treatments-xml/data/03/89/87/03898795FFCDFFA9EEA2FCC69ABDFD2E.xml

<document id="145EDC8EF76C0AD7A774E6CE33907287" ID-DOI="10.5281/zenodo.178296" ID-GBIF-Dataset="0366c4ab-3941-4376-806f-6cf6e3d8c403" ID-ISSN="1175-5326" ID-Zenodo-Dep="178296" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459889844187" checkinUser="plazi" docAuthor="Ng, Peter K. L. &amp; Manuel-Santos, Marivene R." docDate="2007" docId="03898795FFCDFFA9EEA2FCC69ABDFD2E" docLanguage="en" docName="zt01558p068.pdf" docOrigin="Zootaxa 1558" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Vultocinus anfractus Ng &amp; Manuel-Santos, 2007, new species" docType="treatment" docVersion="9" lastPageNumber="52" masterDocId="FFB0FFEDFFC7FFA4EE35FFA49A55FF94" masterDocTitle="Establishment of the Vultocinidae, a new family for an unusual new genus and new species of Indo-West Pacific crab (Crustacea: Decapoda: Brachyura: Goneplacoidea), with comments on the taxonomy of the Goneplacidae" masterLastPageNumber="68" masterPageNumber="39" pageNumber="49" updateTime="1698227947623" updateUser="plazi">
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<mods:title id="8B778F6FF25F4CF0494776EE1C62C16B">Establishment of the Vultocinidae, a new family for an unusual new genus and new species of Indo-West Pacific crab (Crustacea: Decapoda: Brachyura: Goneplacoidea), with comments on the taxonomy of the Goneplacidae</mods:title>
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<mods:namePart id="8A8A9601095EF23115ECDC6F4F00E8A2">Ng, Peter K. L.</mods:namePart>
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<mods:namePart id="728DC10D85FEA2DE4C6D9176D632A201">Manuel-Santos, Marivene R.</mods:namePart>
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<mods:date id="B9D7156D6F7228C66F32BA2AB00C0824">2007</mods:date>
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<treatment id="03898795FFCDFFA9EEA2FCC69ABDFD2E" ID-DOI="http://doi.org/10.5281/zenodo.6250890" ID-GBIF-Taxon="119371812" ID-Zenodo-Dep="6250890" LSID="urn:lsid:plazi:treatment:03898795FFCDFFA9EEA2FCC69ABDFD2E" httpUri="http://treatment.plazi.org/id/03898795FFCDFFA9EEA2FCC69ABDFD2E" lastPageId="13" lastPageNumber="52" pageId="10" pageNumber="49">
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<paragraph id="8B9F3683FFCDFFAEEEA2FCC69867FCE8" blockId="10.[151,620,866,927]" box="[151,562,866,892]" pageId="10" pageNumber="49">
<heading id="D0D781EFFFCDFFAEEEA2FCC69867FCE8" bold="true" box="[151,562,866,892]" fontSize="11" level="1" pageId="10" pageNumber="49" reason="1">
<emphasis id="B954EA91FFCDFFAEEEA2FCC69867FCE8" bold="true" box="[151,562,866,892]" pageId="10" pageNumber="49">
<taxonomicName id="4C204D00FFCDFFAEEEA2FCC69BC1FCE8" ID-CoL="5BSLK" box="[151,404,866,892]" class="Malacostraca" family="Vultocinidae" genus="Vultocinus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="49" phylum="Arthropoda" rank="species" species="anfractus" status="sp. nov.">
<emphasis id="B954EA91FFCDFFAEEEA2FCC69BC1FCE8" bold="true" box="[151,404,866,892]" italics="true" pageId="10" pageNumber="49">Vultocinus anfractus</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A26757EAFFCDFFAEEF96FCC69867FCE8" box="[419,562,866,892]" pageId="10" pageNumber="49" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B9F3683FFCDFFAEEEA2FC219839FC0B" blockId="10.[151,620,866,927]" box="[151,620,901,927]" pageId="10" pageNumber="49">
<figureCitation id="131B2A06FFCDFFAEEEA2FC219B53FC0B" box="[151,262,901,927]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStartId-0="17.[151,255,1912,1936]" captionStartId-1="18.[151,255,1936,1960]" captionStartId-2="19.[151,255,1088,1112]" captionStartId-3="20.[151,255,1144,1168]" captionStartId-4="21.[151,255,1379,1403]" captionTargetBox-0="[226,1360,194,1887]" captionTargetBox-1="[264,1322,182,1924]" captionTargetBox-2="[151,1435,194,1064]" captionTargetBox-3="[151,1435,184,1120]" captionTargetBox-4="[158,1421,201,1348]" captionTargetId-0="figure@17.[226,1360,194,1888]" captionTargetId-1="figure@18.[264,1323,182,1926]" captionTargetId-2="figure@19.[151,1436,194,1064]" captionTargetId-3="figure@20.[151,1436,184,1120]" captionTargetId-4="figure@21.[151,1436,194,1355]" captionTargetPageId-0="17" captionTargetPageId-1="18" captionTargetPageId-2="19" captionTargetPageId-3="20" captionTargetPageId-4="21" captionText-0="FIGURE 1. Vultocinus anfractus, new genus, new species. Fresh colours. A, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC), Philippines, freshly dead, poised on same kind of wood substrate it was reportedly found on; B, paratype male (cl 19.9 mm, cw 22.0 mm) (ZRC), Philippines; C, male (cl 15.6 mm, cw 16.8 mm) (ZRC), Vanuatu." captionText-1="FIGURE 2. Vultocinus anfractus, new genus, new species. Overall dorsal views. A, holotype male (cl 19.5 mm, cw 20.3 mm) (NMCR 19114), Philippines; B, paratype male (cl 19.9 mm, cw 22.0 mm) (ZRC), Philippines; C, male (cl 15.6 mm, cw 16.8 mm) (ZRC), Vanuatu." captionText-2="FIGURE 3. Vultocinus anfractus, new genus, new species. Holotype male (cl 19.5 mm, cw 20.3 mm) (NMCR 19114), Philippines. A, dorsal view of carapace; B, ventral view of carapace; C, buccal area showing third maxillipeds; D, frontal view of carapace and chelipeds." captionText-3="FIGURE 4. Vultocinus anfractus, new genus, new species. Holotype male (cl 19.5 mm, cw 20.3 mm) (NMCR 19114), Philippines. A, outer surface of right chela; B, outer surface of left chela; C, right side of thoracic sternum showing ventral surface of chelipeds and spines on ambulatory meri; D, right P 4." captionText-4="FIGURE 5. Vultocinus anfractus, new genus, new species. Paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC), Philippines. A, left third maxilliped (setae denuded); B, male abdomen (partially denuded); C, left G 1, dorsal view; D, left G 1, ventral view; E, distal part of left G 1, dorsal view; F, distal part of left G 1, ventral view; G, left G 2; H, distal part of left G 2. Scales = 1.0 mm." httpUri-0="https://zenodo.org/record/178297/files/figure.png" httpUri-1="https://zenodo.org/record/178298/files/figure.png" httpUri-2="https://zenodo.org/record/178299/files/figure.png" httpUri-3="https://zenodo.org/record/178300/files/figure.png" httpUri-4="https://zenodo.org/record/178301/files/figure.png" pageId="10" pageNumber="49">Figs. 1–5</figureCitation>
, 
<figureCitation id="131B2A06FFCDFFAEEF27FC219B75FC0B" box="[274,288,901,927]" captionStart="FIGURE 8" captionStartId="24.[151,255,1099,1123]" captionTargetBox="[151,1435,194,1073]" captionTargetId="figure@24.[151,1436,194,1075]" captionTargetPageId="24" captionText="FIGURE 8. Buccal field, showing mouthparts and mandibles. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, Mathildella rubra Ng &amp; Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); C, Progeryon mus Ng &amp; Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC); D, Conleyus defodio Ng &amp; Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH)." httpUri="https://zenodo.org/record/178304/files/figure.png" pageId="10" pageNumber="49">8</figureCitation>
A, 9A, 10A, 11A, 12A, 13A
</paragraph>
</subSubSection>
<subSubSection id="C33A6508FFCDFFAEEEA2FC719968FAA0" pageId="10" pageNumber="49" type="materials_examined">
<paragraph id="8B9F3683FFCDFFAEEEA2FC719968FAA0" blockId="10.[151,1437,981,2012]" pageId="10" pageNumber="49">
<emphasis id="B954EA91FFCDFFAEEEA2FC719BD2FC7B" bold="true" box="[151,391,981,1007]" pageId="10" pageNumber="49">Material examined</emphasis>
. 
<typeStatus id="549B8821FFCDFFAEEFAEFC719859FC7B" box="[411,524,981,1007]" pageId="10" pageNumber="49" type="holotype">Holotype</typeStatus>
: male (cl 
<quantity id="4CD89B66FFCDFFAEECBFFC7198AFFC7B" box="[650,762,981,1007]" metricMagnitude="-2" metricUnit="m" metricValue="1.95" pageId="10" pageNumber="49" unit="mm" value="19.5">19.5 mm</quantity>
, cw 
<quantity id="4CD89B66FFCDFFAEED02FC7199F3FC7B" box="[823,934,981,1007]" metricMagnitude="-2" metricUnit="m" metricValue="2.03" pageId="10" pageNumber="49" unit="mm" value="20.3">20.3 mm</quantity>
) (
<collectionCode id="ED31AE46FFCDFFAEEDF4FC719E4CFC7B" box="[961,1049,981,1007]" pageId="10" pageNumber="49">NMCR</collectionCode>
19114); station M27, Santiago, Maribojoc Bay, Panglao, Bohol, 
<collectingCountry id="F3377613FFCDFFAEEC2CFC5998F5FB83" box="[537,672,1021,1047]" name="Philippines" pageId="10" pageNumber="49">Philippines</collectingCountry>
, 
<geoCoordinate id="EE145044FFCDFFAEEC99FC599976FB83" box="[684,803,1021,1047]" direction="north" orientation="latitude" pageId="10" pageNumber="49" precision="1" value="9.000716">9°0.043'N</geoCoordinate>
<geoCoordinate id="EE145044FFCDFFAEED1FFC5999EBFB83" box="[810,958,1021,1047]" direction="east" orientation="longitude" pageId="10" pageNumber="49" precision="1" value="123.00085">123°0.051'E</geoCoordinate>
, coll. 
<date id="FF9E1043FFCDFFAEEA30FC599ECCFB83" box="[1029,1177,1021,1047]" pageId="10" pageNumber="49" value="2004-06-23">23 Jun 2004</date>
. 
<typeStatus id="549B8821FFCDFFAEEA91FC599F4FFB83" box="[1188,1306,1021,1047]" pageId="10" pageNumber="49" type="paratype">Paratypes</typeStatus>
: 
<specimenCount id="9D26FD0AFFCDFFAEEB13FC599F20FB83" box="[1318,1397,1021,1047]" pageId="10" pageNumber="49" type="male">1 male</specimenCount>
(cl 
<quantity id="4CD89B66FFCDFFAEEEA2FB819B51FBAB" box="[151,260,1061,1087]" metricMagnitude="-2" metricUnit="m" metricValue="1.9899999999999998" pageId="10" pageNumber="49" unit="mm" value="19.9">19.9 mm</quantity>
, cw 22.0 mm) (
<collectionCode id="ED31AE46FFCDFFAEEFF5FB819BAAFBAB" box="[448,511,1061,1087]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="10" pageNumber="49">ZRC</collectionCode>
), station P1, 90– 
<quantity id="4CD89B66FFCDFFAEECFDFB819958FBAB" box="[712,781,1061,1087]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="49" unit="m" value="200.0">200m</quantity>
, Maribojoc Bay, Panglao, Bohol, 
<collectingCountry id="F3377613FFCDFFAEEA95FB819F72FBAB" box="[1184,1319,1061,1087]" name="Philippines" pageId="10" pageNumber="49">Philippines</collectingCountry>
, 
<geoCoordinate id="EE145044FFCDFFAEEB01FB819FCEFBAB" box="[1332,1435,1061,1087]" direction="north" orientation="latitude" pageId="10" pageNumber="49" precision="92" value="9.601666">9°36.1'N</geoCoordinate>
<geoCoordinate id="EE145044FFCDFFAEEEA2FBE99B4EFBF3" box="[151,283,1101,1127]" direction="east" orientation="longitude" pageId="10" pageNumber="49" precision="92" value="123.75">123°45.0'E</geoCoordinate>
, coll. fishermen with tangle nets, 
<date id="FF9E1043FFCDFFAEEC96FBE99960FBF3" box="[675,821,1101,1127]" pageId="10" pageNumber="49" value="2004-06-27">27 Jun 2004</date>
; 
<specimenCount id="9D26FD0AFFCDFFAEED75FBE999DAFBF3" box="[832,911,1101,1127]" pageId="10" pageNumber="49" type="male">1 male</specimenCount>
(cl 
<quantity id="4CD89B66FFCDFFAEED8EFBE99E73FBF3" box="[955,1062,1101,1127]" metricMagnitude="-2" metricUnit="m" metricValue="2.73" pageId="10" pageNumber="49" unit="mm" value="27.3">27.3 mm</quantity>
, cw 
<quantity id="4CD89B66FFCDFFAEEA6FFBE99E91FBF3" box="[1114,1220,1101,1127]" metricMagnitude="-2" metricUnit="m" metricValue="2.53" pageId="10" pageNumber="49" unit="mm" value="25.3">25.3 mm</quantity>
) (
<collectionCode id="ED31AE46FFCDFFAEEAEFFBE99F4DFBF3" box="[1242,1304,1101,1127]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="10" pageNumber="49">ZRC</collectionCode>
, ex 
<collectionCode id="ED31AE46FFCDFFAEEB73FBE99FC8FBF3" box="[1350,1437,1101,1127]" pageId="10" pageNumber="49">NMCR</collectionCode>
19115), Maribojoc Bay, Panglao, Bohol, 
<collectingCountry id="F3377613FFCDFFAEEC4FFBD19954FB1B" box="[634,769,1141,1167]" name="Philippines" pageId="10" pageNumber="49">Philippines</collectingCountry>
, in tangle nets, ca. 
<quantity id="4CD89B66FFCDFFAEEDD7FBD19E72FB1B" box="[994,1063,1141,1167]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="49" unit="m" value="200.0">200m</quantity>
, coll. J. Arbasto, 
<date id="FF9E1043FFCDFFAEEAC0FBD19F34FB1B" box="[1269,1377,1141,1167]" pageId="10" pageNumber="49" value="2004-10">Oct 2004</date>
– 
<date id="FF9E1043FFCDFFAEEB45FBD19A80FB23" pageId="10" pageNumber="49" value="2005-02">Feb 2005</date>
; 
<specimenCount id="9D26FD0AFFCDFFAEEED7FB399B67FB23" box="[226,306,1181,1207]" pageId="10" pageNumber="49" type="male">1 male</specimenCount>
(cl 
<quantity id="4CD89B66FFCDFFAEEF56FB399B85FB23" box="[355,464,1181,1207]" metricMagnitude="-2" metricUnit="m" metricValue="2.7399999999999998" pageId="10" pageNumber="49" unit="mm" value="27.4">27.4 mm</quantity>
, cw 29.0 mm) (
<collectionCode id="ED31AE46FFCDFFAEECB9FB3998B9FB23" LSID="urn:lsid:biocol.org:col:34988" box="[652,748,1181,1207]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="10" pageNumber="49">MNHN</collectionCode>
), Maribojoc Bay, Panglao, Bohol, Visayas, 
<collectingCountry id="F3377613FFCDFFAEEAC7FB399F2CFB23" box="[1266,1401,1181,1207]" name="Philippines" pageId="10" pageNumber="49">Philippines</collectingCountry>
, in tangle nets, 
<quantity id="4CD89B66FFCDFFAEEF12FB619BCDFB4B" box="[295,408,1221,1247]" metricMagnitude="2" metricUnit="m" metricValue="1.1" metricValueMax="1.4" metricValueMin="0.8" pageId="10" pageNumber="49" unit="m" value="110.0" valueMax="140.0" valueMin="80.0">80–140m</quantity>
, coll. J. Arbasto, between 
<date id="FF9E1043FFCDFFAEECE3FB6199DEFB4B" box="[726,907,1221,1247]" pageId="10" pageNumber="49" value="2004" valueMax="2005">2004 and 2005</date>
. Non-type: male (cl 
<quantity id="4CD89B66FFCDFFAEEAB7FB619EA5FB4B" box="[1154,1264,1221,1247]" metricMagnitude="-2" metricUnit="m" metricValue="1.56" pageId="10" pageNumber="49" unit="mm" value="15.6">15.6 mm</quantity>
, cw 
<quantity id="4CD89B66FFCDFFAEEB1DFB619FC1FB4B" box="[1320,1428,1221,1247]" metricMagnitude="-2" metricUnit="m" metricValue="1.68" pageId="10" pageNumber="49" unit="mm" value="16.8">16.8 mm</quantity>
) (
<collectionCode id="ED31AE46FFCDFFAEEEAAFB4B9A8BFA9D" box="[159,222,1263,1289]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="10" pageNumber="49">ZRC</collectionCode>
), station DB16, sandy slope with coral patches, Tutuba off Santo, 
<collectingCountry id="F3377613FFCDFFAEEA24FB4B9E21FA9D" box="[1041,1140,1263,1289]" name="Vanuatu" pageId="10" pageNumber="49">Vanuatu</collectingCountry>
, in tangle nets, ₁₅․₃₆ 
<emphasis id="B954EA91FFCDFFAEEB49FB489FDEFA9D" bold="true" box="[1404,1419,1260,1289]" italics="true" pageId="10" pageNumber="49">°</emphasis>
S 
<geoCoordinate id="EE145044FFCDFFAEEEA2FABE9B50FAA0" box="[151,261,1306,1332]" direction="east" orientation="longitude" pageId="10" pageNumber="49" precision="555" value="167.16">167.16°E</geoCoordinate>
, 32̄ 
<quantity id="4CD89B66FFCDFFAEEF0EFABE9BD9FAA0" box="[315,396,1306,1332]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="10" pageNumber="49" unit="mm" value="40.0">40 mm</quantity>
depth, coll. J. Arbasto, 
<date id="FF9E1043FFCDFFAEEC96FABE996CFAA0" box="[675,825,1306,1332]" pageId="10" pageNumber="49" value="2006-09-14">14 Sep 2006</date>
.
</paragraph>
</subSubSection>
<subSubSection id="C33A6508FFCDFFA8EEF3FAE69B24FBE6" lastPageId="12" lastPageNumber="51" pageId="10" pageNumber="49" type="description">
<paragraph id="8B9F3683FFCDFFAFEEF3FAE698E5FE36" blockId="10.[151,1437,981,2012]" lastBlockId="11.[151,1437,152,2018]" lastPageId="11" lastPageNumber="50" pageId="10" pageNumber="49">
<emphasis id="B954EA91FFCDFFAEEEF3FAE69865FAC8" bold="true" box="[198,560,1346,1372]" pageId="10" pageNumber="49">
Description of male 
<typeStatus id="549B8821FFCDFFAEEFF0FAE69865FAC8" box="[453,560,1346,1372]" pageId="10" pageNumber="49" type="holotype">holotype</typeStatus>
</emphasis>
. Carapace subquadrate, squarish to slightly broader than long; dorsal surface relatively flattened with low convex surfaces; regions well defined, separated by deep, prominent grooves, anterior part more convex than posterior part, marked by prominent areolae, covered with numerous, densely packed very short, stiff, black setae, velvet-like, partially obscuring surface but not completely obscuring margins; 1F divided into 2 cone-shaped lobes; 2F complete, surface uneven; 1M complete; 2M with median depression; 3M not continuous; 4M laterally long, distinct; 1P with small areolae; 2P long, narrow; 2L triangular in shape; 3L elongated, triangular in shape; 6L clearly demarcated, with tubercles of varying sizes; 3R with small areolae (
<figureCitation id="131B2A06FFCDFFAEEF98F9FE9855F9E0" box="[429,512,1626,1652]" captionStart="FIGURE 1" captionStartId="17.[151,255,1912,1936]" captionTargetBox="[226,1360,194,1887]" captionTargetId="figure@17.[226,1360,194,1888]" captionTargetPageId="17" captionText="FIGURE 1. Vultocinus anfractus, new genus, new species. Fresh colours. A, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC), Philippines, freshly dead, poised on same kind of wood substrate it was reportedly found on; B, paratype male (cl 19.9 mm, cw 22.0 mm) (ZRC), Philippines; C, male (cl 15.6 mm, cw 16.8 mm) (ZRC), Vanuatu." httpUri="https://zenodo.org/record/178297/files/figure.png" pageId="10" pageNumber="49">Figs. 1</figureCitation>
, 
<figureCitation id="131B2A06FFCDFFAEEC25F9FE9874F9E0" box="[528,545,1626,1652]" captionStart="FIGURE 2" captionStartId="18.[151,255,1936,1960]" captionTargetBox="[264,1322,182,1924]" captionTargetId="figure@18.[264,1323,182,1926]" captionTargetPageId="18" captionText="FIGURE 2. Vultocinus anfractus, new genus, new species. Overall dorsal views. A, holotype male (cl 19.5 mm, cw 20.3 mm) (NMCR 19114), Philippines; B, paratype male (cl 19.9 mm, cw 22.0 mm) (ZRC), Philippines; C, male (cl 15.6 mm, cw 16.8 mm) (ZRC), Vanuatu." httpUri="https://zenodo.org/record/178298/files/figure.png" pageId="10" pageNumber="49">2</figureCitation>
, 
<figureCitation id="131B2A06FFCDFFAEEC1AF9FE9868F9E0" box="[559,573,1626,1652]" captionStart="FIGURE 3" captionStartId="19.[151,255,1088,1112]" captionTargetBox="[151,1435,194,1064]" captionTargetId="figure@19.[151,1436,194,1064]" captionTargetPageId="19" captionText="FIGURE 3. Vultocinus anfractus, new genus, new species. Holotype male (cl 19.5 mm, cw 20.3 mm) (NMCR 19114), Philippines. A, dorsal view of carapace; B, ventral view of carapace; C, buccal area showing third maxillipeds; D, frontal view of carapace and chelipeds." httpUri="https://zenodo.org/record/178299/files/figure.png" pageId="10" pageNumber="49">3</figureCitation>
A, D). Frontal region almost one-third width of carapace, with a distinct notch formed by 2 conical extensions (
<figureCitation id="131B2A06FFCDFFAEEC6FF92698F8F908" box="[602,685,1666,1692]" captionStart="FIGURE 2" captionStartId="18.[151,255,1936,1960]" captionTargetBox="[264,1322,182,1924]" captionTargetId="figure@18.[264,1323,182,1926]" captionTargetPageId="18" captionText="FIGURE 2. Vultocinus anfractus, new genus, new species. Overall dorsal views. A, holotype male (cl 19.5 mm, cw 20.3 mm) (NMCR 19114), Philippines; B, paratype male (cl 19.9 mm, cw 22.0 mm) (ZRC), Philippines; C, male (cl 15.6 mm, cw 16.8 mm) (ZRC), Vanuatu." httpUri="https://zenodo.org/record/178298/files/figure.png" pageId="10" pageNumber="49">Figs. 2</figureCitation>
, 
<figureCitation id="131B2A06FFCDFFAEEC8CF9269892F908" box="[697,711,1666,1692]" captionStart="FIGURE 3" captionStartId="19.[151,255,1088,1112]" captionTargetBox="[151,1435,194,1064]" captionTargetId="figure@19.[151,1436,194,1064]" captionTargetPageId="19" captionText="FIGURE 3. Vultocinus anfractus, new genus, new species. Holotype male (cl 19.5 mm, cw 20.3 mm) (NMCR 19114), Philippines. A, dorsal view of carapace; B, ventral view of carapace; C, buccal area showing third maxillipeds; D, frontal view of carapace and chelipeds." httpUri="https://zenodo.org/record/178299/files/figure.png" pageId="10" pageNumber="49">3</figureCitation>
A, D). Orbits rounded, eyes fitting tightly into cavity; margins areolate with small flat granules on rim, separated into 3 parts by deep sulci (
<figureCitation id="131B2A06FFCDFFAEEA2EF90E9E0AF950" box="[1051,1119,1706,1732]" captionStart="FIGURE 3" captionStartId="19.[151,255,1088,1112]" captionTargetBox="[151,1435,194,1064]" captionTargetId="figure@19.[151,1436,194,1064]" captionTargetPageId="19" captionText="FIGURE 3. Vultocinus anfractus, new genus, new species. Holotype male (cl 19.5 mm, cw 20.3 mm) (NMCR 19114), Philippines. A, dorsal view of carapace; B, ventral view of carapace; C, buccal area showing third maxillipeds; D, frontal view of carapace and chelipeds." httpUri="https://zenodo.org/record/178299/files/figure.png" pageId="10" pageNumber="49">Fig. 3</figureCitation>
A, D); supra-orbital border tuberculated, each tubercle with smaller granules on surface, with prominent inner orbital tooth and lower external orbital tooth, with 2 smaller teeth between which are separated by a deep narrow fissure, one tooth separated from inner orbital tooth by a cleft, the other separated from outer orbital tooth by a deep narrow fissure (
<figureCitation id="131B2A06FFCDFFAEEEE2F8EE9B49F8F0" box="[215,284,1866,1892]" captionStart="FIGURE 3" captionStartId="19.[151,255,1088,1112]" captionTargetBox="[151,1435,194,1064]" captionTargetId="figure@19.[151,1436,194,1064]" captionTargetPageId="19" captionText="FIGURE 3. Vultocinus anfractus, new genus, new species. Holotype male (cl 19.5 mm, cw 20.3 mm) (NMCR 19114), Philippines. A, dorsal view of carapace; B, ventral view of carapace; C, buccal area showing third maxillipeds; D, frontal view of carapace and chelipeds." httpUri="https://zenodo.org/record/178299/files/figure.png" pageId="10" pageNumber="49">Fig. 3</figureCitation>
A, D); infra-orbital border more or less elevated, with 3 deep notches (
<figureCitation id="131B2A06FFCDFFAEEA61F8EE9ECDF8F0" box="[1108,1176,1866,1892]" captionStart="FIGURE 3" captionStartId="19.[151,255,1088,1112]" captionTargetBox="[151,1435,194,1064]" captionTargetId="figure@19.[151,1436,194,1064]" captionTargetPageId="19" captionText="FIGURE 3. Vultocinus anfractus, new genus, new species. Holotype male (cl 19.5 mm, cw 20.3 mm) (NMCR 19114), Philippines. A, dorsal view of carapace; B, ventral view of carapace; C, buccal area showing third maxillipeds; D, frontal view of carapace and chelipeds." httpUri="https://zenodo.org/record/178299/files/figure.png" pageId="10" pageNumber="49">Fig. 3</figureCitation>
A, D). Anterior dorsal and distal parts of eyestalks fringed with long thick and thin setae. Antennal basal segment almost reaching small ventral prolongation of orbital hiatus; flagellum slender, long, about twice length of diameter of orbit (
<figureCitation id="131B2A06FFCDFFAEEEAAF8669ABFF848" box="[159,234,1986,2012]" captionStart="FIGURE 3" captionStartId="19.[151,255,1088,1112]" captionTargetBox="[151,1435,194,1064]" captionTargetId="figure@19.[151,1436,194,1064]" captionTargetPageId="19" captionText="FIGURE 3. Vultocinus anfractus, new genus, new species. Holotype male (cl 19.5 mm, cw 20.3 mm) (NMCR 19114), Philippines. A, dorsal view of carapace; B, ventral view of carapace; C, buccal area showing third maxillipeds; D, frontal view of carapace and chelipeds." httpUri="https://zenodo.org/record/178299/files/figure.png" pageId="10" pageNumber="49">Fig. 3</figureCitation>
D). Antennules folding obliquely (
<figureCitation id="131B2A06FFCDFFAEECA7F866988EF848" box="[658,731,1986,2012]" captionStart="FIGURE 3" captionStartId="19.[151,255,1088,1112]" captionTargetBox="[151,1435,194,1064]" captionTargetId="figure@19.[151,1436,194,1064]" captionTargetPageId="19" captionText="FIGURE 3. Vultocinus anfractus, new genus, new species. Holotype male (cl 19.5 mm, cw 20.3 mm) (NMCR 19114), Philippines. A, dorsal view of carapace; B, ventral view of carapace; C, buccal area showing third maxillipeds; D, frontal view of carapace and chelipeds." httpUri="https://zenodo.org/record/178299/files/figure.png" pageId="10" pageNumber="49">Fig. 3</figureCitation>
D). Suborbital and pterygostomial surfaces tomentose but smooth underneath; subhepatic region tomentose with small scattered granules. Anterolateral margin with 4 teeth (excluding external orbital tooth), first tooth smallest, areolate, with 3 small flat granules on rim; second tooth bigger, areolate, ringed with small flat granules; third tooth longest, more conical, with 1 flat granule, directed anteriorly; fourth tooth low, rounded, ringed by small flat granules (
<figureCitation id="131B2A06FFCCFFAFEA03FEB49EDFFEBE" box="[1078,1162,272,298]" captionStart="FIGURE 1" captionStartId="17.[151,255,1912,1936]" captionTargetBox="[226,1360,194,1887]" captionTargetId="figure@17.[226,1360,194,1888]" captionTargetPageId="17" captionText="FIGURE 1. Vultocinus anfractus, new genus, new species. Fresh colours. A, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC), Philippines, freshly dead, poised on same kind of wood substrate it was reportedly found on; B, paratype male (cl 19.9 mm, cw 22.0 mm) (ZRC), Philippines; C, male (cl 15.6 mm, cw 16.8 mm) (ZRC), Vanuatu." httpUri="https://zenodo.org/record/178297/files/figure.png" pageId="11" pageNumber="50">Figs. 1</figureCitation>
, 
<figureCitation id="131B2A06FFCCFFAFEAAEFEB49EF9FEBE" box="[1179,1196,272,298]" captionStart="FIGURE 2" captionStartId="18.[151,255,1936,1960]" captionTargetBox="[264,1322,182,1924]" captionTargetId="figure@18.[264,1323,182,1926]" captionTargetPageId="18" captionText="FIGURE 2. Vultocinus anfractus, new genus, new species. Overall dorsal views. A, holotype male (cl 19.5 mm, cw 20.3 mm) (NMCR 19114), Philippines; B, paratype male (cl 19.9 mm, cw 22.0 mm) (ZRC), Philippines; C, male (cl 15.6 mm, cw 16.8 mm) (ZRC), Vanuatu." httpUri="https://zenodo.org/record/178298/files/figure.png" pageId="11" pageNumber="50">2</figureCitation>
, 
<figureCitation id="131B2A06FFCCFFAFEA89FEB49E9FFEBE" box="[1212,1226,272,298]" captionStart="FIGURE 3" captionStartId="19.[151,255,1088,1112]" captionTargetBox="[151,1435,194,1064]" captionTargetId="figure@19.[151,1436,194,1064]" captionTargetPageId="19" captionText="FIGURE 3. Vultocinus anfractus, new genus, new species. Holotype male (cl 19.5 mm, cw 20.3 mm) (NMCR 19114), Philippines. A, dorsal view of carapace; B, ventral view of carapace; C, buccal area showing third maxillipeds; D, frontal view of carapace and chelipeds." httpUri="https://zenodo.org/record/178299/files/figure.png" pageId="11" pageNumber="50">3</figureCitation>
A). Posterolateral margin almost straight, gently converging towards posterior carapace margin, lined with rather 5 large rounded granules that form 3 granulated crests (
<figureCitation id="131B2A06FFCCFFAFECD4FEC49963FEEE" box="[737,822,352,378]" captionStart="FIGURE 1" captionStartId="17.[151,255,1912,1936]" captionTargetBox="[226,1360,194,1887]" captionTargetId="figure@17.[226,1360,194,1888]" captionTargetPageId="17" captionText="FIGURE 1. Vultocinus anfractus, new genus, new species. Fresh colours. A, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC), Philippines, freshly dead, poised on same kind of wood substrate it was reportedly found on; B, paratype male (cl 19.9 mm, cw 22.0 mm) (ZRC), Philippines; C, male (cl 15.6 mm, cw 16.8 mm) (ZRC), Vanuatu." httpUri="https://zenodo.org/record/178297/files/figure.png" pageId="11" pageNumber="50">Figs. 1</figureCitation>
, 
<figureCitation id="131B2A06FFCCFFAFED7DFEC4990CFEEE" box="[840,857,352,378]" captionStart="FIGURE 2" captionStartId="18.[151,255,1936,1960]" captionTargetBox="[264,1322,182,1924]" captionTargetId="figure@18.[264,1323,182,1926]" captionTargetPageId="18" captionText="FIGURE 2. Vultocinus anfractus, new genus, new species. Overall dorsal views. A, holotype male (cl 19.5 mm, cw 20.3 mm) (NMCR 19114), Philippines; B, paratype male (cl 19.9 mm, cw 22.0 mm) (ZRC), Philippines; C, male (cl 15.6 mm, cw 16.8 mm) (ZRC), Vanuatu." httpUri="https://zenodo.org/record/178298/files/figure.png" pageId="11" pageNumber="50">2</figureCitation>
, 
<figureCitation id="131B2A06FFCCFFAFED5CFEC49922FEEE" box="[873,887,352,378]" captionStart="FIGURE 3" captionStartId="19.[151,255,1088,1112]" captionTargetBox="[151,1435,194,1064]" captionTargetId="figure@19.[151,1436,194,1064]" captionTargetPageId="19" captionText="FIGURE 3. Vultocinus anfractus, new genus, new species. Holotype male (cl 19.5 mm, cw 20.3 mm) (NMCR 19114), Philippines. A, dorsal view of carapace; B, ventral view of carapace; C, buccal area showing third maxillipeds; D, frontal view of carapace and chelipeds." httpUri="https://zenodo.org/record/178299/files/figure.png" pageId="11" pageNumber="50">3</figureCitation>
A). Posterior carapace margin gently convex, median part slightly concave (
<figureCitation id="131B2A06FFCCFFAFEFCDFE2C981DFE36" box="[504,584,392,418]" captionStart="FIGURE 3" captionStartId="19.[151,255,1088,1112]" captionTargetBox="[151,1435,194,1064]" captionTargetId="figure@19.[151,1436,194,1064]" captionTargetPageId="19" captionText="FIGURE 3. Vultocinus anfractus, new genus, new species. Holotype male (cl 19.5 mm, cw 20.3 mm) (NMCR 19114), Philippines. A, dorsal view of carapace; B, ventral view of carapace; C, buccal area showing third maxillipeds; D, frontal view of carapace and chelipeds." httpUri="https://zenodo.org/record/178299/files/figure.png" pageId="11" pageNumber="50">Figs. 3</figureCitation>
A, 11A).
</paragraph>
<paragraph id="8B9F3683FFCCFFAFEEF3FE149E69FC9E" blockId="11.[151,1437,152,2018]" pageId="11" pageNumber="50">
Third maxilliped almost completely covered by very short tomentum; merus quadrate, antero-external angle expanded, 3 flat granules lining mid-anteroexternal angle towards upper margin of merus; merus medially concave with a deep sulcus near lateral portion, a shallower sulcus near inner margin; ischium subquadrangular, with a deep median sulcus; exognath straight, inner subdistal angle with prominent triangular tooth, with distinct flagellum that extends beyond width of merus (
<figureCitation id="131B2A06FFCCFFAFED60FDF499F3FDFE" box="[853,934,592,618]" captionStart="FIGURE 3" captionStartId="19.[151,255,1088,1112]" captionTargetBox="[151,1435,194,1064]" captionTargetId="figure@19.[151,1436,194,1064]" captionTargetPageId="19" captionText="FIGURE 3. Vultocinus anfractus, new genus, new species. Holotype male (cl 19.5 mm, cw 20.3 mm) (NMCR 19114), Philippines. A, dorsal view of carapace; B, ventral view of carapace; C, buccal area showing third maxillipeds; D, frontal view of carapace and chelipeds." httpUri="https://zenodo.org/record/178299/files/figure.png" pageId="11" pageNumber="50">Figs. 3</figureCitation>
B, C, 5A, 8A); ischium and basis separated by prominent suture (
<figureCitation id="131B2A06FFCCFFAFEFA6FDDC9B8CFD06" box="[403,473,632,658]" captionStart="FIGURE 5" captionStartId="21.[151,255,1379,1403]" captionTargetBox="[158,1421,201,1348]" captionTargetId="figure@21.[151,1436,194,1355]" captionTargetPageId="21" captionText="FIGURE 5. Vultocinus anfractus, new genus, new species. Paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC), Philippines. A, left third maxilliped (setae denuded); B, male abdomen (partially denuded); C, left G 1, dorsal view; D, left G 1, ventral view; E, distal part of left G 1, dorsal view; F, distal part of left G 1, ventral view; G, left G 2; H, distal part of left G 2. Scales = 1.0 mm." httpUri="https://zenodo.org/record/178301/files/figure.png" pageId="11" pageNumber="50">Fig. 5</figureCitation>
A). Epistome appears sunken; posterior margin divided into 4 lobes, median lobes rectangular, separated by deep narrow fissure, lateral lobes separated from median lobes by uneven cleft (
<figureCitation id="131B2A06FFCCFFAFEB59FD049AF3FD76" captionStart="FIGURE 3" captionStartId="19.[151,255,1088,1112]" captionTargetBox="[151,1435,194,1064]" captionTargetId="figure@19.[151,1436,194,1064]" captionTargetPageId="19" captionText="FIGURE 3. Vultocinus anfractus, new genus, new species. Holotype male (cl 19.5 mm, cw 20.3 mm) (NMCR 19114), Philippines. A, dorsal view of carapace; B, ventral view of carapace; C, buccal area showing third maxillipeds; D, frontal view of carapace and chelipeds." httpUri="https://zenodo.org/record/178299/files/figure.png" pageId="11" pageNumber="50">Fig. 3</figureCitation>
B, D). Endostome with long, prominent ridge running from distal edge of median lobe of posterior margin of epistome to near mandibles, interrupted subdistally, just as epistomal lobe ends.
</paragraph>
<paragraph id="8B9F3683FFCCFFAFEEF3FCBC9EB4FBE6" blockId="11.[151,1437,152,2018]" pageId="11" pageNumber="50">
Both chelipeds large, subequal, laterally flattened, almost entirely covered with dense heavy tomentum (
<figureCitation id="131B2A06FFCCFFAFEEAAFCE49AA1FCCE" box="[159,244,832,858]" captionStart="FIGURE 2" captionStartId="18.[151,255,1936,1960]" captionTargetBox="[264,1322,182,1924]" captionTargetId="figure@18.[264,1323,182,1926]" captionTargetPageId="18" captionText="FIGURE 2. Vultocinus anfractus, new genus, new species. Overall dorsal views. A, holotype male (cl 19.5 mm, cw 20.3 mm) (NMCR 19114), Philippines; B, paratype male (cl 19.9 mm, cw 22.0 mm) (ZRC), Philippines; C, male (cl 15.6 mm, cw 16.8 mm) (ZRC), Vanuatu." httpUri="https://zenodo.org/record/178298/files/figure.png" pageId="11" pageNumber="50">Figs. 2</figureCitation>
, 
<figureCitation id="131B2A06FFCCFFAFEF37FCE49B45FCCE" box="[258,272,832,858]" captionStart="FIGURE 3" captionStartId="19.[151,255,1088,1112]" captionTargetBox="[151,1435,194,1064]" captionTargetId="figure@19.[151,1436,194,1064]" captionTargetPageId="19" captionText="FIGURE 3. Vultocinus anfractus, new genus, new species. Holotype male (cl 19.5 mm, cw 20.3 mm) (NMCR 19114), Philippines. A, dorsal view of carapace; B, ventral view of carapace; C, buccal area showing third maxillipeds; D, frontal view of carapace and chelipeds." httpUri="https://zenodo.org/record/178299/files/figure.png" pageId="11" pageNumber="50">3</figureCitation>
D, 4A, B). Merus of cheliped with 2 large granules on proximal portion, 2 spinules on distal part, forming 2 granulated crests; carpus with 2 large tubercles, several nodules, granules on outer dorsal surface. Ischio-merus fused but suture still discernible. Chela somewhat flattened laterally; outer surface covered with numerous granules and tubercles of various sizes, with one median longitudinal ridge on outer surface evident, inner subventral surface with a curved row of sharp, orange-tipped tubercles; fingers shorter than palm; propodal finger with 3 nodules on inner margin, immovable finger with lower margin covered with dense thick tomentum, elevated, forming stiff fringe; dactylus with 2 deep sulci filled with short thick tomentum; dark brown pigmentation present only on distal part of fingers and cutting edges (
<figureCitation id="131B2A06FFCCFFAFEA66FBFC9ECCFBE6" box="[1107,1177,1112,1138]" captionStart="FIGURE 4" captionStartId="20.[151,255,1144,1168]" captionTargetBox="[151,1435,184,1120]" captionTargetId="figure@20.[151,1436,184,1120]" captionTargetPageId="20" captionText="FIGURE 4. Vultocinus anfractus, new genus, new species. Holotype male (cl 19.5 mm, cw 20.3 mm) (NMCR 19114), Philippines. A, outer surface of right chela; B, outer surface of left chela; C, right side of thoracic sternum showing ventral surface of chelipeds and spines on ambulatory meri; D, right P 4." httpUri="https://zenodo.org/record/178300/files/figure.png" pageId="11" pageNumber="50">Fig. 4</figureCitation>
A, B).
</paragraph>
<paragraph id="8B9F3683FFCCFFAFEEF3FB2498A1F9C6" blockId="11.[151,1437,152,2018]" pageId="11" pageNumber="50">
Ambulatory legs relatively long, relatively stout, thickly fringed by dense, stiff setae; P3 longest, P5 shortest, distinctly upturned (
<figureCitation id="131B2A06FFCCFFAFEF87FB0C9851FB56" box="[434,516,1192,1218]" captionStart="FIGURE 1" captionStartId="17.[151,255,1912,1936]" captionTargetBox="[226,1360,194,1887]" captionTargetId="figure@17.[226,1360,194,1888]" captionTargetPageId="17" captionText="FIGURE 1. Vultocinus anfractus, new genus, new species. Fresh colours. A, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC), Philippines, freshly dead, poised on same kind of wood substrate it was reportedly found on; B, paratype male (cl 19.9 mm, cw 22.0 mm) (ZRC), Philippines; C, male (cl 15.6 mm, cw 16.8 mm) (ZRC), Vanuatu." httpUri="https://zenodo.org/record/178297/files/figure.png" pageId="11" pageNumber="50">Figs. 1</figureCitation>
, 
<figureCitation id="131B2A06FFCCFFAFEC27FB0C9876FB56" box="[530,547,1192,1218]" captionStart="FIGURE 2" captionStartId="18.[151,255,1936,1960]" captionTargetBox="[264,1322,182,1924]" captionTargetId="figure@18.[264,1323,182,1926]" captionTargetPageId="18" captionText="FIGURE 2. Vultocinus anfractus, new genus, new species. Overall dorsal views. A, holotype male (cl 19.5 mm, cw 20.3 mm) (NMCR 19114), Philippines; B, paratype male (cl 19.9 mm, cw 22.0 mm) (ZRC), Philippines; C, male (cl 15.6 mm, cw 16.8 mm) (ZRC), Vanuatu." httpUri="https://zenodo.org/record/178298/files/figure.png" pageId="11" pageNumber="50">2</figureCitation>
). Merus of P2–P4 with 2 longitudinal ridges (anterior one larger), distinctly meeting near distal segment on outer surface, margins fringed by dense stiff thick setae, margins somewhat carinate, dorso-subdistal area with deep cleft that forms prominent subdistal tooth with rounded tip and larger sharp sublamellate distal tooth which joins distinctly carinate proximal margin, area appears cup-like; merus of P5 with 1 longitudinal bony ridge, cleft, 2 teeth, cup-like part on dorso-subdistal area less prominent (
<figureCitation id="131B2A06FFCCFFAFEB54FAEC9AFDFA1E" captionStart="FIGURE 2" captionStartId="18.[151,255,1936,1960]" captionTargetBox="[264,1322,182,1924]" captionTargetId="figure@18.[264,1323,182,1926]" captionTargetPageId="18" captionText="FIGURE 2. Vultocinus anfractus, new genus, new species. Overall dorsal views. A, holotype male (cl 19.5 mm, cw 20.3 mm) (NMCR 19114), Philippines; B, paratype male (cl 19.9 mm, cw 22.0 mm) (ZRC), Philippines; C, male (cl 15.6 mm, cw 16.8 mm) (ZRC), Vanuatu." httpUri="https://zenodo.org/record/178298/files/figure.png" pageId="11" pageNumber="50">Figs. 2</figureCitation>
, 
<figureCitation id="131B2A06FFCCFFAFEE80FAD49A96FA1E" box="[181,195,1392,1418]" captionStart="FIGURE 3" captionStartId="19.[151,255,1088,1112]" captionTargetBox="[151,1435,194,1064]" captionTargetId="figure@19.[151,1436,194,1064]" captionTargetPageId="19" captionText="FIGURE 3. Vultocinus anfractus, new genus, new species. Holotype male (cl 19.5 mm, cw 20.3 mm) (NMCR 19114), Philippines. A, dorsal view of carapace; B, ventral view of carapace; C, buccal area showing third maxillipeds; D, frontal view of carapace and chelipeds." httpUri="https://zenodo.org/record/178299/files/figure.png" pageId="11" pageNumber="50">3</figureCitation>
A, 4C, D); ventral surface of merus of P2–P4 with 2 or 3 long perpendicular spines on posterior margin, scattered granules on anterior margin; ventral surface of merus of P5 without spines, only granules on posterior margin, scattered much smaller granules on anterior margin (
<figureCitation id="131B2A06FFCCFFAFEDACFA6499B8FA4E" box="[921,1005,1472,1498]" captionStart="FIGURE 2" captionStartId="18.[151,255,1936,1960]" captionTargetBox="[264,1322,182,1924]" captionTargetId="figure@18.[264,1323,182,1926]" captionTargetPageId="18" captionText="FIGURE 2. Vultocinus anfractus, new genus, new species. Overall dorsal views. A, holotype male (cl 19.5 mm, cw 20.3 mm) (NMCR 19114), Philippines; B, paratype male (cl 19.9 mm, cw 22.0 mm) (ZRC), Philippines; C, male (cl 15.6 mm, cw 16.8 mm) (ZRC), Vanuatu." httpUri="https://zenodo.org/record/178298/files/figure.png" pageId="11" pageNumber="50">Figs. 2</figureCitation>
, 
<figureCitation id="131B2A06FFCCFFAFEDCCFA649E52FA4E" box="[1017,1031,1472,1498]" captionStart="FIGURE 3" captionStartId="19.[151,255,1088,1112]" captionTargetBox="[151,1435,194,1064]" captionTargetId="figure@19.[151,1436,194,1064]" captionTargetPageId="19" captionText="FIGURE 3. Vultocinus anfractus, new genus, new species. Holotype male (cl 19.5 mm, cw 20.3 mm) (NMCR 19114), Philippines. A, dorsal view of carapace; B, ventral view of carapace; C, buccal area showing third maxillipeds; D, frontal view of carapace and chelipeds." httpUri="https://zenodo.org/record/178299/files/figure.png" pageId="11" pageNumber="50">3</figureCitation>
A, 4C, D). Carpus with prominent median ridge lined with several flat granules (
<figureCitation id="131B2A06FFCCFFAFECFBFA4C9970F996" box="[718,805,1512,1538]" captionStart="FIGURE 2" captionStartId="18.[151,255,1936,1960]" captionTargetBox="[264,1322,182,1924]" captionTargetId="figure@18.[264,1323,182,1926]" captionTargetPageId="18" captionText="FIGURE 2. Vultocinus anfractus, new genus, new species. Overall dorsal views. A, holotype male (cl 19.5 mm, cw 20.3 mm) (NMCR 19114), Philippines; B, paratype male (cl 19.9 mm, cw 22.0 mm) (ZRC), Philippines; C, male (cl 15.6 mm, cw 16.8 mm) (ZRC), Vanuatu." httpUri="https://zenodo.org/record/178298/files/figure.png" pageId="11" pageNumber="50">Figs. 2</figureCitation>
, 
<figureCitation id="131B2A06FFCCFFAFED03FA4C9911F996" box="[822,836,1512,1538]" captionStart="FIGURE 3" captionStartId="19.[151,255,1088,1112]" captionTargetBox="[151,1435,194,1064]" captionTargetId="figure@19.[151,1436,194,1064]" captionTargetPageId="19" captionText="FIGURE 3. Vultocinus anfractus, new genus, new species. Holotype male (cl 19.5 mm, cw 20.3 mm) (NMCR 19114), Philippines. A, dorsal view of carapace; B, ventral view of carapace; C, buccal area showing third maxillipeds; D, frontal view of carapace and chelipeds." httpUri="https://zenodo.org/record/178299/files/figure.png" pageId="11" pageNumber="50">3</figureCitation>
A, 4C, D). Propodus of P2–P5 with 2 prominent median granular ridges (
<figureCitation id="131B2A06FFCCFFAFEF89F9B49847F9BE" box="[444,530,1552,1578]" captionStart="FIGURE 2" captionStartId="18.[151,255,1936,1960]" captionTargetBox="[264,1322,182,1924]" captionTargetId="figure@18.[264,1323,182,1926]" captionTargetPageId="18" captionText="FIGURE 2. Vultocinus anfractus, new genus, new species. Overall dorsal views. A, holotype male (cl 19.5 mm, cw 20.3 mm) (NMCR 19114), Philippines; B, paratype male (cl 19.9 mm, cw 22.0 mm) (ZRC), Philippines; C, male (cl 15.6 mm, cw 16.8 mm) (ZRC), Vanuatu." httpUri="https://zenodo.org/record/178298/files/figure.png" pageId="11" pageNumber="50">Figs. 2</figureCitation>
, 
<figureCitation id="131B2A06FFCCFFAFEC14F9B49864F9BE" box="[545,561,1552,1578]" captionStart="FIGURE 4" captionStartId="20.[151,255,1144,1168]" captionTargetBox="[151,1435,184,1120]" captionTargetId="figure@20.[151,1436,184,1120]" captionTargetPageId="20" captionText="FIGURE 4. Vultocinus anfractus, new genus, new species. Holotype male (cl 19.5 mm, cw 20.3 mm) (NMCR 19114), Philippines. A, outer surface of right chela; B, outer surface of left chela; C, right side of thoracic sternum showing ventral surface of chelipeds and spines on ambulatory meri; D, right P 4." httpUri="https://zenodo.org/record/178300/files/figure.png" pageId="11" pageNumber="50">4</figureCitation>
D). Dactylus long, as long as merus, covered by dense thick setae which hides tip, distal part not pigmented (
<figureCitation id="131B2A06FFCCFFAFEC0BF99C98C4F9C6" box="[574,657,1592,1618]" captionStart="FIGURE 2" captionStartId="18.[151,255,1936,1960]" captionTargetBox="[264,1322,182,1924]" captionTargetId="figure@18.[264,1323,182,1926]" captionTargetPageId="18" captionText="FIGURE 2. Vultocinus anfractus, new genus, new species. Overall dorsal views. A, holotype male (cl 19.5 mm, cw 20.3 mm) (NMCR 19114), Philippines; B, paratype male (cl 19.9 mm, cw 22.0 mm) (ZRC), Philippines; C, male (cl 15.6 mm, cw 16.8 mm) (ZRC), Vanuatu." httpUri="https://zenodo.org/record/178298/files/figure.png" pageId="11" pageNumber="50">Figs. 2</figureCitation>
, 
<figureCitation id="131B2A06FFCCFFAFECA8F99C98F8F9C6" box="[669,685,1592,1618]" captionStart="FIGURE 4" captionStartId="20.[151,255,1144,1168]" captionTargetBox="[151,1435,184,1120]" captionTargetId="figure@20.[151,1436,184,1120]" captionTargetPageId="20" captionText="FIGURE 4. Vultocinus anfractus, new genus, new species. Holotype male (cl 19.5 mm, cw 20.3 mm) (NMCR 19114), Philippines. A, outer surface of right chela; B, outer surface of left chela; C, right side of thoracic sternum showing ventral surface of chelipeds and spines on ambulatory meri; D, right P 4." httpUri="https://zenodo.org/record/178300/files/figure.png" pageId="11" pageNumber="50">4</figureCitation>
C, D).
</paragraph>
<paragraph id="8B9F3683FFCCFFA8EEF3F9C49AEEFEBE" blockId="11.[151,1437,152,2018]" lastBlockId="12.[151,1437,152,1978]" lastPageId="12" lastPageNumber="51" pageId="11" pageNumber="50">
Thoracic sternum relatively narrow transversely, entire surface covered with very short tomentum, small granules (
<figureCitation id="131B2A06FFCCFFAFEF3CF92C9B0FF936" box="[265,346,1672,1698]" captionStart="FIGURE 3" captionStartId="19.[151,255,1088,1112]" captionTargetBox="[151,1435,194,1064]" captionTargetId="figure@19.[151,1436,194,1064]" captionTargetPageId="19" captionText="FIGURE 3. Vultocinus anfractus, new genus, new species. Holotype male (cl 19.5 mm, cw 20.3 mm) (NMCR 19114), Philippines. A, dorsal view of carapace; B, ventral view of carapace; C, buccal area showing third maxillipeds; D, frontal view of carapace and chelipeds." httpUri="https://zenodo.org/record/178299/files/figure.png" pageId="11" pageNumber="50">Figs. 3</figureCitation>
B, C, 9A, 10A, 12A). Sternite 3 with shallow but distinct median longitudinal groove; sternite 4 as long as sternites 5 and 6 combined; sternite 7 very narrow longitudinally, posterior part tapering abruptly (
<figureCitation id="131B2A06FFCCFFAFEEAAF97C9AA0F966" box="[159,245,1752,1778]" captionStart="FIGURE 10" captionStartId="26.[151,255,1725,1749]" captionTargetBox="[151,1436,194,1701]" captionTargetId="figure@26.[151,1436,194,1701]" captionTargetPageId="26" captionText="FIGURE 10. Posterior part of thoracic sternum and male abdominal somites 3 – 5. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu &amp; Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng &amp; Ho, 2003: male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot &amp; Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng &amp; Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng &amp; Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: t = telson, a 2 – 5 = male abdominal somites 2 to 5 respectively, st 7 = sternites 7, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178306/files/figure.png" pageId="11" pageNumber="50">Fig. 10</figureCitation>
A), with posterior part of episternite 7 prolonged posteriorly to form spur which just touches coxa of P5 and covers penis (
<figureCitation id="131B2A06FFCCFFAFEFA5F8A49BB1F88E" box="[400,484,1792,1818]" captionStart="FIGURE 13" captionStartId="29.[151,255,1377,1401]" captionTargetBox="[264,1323,194,1353]" captionTargetId="figure@29.[264,1323,194,1353]" captionTargetPageId="29" captionText="FIGURE 13. Structure and position of penis, coxa of P 5 and episternite 7 (position of penis indicated by arrow). A, B, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); C, “ Goneplax ” marivenae Komatsu &amp; Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); D, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); E, Intesius pilosus Guinot &amp; Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Conleyus defodio Ng &amp; Ng, 2003, holotype male (cl, 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: p = penis, st 7 and 8 = sternites 7 and 8 respectively, es 7 = episternite 7, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178309/files/figure.png" pageId="11" pageNumber="50">Fig. 13</figureCitation>
A, B); sternite 8 not exposed when abdomen closed (
<figureCitation id="131B2A06FFCCFFAFEA7AF8A49EF1F88E" box="[1103,1188,1792,1818]" captionStart="FIGURE 10" captionStartId="26.[151,255,1725,1749]" captionTargetBox="[151,1436,194,1701]" captionTargetId="figure@26.[151,1436,194,1701]" captionTargetPageId="26" captionText="FIGURE 10. Posterior part of thoracic sternum and male abdominal somites 3 – 5. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu &amp; Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng &amp; Ho, 2003: male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot &amp; Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng &amp; Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng &amp; Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: t = telson, a 2 – 5 = male abdominal somites 2 to 5 respectively, st 7 = sternites 7, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178306/files/figure.png" pageId="11" pageNumber="50">Fig. 10</figureCitation>
A). Longitudinal line running through median part of sternites 7 and 8, those on sternites 4 and 6 incomplete, that of sternite 5 undiscernible (
<figureCitation id="131B2A06FFCCFFAFEF72F8F49BCAF8FE" box="[327,415,1872,1898]" captionStart="FIGURE 12" captionStartId="28.[151,255,1724,1748]" captionTargetBox="[227,1361,180,1710]" captionTargetId="figure@28.[227,1361,180,1710]" captionTargetPageId="28" captionText="FIGURE 12. Sterno-abdominal cavity, showing position of press button of abdominal locking mechanism (indicated by arrows) and sternal sutures. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu &amp; Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng &amp; Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot &amp; Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng &amp; Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng &amp; Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: pb = press button of abdominal locking mechanism, st 4 – 7 = sternites 4 to 7 respectively, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178308/files/figure.png" pageId="11" pageNumber="50">Fig. 12</figureCitation>
A). Suture 1/2 absent; suture 2/3 complete; suture 3/4 distinct with median part relatively shallower; sutures 4/5 and 5/6 medially interrupted (
<figureCitation id="131B2A06FFCCFFAFED73F8DC99CEF806" box="[838,923,1912,1938]" captionStart="FIGURE 12" captionStartId="28.[151,255,1724,1748]" captionTargetBox="[227,1361,180,1710]" captionTargetId="figure@28.[227,1361,180,1710]" captionTargetPageId="28" captionText="FIGURE 12. Sterno-abdominal cavity, showing position of press button of abdominal locking mechanism (indicated by arrows) and sternal sutures. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu &amp; Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng &amp; Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot &amp; Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng &amp; Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng &amp; Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: pb = press button of abdominal locking mechanism, st 4 – 7 = sternites 4 to 7 respectively, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178308/files/figure.png" pageId="11" pageNumber="50">Fig. 12</figureCitation>
A); suture 6/7 appears complete but median portion very shallow (
<figureCitation id="131B2A06FFCCFFAFEFADF8049BB8F82E" box="[408,493,1952,1978]" captionStart="FIGURE 12" captionStartId="28.[151,255,1724,1748]" captionTargetBox="[227,1361,180,1710]" captionTargetId="figure@28.[227,1361,180,1710]" captionTargetPageId="28" captionText="FIGURE 12. Sterno-abdominal cavity, showing position of press button of abdominal locking mechanism (indicated by arrows) and sternal sutures. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu &amp; Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng &amp; Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot &amp; Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng &amp; Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng &amp; Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: pb = press button of abdominal locking mechanism, st 4 – 7 = sternites 4 to 7 respectively, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178308/files/figure.png" pageId="11" pageNumber="50">Fig. 12</figureCitation>
A); suture 7/8 distinctly complete with median part deep (
<figureCitation id="131B2A06FFCCFFAFEAA5F8049EB0F82E" box="[1168,1253,1952,1978]" captionStart="FIGURE 12" captionStartId="28.[151,255,1724,1748]" captionTargetBox="[227,1361,180,1710]" captionTargetId="figure@28.[227,1361,180,1710]" captionTargetPageId="28" captionText="FIGURE 12. Sterno-abdominal cavity, showing position of press button of abdominal locking mechanism (indicated by arrows) and sternal sutures. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu &amp; Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng &amp; Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot &amp; Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng &amp; Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng &amp; Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: pb = press button of abdominal locking mechanism, st 4 – 7 = sternites 4 to 7 respectively, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178308/files/figure.png" pageId="11" pageNumber="50">Fig. 12</figureCitation>
A). Episternites 4–7 not prominently expanded laterally (
<figureCitation id="131B2A06FFCCFFAFECBEF86C98B4F876" box="[651,737,1992,2018]" captionStart="FIGURE 9" captionStartId="25.[151,255,1726,1750]" captionTargetBox="[224,1358,194,1697]" captionTargetId="figure@25.[224,1358,194,1697]" captionTargetPageId="25" captionText="FIGURE 9. Anterior part of thoracic sternum and male abdominal somites 5, 6 and telson. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu &amp; Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng &amp; Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot &amp; Richer de Forges, 1981: male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng &amp; Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng &amp; Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: t = telson, a 4 – 6 = male abdominal somites 4 to 6 respectively, st 4 – 7 = sternites 4 to 7 respectively." httpUri="https://zenodo.org/record/178305/files/figure.png" pageId="11" pageNumber="50">Figs. 9</figureCitation>
A, 10A, 12A). Sterno-abdominal cavity relatively narrow transversely, almost reaches anterior edge of sternite 4 (
<figureCitation id="131B2A06FFCBFFA8ED0DFF3C99D9FF26" box="[824,908,152,178]" captionStart="FIGURE 9" captionStartId="25.[151,255,1726,1750]" captionTargetBox="[224,1358,194,1697]" captionTargetId="figure@25.[224,1358,194,1697]" captionTargetPageId="25" captionText="FIGURE 9. Anterior part of thoracic sternum and male abdominal somites 5, 6 and telson. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu &amp; Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng &amp; Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot &amp; Richer de Forges, 1981: male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng &amp; Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng &amp; Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: t = telson, a 4 – 6 = male abdominal somites 4 to 6 respectively, st 4 – 7 = sternites 4 to 7 respectively." httpUri="https://zenodo.org/record/178305/files/figure.png" pageId="12" pageNumber="51">Figs. 9</figureCitation>
A, 12A). Press button of abdominal locking mechanism distinct, on posterior outer edge of sternite 5 of sterno-abdominal cavity (
<figureCitation id="131B2A06FFCBFFA8EAACFF649EA5FF4E" box="[1177,1264,192,218]" captionStart="FIGURE 12" captionStartId="28.[151,255,1724,1748]" captionTargetBox="[227,1361,180,1710]" captionTargetId="figure@28.[227,1361,180,1710]" captionTargetPageId="28" captionText="FIGURE 12. Sterno-abdominal cavity, showing position of press button of abdominal locking mechanism (indicated by arrows) and sternal sutures. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu &amp; Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng &amp; Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot &amp; Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng &amp; Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng &amp; Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: pb = press button of abdominal locking mechanism, st 4 – 7 = sternites 4 to 7 respectively, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178308/files/figure.png" pageId="12" pageNumber="51">Fig. 12</figureCitation>
A). Penis soft, effectively coxal, exits at base of coxa of P5 but covered by posterior prolongation of episternite 7 (
<figureCitation id="131B2A06FFCBFFA8EB15FF4C9F2AFE96" box="[1312,1407,232,258]" captionStart="FIGURE 13" captionStartId="29.[151,255,1377,1401]" captionTargetBox="[264,1323,194,1353]" captionTargetId="figure@29.[264,1323,194,1353]" captionTargetPageId="29" captionText="FIGURE 13. Structure and position of penis, coxa of P 5 and episternite 7 (position of penis indicated by arrow). A, B, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); C, “ Goneplax ” marivenae Komatsu &amp; Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); D, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); E, Intesius pilosus Guinot &amp; Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Conleyus defodio Ng &amp; Ng, 2003, holotype male (cl, 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: p = penis, st 7 and 8 = sternites 7 and 8 respectively, es 7 = episternite 7, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178309/files/figure.png" pageId="12" pageNumber="51">Figs. 13</figureCitation>
A, B).
</paragraph>
<paragraph id="8B9F3683FFCBFFA8EEF3FE9C9E1AFDFE" blockId="12.[151,1437,152,1978]" pageId="12" pageNumber="51">
Abdomen relatively narrow transversely, outer surfaces mostly covered with small low granules, very short, dense tomentum (
<figureCitation id="131B2A06FFCBFFA8EF82FEC4985EFEEE" box="[439,523,352,378]" captionStart="FIGURE 5" captionStartId="21.[151,255,1379,1403]" captionTargetBox="[158,1421,201,1348]" captionTargetId="figure@21.[151,1436,194,1355]" captionTargetPageId="21" captionText="FIGURE 5. Vultocinus anfractus, new genus, new species. Paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC), Philippines. A, left third maxilliped (setae denuded); B, male abdomen (partially denuded); C, left G 1, dorsal view; D, left G 1, ventral view; E, distal part of left G 1, dorsal view; F, distal part of left G 1, ventral view; G, left G 2; H, distal part of left G 2. Scales = 1.0 mm." httpUri="https://zenodo.org/record/178301/files/figure.png" pageId="12" pageNumber="51">Figs. 5</figureCitation>
B, 9A, 11A). Telson triangular with gently concave lateral margins, rounded apex (
<figureCitation id="131B2A06FFCBFFA8EEE8FE2C9B7BFE36" box="[221,302,392,418]" captionStart="FIGURE 5" captionStartId="21.[151,255,1379,1403]" captionTargetBox="[158,1421,201,1348]" captionTargetId="figure@21.[151,1436,194,1355]" captionTargetPageId="21" captionText="FIGURE 5. Vultocinus anfractus, new genus, new species. Paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC), Philippines. A, left third maxilliped (setae denuded); B, male abdomen (partially denuded); C, left G 1, dorsal view; D, left G 1, ventral view; E, distal part of left G 1, dorsal view; F, distal part of left G 1, ventral view; G, left G 2; H, distal part of left G 2. Scales = 1.0 mm." httpUri="https://zenodo.org/record/178301/files/figure.png" pageId="12" pageNumber="51">Figs. 5</figureCitation>
B, 9A); somite 1 longitudinally and transversely broad, trapezoidal, exposed, not covered by posterior carapace margin (
<figureCitation id="131B2A06FFCBFFA8EF9BFE149854FE5E" box="[430,513,432,458]" captionStart="FIGURE 11" captionStartId="27.[151,255,1445,1469]" captionTargetBox="[151,1435,194,1420]" captionTargetId="figure@27.[151,1436,194,1421]" captionTargetPageId="27" captionText="FIGURE 11. Posterior part of carapace showing posterior carapace margin, male abdominal somites 1 and 2, and coxa of last ambulatory legs. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu &amp; Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng &amp; Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot &amp; Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng &amp; Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng &amp; Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: t = telson, a 1 – 3 = male abdominal somites 1 to 3 respectively, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178307/files/figure.png" pageId="12" pageNumber="51">Fig. 11</figureCitation>
A), somites 2–6 relatively narrow transversely, somite 3 broadest transversely; somite 2 with convex lateral margins, subrectangular in shape; somite 3 with gently convex lateral margins, trapezoidal; somites 4 larger than somite 5, both trapezoidal with gently concave lateral margins; somite 6 as long as broad, lateral margins concave (
<figureCitation id="131B2A06FFCBFFA8EC40FD8C989DFDD6" box="[629,712,552,578]" captionStart="FIGURE 9" captionStartId="25.[151,255,1726,1750]" captionTargetBox="[224,1358,194,1697]" captionTargetId="figure@25.[224,1358,194,1697]" captionTargetPageId="25" captionText="FIGURE 9. Anterior part of thoracic sternum and male abdominal somites 5, 6 and telson. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu &amp; Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng &amp; Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot &amp; Richer de Forges, 1981: male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng &amp; Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng &amp; Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: t = telson, a 4 – 6 = male abdominal somites 4 to 6 respectively, st 4 – 7 = sternites 4 to 7 respectively." httpUri="https://zenodo.org/record/178305/files/figure.png" pageId="12" pageNumber="51">Figs. 9</figureCitation>
A, 10A); sutures between all somites and telson distinct, but somites 3 and 4 immovable, somites 4 and 5 slightly mobile (
<figureCitation id="131B2A06FFCBFFA8ED53FDF499E2FDFE" box="[870,951,592,618]" captionStart="FIGURE 5" captionStartId="21.[151,255,1379,1403]" captionTargetBox="[158,1421,201,1348]" captionTargetId="figure@21.[151,1436,194,1355]" captionTargetPageId="21" captionText="FIGURE 5. Vultocinus anfractus, new genus, new species. Paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC), Philippines. A, left third maxilliped (setae denuded); B, male abdomen (partially denuded); C, left G 1, dorsal view; D, left G 1, ventral view; E, distal part of left G 1, dorsal view; F, distal part of left G 1, ventral view; G, left G 2; H, distal part of left G 2. Scales = 1.0 mm." httpUri="https://zenodo.org/record/178301/files/figure.png" pageId="12" pageNumber="51">Figs. 5</figureCitation>
B, 9A, 11A).
</paragraph>
<paragraph id="8B9F3683FFCBFFA8EEF3FDDC9B4EFC16" blockId="12.[151,1437,152,1978]" pageId="12" pageNumber="51">
Penis effectively coxosternal although it exits clearly from coxa, enters directly into sterno-abdominal cavity, not protected by sternal plates except for posterior part of sternite 7 which forms a spur-like structure that covers it (
<figureCitation id="131B2A06FFCBFFA8EF09FD6C9BCFFD76" box="[316,410,712,738]" captionStart="FIGURE 13" captionStartId="29.[151,255,1377,1401]" captionTargetBox="[264,1323,194,1353]" captionTargetId="figure@29.[264,1323,194,1353]" captionTargetPageId="29" captionText="FIGURE 13. Structure and position of penis, coxa of P 5 and episternite 7 (position of penis indicated by arrow). A, B, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); C, “ Goneplax ” marivenae Komatsu &amp; Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); D, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); E, Intesius pilosus Guinot &amp; Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Conleyus defodio Ng &amp; Ng, 2003, holotype male (cl, 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: p = penis, st 7 and 8 = sternites 7 and 8 respectively, es 7 = episternite 7, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178309/files/figure.png" pageId="12" pageNumber="51">Figs. 13</figureCitation>
A, B). G1 relatively long, slender, with the rounded tip reaching suture of sternite 4, distal surfaces lined with numerous small spinules, tip open (
<figureCitation id="131B2A06FFCBFFA8ED72FD5499CDFC9E" box="[839,920,752,778]" captionStart="FIGURE 5" captionStartId="21.[151,255,1379,1403]" captionTargetBox="[158,1421,201,1348]" captionTargetId="figure@21.[151,1436,194,1355]" captionTargetPageId="21" captionText="FIGURE 5. Vultocinus anfractus, new genus, new species. Paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC), Philippines. A, left third maxilliped (setae denuded); B, male abdomen (partially denuded); C, left G 1, dorsal view; D, left G 1, ventral view; E, distal part of left G 1, dorsal view; F, distal part of left G 1, ventral view; G, left G 2; H, distal part of left G 2. Scales = 1.0 mm." httpUri="https://zenodo.org/record/178301/files/figure.png" pageId="12" pageNumber="51">Figs. 5</figureCitation>
C–F). G2 as long as G1, very slender, long, with tip reaching beyond mid-portion of suture of sternite 5, distal segment ca. 0.2 times length of subdistal segment, base of distal segment forming cup-like structure lined with several spinules (
<figureCitation id="131B2A06FFCBFFA8EAAFFCE49EBFFCCE" box="[1178,1258,832,858]" captionStart="FIGURE 5" captionStartId="21.[151,255,1379,1403]" captionTargetBox="[158,1421,201,1348]" captionTargetId="figure@21.[151,1436,194,1355]" captionTargetPageId="21" captionText="FIGURE 5. Vultocinus anfractus, new genus, new species. Paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC), Philippines. A, left third maxilliped (setae denuded); B, male abdomen (partially denuded); C, left G 1, dorsal view; D, left G 1, ventral view; E, distal part of left G 1, dorsal view; F, distal part of left G 1, ventral view; G, left G 2; H, distal part of left G 2. Scales = 1.0 mm." httpUri="https://zenodo.org/record/178301/files/figure.png" pageId="12" pageNumber="51">Figs. 5</figureCitation>
G, H). Females not known.
</paragraph>
<paragraph id="8B9F3683FFCBFFA8EEF3FC2B9B24FBE6" blockId="12.[151,1437,152,1978]" pageId="12" pageNumber="51">
<emphasis id="B954EA91FFCBFFA8EEF3FC2B9B49FC3D" bold="true" box="[198,284,911,937]" pageId="12" pageNumber="51">Colour</emphasis>
. The carapace and pereiopods are white without any distinct pigmentation, but this is barely discernible as the dark brown or orangish-brown tomentum and setae covers most of the animal (
<figureCitation id="131B2A06FFCBFFA8EB39FC1C9F06FC46" box="[1292,1363,952,978]" captionStart="FIGURE 1" captionStartId="17.[151,255,1912,1936]" captionTargetBox="[226,1360,194,1887]" captionTargetId="figure@17.[226,1360,194,1888]" captionTargetPageId="17" captionText="FIGURE 1. Vultocinus anfractus, new genus, new species. Fresh colours. A, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC), Philippines, freshly dead, poised on same kind of wood substrate it was reportedly found on; B, paratype male (cl 19.9 mm, cw 22.0 mm) (ZRC), Philippines; C, male (cl 15.6 mm, cw 16.8 mm) (ZRC), Vanuatu." httpUri="https://zenodo.org/record/178297/files/figure.png" pageId="12" pageNumber="51">Fig. 1</figureCitation>
). The mandibles are a dark brown, like dark-coloured chitin, and this colour stays even after many months and many changes of alcohol (
<figureCitation id="131B2A06FFCBFFA8EFB7FBAC9B92FBB6" box="[386,455,1032,1058]" captionStart="FIGURE 8" captionStartId="24.[151,255,1099,1123]" captionTargetBox="[151,1435,194,1073]" captionTargetId="figure@24.[151,1436,194,1075]" captionTargetPageId="24" captionText="FIGURE 8. Buccal field, showing mouthparts and mandibles. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, Mathildella rubra Ng &amp; Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); C, Progeryon mus Ng &amp; Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC); D, Conleyus defodio Ng &amp; Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH)." httpUri="https://zenodo.org/record/178304/files/figure.png" pageId="12" pageNumber="51">Fig. 8</figureCitation>
A). The posterior margin of the epistome is lined with small orange and white granules when fresh. Some of the the larger granules on the carapace and pereiopods (especially those on the chelipeds) are orange.
</paragraph>
</subSubSection>
<subSubSection id="C33A6508FFCBFFA8EEF3FBDB9B9DFB56" pageId="12" pageNumber="51" type="etymology">
<paragraph id="8B9F3683FFCBFFA8EEF3FBDB9B9DFB56" blockId="12.[151,1437,152,1978]" pageId="12" pageNumber="51">
<emphasis id="B954EA91FFCBFFA8EEF3FBDB9B1EFB0D" bold="true" box="[198,331,1151,1177]" pageId="12" pageNumber="51">Etymology</emphasis>
. The name is derived from the Latin 
<taxonomicName id="4C204D00FFCBFFA8ECCAFB249938FB0D" box="[767,877,1152,1177]" class="Malacostraca" family="Vultocinidae" genus="Vultocinus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="51" phylum="Arthropoda" rank="species" species="anfractus">
<emphasis id="B954EA91FFCBFFA8ECCAFB249938FB0D" box="[767,877,1152,1177]" italics="true" pageId="12" pageNumber="51">anfractus</emphasis>
</taxonomicName>
for having many turns, alluding to the complex sculpture on the carapace.
</paragraph>
</subSubSection>
<subSubSection id="C33A6508FFCBFFA9EEF3FB6B9ABDFD2E" lastPageId="13" lastPageNumber="52" pageId="12" pageNumber="51" type="discussion">
<paragraph id="8B9F3683FFCBFFA8EEF3FB6B9B9AF996" blockId="12.[151,1437,152,1978]" pageId="12" pageNumber="51">
<emphasis id="B954EA91FFCBFFA8EEF3FB6B9B6DFB7D" bold="true" box="[198,312,1231,1257]" pageId="12" pageNumber="51">Remarks</emphasis>
. The unusual carapace and pereiopods of 
<taxonomicName id="4C204D00FFCBFFA8ED1DFB7499E1FB7D" box="[808,948,1232,1257]" class="Malacostraca" family="Vultocinidae" genus="Vultocinus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="51" phylum="Arthropoda" rank="species" species="anfractus">
<emphasis id="B954EA91FFCBFFA8ED1DFB7499E1FB7D" box="[808,948,1232,1257]" italics="true" pageId="12" pageNumber="51">V. anfractus</emphasis>
</taxonomicName>
are unique in the Goneplacoidea as presently understood. The complex labyrinth-like setal pattern is reminiscent of pilumnids like 
<taxonomicName id="4C204D00FFCBFFA8EAF2FB5C9B2EFAAE" authority="Miers, 1884" authorityName="Miers" authorityYear="1884" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="51" phylum="Arthropoda" rank="species" species="labyrinthicus">
<emphasis id="B954EA91FFCBFFA8EAF2FB5C9A88FAAD" italics="true" pageId="12" pageNumber="51">Pilumnus labyrinthicus</emphasis>
<bibRefCitation id="EFB14B72FFCBFFA8EEDDFA849B2EFAAE" author="Miers" box="[232,379,1312,1338]" pageId="12" pageNumber="54" refString="Miers, E. J. (1884) Crustacea. In Report on the Zoological Collections made in the Indo-Pacific Ocean during the voyage of the H. M. S. &quot; Alert &quot;, 1881. London, pp. i-xxv, 1 - 684, pls. I - LIV." type="journal article" year="1884">Miers, 1884</bibRefCitation>
</taxonomicName>
, or pseudoziids like 
<taxonomicName id="4C204D00FFCBFFA8EC4EFA849E31FAAE" authority="Balss, 1933" authorityName="Balss" authorityYear="1933" box="[635,1124,1312,1338]" class="Malacostraca" family="Planopilumnidae" genus="Planopilumnus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="51" phylum="Arthropoda" rank="species" species="spongiosus">
<emphasis id="B954EA91FFCBFFA8EC4EFA8499ECFAAD" box="[635,953,1312,1337]" italics="true" pageId="12" pageNumber="51">Planopilumnus spongiosus</emphasis>
(
<bibRefCitation id="EFB14B72FFCBFFA8EDF9FA849E0EFAAE" author="Balss" box="[972,1115,1312,1338]" pageId="12" pageNumber="52" refString="Balss, H. (1933) Beitrage zur Kenntnis der Gattungen Pilumnus (Crustacea Dekapoda) und verwandter Gattungen. Capita Zoologica, 4, 1 - 47, pls. 1 - 7." type="journal article" year="1933">Balss, 1933</bibRefCitation>
)
</taxonomicName>
although they are clearly unrelated (see discussion above). Certainly none possess such a complex setal pattern and the ambulatory legs are not armed with spines. The dense carapace and pereiopod setation of 
<taxonomicName id="4C204D00FFCBFFA8EDCDFAD49ED1FA1D" box="[1016,1156,1392,1417]" class="Malacostraca" family="Vultocinidae" genus="Vultocinus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="51" phylum="Arthropoda" rank="species" species="anfractus">
<emphasis id="B954EA91FFCBFFA8EDCDFAD49ED1FA1D" box="[1016,1156,1392,1417]" italics="true" pageId="12" pageNumber="51">V. anfractus</emphasis>
</taxonomicName>
superficially resembles those of the mathildellid, 
<taxonomicName id="4C204D00FFCBFFA8EFF1FA3C9913FA26" authority="Guinot &amp; Richer" authorityName="Guinot &amp; Richer" box="[452,838,1432,1458]" class="Malacostraca" family="Mathildellidae" genus="Intesius" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="51" phylum="Arthropoda" rank="species" species="pilosus">
<emphasis id="B954EA91FFCBFFA8EFF1FA3C982CFA25" box="[452,633,1432,1457]" italics="true" pageId="12" pageNumber="51">Intesius pilosus</emphasis>
Guinot &amp; Richer
</taxonomicName>
de 
<bibRefCitation id="EFB14B72FFCBFFA8ED44FA3C9E59FA26" author="Forges" box="[881,1036,1432,1458]" pageId="12" pageNumber="53" refString="Guinot, D. &amp; Richer de Forges, B. (1981) Crabes de profondeur, nouveaux ou rares, de l'Indo-Pacifique (Crustacea, Decapoda, Brachyura) (Deuxieme partie). Bulletin du Museum National d'Histoire Naturelle, Paris, 1981, (4) 3 (A 1), 227 - 260." type="journal article" year="1981">Forges, 1981</bibRefCitation>
, but in the latter species, the setae are more scattered and the carapace lacks the dense setae and distinct setal arrangement of 
<taxonomicName id="4C204D00FFCBFFA8EAE2FA649F31FA4D" box="[1239,1380,1472,1497]" class="Malacostraca" family="Vultocinidae" genus="Vultocinus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="51" phylum="Arthropoda" rank="species" species="anfractus">
<emphasis id="B954EA91FFCBFFA8EAE2FA649F31FA4D" box="[1239,1380,1472,1497]" italics="true" pageId="12" pageNumber="51">V. anfractus</emphasis>
</taxonomicName>
(see also 
<bibRefCitation id="EFB14B72FFCBFFA8EEF8FA4C9B96F996" author="Crosnier" box="[205,451,1512,1538]" pageId="12" pageNumber="53" refString="Crosnier, A. &amp; Ng, P. K. L. (2004) Remarques sur le genre Intesius (Crustacea, Decapoda, Brachyura, Goneplacidae) et description de deux especes nouvelles. Zoosystema, 26 (2), 263 - 277." type="journal article" year="2004">Crosnier &amp; Ng 2004</bibRefCitation>
).
</paragraph>
<paragraph id="8B9F3683FFCBFFA8EEF3F9B499FDF82E" blockId="12.[151,1437,152,1978]" pageId="12" pageNumber="51">
The 
<typeStatus id="549B8821FFCBFFA8EECEF9B49B37F9BE" box="[251,354,1552,1578]" pageId="12" pageNumber="51" type="holotype">holotype</typeStatus>
male of 
<taxonomicName id="4C204D00FFCBFFA8EFFCF9B49801F9BD" box="[457,596,1552,1577]" class="Malacostraca" family="Vultocinidae" genus="Vultocinus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="51" phylum="Arthropoda" rank="species" species="anfractus">
<emphasis id="B954EA91FFCBFFA8EFFCF9B49801F9BD" box="[457,596,1552,1577]" italics="true" pageId="12" pageNumber="51">V. anfractus</emphasis>
</taxonomicName>
was collected from the rocky shoreline outside a Nipa mangrove near a river. It was crawling along on the shore in the afternoon when it was caught by the second author. We believe it had been actually caught by fishermen with tangle nets in the waters offshore as they had been observed cleaning their nets there earlier. In addition, there is a small piece of net stuck at the base of the chela of the specimen. It was almost certainly thrown onto the shore just before it was collected. The larger 
<typeStatus id="549B8821FFCBFFA8EAC2F9149F09F95E" box="[1271,1372,1712,1738]" pageId="12" pageNumber="51" type="paratype">paratype</typeStatus>
male was collected from about 
<quantity id="4CD89B66FFCBFFA8EFF6F97C9853F966" box="[451,518,1752,1778]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="12" pageNumber="51" unit="m" value="200.0">200m</quantity>
depth hiding inside a piece of sunken wood about 
<quantity id="4CD89B66FFCBFFA8EA60F97C9EC2F966" box="[1109,1175,1752,1778]" metricMagnitude="-1" metricUnit="m" metricValue="7.0" pageId="12" pageNumber="51" unit="m" value="0.7">0.7 m</quantity>
in length and 
<quantity id="4CD89B66FFCBFFA8EB09F97C9F2AF966" box="[1340,1407,1752,1778]" metricMagnitude="-1" metricUnit="m" metricValue="3.0" pageId="12" pageNumber="51" unit="m" value="0.3">0.3 m</quantity>
in diameter which was stuck in a tangle net (J. Arbasto, pers. comm.). The colour of the body and legs (dark brown) matches the wood substrate where the animal was found and it uses the spiniform ventral margins of the ambulatory meri to help anchor itself to the wood (J. Arbasto, pers. comm.). Freshly caught specimens were very dirty, being covered with detritus and mud, clearly a consequence of its setose body and pereiopods. This probably aids in camouflaging them against the wood as well.
</paragraph>
<paragraph id="8B9F3683FFCAFFA9EEF3FF3C9B72FDFE" blockId="13.[151,1436,152,698]" pageId="13" pageNumber="52">
The single male specimen of 
<taxonomicName id="4C204D00FFCAFFA9EC19FF3C9975FF25" box="[556,800,152,177]" class="Malacostraca" family="Vultocinidae" genus="Vultocinus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="52" phylum="Arthropoda" rank="species" species="anfractus">
<emphasis id="B954EA91FFCAFFA9EC19FF3C9975FF25" box="[556,800,152,177]" italics="true" pageId="13" pageNumber="52">Vultocinus anfractus</emphasis>
</taxonomicName>
collected from 
<collectingCountry id="F3377613FFCAFFA9EDD4FF3C9E14FF26" box="[993,1089,152,178]" name="Vanuatu" pageId="13" pageNumber="52">Vanuatu</collectingCountry>
was from a tangle net set at about ₃₂ ɭΟ ɬ0 m depth. The specimen was obtained alive but we have no indication if it had been living on wood. This specimen differs from the 
<typeStatus id="549B8821FFCAFFA9EC52FF4C98CFFE96" box="[615,666,232,258]" pageId="13" pageNumber="52">type</typeStatus>
series of the species in having an orangish-brown body (versus dark brown), somewhat rounder carapace, and proportionately slightly longer ambulatory legs (
<figureCitation id="131B2A06FFCAFFA9EAC0FEB49F17FEBE" box="[1269,1346,272,298]" captionStart="FIGURE 1" captionStartId="17.[151,255,1912,1936]" captionTargetBox="[226,1360,194,1887]" captionTargetId="figure@17.[226,1360,194,1888]" captionTargetPageId="17" captionText="FIGURE 1. Vultocinus anfractus, new genus, new species. Fresh colours. A, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC), Philippines, freshly dead, poised on same kind of wood substrate it was reportedly found on; B, paratype male (cl 19.9 mm, cw 22.0 mm) (ZRC), Philippines; C, male (cl 15.6 mm, cw 16.8 mm) (ZRC), Vanuatu." httpUri="https://zenodo.org/record/178297/files/figure.png" pageId="13" pageNumber="52">Figs. 1</figureCitation>
C, 2C). It agrees with the 
<typeStatus id="549B8821FFCAFFA9EF59FE9C9BCAFEC6" box="[364,415,312,338]" pageId="13" pageNumber="52">type</typeStatus>
material, in almost all other aspects, and we prefer to regard it as conspecific with the 
<taxonomicName id="4C204D00FFCAFFA9EEA2FEC49BDCFEED" box="[151,393,352,377]" class="Malacostraca" family="Vultocinidae" genus="Vultocinus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="52" phylum="Arthropoda" rank="species" species="anfractus">
<emphasis id="B954EA91FFCAFFA9EEA2FEC49BDCFEED" box="[151,393,352,377]" italics="true" pageId="13" pageNumber="52">Vultocinus anfractus</emphasis>
</taxonomicName>
material from the 
<collectingCountry id="F3377613FFCAFFA9EC50FEC498BCFEEE" box="[613,745,352,378]" name="Philippines" pageId="13" pageNumber="52">Philippines</collectingCountry>
for the moment. Although mature, it is the smallest specimen of 
<taxonomicName id="4C204D00FFCAFFA9EEC4FE2C9B38FE35" box="[241,365,392,417]" class="Malacostraca" family="Vultocinidae" genus="Vultocinus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="B954EA91FFCAFFA9EEC4FE2C9B38FE35" box="[241,365,392,417]" italics="true" pageId="13" pageNumber="52">Vultocinus</emphasis>
</taxonomicName>
available, and the differences observed may well be due to size and habitat. A larger series of specimens from 
<collectingCountry id="F3377613FFCAFFA9EF4EFE149B8EFE5E" box="[379,475,432,458]" name="Vanuatu" pageId="13" pageNumber="52">Vanuatu</collectingCountry>
or other areas will be needed to ascertain this. The specimen of 
<taxonomicName id="4C204D00FFCAFFA9EAEFFE149F33FE5D" box="[1242,1382,432,457]" class="Malacostraca" family="Vultocinidae" genus="Vultocinus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="52" phylum="Arthropoda" rank="species" species="anfractus">
<emphasis id="B954EA91FFCAFFA9EAEFFE149F33FE5D" box="[1242,1382,432,457]" italics="true" pageId="13" pageNumber="52">V. anfractus</emphasis>
</taxonomicName>
was observed alive for several hours before being preserved. It moved slowly and was not aggressive when handled, but that may have been due to it being tangled in the nets for many hours. When placed against a hard substrate, the ambulatory legs invariably clamp backwards, confirming what we believe the spines on the meri are used for.
</paragraph>
<paragraph id="8B9F3683FFCAFFA9EEF3FDDC9ABDFD2E" blockId="13.[151,1436,152,698]" pageId="13" pageNumber="52">It is unfortunate that all the specimens collected were males, so key female characters could not be determined.</paragraph>
</subSubSection>
</treatment>
</document>