treatments-xml/data/8D/7A/87/8D7A87A6FFE7FFA51F50B8B2AFB2824C.xml

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<mods:title id="6BBA096AF77E63CBDC349BEA8B481ABC">Review of the Hemiosus Sharp and Derallus Sharp of North America, Central America, and the Caribbean (Coleoptera: Hydrophilidae: Berosini)</mods:title>
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<mods:namePart id="EDFA41E5CF86C9F481D33F01AA11B22A">Short, Andrew Edward Z.</mods:namePart>
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<mods:namePart id="8A73A0A906FEC351F3525262ACC530C3">Torres, Phillip J.</mods:namePart>
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<mods:date id="C745EF3168AF4D1593E8454981B56D55">2006</mods:date>
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<subSubSection id="4DC9653BFFE7FFAA1F50B8B2AA6884BC" pageId="7" pageNumber="8" type="nomenclature">
<paragraph id="056C36B0FFE7FFAA1F50B8B2A99F8499" blockId="7.[264,663,1153,1214]" box="[264,663,1153,1179]" pageId="7" pageNumber="8">
<heading id="5E2481DCFFE7FFAA1F50B8B2A99F8499" bold="true" box="[264,663,1153,1179]" fontSize="11" level="1" pageId="7" pageNumber="8" reason="1">
<taxonomicName id="C2D34D33FFE7FFAA1F50B8B2A99F8499" ID-CoL="3KR6Z" authority="LeConte, 1852" authorityName="LeConte" authorityYear="1852" box="[264,663,1153,1179]" class="Insecta" family="Hydrophilidae" genus="Hemiosus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="exilis">
<emphasis id="37A7EAA2FFE7FFAA1F50B8B2A99F8499" bold="true" box="[264,663,1153,1179]" pageId="7" pageNumber="8">
<emphasis id="37A7EAA2FFE7FFAA1F50B8B2AACF8499" bold="true" box="[264,455,1153,1179]" italics="true" pageId="7" pageNumber="8">Hemiosus exilis</emphasis>
(
<bibRefCitation id="61424B41FFE7FFAA1F8FB8B2A9878499" author="LeConte" box="[471,655,1153,1179]" pageId="7" pageNumber="16" refString="LeConte, J. L. (1852). Descriptions of New Species of Coleoptera, from California. Annals of the Lyceum of natural History of New York, 5, 125 - 216." type="journal article" year="1852">
LeConte, 
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)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="056C36B0FFE7FFAA1F50B897AA6884BC" blockId="7.[264,663,1153,1214]" box="[264,352,1188,1214]" pageId="7" pageNumber="8">
(
<figureCitation id="9DE82A35FFE7FFAA1F49B897AA5F84BC" box="[273,343,1188,1214]" captionStart="FIGURES 3 – 5" captionStartId="7.[264,382,1004,1028]" captionTargetBox="[418,1160,325,975]" captionTargetId="figure@7.[409,1177,304,994]" captionTargetPageId="7" captionText="FIGURES 3 – 5. Details of Hemiosus selva Short, sp. n. (3, 5) and H. exilis (LeConte) (4). 3 – 4: pronotal coloration, scale bar 0.18 mm; 5: aedeagus, scale bar 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.174868" httpUri="https://zenodo.org/record/174868/files/figure.png" pageId="7" pageNumber="8">
Fig. 
<number id="B31B2497FFE7FFAA1F1FB897AA5F84BC" box="[327,343,1188,1214]" pageId="7" pageNumber="8" value="4.0">4</number>
</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="4DC9653BFFE7FFAA1F50B8DFA8718541" pageId="7" pageNumber="8" type="reference_group">
<paragraph id="056C36B0FFE7FFAA1F50B8DFA98C8501" blockId="7.[264,930,1260,1347]" box="[264,644,1260,1283]" pageId="7" pageNumber="8">
<treatmentCitationGroup id="25C3119EFFE7FFAA1F50B8DFA98C8501" box="[264,644,1260,1283]" pageId="7" pageNumber="8">
<taxonomicName id="C2D34D33FFE7FFAA1F50B8DFA9888501" ID-CoL="68JVB" authority="LeConte, 1852: 211" authorityName="LeConte" authorityPageNumber="211" authorityYear="1852" box="[264,640,1260,1283]" class="Insecta" family="Hydrophilidae" genus="Berosus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="exilis">
<emphasis id="37A7EAA2FFE7FFAA1F50B8DFAA948501" box="[264,412,1260,1283]" italics="true" pageId="7" pageNumber="8">Berosus exilis</emphasis>
<treatmentCitation id="847210A1FFE7FFAA1FFBB8DFA9888501" author="LeConte" box="[419,640,1260,1283]" page="211" pageId="7" pageNumber="8" year="1852">
<bibRefCitation id="61424B41FFE7FFAA1FFBB8DFA94D8501" author="LeConte" box="[419,581,1260,1283]" pageId="7" pageNumber="16" refString="LeConte, J. L. (1852). Descriptions of New Species of Coleoptera, from California. Annals of the Lyceum of natural History of New York, 5, 125 - 216." type="journal article" year="1852">
LeConte, 
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</bibRefCitation>
: 
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</taxonomicName>
.
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</paragraph>
<paragraph id="056C36B0FFE7FFAA1F50B93FA8718541" blockId="7.[264,930,1260,1347]" pageId="7" pageNumber="8">
<treatmentCitationGroup id="25C3119EFFE7FFAA1F50B93FA8718541" pageId="7" pageNumber="8">
<taxonomicName id="C2D34D33FFE7FFAA1F50B93FA9D78521" ID-CoL="6LP93" authority="d'Orchymont, 1940: 181" authorityName="d'Orchymont" authorityPageNumber="181" authorityYear="1940" box="[264,735,1292,1315]" class="Insecta" family="Hydrophilidae" genus="Hemiosus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="toxillus">
<emphasis id="37A7EAA2FFE7FFAA1F50B93FAACF8521" box="[264,455,1292,1315]" italics="true" pageId="7" pageNumber="8">Hemiosus toxillus</emphasis>
d’Orchymont, 
<number id="B31B2497FFE7FFAA1C32B93FA9AB8521" box="[618,675,1292,1315]" pageId="7" pageNumber="8" value="1940.0">1940</number>
: 
<number id="B31B2497FFE7FFAA1CECB93FA9D78521" box="[692,735,1292,1315]" pageId="7" pageNumber="8" value="181.0">181</number>
</taxonomicName>
. 
<emphasis id="37A7EAA2FFE7FFAA1CA8B93FA8AA8526" bold="true" box="[752,930,1292,1316]" pageId="7" pageNumber="8">New synonomy.</emphasis>
<taxonomicName id="C2D34D33FFE7FFAA1F50B91FA9738541" authority="Leech, 1943: 61" authorityName="Leech" authorityPageNumber="61" authorityYear="1943" box="[264,635,1324,1347]" class="Insecta" family="Hydrophilidae" genus="Hemiosus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="exilis">
<emphasis id="37A7EAA2FFE7FFAA1F50B91FAAB88541" box="[264,432,1324,1347]" italics="true" pageId="7" pageNumber="8">Hemiosus exilis</emphasis>
: 
<treatmentCitation id="847210A1FFE7FFAA1F9DB91FA9738541" author="Leech" box="[453,635,1324,1347]" page="61" pageId="7" pageNumber="8" year="1943">
<bibRefCitation id="61424B41FFE7FFAA1F9DB91FA9458541" author="Leech" box="[453,589,1324,1347]" pageId="7" pageNumber="16" refString="Leech, H. B. (1943). Berosus exilis LeConte a Hemiosus (Coleoptera: Hydrophilidae). Pan-Pacific Entomologist, 19, 61 - 62." type="journal article" year="1943">
Leech, 
<number id="B31B2497FFE7FFAA1C4CB91FA9458541" box="[532,589,1324,1347]" pageId="7" pageNumber="8" value="1943.0">1943</number>
</bibRefCitation>
: 
<number id="B31B2497FFE7FFAA1C06B91FA9738541" box="[606,635,1324,1347]" pageId="7" pageNumber="8" value="61.0">61</number>
</treatmentCitation>
</taxonomicName>
; 
<treatmentCitation id="847210A1FFE7FFAA1CDEB91FA87D8541" author="Van" box="[646,885,1324,1347]" page="54" pageId="7" pageNumber="8" year="1964">
<bibRefCitation id="61424B41FFE7FFAA1CDEB91FA84E8541" author="Van" box="[646,838,1324,1347]" pageId="7" pageNumber="17" refString="Van Tassell, E. R. (1964). A note on Hemiosus exilis LeConte (Coleoptera: Hydrophilidae). Coleopterists Bulletin, 18, 53 - 56." type="journal article" year="1964">
Van Tassell, 
<number id="B31B2497FFE7FFAA1D55B91FA84E8541" box="[781,838,1324,1347]" pageId="7" pageNumber="8" value="1964.0">1964</number>
</bibRefCitation>
: 
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</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="4DC9653BFFE7FFAA1F50B947A8AA871C" pageId="7" pageNumber="8" type="materials_examined">
<paragraph id="056C36B0FFE7FFAA1F50B947A9E085B4" blockId="7.[264,1323,1396,1902]" pageId="7" pageNumber="8">
<emphasis id="37A7EAA2FFE7FFAA1F50B947A937858C" bold="true" box="[264,575,1396,1422]" pageId="7" pageNumber="8">
<typeStatus id="DA688812FFE7FFAA1F50B947AA4D858C" box="[264,325,1396,1422]" pageId="7" pageNumber="8">Type</typeStatus>
material examined.
</emphasis>
<taxonomicName id="C2D34D33FFE7FFAA1C17B947A86C858C" authority="LeConte" authorityName="LeConte" box="[591,868,1396,1422]" class="Insecta" family="Hydrophilidae" genus="Berosus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="exilis">
<emphasis id="37A7EAA2FFE7FFAA1C17B947A9FA858F" box="[591,754,1396,1421]" italics="true" pageId="7" pageNumber="8">Berosus exilis</emphasis>
LeConte
</taxonomicName>
: 
<typeStatus id="DA688812FFE7FFAA1D2FB947A8EC858C" box="[887,996,1396,1422]" pageId="7" pageNumber="8">Holotype</typeStatus>
(male): [colored disc], “ 
<taxonomicName id="C2D34D33FFE7FFAA1B57B947AA4E85B4" class="Insecta" family="Hydrophilidae" genus="Berosus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="exilis">B. exilis</taxonomicName>
/ Lec./ Gila”, “ 
<typeStatus id="DA688812FFE7FFAA1FA9B9AFA92585B4" box="[497,557,1436,1462]" pageId="7" pageNumber="8">Type</typeStatus>
/ 
<number id="B31B2497FFE7FFAA1C66B9AFA97385B4" box="[574,635,1436,1462]" pageId="7" pageNumber="8" value="3081.0">3081</number>
” (
<collectionCode id="63C2AE75FFE7FFAA1CCDB9AFA9D385B4" LSID="urn:lsid:biocol.org:col:33791" box="[661,731,1436,1462]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="7" pageNumber="8">MCZ</collectionCode>
).
</paragraph>
<paragraph id="056C36B0FFE7FFAA1F60B9F7AF6D8654" blockId="7.[264,1323,1396,1902]" pageId="7" pageNumber="8">
<taxonomicName id="C2D34D33FFE7FFAA1F60B9F7A9BA85DC" authority="d'Orchymont" authorityName="d'Orchymont" box="[312,690,1476,1502]" class="Insecta" family="Hydrophilidae" genus="Hemiosus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="toxillus">
<emphasis id="37A7EAA2FFE7FFAA1F60B9F7A90085DF" box="[312,520,1476,1501]" italics="true" pageId="7" pageNumber="8">Hemiosus toxillus</emphasis>
d’Orchymont
</taxonomicName>
: 
<typeStatus id="DA688812FFE7FFAA1C9BB9F7A82185DC" box="[707,809,1476,1502]" pageId="7" pageNumber="8">holotype</typeStatus>
(male): “[male symbol]”, “Mex. Mazatlan/ brack, pools 
<date id="716D1070FFE7FFAA1FF5B9DFA9388604" box="[429,560,1516,1542]" pageId="7" pageNumber="8" value="1934-05-25">
<number id="B31B2497FFE7FFAA1FF5B9DFAAC18604" box="[429,457,1516,1542]" pageId="7" pageNumber="8" value="25.0">25</number>
/ v. 
<number id="B31B2497FFE7FFAA1FACB9DFA9388604" box="[500,560,1516,1542]" pageId="7" pageNumber="8" value="1934.0">1934</number>
</date>
H. Hinton”, “A. d’Orchymont det./ 
<taxonomicName id="C2D34D33FFE7FFAA1A5DB9DFAF718604" box="[1029,1145,1516,1542]" class="Insecta" family="Hydrophilidae" genus="Hemiosus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">Hemiosus</taxonomicName>
/ 
<taxonomicName id="C2D34D33FFE7FFAA1AC9B9DFAFE18604" box="[1169,1257,1516,1542]" class="Insecta" family="Hydrophilidae" genus="Hemiosus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="toxillus">toxillus</taxonomicName>
m.”, “ 
<typeStatus id="DA688812FFE7FFAA1F4ABA27AA56862C" box="[274,350,1556,1582]" pageId="7" pageNumber="8">
<collectionCode id="63C2AE75FFE7FFAA1F4ABA27AA56862C" box="[274,350,1556,1582]" pageId="7" pageNumber="8">TYPE</collectionCode>
</typeStatus>
[red card]” (
<collectionCode id="63C2AE75FFE7FFAA1FA4BA27A951862C" box="[508,601,1556,1582]" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="7" pageNumber="8">IRSNB</collectionCode>
). 
<typeStatus id="DA688812FFE7FFAA1C20BA27A9EB862C" box="[632,739,1556,1582]" pageId="7" pageNumber="8">Paratype</typeStatus>
: “Female symbol”, “Sinaloa/ Mex”, “Para-/ 
<typeStatus id="DA688812FFE7FFAA1AAFBA27AE22862C" box="[1271,1322,1556,1582]" pageId="7" pageNumber="8">type</typeStatus>
[orange card]”, “ 
<taxonomicName id="C2D34D33FFE7FFAA1F93BA0FA9A98654" box="[459,673,1596,1622]" class="Insecta" family="Hydrophilidae" genus="Hemiosus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="toxillus">Hemiosus toxillus</taxonomicName>
/ Orchymont/ Tete Manque” (
<collectionCode id="63C2AE75FFE7FFAA1DA5BA0FAF518654" box="[1021,1113,1596,1622]" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="7" pageNumber="8">IRSNB</collectionCode>
).
</paragraph>
<paragraph id="056C36B0FFE7FFAA1F60BA57A8AA871C" blockId="7.[264,1323,1396,1902]" pageId="7" pageNumber="8">
<emphasis id="37A7EAA2FFE7FFAA1F60BA57AF06867C" bold="true" box="[312,1038,1636,1662]" pageId="7" pageNumber="8">
Additional material examined (
<number id="B31B2497FFE7FFAA1C90BA57A9E0867C" box="[712,744,1636,1662]" pageId="7" pageNumber="8" value="69.0">69</number>
). 
<collectingCountry id="7DC47620FFE7FFAA1D55BA57A885867C" box="[781,909,1636,1662]" name="Mexico" pageId="7" pageNumber="8">MEXICO</collectingCountry>
: Sinaloa:
</emphasis>
Culiacan, 
<date id="716D1070FFE7FFAA1AC6BA57AE2E867C" box="[1182,1318,1636,1662]" pageId="7" pageNumber="8" value="1963-07-16">
<number id="B31B2497FFE7FFAA1AC6BA57AFB4867C" box="[1182,1212,1636,1662]" pageId="7" pageNumber="8" value="16.0">16</number>
.vii. 
<number id="B31B2497FFE7FFAA1ABEBA57AE2E867C" box="[1254,1318,1636,1662]" pageId="7" pageNumber="8" value="1963.0">1963</number>
</date>
, P.J. Spangler leg. (
<number id="B31B2497FFE7FFAA1FB1BABFA90386A4" box="[489,523,1676,1702]" pageId="7" pageNumber="8" value="60.0">60</number>
: 
<collectionCode id="63C2AE75FFE7FFAA1C40BABFA96C86A4" box="[536,612,1676,1702]" pageId="7" pageNumber="8">AEZS</collectionCode>
, 
<collectionCode id="63C2AE75FFE7FFAA1C2BBABFA9B686A4" box="[627,702,1676,1702]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="7" pageNumber="8">NMW</collectionCode>
, 
<collectionCode id="63C2AE75FFE7FFAA1C95BABFA82F86A4" LSID="urn:lsid:biocol.org:col:34871" box="[717,807,1676,1702]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="7" pageNumber="8">USNM</collectionCode>
); 
<emphasis id="37A7EAA2FFE7FFAA1D65BABFA89386A4" bold="true" box="[829,923,1676,1702]" pageId="7" pageNumber="8">Jalisco:</emphasis>
Rio Cuitzmala, SE of Chamela, 
<quantity id="C22B9B55FFE7FFAA1F50BA87AA5B86CC" box="[264,339,1716,1742]" metricMagnitude="1" metricUnit="m" metricValue="5.4864" pageId="7" pageNumber="8" unit="ft" value="180.0">180 ft</quantity>
. asl, 
<date id="716D1070FFE7FFAA1FC8BA87AAFA86CC" box="[400,498,1716,1742]" pageId="7" pageNumber="8" value="2005-01-06">
<number id="B31B2497FFE7FFAA1FC8BA87AA9786CC" box="[400,415,1716,1742]" pageId="7" pageNumber="8" value="6.0">6</number>
.i. 
<number id="B31B2497FFE7FFAA1FF7BA87AAFA86CC" box="[431,498,1716,1742]" pageId="7" pageNumber="8" value="2005.0">2005</number>
</date>
, W.D. Shepard leg. (
<number id="B31B2497FFE7FFAA1CB6BA87A80886CC" box="[750,768,1716,1742]" pageId="7" pageNumber="8" value="7.0">7</number>
: 
<collectionCode id="63C2AE75FFE7FFAA1D55BA87A85386CC" box="[781,859,1716,1742]" pageId="7" pageNumber="8">AEZS</collectionCode>
); 
<quantity id="C22B9B55FFE7FFAA1D2ABA87A8CD86CC" box="[882,965,1716,1742]" metricMagnitude="3" metricUnit="m" metricValue="6.6" pageId="7" pageNumber="8" unit="km" value="6.6">6.6 km</quantity>
SE Villa Purificacion, Puente El Amorin, 
<date id="716D1070FFE7FFAA1FCDBAEFAAFF86F4" box="[405,503,1756,1782]" pageId="7" pageNumber="8" value="2005-01-08">
<number id="B31B2497FFE7FFAA1FCDBAEFAAAB86F4" box="[405,419,1756,1782]" pageId="7" pageNumber="8" value="8.0">8</number>
.i. 
<number id="B31B2497FFE7FFAA1FEBBAEFAAFF86F4" box="[435,503,1756,1782]" pageId="7" pageNumber="8" value="2005.0">2005</number>
</date>
, W.D. Shepard leg. (
<number id="B31B2497FFE7FFAA1CAABAEFA80886F4" box="[754,768,1756,1782]" pageId="7" pageNumber="8" value="1.0">1</number>
: 
<collectionCode id="63C2AE75FFE7FFAA1D56BAEFA85486F4" box="[782,860,1756,1782]" pageId="7" pageNumber="8">AEZS</collectionCode>
); 
<emphasis id="37A7EAA2FFE7FFAA1D2ABAEFA8E386F4" bold="true" box="[882,1003,1756,1782]" pageId="7" pageNumber="8">Veracruz:</emphasis>
<quantity id="C22B9B55FFE7FFAA1DA6BAEFAF4F86F4" box="[1022,1095,1756,1782]" metricMagnitude="4" metricUnit="m" metricValue="4.6670976" pageId="7" pageNumber="8" unit="mi" value="29.0">29 mi</quantity>
. SE Jalapa, 
<quantity id="C22B9B55FFE7FFAA1A8EBAEFAE2A86F4" box="[1238,1314,1756,1782]" metricMagnitude="2" metricUnit="m" metricValue="2.7432" pageId="7" pageNumber="8" unit="ft" value="900.0">900 ft</quantity>
., 
<date id="716D1070FFE7FFAA1F50BB37AA98871C" box="[264,400,1796,1822]" pageId="7" pageNumber="8" value="1963-12-26">
<number id="B31B2497FFE7FFAA1F50BB37AA20871C" box="[264,296,1796,1822]" pageId="7" pageNumber="8" value="26.0">26</number>
.xii. 
<number id="B31B2497FFE7FFAA1F0ABB37AA98871C" box="[338,400,1796,1822]" pageId="7" pageNumber="8" value="1963.0">1963</number>
</date>
, M.J. Tauber &amp; C.A. Tauber leg. (
<number id="B31B2497FFE7FFAA1D7CBB37A83A871C" box="[804,818,1796,1822]" pageId="7" pageNumber="8" value="1.0">1</number>
: 
<collectionCode id="63C2AE75FFE7FFAA1D67BB37A89E871C" LSID="urn:lsid:biocol.org:col:1013" box="[831,918,1796,1822]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:1013" name="Essig Museum of Entomology" pageId="7" pageNumber="8">EMEC</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="4DC9653BFFE7FFAA1F60BB1FAFC5876C" pageId="7" pageNumber="8" type="diagnosis">
<paragraph id="056C36B0FFE7FFAA1F60BB1FAFC5876C" blockId="7.[264,1323,1396,1902]" pageId="7" pageNumber="8">
<emphasis id="37A7EAA2FFE7FFAA1F60BB1FAABF8744" bold="true" box="[312,439,1836,1862]" pageId="7" pageNumber="8">Diagnosis.</emphasis>
Total length: 
<quantity id="C22B9B55FFE7FFAA1C22BB1FA8078744" box="[634,783,1836,1862]" metricMagnitude="-3" metricUnit="m" metricValue="2.55" metricValueMax="2.8" metricValueMin="2.3" pageId="7" pageNumber="8" unit="mm" value="2.55" valueMax="2.8" valueMin="2.3">2.3–2.8 mm</quantity>
. The broadly testaceous anterior and lateral margins of the pronotum (
<figureCitation id="9DE82A35FFE7FFAA1C62BB67A989876C" box="[570,641,1876,1902]" captionStart="FIGURES 3 – 5" captionStartId="7.[264,382,1004,1028]" captionTargetBox="[418,1160,325,975]" captionTargetId="figure@7.[409,1177,304,994]" captionTargetPageId="7" captionText="FIGURES 3 – 5. Details of Hemiosus selva Short, sp. n. (3, 5) and H. exilis (LeConte) (4). 3 – 4: pronotal coloration, scale bar 0.18 mm; 5: aedeagus, scale bar 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.174868" httpUri="https://zenodo.org/record/174868/files/figure.png" pageId="7" pageNumber="8">
Fig. 
<number id="B31B2497FFE7FFAA1C29BB67A989876C" box="[625,641,1876,1902]" pageId="7" pageNumber="8" value="4.0">4</number>
</figureCitation>
) serve to distinguish this species in Mesoamerica.
</paragraph>
</subSubSection>
<subSubSection id="4DC9653BFFE8FFA51F60BD28A8B78134" box="[312,959,283,310]" pageId="8" pageNumber="9" type="distribution">
<paragraph id="056C36B0FFE8FFA51F60BD28A8B78134" blockId="8.[264,1323,283,590]" box="[312,959,283,310]" pageId="8" pageNumber="9">
<emphasis id="37A7EAA2FFE8FFA51F60BD28AAD08137" bold="true" box="[312,472,283,309]" pageId="8" pageNumber="9">Distribution.</emphasis>
<collectingCountry id="7DC47620FFE8FFA51FBFBD2FA94B8134" box="[487,579,284,310]" name="Mexico" pageId="8" pageNumber="9">Mexico</collectingCountry>
, 
<collectingCountry id="7DC47620FFE8FFA51C17BD2FA9E38134" box="[591,747,284,310]" name="United States of America" pageId="8" pageNumber="9">United States</collectingCountry>
(Arizona, Texas).
</paragraph>
</subSubSection>
<subSubSection id="4DC9653BFFE8FFA51F60BD70AFB2824C" pageId="8" pageNumber="9" type="discussion">
<paragraph id="056C36B0FFE8FFA51F60BD70AFB2824C" blockId="8.[264,1323,283,590]" pageId="8" pageNumber="9">
<emphasis id="37A7EAA2FFE8FFA51F60BD70AAB9815F" bold="true" box="[312,433,323,349]" pageId="8" pageNumber="9">Remarks.</emphasis>
See 
<bibRefCitation id="61424B41FFE8FFA51FAEBD77A9DC815C" box="[502,724,324,350]" pageId="8" pageNumber="17" refString="Van Tassell, E. R. (1964). A note on Hemiosus exilis LeConte (Coleoptera: Hydrophilidae). Coleopterists Bulletin, 18, 53 - 56." type="journal article">
Van Tassell (
<number id="B31B2497FFE8FFA51CD7BD77A9C4815C" box="[655,716,324,350]" pageId="8" pageNumber="9" value="1964.0">1964</number>
)
</bibRefCitation>
for a detailed discussion of the taxonomic history of the species. The synonymy of 
<taxonomicName id="C2D34D33FFE8FFA51CFBBD5FA8CF8184" authority="d'Orchymont" authorityName="d'Orchymont" box="[675,967,364,390]" class="Insecta" family="Hydrophilidae" genus="Hemiosus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="toxillus">
<emphasis id="37A7EAA2FFE8FFA51CFBBD5FA8178187" box="[675,799,364,389]" italics="true" pageId="8" pageNumber="9">H. toxillus</emphasis>
d’Orchymont
</taxonomicName>
with 
<taxonomicName id="C2D34D33FFE8FFA51A56BD5FAFFC8184" authority="LeConte" authorityName="LeConte" box="[1038,1268,364,390]" class="Insecta" family="Hydrophilidae" genus="Berosus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="exilis">
<emphasis id="37A7EAA2FFE8FFA51A56BD5FAF7A8187" box="[1038,1138,364,389]" italics="true" pageId="8" pageNumber="9">H. exilis</emphasis>
(LeConte)
</taxonomicName>
was first recognized but unpublished by 
<bibRefCitation id="61424B41FFE8FFA51CEBBDA7A89C81AC" box="[691,916,404,430]" pageId="8" pageNumber="17" refString="Van Tassell, E. R. (1966). Taxonomy and biology of the subfamily Berosinae of North and Central America and the West Indies (Coleoptera: Hydrophilidae). Unpublished Ph. D. dissertation, The Catholic University of America, Washington D. C., 329 pp." type="book">
Van Tassell (
<number id="B31B2497FFE8FFA51D13BDA7A88381AC" box="[843,907,404,430]" pageId="8" pageNumber="9" value="1966.0">1966</number>
)
</bibRefCitation>
. I have examined the 
<typeStatus id="DA688812FFE8FFA51AC0BDA7AE0281AC" box="[1176,1290,404,430]" pageId="8" pageNumber="9">holotypes</typeStatus>
of both species and concur that 
<taxonomicName id="C2D34D33FFE8FFA51CD0BD8FA8B181D4" authority="d'Orchymont" authorityName="d'Orchymont" box="[648,953,444,470]" class="Insecta" family="Hydrophilidae" genus="Hemiosus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="toxillus">
<emphasis id="37A7EAA2FFE8FFA51CD0BD8FA80381D7" box="[648,779,444,469]" italics="true" pageId="8" pageNumber="9">H. toxillus</emphasis>
d’Orchymont
</taxonomicName>
and 
<taxonomicName id="C2D34D33FFE8FFA51A5CBD8FAFFE81D4" authority="LeConte" authorityName="LeConte" box="[1028,1270,444,470]" class="Insecta" family="Hydrophilidae" genus="Berosus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="exilis">
<emphasis id="37A7EAA2FFE8FFA51A5CBD8FAF6681D7" box="[1028,1134,444,469]" italics="true" pageId="8" pageNumber="9">H. exilis</emphasis>
(LeConte)
</taxonomicName>
are conspecific. This likely occurred because 
<taxonomicName id="C2D34D33FFE8FFA51D58BDD7A86C81FF" box="[768,868,484,509]" class="Insecta" family="Hydrophilidae" genus="Berosus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="exilis">
<emphasis id="37A7EAA2FFE8FFA51D58BDD7A86C81FF" box="[768,868,484,509]" italics="true" pageId="8" pageNumber="9">H. exilis</emphasis>
</taxonomicName>
was still placed in the genus 
<taxonomicName id="C2D34D33FFE8FFA51A95BDD7AE2281FF" box="[1229,1322,484,509]" class="Insecta" family="Hydrophilidae" genus="Berosus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="37A7EAA2FFE8FFA51A95BDD7AE2281FF" box="[1229,1322,484,509]" italics="true" pageId="8" pageNumber="9">Berosus</emphasis>
</taxonomicName>
when d’Orchymont (
<number id="B31B2497FFE8FFA51C55BE3FA9428224" box="[525,586,524,550]" pageId="8" pageNumber="9" value="1940.0">1940</number>
) described 
<emphasis id="37A7EAA2FFE8FFA51CBBBE3FA8658227" box="[739,877,524,549]" italics="true" pageId="8" pageNumber="9">
<taxonomicName id="C2D34D33FFE8FFA51CBBBE3FA8618227" box="[739,873,524,549]" class="Insecta" family="Hydrophilidae" genus="Hemiosus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="toxillus">H. toxillus</taxonomicName>
.
</emphasis>
A full description and additional illustrations of this species are given by 
<bibRefCitation id="61424B41FFE8FFA51C87BE07A87E824C" author="Oliva" box="[735,886,564,590]" pageId="8" pageNumber="16" refString="Oliva, A. (1991). The types of Hemiosus Sharp, 1882 (Coleoptera: Hydrophilidae) in the Orchymont and Mouchamps collections. Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, 61, 167 - 181." type="journal article" year="1991">
Oliva (
<number id="B31B2497FFE8FFA51D68BE07A865824C" box="[816,877,564,590]" pageId="8" pageNumber="9" value="1991.0">1991</number>
)
</bibRefCitation>
under the name 
<taxonomicName id="C2D34D33FFE8FFA51A61BE07AFBC824F" box="[1081,1204,564,589]" class="Insecta" family="Hydrophilidae" genus="Hemiosus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="toxillus">
<emphasis id="37A7EAA2FFE8FFA51A61BE07AFBC824F" box="[1081,1204,564,589]" italics="true" pageId="8" pageNumber="9">H. toxillus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>