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<document id="65770696F08E550E3A59E883F858ECA8" ID-CLB-Dataset="31619" ID-DOI="10.11646/zootaxa.4344.1.4" ID-GBIF-Dataset="3c782ba4-ed7e-4064-a613-2c95aae16bbe" ID-ISSN="1175-5326" ID-Zenodo-Dep="1042317" ID-ZooBank="E87548A3-3CDF-43DA-A5F2-5BE87F8F1910" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1509955301820" checkinUser="plazi" docAuthor="Tahami, Mohadeseh Sadat, Muilwijk, Jan, Lohaj, Roman &amp; Sadeghi, Saber" docDate="2017" docId="A71687A9E44EFFAC848019AE2F26F924" docLanguage="en" docName="zootaxa.4344.1.4.pdf" docOrigin="Zootaxa 4344 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Laemostenus (Iranosphodrus) troglophilus Tahami &amp; Muilwijk &amp; Lohaj &amp; Sadeghi 2017, sp. nov." docType="treatment" docVersion="7" lastPageNumber="131" masterDocId="5B2FFFD1E441FFBC841719392D53FFC1" masterDocTitle="Study of Laemostenus species across Zagros and Central zone of Iran, with the description of seven new cavernicolous species and notes on subgenus Iranosphodrus (Coleoptera: Carabidae: Sphodrini)" masterLastPageNumber="136" masterPageNumber="115" pageNumber="130" updateTime="1698480792691" updateUser="ExternalLinkService">
<mods:mods id="C6898826195452DF7576A81F6A8012EF" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="AF13303A94B69BC6956483E6C0A4494D">
<mods:title id="D6AA9077662B56E0BD88C75B7E0EDBE7">Study of Laemostenus species across Zagros and Central zone of Iran, with the description of seven new cavernicolous species and notes on subgenus Iranosphodrus (Coleoptera: Carabidae: Sphodrini)</mods:title>
</mods:titleInfo>
<mods:name id="8475EAFEEA87BA2663A3F185A123B627" type="personal">
<mods:role id="D2AA54885D08BC6167A1913B8C753C9B">
<mods:roleTerm id="9E68D6DF335F20178AD7729D8F70F916">Author</mods:roleTerm>
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<mods:namePart id="11A94D981C0CE2CE2C743D4ADF381341">Tahami, Mohadeseh Sadat</mods:namePart>
</mods:name>
<mods:name id="07183C5AA6CD4396672C3C319084EAF4" type="personal">
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<mods:namePart id="098B5D3372C49D7A7838A968928FEED0">Muilwijk, Jan</mods:namePart>
</mods:name>
<mods:name id="5F498FE8233F5C3361919A0BFD44F534" type="personal">
<mods:role id="4E86A93204E0CA50A2EB3B478653B2F2">
<mods:roleTerm id="8B343FC0FC191F5C0EE9B06EADC0F667">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E98F766DE64F98CCC679D1E8EDBDC331">Lohaj, Roman</mods:namePart>
</mods:name>
<mods:name id="6B3108E5B5D354AA0E75268508155414" type="personal">
<mods:role id="023AFF127C68BABFB3D3FE581F4111B2">
<mods:roleTerm id="3904D3173E9247AD70DE3C70A66EFBF1">Author</mods:roleTerm>
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<mods:namePart id="96F5D2D1BE1FEC57D43CEE098C08174D">Sadeghi, Saber</mods:namePart>
</mods:name>
<mods:typeOfResource id="4D58EB0E068A3B8BC0C1471522A5AA70">text</mods:typeOfResource>
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<mods:titleInfo id="CA9DEBCDCC16A2F8E275434D5CABC9C9">
<mods:title id="07E0388C83CE2B1E98E3D9EB3E296CC6">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="FDCBF217EA628258A3E8C05667AF6A7B">
<mods:date id="6DE446FA9D00709FBAFF155BC73A4949">2017</mods:date>
<mods:detail id="B60AC6EED38BF1274DAD9EA857119A3F" type="pubDate">
<mods:number id="B334A9A7BAD2866492353F6A4B1A855B">2017-11-06</mods:number>
</mods:detail>
<mods:detail id="143AEE289F4AEF86C6266019640E9E6E" type="volume">
<mods:number id="53DF5822D9294286E80FEA7E36AFEC3D">4344</mods:number>
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<mods:detail id="9369C619D722050217D2064E2540D6DC" type="issue">
<mods:number id="997E3841592E1D72395C33FA81ECA558">1</mods:number>
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<mods:classification id="66D27FC58122FA6520EACB7F9C9552FE">journal article</mods:classification>
<mods:identifier id="3BD1FA0D8F5BC6F72B5E883EBE4001F3" type="CLB-Dataset">31619</mods:identifier>
<mods:identifier id="78185E1B06AF8DFDE4691F60865BDD0B" type="DOI">10.11646/zootaxa.4344.1.4</mods:identifier>
<mods:identifier id="05B42AA38D8F3562BE165F08A9175CC7" type="GBIF-Dataset">3c782ba4-ed7e-4064-a613-2c95aae16bbe</mods:identifier>
<mods:identifier id="2DAF432AFBE03BF13BD7202773C712EC" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="D3C256B1BC8E1F5B531BFB59B007E1FF" type="Zenodo-Dep">1042317</mods:identifier>
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<treatment id="A71687A9E44EFFAC848019AE2F26F924" ID-DOI="http://doi.org/10.5281/zenodo.6020140" ID-GBIF-Taxon="137239342" ID-Zenodo-Dep="6020140" LSID="urn:lsid:plazi:treatment:A71687A9E44EFFAC848019AE2F26F924" httpUri="http://treatment.plazi.org/id/A71687A9E44EFFAC848019AE2F26F924" lastPageId="16" lastPageNumber="131" pageId="15" pageNumber="130">
<subSubSection id="67A56534E44EFFB3848019AE2CFFFF12" pageId="15" pageNumber="130" type="nomenclature">
<paragraph id="2F0036BFE44EFFB3848019AE2936FF73" blockId="15.[151,1125,151,211]" box="[151,1125,151,178]" pageId="15" pageNumber="130">
<heading id="744881D3E44EFFB3848019AE2936FF73" bold="true" box="[151,1125,151,178]" fontSize="11" level="1" pageId="15" pageNumber="130" reason="1">
<emphasis id="1DCBEAADE44EFFB3848019AE2936FF73" bold="true" box="[151,1125,151,178]" pageId="15" pageNumber="130">
<taxonomicName id="E8BF4D3CE44EFFB3848019AE2EA8FF73" ID-CoL="8M7QS" authority="Muilwijk, Tahami &amp; Lohaj" authorityName="Tahami &amp; Muilwijk &amp; Lohaj &amp; Sadeghi" authorityYear="2017" box="[151,1019,151,178]" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="130" phylum="Arthropoda" rank="species" species="troglophilus" status="sp. nov." subGenus="Iranosphodrus">
<emphasis id="1DCBEAADE44EFFB3848019AE2FCCFF70" bold="true" box="[151,671,151,177]" italics="true" pageId="15" pageNumber="130">Laemostenus (Iranosphodrus) troglophilus</emphasis>
Muilwijk, Tahami &amp; Lohaj
</taxonomicName>
,
<taxonomicNameLabel id="06F857D6E44EFFB3801019A12936FF73" box="[1031,1125,152,178]" pageId="15" pageNumber="130" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="2F0036BFE44EFFB3848019832CFFFF12" blockId="15.[151,1125,151,211]" box="[151,428,186,211]" pageId="15" pageNumber="130">
(
<figureCitation id="B7842A3AE44EFFB3848819832DAFFF12" box="[159,252,186,211]" captionStart="FIGURES 22 – 23" captionStartId="13.[151,264,1821,1844]" captionTargetBox="[197,1297,967,1777]" captionTargetId="figure@13.[158,1428,914,1797]" captionTargetPageId="13" captionText="FIGURES 22 – 23. Habitus in dorsal view of Laemostenus yazdensis (holotype) (22), Laemostenus troglophilus (holotype) (23)." httpUri="https://zenodo.org/record/1042331/files/figure.png" pageId="15" pageNumber="130">Figs. 23</figureCitation>
,
<figureCitation id="B7842A3AE44EFFB3851F19832C75FF12" box="[264,294,186,211]" captionStart="FIGURES 24 – 25" captionStartId="14.[151,264,1051,1074]" captionTargetBox="[249,1277,230,982]" captionTargetId="figure@14.[151,1436,193,1062]" captionTargetPageId="14" captionText="FIGURES 24 – 25. Mesosternum of Laemostenus yazdensis, (24), Laemostenus troglophilus (25)." httpUri="https://zenodo.org/record/1042333/files/figure.png" pageId="15" pageNumber="130">25</figureCitation>
,
<figureCitation id="B7842A3AE44EFFB3852519822C28FF12" box="[306,379,187,211]" captionStart="FIGURES 26 – 29" captionStartId="15.[151,264,1898,1921]" captionTargetBox="[190,1423,1111,1837]" captionTargetId="figure@15.[151,1436,1022,1874]" captionTargetPageId="15" captionText="FIGURES 26 – 29. Aedeagus. Iin lateral view, Laemostenus yazdensis (26), troglophilus speleophilus (27). In dorsal view,. Laemostenus yazdensis (28), Laemostenus troglophilus (29)." httpUri="https://zenodo.org/record/1042335/files/figure.png" pageId="15" pageNumber="130">27, 29</figureCitation>
,
<figureCitation id="B7842A3AE44EFFB3859119822CF0FF12" box="[390,419,187,211]" captionStart="FIGURES 30 – 31" captionStartId="16.[151,264,1169,1192]" captionTargetBox="[252,1272,220,1090]" captionTargetId="figure@16.[219,1367,131,1142]" captionTargetPageId="16" captionText="FIGURES 30 – 31. Ovipositor. Laemostenus yazdensis (30), Laemostenus troglophilus (31)." httpUri="https://zenodo.org/record/1042337/files/figure.png" pageId="15" pageNumber="130">31</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="67A56534E44EFFB38480183B2CDFFE6B" pageId="15" pageNumber="130" type="materials_examined">
<paragraph id="2F0036BFE44EFFB38480183B2CDFFE6B" blockId="15.[151,1437,258,967]" pageId="15" pageNumber="130">
<emphasis id="1DCBEAADE44EFFB38480183B2C15FEDA" bold="true" box="[151,326,258,283]" pageId="15" pageNumber="130">Type material.</emphasis>
Holotype male, labelled: “Iran, Yazd, Tange Chenar, Nafas Cave;
<date id="5B01107FE44EFFB3802D183B2996FEDA" box="[1082,1221,258,283]" pageId="15" pageNumber="130" value="2014-08-28">28.viii.2014</date>
; leg. M.S. Tahami, H. Entezari, M. Sheykh Rasti, Meshki”/
<taxonomicName id="E8BF4D3CE44EFFB38675181E2ED0FEFE" authority="Tahami &amp; Muilwijk &amp; Lohaj &amp; Sadeghi, 2017" authorityName="Tahami &amp; Muilwijk &amp; Lohaj &amp; Sadeghi" authorityYear="2017" box="[610,899,294,319]" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="130" phylum="Arthropoda" rank="species" species="troglophilus" status="sp. nov.">
<emphasis id="1DCBEAADE44EFFB38675181E2ED0FEFE" box="[610,899,294,319]" italics="true" pageId="15" pageNumber="130">Laemostenus troglophilus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="06F857D6E44EFFB3879B181F2EB5FEFE" box="[908,998,294,319]" pageId="15" pageNumber="130" rank="species">
<emphasis id="1DCBEAADE44EFFB3879B181F2EB5FEFE" bold="true" box="[908,998,294,319]" pageId="15" pageNumber="130">sp. nov.</emphasis>
</taxonomicNameLabel>
Muilwijk, Tahami &amp; Lohaj; 2016 (red label Holotype). Paratypes: one male and one female, the same data as holotype /
<taxonomicName id="E8BF4D3CE44EFFB3805A18752822FEA2" authority="Tahami &amp; Muilwijk &amp; Lohaj &amp; Sadeghi, 2017" authorityName="Tahami &amp; Muilwijk &amp; Lohaj &amp; Sadeghi" authorityYear="2017" box="[1101,1393,330,355]" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="130" phylum="Arthropoda" rank="species" species="troglophilus" status="sp. nov.">
<emphasis id="1DCBEAADE44EFFB3805A18752822FEA2" box="[1101,1393,330,355]" italics="true" pageId="15" pageNumber="130">Laemostenus troglophilus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="06F857D6E44EFFB3816D18732D9BFE46" pageId="15" pageNumber="130" rank="species">
<emphasis id="1DCBEAADE44EFFB3816D18732D9BFE46" bold="true" pageId="15" pageNumber="130">sp. nov.</emphasis>
</taxonomicNameLabel>
Muilwijk, Tahami &amp; Lohaj 2016 (red label Paratype). Holotype and one paratype deposited in ZM–CBSU, one paratype in cMU.
</paragraph>
</subSubSection>
<subSubSection id="67A56534E44EFFB384D0188F2E2DFDD6" pageId="15" pageNumber="130" type="diagnosis">
<paragraph id="2F0036BFE44EFFB384D0188F2E2DFDD6" blockId="15.[151,1437,258,967]" pageId="15" pageNumber="130">
<emphasis id="1DCBEAADE44EFFB384D0188F2C12FE0E" bold="true" box="[199,321,438,463]" pageId="15" pageNumber="130">Diagnosis.</emphasis>
A medium sized, brachypterous
<taxonomicName id="E8BF4D3CE44EFFB386A4188E2E16FE0F" authorityName="Tahami &amp; Muilwijk &amp; Lohaj &amp; Sadeghi" authorityYear="2017" box="[691,837,439,462]" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="130" phylum="Arthropoda" rank="genus">
<emphasis id="1DCBEAADE44EFFB386A4188E2E16FE0F" box="[691,837,439,462]" italics="true" pageId="15" pageNumber="130">Laemostenus</emphasis>
</taxonomicName>
sg.
<emphasis id="1DCBEAADE44EFFB38764188F294BFE0E" box="[883,1048,438,463]" italics="true" pageId="15" pageNumber="130">Iranosphodrus</emphasis>
species. Upper side including legs and antennae dull black, female
<typeStatus id="F004881DE44EFFB3863318E22FD7FE32" box="[548,644,475,499]" pageId="15" pageNumber="130" type="paratype">paratype</typeStatus>
with bluish shine on elytra. Pronotum and head shiny, with fine microsculpture, elytra matt, with distinct microsculpture (
<figureCitation id="B7842A3AE44EFFB3870A18C72E22FDD6" box="[797,881,510,535]" captionStart="FIGURES 22 – 23" captionStartId="13.[151,264,1821,1844]" captionTargetBox="[197,1297,967,1777]" captionTargetId="figure@13.[158,1428,914,1797]" captionTargetPageId="13" captionText="FIGURES 22 – 23. Habitus in dorsal view of Laemostenus yazdensis (holotype) (22), Laemostenus troglophilus (holotype) (23)." httpUri="https://zenodo.org/record/1042331/files/figure.png" pageId="15" pageNumber="130">Fig. 23</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="67A56534E44EFFAC84D01B1B2CB6FA29" lastPageId="16" lastPageNumber="131" pageId="15" pageNumber="130" type="description">
<paragraph id="2F0036BFE44EFFB384D01B1B290CFD0A" blockId="15.[151,1437,258,967]" pageId="15" pageNumber="130">
<emphasis id="1DCBEAADE44EFFB384D01B1B2C0AFDFA" bold="true" box="[199,345,546,571]" pageId="15" pageNumber="130">Description.</emphasis>
BL 15.8–16.0 (
<typeStatus id="F004881DE44EFFB3861E1B1B2F3DFDFA" box="[521,622,546,571]" pageId="15" pageNumber="130" type="holotype">holotype</typeStatus>
) mm. Head narrow, parallel-sided, distinctly longer than wide (HL/HW: 1.1), with only posterior pair of supraorbital setae; frontal furrows short, but distinct; eyes flat, reduced (
<quantity id="E8479B5AE44EFFB381231B7F28C3FD9E" box="[1332,1424,582,607]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="15" pageNumber="130" unit="mm" value="0.5">0.5 mm</quantity>
), shorter than temples (1.0 mm). Clypeus with two long setae; labrum with six setae. Antennae in
<typeStatus id="F004881DE44EFFB380C21B532864FD42" box="[1237,1335,618,643]" pageId="15" pageNumber="130" type="holotype">holotype</typeStatus>
missing, antennae in
<typeStatus id="F004881DE44EFFB385321BB62CC3FD66" box="[293,400,655,679]" pageId="15" pageNumber="130" type="paratype">paratypes</typeStatus>
rather short, reaching anterior eighth of elytral length, antennomere 3 without accessory pubescence except a few apical setae, antennomeres 4–11 pubescent. Mandibles short.
</paragraph>
<paragraph id="2F0036BFE44EFFB384D01BEE29D2FCF6" blockId="15.[151,1437,258,967]" pageId="15" pageNumber="130">Pronotum cordiform, distinctly wider than head, as long as wide (PL/ PW: 1.0). Basal impressions short, sparsely punctuated. Lateral bead with superficial punctuation, two antero-lateral setae present, basolateral setae missing. Anterior angles prominent, at apex rounded, posterior angles almost rectangular.</paragraph>
<paragraph id="2F0036BFE44EFFB384D01A7B280DFC07" blockId="15.[151,1437,258,967]" pageId="15" pageNumber="130">Elytra ovate elongate, relatively broad (EL/EW: 1.6–1.7), with maximum width at apical third; disc with small antero-medial depression; base wider than pronotal base. Humeri moderately rounded, with a small tooth. Striae fine, with punctures, intervals flat. Scutellar striae present; scutellar setiferous punctures situated in striae 1. Umbilicate series consists of 18 setiferous punctures, more interrupted in middle; two setae at apex of stria 7.</paragraph>
<caption id="7BC06637E44EFFB384801E532FAEF861" httpUri="https://zenodo.org/record/1042335/files/figure.png" pageId="15" pageNumber="130" targetBox="[190,1423,1111,1837]" targetPageId="15">
<paragraph id="2F0036BFE44EFFB384801E532FAEF861" blockId="15.[151,1436,1898,1952]" pageId="15" pageNumber="130">
<emphasis id="1DCBEAADE44EFFB384801E532C05F841" bold="true" box="[151,342,1898,1921]" pageId="15" pageNumber="130">FIGURES 26–29.</emphasis>
Aedeagus. Iin lateral view,
<taxonomicName id="E8BF4D3CE44EFFB386961E552E3CF840" authorityName="Tahami &amp; Muilwijk &amp; Lohaj &amp; Sadeghi" authorityYear="2017" box="[641,879,1899,1921]" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="130" phylum="Arthropoda" rank="species" species="yazdensis">
<emphasis id="1DCBEAADE44EFFB386961E552E3CF840" box="[641,879,1899,1921]" italics="true" pageId="15" pageNumber="130">Laemostenus yazdensis</emphasis>
</taxonomicName>
(26),
<emphasis id="1DCBEAADE44EFFB387A51E5229E6F840" box="[946,1205,1899,1921]" italics="true" pageId="15" pageNumber="130">
<taxonomicName id="E8BF4D3CE44EFFB387A51E52297DF840" authorityName="Tahami &amp; Muilwijk &amp; Lohaj &amp; Sadeghi" authorityYear="2017" box="[946,1070,1899,1921]" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="130" phylum="Arthropoda" rank="species" species="troglophilus">troglophilus</taxonomicName>
<taxonomicName id="E8BF4D3CE44EFFB380201E5229E6F840" authorityName="Tahami &amp; Muilwijk &amp; Lohaj &amp; Sadeghi" authorityYear="2017" box="[1079,1205,1899,1921]" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="130" phylum="Arthropoda" rank="species" species="speleophilus">speleophilus</taxonomicName>
</emphasis>
(27). In dorsal view,.
<taxonomicName id="E8BF4D3CE44EFFB384801EB22CD1F861" authorityName="Tahami &amp; Muilwijk &amp; Lohaj &amp; Sadeghi" authorityYear="2017" box="[151,386,1930,1952]" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="130" phylum="Arthropoda" rank="species" species="yazdensis">
<emphasis id="1DCBEAADE44EFFB384801EB22CD1F861" box="[151,386,1930,1952]" italics="true" pageId="15" pageNumber="130">Laemostenus yazdensis</emphasis>
</taxonomicName>
(28),
<taxonomicName id="E8BF4D3CE44EFFB385A81EB22F95F861" authorityName="Tahami &amp; Muilwijk &amp; Lohaj &amp; Sadeghi" authorityYear="2017" box="[447,710,1930,1952]" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="130" phylum="Arthropoda" rank="species" species="troglophilus">
<emphasis id="1DCBEAADE44EFFB385A81EB22F95F861" box="[447,710,1930,1952]" italics="true" pageId="15" pageNumber="130">Laemostenus troglophilus</emphasis>
</taxonomicName>
(29).
</paragraph>
</caption>
<caption id="7BC06637E451FFAC84801DA8296AFB66" box="[151,1081,1169,1192]" httpUri="https://zenodo.org/record/1042337/files/figure.png" pageId="16" pageNumber="131" targetBox="[252,1272,220,1090]" targetPageId="16">
<paragraph id="2F0036BFE451FFAC84801DA8296AFB66" blockId="16.[151,1081,1169,1192]" box="[151,1081,1169,1192]" pageId="16" pageNumber="131">
<emphasis id="1DCBEAADE451FFAC84801DA82C00FB67" bold="true" box="[151,339,1169,1192]" pageId="16" pageNumber="131">FIGURES 30–31.</emphasis>
Ovipositor.
<taxonomicName id="E8BF4D3CE451FFAC85C41DAB2FEDFB66" authorityName="Tahami &amp; Muilwijk &amp; Lohaj &amp; Sadeghi" authorityYear="2017" box="[467,702,1169,1191]" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="131" phylum="Arthropoda" rank="species" species="yazdensis">
<emphasis id="1DCBEAADE451FFAC85C41DAB2FEDFB66" box="[467,702,1169,1191]" italics="true" pageId="16" pageNumber="131">Laemostenus yazdensis</emphasis>
</taxonomicName>
(30),
<taxonomicName id="E8BF4D3CE451FFAC86EC1DAB2952FB66" authorityName="Tahami &amp; Muilwijk &amp; Lohaj &amp; Sadeghi" authorityYear="2017" box="[763,1025,1169,1191]" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="131" phylum="Arthropoda" rank="species" species="troglophilus">
<emphasis id="1DCBEAADE451FFAC86EC1DAB2952FB66" box="[763,1025,1169,1191]" italics="true" pageId="16" pageNumber="131">Laemostenus troglophilus</emphasis>
</taxonomicName>
(31).
</paragraph>
</caption>
<paragraph id="2F0036BFE451FFAC84D01DEC292DFAD0" blockId="16.[151,1437,1236,1765]" pageId="16" pageNumber="131">
Mesosternum with small teeth before mesocoxae, in male
<typeStatus id="F004881DE451FFAC874B1DEC2EEFFB2C" box="[860,956,1237,1261]" pageId="16" pageNumber="131" type="paratype">paratype</typeStatus>
reduced. Metepisternum longer than wide (
<figureCitation id="B7842A3AE451FFAC84881DC12DA0FAD0" box="[159,243,1272,1297]" captionStart="FIGURES 24 – 25" captionStartId="14.[151,264,1051,1074]" captionTargetBox="[249,1277,230,982]" captionTargetId="figure@14.[151,1436,193,1062]" captionTargetPageId="14" captionText="FIGURES 24 – 25. Mesosternum of Laemostenus yazdensis, (24), Laemostenus troglophilus (25)." httpUri="https://zenodo.org/record/1042333/files/figure.png" pageId="16" pageNumber="131">Fig. 25</figureCitation>
). Abdominal sternites black, with superficial wrinkles and dense microsculpture.
</paragraph>
<paragraph id="2F0036BFE451FFAC84D01C24280FFABC" blockId="16.[151,1437,1236,1765]" pageId="16" pageNumber="131">Legs short. Meso- and metatibiae almost straight, not curved, with short ventral brush of setae near apex. Profemori without teeth at ventral sides. Protibiae with sparse, short hairs between cleansing apparatus and apical setae. Tarsi with decumbent pubescence at dorsal sides; claws with small teeth, male protarsi slightly dilated.</paragraph>
<paragraph id="2F0036BFE451FFAC84D01CB12E8BFA04" blockId="16.[151,1437,1236,1765]" pageId="16" pageNumber="131">
Aedeagus: (
<figureCitation id="B7842A3AE451FFAC855B1CB12C81FA61" box="[332,466,1416,1441]" captionStart="FIGURES 26 – 29" captionStartId="15.[151,264,1898,1921]" captionTargetBox="[190,1423,1111,1837]" captionTargetId="figure@15.[151,1436,1022,1874]" captionTargetPageId="15" captionText="FIGURES 26 – 29. Aedeagus. Iin lateral view, Laemostenus yazdensis (26), troglophilus speleophilus (27). In dorsal view,. Laemostenus yazdensis (28), Laemostenus troglophilus (29)." httpUri="https://zenodo.org/record/1042335/files/figure.png" pageId="16" pageNumber="131">Figs. 27, 29</figureCitation>
) median lobe of aedeagus short (
<quantity id="E8479B5AE451FFAC87541CB12EFFFA61" box="[835,940,1416,1440]" metricMagnitude="-3" metricUnit="m" metricValue="1.93" pageId="16" pageNumber="131" unit="mm" value="1.93">1.93 mm</quantity>
), slightly arcuate, regularly narrowing to the apex in lateral view; dorsal view, apex moderately rounded in dorsal view.
</paragraph>
<paragraph id="2F0036BFE451FFAC84D01CE92CB6FA29" blockId="16.[151,1437,1236,1765]" box="[199,485,1488,1513]" pageId="16" pageNumber="131">
Ovipositor: as on
<figureCitation id="B7842A3AE451FFAC85981CE92CB2FA29" box="[399,481,1488,1513]" captionStart="FIGURES 30 – 31" captionStartId="16.[151,264,1169,1192]" captionTargetBox="[252,1272,220,1090]" captionTargetId="figure@16.[219,1367,131,1142]" captionTargetPageId="16" captionText="FIGURES 30 – 31. Ovipositor. Laemostenus yazdensis (30), Laemostenus troglophilus (31)." httpUri="https://zenodo.org/record/1042337/files/figure.png" pageId="16" pageNumber="131">Fig. 31</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="67A56534E451FFAC84D01CCD2C9EF9B8" pageId="16" pageNumber="131" type="diagnosis">
<paragraph id="2F0036BFE451FFAC84D01CCD2C9EF9B8" blockId="16.[151,1437,1236,1765]" pageId="16" pageNumber="131">
<emphasis id="1DCBEAADE451FFAC84D01CCD2C9DF9CC" bold="true" box="[199,462,1524,1549]" pageId="16" pageNumber="131">Differential diagnosis.</emphasis>
<taxonomicName id="E8BF4D3CE451FFAC85C31CCC2FF7F9CC" authority="Tahami &amp; Muilwijk &amp; Lohaj &amp; Sadeghi, 2017" authorityName="Tahami &amp; Muilwijk &amp; Lohaj &amp; Sadeghi" authorityYear="2017" box="[468,676,1524,1549]" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="131" phylum="Arthropoda" rank="species" species="troglophilus" status="sp. nov." subGenus="Iranosphodrus">
<emphasis id="1DCBEAADE451FFAC85C31CCC2FF7F9CC" box="[468,676,1524,1549]" italics="true" pageId="16" pageNumber="131">L. (I.) troglophilus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="06F857D6E451FFAC86BB1CCD2E56F9CC" box="[684,773,1524,1549]" pageId="16" pageNumber="131" rank="species">
<emphasis id="1DCBEAADE451FFAC86BB1CCD2E56F9CC" bold="true" box="[684,773,1524,1549]" pageId="16" pageNumber="131">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from
<taxonomicName id="E8BF4D3CE451FFAC878D1CCC2971F9CC" authority="Tahami &amp; Muilwijk &amp; Lohaj &amp; Sadeghi, 2017" authorityName="Tahami &amp; Muilwijk &amp; Lohaj &amp; Sadeghi" authorityYear="2017" box="[922,1058,1524,1549]" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="131" phylum="Arthropoda" rank="species" species="yazdensis" status="sp. nov.">
<emphasis id="1DCBEAADE451FFAC878D1CCC2971F9CC" box="[922,1058,1524,1549]" italics="true" pageId="16" pageNumber="131">L. yazdensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="06F857D6E451FFAC803D1CCD29D7F9CC" box="[1066,1156,1524,1549]" pageId="16" pageNumber="131" rank="species">
<emphasis id="1DCBEAADE451FFAC803D1CCD29D7F9CC" bold="true" box="[1066,1156,1524,1549]" pageId="16" pageNumber="131">sp. nov.</emphasis>
</taxonomicNameLabel>
by the presence of small mesosternal tooth, absence of tooth on profemori, by the shape of pronotum and metepisternum as well as median lobe of aedeagus. Both new species differs from
<taxonomicName id="E8BF4D3CE451FFAC86AF1F042E30F994" authorityName="Tahami &amp; Muilwijk &amp; Lohaj &amp; Sadeghi" authorityYear="2017" box="[696,867,1596,1621]" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="131" phylum="Arthropoda" rank="species" species="rudichae" subGenus="Iranosphodrus">
<emphasis id="1DCBEAADE451FFAC86AF1F042E30F994" box="[696,867,1596,1621]" italics="true" pageId="16" pageNumber="131">L. (I.) rudichae</emphasis>
</taxonomicName>
by straight mesotibiae, more oval elytra, and more short and stout appendages.
</paragraph>
</subSubSection>
<subSubSection id="67A56534E451FFAC84D01FBD2862F95D" box="[199,1329,1668,1693]" pageId="16" pageNumber="131" type="distribution">
<paragraph id="2F0036BFE451FFAC84D01FBD2862F95D" blockId="16.[151,1437,1236,1765]" box="[199,1329,1668,1693]" pageId="16" pageNumber="131">
<emphasis id="1DCBEAADE451FFAC84D01FBD2CA2F95C" bold="true" box="[199,497,1668,1693]" pageId="16" pageNumber="131">Distribution and habitat.</emphasis>
So far known only from the twilight zone of the
<typeStatus id="F004881DE451FFAC80011FBC2915F95C" box="[1046,1094,1669,1693]" pageId="16" pageNumber="131">type</typeStatus>
locality, Nafas cave.
</paragraph>
</subSubSection>
<subSubSection id="67A56534E451FFAC84D01F912F26F924" pageId="16" pageNumber="131" type="etymology">
<paragraph id="2F0036BFE451FFAC84D01F912F26F924" blockId="16.[151,1437,1236,1765]" pageId="16" pageNumber="131">
<emphasis id="1DCBEAADE451FFAC84D01F912C1DF900" bold="true" box="[199,334,1704,1729]" pageId="16" pageNumber="131">Etymology.</emphasis>
The name is taken from the Greek origin, “
<emphasis id="1DCBEAADE451FFAC872B1F912ED3F900" box="[828,896,1704,1729]" italics="true" pageId="16" pageNumber="131">trōglo</emphasis>
” means “cave”, and “
<emphasis id="1DCBEAADE451FFAC806C1F9129ECF900" box="[1147,1215,1704,1729]" italics="true" pageId="16" pageNumber="131">philus</emphasis>
” means “liking” or “attracted to”, generally means cave-lover.
</paragraph>
</subSubSection>
</treatment>
</document>
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