), broadly triangular, with a macrosetae near sensory field, and a distal setal field, one setae distally spatulate; sensory fields with 28 trichobothria on basal rosettes. Epiproct (
<figureCitation id="13563A83FFEFFFB1581DFA92B35DFACC" box="[263,331,1286,1313]" captionStart="FIGURES 1‒6" captionStartId="5.[151,264,1918,1942]" captionTargetBox="[156,1432,197,1866]" captionTargetId="figure-17@5.[151,1435,181,1894]" captionTargetPageId="5" captionText="FIGURES 1‒6. G. caldasensis n. sp. Male. 1. Forewing. 2. Hindwing. 3. Front view of head. 4. Left paraproct and epiproct. 5. Hypandrium. 6. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.4431313" httpUri="https://zenodo.org/record/4431313/files/figure.png" pageId="4" pageNumber="505">Fig. 4</figureCitation>
) wide, semioval, rounded posteriorly, with two lateral macrosetae, setal field on distal fourth as illustrated, one macrosetae mesally.
</paragraph>
</subSubSection>
<subSubSection id="C377758DFFEFFFB159DDFADBB73BFA39" pageId="4" pageNumber="505" type="description">
<paragraph id="8BD22606FFEFFFB159DDFADBB081FA60" blockId="4.[151,1437,566,1529]" pageId="4" pageNumber="505">
<emphasis id="B919FA14FFEFFFB159DDFADBB368FA84" bold="true" box="[199,382,1359,1385]" pageId="4" pageNumber="505">Measurements.</emphasis>
FW: 3737.5, HW: 2875, F: 950, T: 1560, t1: 710, t2: 240, ctt1: 26, f1: 700, f2: 550, Mx4: 260, IO: 400, D: 243, d: 180, IO/d: 2.22, PO: 0.74.
</paragraph>
<paragraph id="8BD22606FFEFFFB159DDFA02B73BFA39" blockId="4.[151,1437,566,1529]" pageId="4" pageNumber="505">
<emphasis id="B919FA14FFEFFFB159DDFA02B38EFA5D" bold="true" box="[199,408,1430,1456]" pageId="4" pageNumber="505">Specimen studied</emphasis>
.
<materialsCitation id="3B052C5BFFEFFFB158BFFA02B73BFA39" ID-GBIF-Occurrence="3014907325" collectingDate="2011-01-15" collectionCode="R" collectorName="Gonzalez &amp; A. N. Garcia Aldrete. Beating" country="Colombia" elevation="2592" latitude="5.0734167" location="Rio Blanco Natural Reserve" longLatPrecision="1" longitude="-75.437836" municipality="Manizales" pageId="4" pageNumber="505" specimenCount="1" stateProvince="Caldas" typeStatus="holotype">
<typeStatus id="54D698A4FFEFFFB158BFFA02B01BFA5D" box="[421,525,1430,1456]" pageId="4" pageNumber="505" type="holotype">Holotype</typeStatus>
male.
<collectingCountry id="F37A6696FFEFFFB15B42FA03B0E7FA5C" box="[600,753,1431,1457]" name="Colombia" pageId="4" pageNumber="505">COLOMBIA</collectingCountry>
.
<collectingRegion id="49A9E8E4FFEFFFB15BE1FA02B15CFA5D" box="[763,842,1430,1456]" country="Colombia" name="Caldas" pageId="4" pageNumber="505">Caldas</collectingRegion>
,
<collectingMunicipality id="6BB6BC7CFFEFFFB15A4EFA02B1DCFA5D" box="[852,970,1430,1456]" pageId="4" pageNumber="505">Manizales</collectingMunicipality>
,
<location id="8EB270DDFFEFFFB15ACFFA03B718FA5C" LSID="urn:lsid:plazi:treatment:03C49710FFEFFFB1598DFE5EB791FA15:8EB270DDFFEFFFB15ACFFA03B718FA5C" box="[981,1294,1430,1457]" country="Colombia" latitude="5.0734167" longLatPrecision="1" longitude="-75.437836" municipality="Manizales" name="Rio Blanco Natural Reserve" pageId="4" pageNumber="505" stateProvince="Caldas">Río Blanco Natural Reserve</location>
<geoCoordinate id="EE5940C1FFEFFFB15C0FFA02B78FFA5C" box="[1301,1433,1430,1457]" degrees="5" direction="north" minutes="4" orientation="latitude" pageId="4" pageNumber="505" precision="1" seconds="24.3" value="5.0734167">5º4’24.3”N</geoCoordinate>
:
<geoCoordinate id="EE5940C1FFEFFFB1598DFA2EB32FFA38" box="[151,313,1466,1493]" degrees="75" direction="west" minutes="26" orientation="longitude" pageId="4" pageNumber="505" precision="1" seconds="16.2" value="-75.437836">75º26’16.2”W</geoCoordinate>
,
<quantity id="4C958BE3FFEFFFB1585EFA2EB388FA38" box="[324,414,1466,1493]" metricMagnitude="3" metricUnit="m" metricValue="2.5919999999999996" pageId="4" pageNumber="505" unit="m" value="2592.0">
<elevation id="0040C135FFEFFFB1585EFA2EB388FA38" box="[324,414,1466,1493]" metricMagnitude="3" metricUnit="m" metricValue="2.5919999999999996" pageId="4" pageNumber="505" unit="m" value="2592.0">2592 m</elevation>
</quantity>
.,
<date id="FFD300C6FFEFFFB158B4FA2EB00AFA38" box="[430,540,1466,1493]" pageId="4" pageNumber="505" value="2011-01-15">
<collectingDate id="EF97F92EFFEFFFB158B4FA2EB00AFA38" box="[430,540,1466,1493]" pageId="4" pageNumber="505" value="2011-01-15">15.I.2011</collectingDate>
</date>
.
<collectionCode id="ED7CBEC3FFEFFFB15B3DFA2FB02BFA38" box="[551,573,1467,1493]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="4" pageNumber="505">R</collectionCode>
.
<collectorName id="269843D0FFEFFFB15B5DFA2EB0A7FA39" box="[583,689,1466,1492]" pageId="4" pageNumber="505">González</collectorName>
&amp;
<collectorName id="269843D0FFEFFFB15BC9FA2FB637FA39" box="[723,1057,1466,1493]" pageId="4" pageNumber="505">A. N. García Aldrete. Beating</collectorName>
branches of vegetation.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C377758DFFEFFFB159DDFA4BB791FA15" box="[199,1415,1502,1529]" pageId="4" pageNumber="505" type="etymology">
<paragraph id="8BD22606FFEFFFB159DDFA4BB791FA15" blockId="4.[151,1437,566,1529]" box="[199,1415,1502,1529]" pageId="4" pageNumber="505">
<emphasis id="B919FA14FFEFFFB159DDFA4BB358FA14" bold="true" box="[199,334,1503,1529]" pageId="4" pageNumber="505">Etymology.</emphasis>
The specific epithet refers to
<collectingRegion id="49A9E8E4FFEFFFB15B86FA4AB0FDFA15" box="[668,747,1502,1528]" country="Colombia" name="Caldas" pageId="4" pageNumber="505">Caldas</collectingRegion>
, the Colombian Department where the
<typeStatus id="54D698A4FFEFFFB15DB8FA4AB712FA15" box="[1186,1284,1502,1528]" pageId="4" pageNumber="505" type="holotype">holotype</typeStatus>
was found.
</paragraph>
</subSubSection>
</treatment>
</document>
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