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<document id="8D840EA803168A2F8CE5D7F1BFFBB1A9" ID-DOI="10.11646/zootaxa.4790.3.13" ID-ISSN="1175-5326" ID-Zenodo-Dep="3891267" ID-ZooBank="AC939AA2-D8D6-4F5A-85A8-6DE7B316FC09" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1591959379214" checkinUser="plazi" docAuthor="Jiang, Ri-Xin, Li, Bo-Yan &amp; Wang, Shuo" docDate="2020" docId="386D2161DE59FFE7FF757748DEE2AEF8" docLanguage="en" docName="zootaxa.4790.3.13.pdf" docOrigin="Zootaxa 4790 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Orphnebius lilizheni Jiang &amp; Li &amp; Wang 2020, sp. nov." docType="treatment" docVersion="3" lastPageNumber="598" masterDocId="C4545919DE58FFE2FFE27501DA21AF57" masterDocTitle="Description a new species of the genus Orphnebius Motschulsky, 1858 (Coleoptera: Staphylinidae: Aleocharinae) from Xizang, China" masterLastPageNumber="599" masterPageNumber="593" pageNumber="594" updateTime="1700276635944" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="2CFAFCCAF3C5F13E7AF9C50E3354DBA3" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="1AB2ED1A75926308B8086F440A59A490">
<mods:title id="FBA19BFE1ECE99460D50BF974105833C">Description a new species of the genus Orphnebius Motschulsky, 1858 (Coleoptera: Staphylinidae: Aleocharinae) from Xizang, China</mods:title>
</mods:titleInfo>
<mods:name id="C403CB6BD31AB66E19A7359D4249A6C7" type="personal">
<mods:role id="AE2924734AD71195AD1F3851630003DC">
<mods:roleTerm id="4CEF76DF0E3288C097F5B5B624AED78A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EEF489276AABD0741C0BEE695B43A5FA">Jiang, Ri-Xin</mods:namePart>
<mods:affiliation id="34E64E57177266CC7DCCD89E95DF2E6F">maoshuwuyouzhi @ 163. com; https: // orcid. org / 0000 - 0002 - 5339 - 853 X</mods:affiliation>
</mods:name>
<mods:name id="FC75C58BAE3B88B124AF98866FC2CA4C" type="personal">
<mods:role id="A89DBD1877560806D6BFA62C1BFBF106">
<mods:roleTerm id="5F6B31A4493329D4DDC75B3D9375273E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0C7E079C64A0C3CA5022CD494A6C7371">Li, Bo-Yan</mods:namePart>
<mods:affiliation id="D6C46BDD4864A8A64A480B7237848EEC">Institute of Entomology, Guizhou University, Guizhou Provincial Key Laboratory for Agricultural Pest Management of the Mountainous Region, Guiyang, 550025, P. R. China.</mods:affiliation>
</mods:name>
<mods:name id="3F4F3E90700A135AA182109D43F26E01" type="personal">
<mods:role id="4E3B896ED2FE28F47B7AFD92F17C669F">
<mods:roleTerm id="0A0128F9C24BD9AA418B78A0204630D3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F4A87E7174A1619499F662B763A038CE">Wang, Shuo</mods:namePart>
</mods:name>
<mods:typeOfResource id="71BF3672297369E46C6D6C0D42583E75">text</mods:typeOfResource>
<mods:relatedItem id="C706EE984D4753B67AA174B214490AA2" type="host">
<mods:titleInfo id="E12ECA95729C85BDC0AD0FBFEC15168D">
<mods:title id="2528D78643F6AE72A49DF1F3215A5E06">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="5DD9728ACCEACC82F76A8EEB9CC73985">
<mods:date id="452AC5D09950779EED24AA57FCD1092C">2020</mods:date>
<mods:detail id="3EDCD5B532E153F37D1E2CC2070DAE3C" type="pubDate">
<mods:number id="9A8B859A76F3D2E60E641DE09F5A4CA6">2020-06-12</mods:number>
</mods:detail>
<mods:detail id="19BF728A171463DCEFC84077AB5F92B5" type="volume">
<mods:number id="86087FE258B10029DE446E48DE0CD0F5">4790</mods:number>
</mods:detail>
<mods:detail id="DA807887D11F8D71C7917CA5642B80D3" type="issue">
<mods:number id="E32135676C94EA160A21247DD02574A8">3</mods:number>
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<mods:start id="4853883A46BDDE187C838B0071EC13F7">593</mods:start>
<mods:end id="14B7A2E196AFF81E4EFA0624CA6DFD51">599</mods:end>
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<mods:classification id="0F0DC4EA86154677D1BE9E28F198B2E5">journal article</mods:classification>
<mods:identifier id="2FDFF112A834D5BF95D677EBD5A0C17A" type="DOI">10.11646/zootaxa.4790.3.13</mods:identifier>
<mods:identifier id="78519C6246D27FCA8090D34B5358B2E6" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="A2D30C0107DAC738BABD89A3B40CA06A" type="Zenodo-Dep">3891267</mods:identifier>
<mods:identifier id="83107F5F7FCF22A39BA737C75963FD06" type="ZooBank">AC939AA2-D8D6-4F5A-85A8-6DE7B316FC09</mods:identifier>
</mods:mods>
<treatment id="386D2161DE59FFE7FF757748DEE2AEF8" LSID="urn:lsid:plazi:treatment:386D2161DE59FFE7FF757748DEE2AEF8" httpUri="http://treatment.plazi.org/id/386D2161DE59FFE7FF757748DEE2AEF8" lastPageId="5" lastPageNumber="598" pageId="1" pageNumber="594">
<subSubSection id="F8DEC3FCDE59FFE3FF757748DE34AD08" box="[151,1045,583,612]" pageId="1" pageNumber="594" type="nomenclature">
<paragraph id="B07B9077DE59FFE3FF757748DE34AD08" blockId="1.[151,1045,583,647]" box="[151,1045,583,612]" pageId="1" pageNumber="594">
<heading id="EB33271BDE59FFE3FF757748DE34AD08" bold="true" box="[151,1045,583,612]" fontSize="11" level="1" pageId="1" pageNumber="594" reason="1">
<emphasis id="82B04C65DE59FFE3FF757748DE34AD08" bold="true" box="[151,1045,583,612]" pageId="1" pageNumber="594">
<taxonomicName id="77C4EBF4DE59FFE3FF757748DBB1AD33" ID-CoL="8Y7BY" authority="Jiang &amp; Li &amp; Wang, 2020" authorityName="Jiang &amp; Li &amp; Wang" authorityYear="2020" box="[151,400,585,612]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="594" phylum="Arthropoda" rank="species" species="lilizheni" status="sp. nov.">
<emphasis id="82B04C65DE59FFE3FF757748DBB1AD33" bold="true" box="[151,400,585,612]" italics="true" pageId="1" pageNumber="594">Orphnebius lilizheni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9983F11EDE59FFE3FE757748DBD4AD34" box="[407,501,585,611]" pageId="1" pageNumber="594" rank="species">sp. nov.</taxonomicNameLabel>
Jiang, Li &amp; Wang sp. nov. ÷KHẘDaem
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="F8DEC3FCDE59FFE3FF75776CDB33ADD0" box="[151,274,621,647]" pageId="1" pageNumber="594" type="description">
<paragraph id="B07B9077DE59FFE3FF75776CDB33ADD0" blockId="1.[151,1045,583,647]" box="[151,274,621,647]" pageId="1" pageNumber="594">
(
<figureCitation id="28FF8CF2DE59FFE3FF7D776CDB2BADD0" box="[159,266,621,647]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="1.[151,250,2017,2042]" captionStartId-1="2.[151,250,1902,1927]" captionStartId-2="3.[151,250,1941,1966]" captionStartId-3="4.[151,250,1734,1759]" captionTargetBox-0="[186,1398,819,1981]" captionTargetBox-1="[160,1424,841,1876]" captionTargetBox-2="[151,1436,453,1916]" captionTargetBox-3="[151,1436,309,1709]" captionTargetId-0="figure@1.[176,1412,813,1992]" captionTargetId-1="figure@2.[160,1427,813,1878]" captionTargetId-2="figure@3.[151,1436,453,1916]" captionTargetId-3="figure@4.[151,1436,309,1709]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionTargetPageId-3="4" captionText-0="FIGURE 1. Habitus of Orphnebius lilizheni sp. nov. A. Male; B. Female." captionText-1="FIGURE 2. Diagnostic features of Orphnebius lilizheni sp. nov. A. Head and pronotum, in dorsal view; B. Antenna; C. Spermathecal; D. Sternite VIII; E. Tergite VIII; F. Paramere; G. Male segments IX–X, dorsal view; H. Same, ventral view; I. Female segments IX–X, dorsal view; J. Same, ventral view; K. Media lobe of aedegus, ventral view; L. Same, lateral view; M. Same, dorsal view." captionText-2="FIGURE 3. A. Larva of Orphnebius lilizheni sp. nov., dorsal view; B. Same, lateral view; C. Same, ventral view; D. Myrmica sp., dorsal view; E. Same, lateral view; F. Larva of Myrmica sp." captionText-3="FIGURE 4. Habitat of Orphnebius lilizheni sp. nov. A. Adults and larvae in a nest of Myrmica sp.; B. A Myrmica sp. ant in nest; C. General environment of the type locality: Baotianman (Xizang, China). Laa—Larva of the host ant; La—Larva of Orphnebius lilizheni sp. nov.; ad—Adults of Orphnebius lilizheni sp. nov." figureDoi-0="http://doi.org/10.5281/zenodo.3891269" figureDoi-1="http://doi.org/10.5281/zenodo.3891271" figureDoi-2="http://doi.org/10.5281/zenodo.3891273" figureDoi-3="http://doi.org/10.5281/zenodo.3891275" httpUri-0="https://zenodo.org/record/3891269/files/figure.png" httpUri-1="https://zenodo.org/record/3891271/files/figure.png" httpUri-2="https://zenodo.org/record/3891273/files/figure.png" httpUri-3="https://zenodo.org/record/3891275/files/figure.png" pageId="1" pageNumber="594">Figs. 1–4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="F8DEC3FCDE59FFE3FF7577B4DF76AC40" pageId="1" pageNumber="594" type="materials_examined">
<paragraph id="B07B9077DE59FFE3FF7577B4DF76AC40" blockId="1.[151,1437,691,791]" pageId="1" pageNumber="594">
<emphasis id="82B04C65DE59FFE3FF7577B4DBB1AD98" bold="true" box="[151,400,692,719]" pageId="1" pageNumber="594">Type material (9 exs,</emphasis>
<specimenCount id="A6C25BFEDE59FFE3FE7577B4DBF9AD98" box="[407,472,693,719]" pageId="1" pageNumber="594" type="male">3 ♂♂</specimenCount>
,
<specimenCount id="A6C25BFEDE59FFE3FE0677B4D802AD98" box="[484,547,693,719]" pageId="1" pageNumber="594" type="female">6 ♀♀</specimenCount>
<emphasis id="82B04C65DE59FFE3FDC877B5D899AD98" bold="true" box="[554,696,692,719]" pageId="1" pageNumber="594">
).
<typeStatus id="6F7F2ED5DE59FFE3FDA077B4D892AD98" box="[578,691,693,719]" pageId="1" pageNumber="594" type="holotype">Holotype</typeStatus>
:
</emphasis>
<materialsCitation id="00AC9A2ADE59FFE3FD5D77B5D980AD98" box="[703,929,692,719]" country="China" location="CHINA" pageId="1" pageNumber="594" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="82B04C65DE59FFE3FD5D77B5D907AD98" bold="true" box="[703,806,692,719]" pageId="1" pageNumber="594">
<collectingCountry id="C8D3D0E7DE59FFE3FD5D77B5D900AD98" box="[703,801,692,719]" name="China" pageId="1" pageNumber="594">CHINA</collectingCountry>
:
</emphasis>
<specimenCount id="A6C25BFEDE59FFE3FCCF77B4D965AD98" box="[813,836,693,719]" pageId="1" pageNumber="594" type="male">♂</specimenCount>
, labeled
</materialsCitation>
‘
<materialsCitation id="00AC9A2ADE59FFE3FC4E77B4DFB8ADA4" collectingDate="2019-07-19" collectionCode="SHNU" collectorName="Li Bo-yan" country="China" county="Milin County" elevation="3166" location="Nanyigou" municipality="Linzhi City" pageId="1" pageNumber="594" specimenCount="1" stateProvince="Xizang" typeStatus="holotype">
<collectingCountry id="C8D3D0E7DE59FFE3FC4E77B4D9D9AD98" box="[940,1016,693,719]" name="China" pageId="1" pageNumber="594">China</collectingCountry>
:
<collectingRegion id="72005E95DE59FFE3FBE177B4DE76AD98" box="[1027,1111,693,719]" country="China" name="Xizang" pageId="1" pageNumber="594">Xizang</collectingRegion>
,
<collectingMunicipality id="501F0A0DDE59FFE3FB8077B4DEC3AD98" box="[1122,1250,693,719]" pageId="1" pageNumber="594">Linzhi City</collectingMunicipality>
(
<emphasis id="82B04C65DE59FFE3FB1177B2DF66AD9D" bold="true" box="[1267,1351,691,714]" pageId="1" pageNumber="594">ẇźṃ</emphasis>
),
<collectingCounty id="591AE8FBDE59FFE3FABC77B4DAC8ADA4" pageId="1" pageNumber="594">Milin County</collectingCounty>
(
<emphasis id="82B04C65DE59FFE3FF1B77D6DB6CADB9" bold="true" box="[249,333,727,750]" pageId="1" pageNumber="594">*ẇu</emphasis>
),
<location id="B51BC6ACDE59FFE3FE8177D8DBF0ADA4" LSID="urn:lsid:plazi:treatment:386D2161DE59FFE7FF757748DEE2AEF8:B51BC6ACDE59FFE3FE8177D8DBF0ADA4" box="[355,465,729,755]" country="China" county="Milin County" municipality="Linzhi City" name="Nanyigou" pageId="1" pageNumber="594" stateProvince="Xizang">Nanyigou</location>
(
<emphasis id="82B04C65DE59FFE3FE0277D6D815ADB9" bold="true" box="[480,564,727,750]" pageId="1" pageNumber="594">ṁDzḿ</emphasis>
), H:
<quantity id="773C3D92DE59FFE3FD8E77D8D8E3ADA4" box="[620,706,729,755]" metricMagnitude="3" metricUnit="m" metricValue="3.166" pageId="1" pageNumber="594" unit="m" value="3166.0">
<elevation id="3BE97744DE59FFE3FD8E77D8D8E3ADA4" box="[620,706,729,755]" metricMagnitude="3" metricUnit="m" metricValue="3.166" pageId="1" pageNumber="594" unit="m" value="3166.0">3166 m</elevation>
</quantity>
,
<date id="C47AB6B7DE59FFE3FD2E77D8D979ADA4" box="[716,856,729,755]" pageId="1" pageNumber="594" value="2019-07-19">
<collectingDate id="D43E4F5FDE59FFE3FD2E77D8D979ADA4" box="[716,856,729,755]" pageId="1" pageNumber="594" value="2019-07-19">19.VII.2019</collectingDate>
</date>
,
<collectorName id="1D31F5A1DE59FFE3FC8077D8D9F2ADA4" box="[866,979,729,755]" pageId="1" pageNumber="594">Li Bo-yan</collectorName>
leg., in nest of
<taxonomicName id="77C4EBF4DE59FFE3FB9877DBDF22ADA4" authorityName="Smith" authorityYear="1852" box="[1146,1283,730,755]" class="Insecta" family="Formicidae" genus="Tetraponera" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="594" phylum="Arthropoda" rank="genus">
<emphasis id="82B04C65DE59FFE3FB9877DBDF22ADA4" box="[1146,1283,730,755]" italics="true" pageId="1" pageNumber="594">Tetraponera</emphasis>
</taxonomicName>
sp.’ (
<collectionCode id="D6D508B2DE59FFE3FADC77D8DFB1ADA4" box="[1342,1424,729,755]" pageId="1" pageNumber="594">SHNU</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="00AC9A2ADE59FFE3FF7577FCDF75AC40" box="[151,1364,764,791]" collectingDate="2019-07-19" collectionCode="SHNU, GUGC" collectorName="Li Bo-yan" country="China" county="Milin County" elevation="3166" location="Same" municipality="Linzhi City" pageId="1" pageNumber="594" specimenCount="16" specimenCount-female="12" specimenCount-male="4" stateProvince="Xizang" typeStatus="paratype">
<emphasis id="82B04C65DE59FFE3FF7577FCDBA7AC40" bold="true" box="[151,390,764,791]" pageId="1" pageNumber="594">
<typeStatus id="6F7F2ED5DE59FFE3FF7577FCDB32AC40" box="[151,275,765,791]" pageId="1" pageNumber="594" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="C8D3D0E7DE59FFE3FEFD77FDDBA0AC40" box="[287,385,764,791]" name="China" pageId="1" pageNumber="594">CHINA</collectingCountry>
:
</emphasis>
<specimenCount id="A6C25BFEDE59FFE3FE6F77FCDBEFAC40" box="[397,462,765,791]" pageId="1" pageNumber="594" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="A6C25BFEDE59FFE3FE3877FCD83AAC40" box="[474,539,765,791]" pageId="1" pageNumber="594" type="female">6 ♀♀</specimenCount>
.
<location id="B51BC6ACDE59FFE3FDC577FCD844AC40" LSID="urn:lsid:plazi:treatment:386D2161DE59FFE7FF757748DEE2AEF8:B51BC6ACDE59FFE3FDC577FCD844AC40" box="[551,613,765,791]" country="China" name="Same" pageId="1" pageNumber="594">Same</location>
label data as the holotype (
<specimenCount id="A6C25BFEDE59FFE3FC4277FCD9EDAC40" box="[928,972,765,791]" pageId="1" pageNumber="594" type="male">1 ♂</specimenCount>
,
<specimenCount id="A6C25BFEDE59FFE3FC3A77FCDE36AC40" box="[984,1047,765,791]" pageId="1" pageNumber="594" type="female">2 ♀♀</specimenCount>
<collectionCode id="D6D508B2DE59FFE3FBFC77FCDE4FAC40" box="[1054,1134,765,791]" pageId="1" pageNumber="594">SHNU</collectionCode>
;
<specimenCount id="A6C25BFEDE59FFE3FB9B77FCDE84AC40" box="[1145,1189,765,791]" pageId="1" pageNumber="594" type="male">1 ♂</specimenCount>
,
<specimenCount id="A6C25BFEDE59FFE3FB5377FCDED1AC40" box="[1201,1264,765,791]" pageId="1" pageNumber="594" type="female">4 ♀♀</specimenCount>
<collectionCode id="D6D508B2DE59FFE3FB1577FCDF6AAC40" box="[1271,1355,765,791]" pageId="1" pageNumber="594">GUGC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="E4BBC0FFDE59FFE3FF7572E0D9A6A8AD" ID-DOI="http://doi.org/10.5281/zenodo.3891269" ID-Zenodo-Dep="3891269" box="[151,903,2017,2042]" httpUri="https://zenodo.org/record/3891269/files/figure.png" pageId="1" pageNumber="594" startId="1.[151,250,2017,2042]" targetBox="[186,1398,819,1981]" targetPageId="1">
<paragraph id="B07B9077DE59FFE3FF7572E0D9A6A8AD" blockId="1.[151,903,2017,2042]" box="[151,903,2017,2042]" pageId="1" pageNumber="594">
<emphasis id="82B04C65DE59FFE3FF7572E0DB32A8AD" bold="true" box="[151,275,2017,2042]" pageId="1" pageNumber="594">FIGURE 1.</emphasis>
Habitus of
<taxonomicName id="77C4EBF4DE59FFE3FE6872E0D87BA8AE" authority="Jiang &amp; Li &amp; Wang, 2020" authorityName="Jiang &amp; Li &amp; Wang" authorityYear="2020" box="[394,602,2017,2041]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="594" phylum="Arthropoda" rank="species" species="lilizheni" status="sp. nov.">
<emphasis id="82B04C65DE59FFE3FE6872E0D87BA8AE" box="[394,602,2017,2041]" italics="true" pageId="1" pageNumber="594">Orphnebius lilizheni</emphasis>
</taxonomicName>
<emphasis id="82B04C65DE59FFE3FD8272E3D8F1A8AD" bold="true" box="[608,720,2017,2042]" pageId="1" pageNumber="594">
<taxonomicNameLabel id="9983F11EDE59FFE3FD8272E3D891A8AD" box="[608,688,2018,2042]" pageId="1" pageNumber="594" rank="species">sp. nov.</taxonomicNameLabel>
A.
</emphasis>
Male;
<emphasis id="82B04C65DE59FFE3FCFA72E3D90EA8AD" bold="true" box="[792,815,2018,2042]" pageId="1" pageNumber="594">B.</emphasis>
Female.
</paragraph>
</caption>
<subSubSection id="F8DEC3FCDE5AFFE7FF257599DEACAFAC" lastPageId="5" lastPageNumber="598" pageId="2" pageNumber="595" type="description">
<paragraph id="B07B9077DE5AFFE0FF257599D92AAF80" blockId="2.[151,1437,152,791]" pageId="2" pageNumber="595">
<emphasis id="82B04C65DE5AFFE0FF257599DB78AFE4" bold="true" box="[199,345,152,179]" pageId="2" pageNumber="595">Description.</emphasis>
Male (
<figureCitation id="28FF8CF2DE5AFFE0FE487598D829AFE4" box="[426,520,153,179]" captionStart="FIGURE 1" captionStartId="1.[151,250,2017,2042]" captionTargetBox="[186,1398,819,1981]" captionTargetId="figure@1.[176,1412,813,1992]" captionTargetPageId="1" captionText="FIGURE 1. Habitus of Orphnebius lilizheni sp. nov. A. Male; B. Female." figureDoi="http://doi.org/10.5281/zenodo.3891269" httpUri="https://zenodo.org/record/3891269/files/figure.png" pageId="2" pageNumber="595">Figs 1A</figureCitation>
,
<figureCitation id="28FF8CF2DE5AFFE0FDF77598D802AFE4" box="[533,547,153,179]" captionStart="FIGURE 2" captionStartId="2.[151,250,1902,1927]" captionTargetBox="[160,1424,841,1876]" captionTargetId="figure@2.[160,1427,813,1878]" captionTargetPageId="2" captionText="FIGURE 2. Diagnostic features of Orphnebius lilizheni sp. nov. A. Head and pronotum, in dorsal view; B. Antenna; C. Spermathecal; D. Sternite VIII; E. Tergite VIII; F. Paramere; G. Male segments IX–X, dorsal view; H. Same, ventral view; I. Female segments IX–X, dorsal view; J. Same, ventral view; K. Media lobe of aedegus, ventral view; L. Same, lateral view; M. Same, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3891271" httpUri="https://zenodo.org/record/3891271/files/figure.png" pageId="2" pageNumber="595">2</figureCitation>
A–B, D–H, K–M). Body bicolourable, head, pronotum and elytra black, abdomen brown, dorsal surface shiny and without punctures.
</paragraph>
<paragraph id="B07B9077DE5AFFE0FF2575E0DB6BAEDC" blockId="2.[151,1437,152,791]" pageId="2" pageNumber="595">
Head (
<figureCitation id="28FF8CF2DE5AFFE0FEF175E0DB4DAFAC" box="[275,364,225,251]" captionStart="FIGURE 2" captionStartId="2.[151,250,1902,1927]" captionTargetBox="[160,1424,841,1876]" captionTargetId="figure@2.[160,1427,813,1878]" captionTargetPageId="2" captionText="FIGURE 2. Diagnostic features of Orphnebius lilizheni sp. nov. A. Head and pronotum, in dorsal view; B. Antenna; C. Spermathecal; D. Sternite VIII; E. Tergite VIII; F. Paramere; G. Male segments IX–X, dorsal view; H. Same, ventral view; I. Female segments IX–X, dorsal view; J. Same, ventral view; K. Media lobe of aedegus, ventral view; L. Same, lateral view; M. Same, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3891271" httpUri="https://zenodo.org/record/3891271/files/figure.png" pageId="2" pageNumber="595">Fig. 2A</figureCitation>
) slightly wider than long, shiny and covered with sparse long hairs. Antenna (
<figureCitation id="28FF8CF2DE5AFFE0FB0675E0DF1AAFAC" box="[1252,1339,225,251]" captionStart="FIGURE 2" captionStartId="2.[151,250,1902,1927]" captionTargetBox="[160,1424,841,1876]" captionTargetId="figure@2.[160,1427,813,1878]" captionTargetPageId="2" captionText="FIGURE 2. Diagnostic features of Orphnebius lilizheni sp. nov. A. Head and pronotum, in dorsal view; B. Antenna; C. Spermathecal; D. Sternite VIII; E. Tergite VIII; F. Paramere; G. Male segments IX–X, dorsal view; H. Same, ventral view; I. Female segments IX–X, dorsal view; J. Same, ventral view; K. Media lobe of aedegus, ventral view; L. Same, lateral view; M. Same, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3891271" httpUri="https://zenodo.org/record/3891271/files/figure.png" pageId="2" pageNumber="595">Fig. 2B</figureCitation>
) moderately long and apically asymmetric; antennomere I–V longer than wide, I expanded near apex; II about as long as IV; III long, about 1.5 times as long as II; V shorter than IV; VI about as long as wide; VII–X similar, obviously wider than long, VII–X similar, obviously wider than long; XI of ovoid shape and approximately as long as the combined length of IX–X.
</paragraph>
<paragraph id="B07B9077DE5AFFE0FF257494D8B8AE84" blockId="2.[151,1437,152,791]" pageId="2" pageNumber="595">
Pronotum (
<figureCitation id="28FF8CF2DE5AFFE0FEA57494DB83AEF8" box="[327,418,405,431]" captionStart="FIGURE 2" captionStartId="2.[151,250,1902,1927]" captionTargetBox="[160,1424,841,1876]" captionTargetId="figure@2.[160,1427,813,1878]" captionTargetPageId="2" captionText="FIGURE 2. Diagnostic features of Orphnebius lilizheni sp. nov. A. Head and pronotum, in dorsal view; B. Antenna; C. Spermathecal; D. Sternite VIII; E. Tergite VIII; F. Paramere; G. Male segments IX–X, dorsal view; H. Same, ventral view; I. Female segments IX–X, dorsal view; J. Same, ventral view; K. Media lobe of aedegus, ventral view; L. Same, lateral view; M. Same, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3891271" httpUri="https://zenodo.org/record/3891271/files/figure.png" pageId="2" pageNumber="595">Fig. 2A</figureCitation>
), about 1.4 times as wide as long; dorsal surface shiny and without punctures and pubescence; lateral margins with several long hairs.
</paragraph>
<paragraph id="B07B9077DE5AFFE0FF2574DCD982AD4C" blockId="2.[151,1437,152,791]" pageId="2" pageNumber="595">Elytra wider than long; near trapezoid shape; dorsal surface shiny, covered with sparse hairs and small punctures; lateral margins with erect long black hairs. Hind wings present.</paragraph>
<paragraph id="B07B9077DE5AFFE0FF257724DE3CAD34" blockId="2.[151,1437,152,791]" pageId="2" pageNumber="595">Legs simple. All femora black, without modification. Tibiae dark brown, protibiae covered with very dense short hairs, and mesotibiae and metatibiae covered with much sparse short hairs.</paragraph>
<paragraph id="B07B9077DE5AFFE0FF25776CDF00AC40" blockId="2.[151,1437,152,791]" pageId="2" pageNumber="595">
Abdominal segments III–VIII reddish brown, III–VII with weakly tapering posteriad, III widest. Posterior margin of tergites III–VI with moderately long black hairs. Tergite VII weakly impressed and in basal 1/3 with transverse row of regular striae, posterior margin of tergite VII with pronounced palisade fringe. Tergite VIII (
<figureCitation id="28FF8CF2DE5AFFE0FADD77B4DFB5AD98" box="[1343,1428,693,719]" captionStart="FIGURE 2" captionStartId="2.[151,250,1902,1927]" captionTargetBox="[160,1424,841,1876]" captionTargetId="figure@2.[160,1427,813,1878]" captionTargetPageId="2" captionText="FIGURE 2. Diagnostic features of Orphnebius lilizheni sp. nov. A. Head and pronotum, in dorsal view; B. Antenna; C. Spermathecal; D. Sternite VIII; E. Tergite VIII; F. Paramere; G. Male segments IX–X, dorsal view; H. Same, ventral view; I. Female segments IX–X, dorsal view; J. Same, ventral view; K. Media lobe of aedegus, ventral view; L. Same, lateral view; M. Same, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3891271" httpUri="https://zenodo.org/record/3891271/files/figure.png" pageId="2" pageNumber="595">Fig. 2E</figureCitation>
) transverse with convex posterior margins and covered with sparse strong setae. Sternite VIII (
<figureCitation id="28FF8CF2DE5AFFE0FB5777D8DF2CADA4" box="[1205,1293,729,755]" captionStart="FIGURE 2" captionStartId="2.[151,250,1902,1927]" captionTargetBox="[160,1424,841,1876]" captionTargetId="figure@2.[160,1427,813,1878]" captionTargetPageId="2" captionText="FIGURE 2. Diagnostic features of Orphnebius lilizheni sp. nov. A. Head and pronotum, in dorsal view; B. Antenna; C. Spermathecal; D. Sternite VIII; E. Tergite VIII; F. Paramere; G. Male segments IX–X, dorsal view; H. Same, ventral view; I. Female segments IX–X, dorsal view; J. Same, ventral view; K. Media lobe of aedegus, ventral view; L. Same, lateral view; M. Same, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3891271" httpUri="https://zenodo.org/record/3891271/files/figure.png" pageId="2" pageNumber="595">Fig. 2D</figureCitation>
) with sparse long setae at posterior margin. IX–X (
<figureCitation id="28FF8CF2DE5AFFE0FDA277FCD8A7AC40" box="[576,646,765,791]" captionStart="FIGURE 2" captionStartId="2.[151,250,1902,1927]" captionTargetBox="[160,1424,841,1876]" captionTargetId="figure@2.[160,1427,813,1878]" captionTargetPageId="2" captionText="FIGURE 2. Diagnostic features of Orphnebius lilizheni sp. nov. A. Head and pronotum, in dorsal view; B. Antenna; C. Spermathecal; D. Sternite VIII; E. Tergite VIII; F. Paramere; G. Male segments IX–X, dorsal view; H. Same, ventral view; I. Female segments IX–X, dorsal view; J. Same, ventral view; K. Media lobe of aedegus, ventral view; L. Same, lateral view; M. Same, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3891271" httpUri="https://zenodo.org/record/3891271/files/figure.png" pageId="2" pageNumber="595">Figs 2</figureCitation>
G–H) strongly modified, covered with dense and long hairs
</paragraph>
<caption id="E4BBC0FFDE5AFFE0FF75726FDB33A8A4" ID-DOI="http://doi.org/10.5281/zenodo.3891271" ID-Zenodo-Dep="3891271" httpUri="https://zenodo.org/record/3891271/files/figure.png" pageId="2" pageNumber="595" startId="2.[151,250,1902,1927]" targetBox="[160,1424,841,1876]" targetPageId="2">
<paragraph id="B07B9077DE5AFFE0FF75726FDB33A8A4" blockId="2.[151,1437,1902,2035]" pageId="2" pageNumber="595">
<emphasis id="82B04C65DE5AFFE0FF75726FDB32A8D0" bold="true" box="[151,275,1902,1927]" pageId="2" pageNumber="595">FIGURE 2.</emphasis>
Diagnostic features of
<taxonomicName id="77C4EBF4DE5AFFE0FDE0726ED8F3A8D0" authority="Jiang &amp; Li &amp; Wang, 2020" authorityName="Jiang &amp; Li &amp; Wang" authorityYear="2020" box="[514,722,1903,1927]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="595" phylum="Arthropoda" rank="species" species="lilizheni" status="sp. nov.">
<emphasis id="82B04C65DE5AFFE0FDE0726ED8F3A8D0" box="[514,722,1903,1927]" italics="true" pageId="2" pageNumber="595">Orphnebius lilizheni</emphasis>
</taxonomicName>
<emphasis id="82B04C65DE5AFFE0FD3B726ED96BA8D0" bold="true" box="[729,842,1902,1927]" pageId="2" pageNumber="595">
<taxonomicNameLabel id="9983F11EDE5AFFE0FD3B726ED90BA8D0" box="[729,810,1903,1927]" pageId="2" pageNumber="595" rank="species">sp. nov.</taxonomicNameLabel>
A.
</emphasis>
Head and pronotum, in dorsal view;
<emphasis id="82B04C65DE5AFFE0FB24726EDEFCA8D0" bold="true" box="[1222,1245,1903,1927]" pageId="2" pageNumber="595">B.</emphasis>
Antenna;
<emphasis id="82B04C65DE5AFFE0FAA4726FDF7EA8D0" bold="true" box="[1350,1375,1902,1927]" pageId="2" pageNumber="595">C.</emphasis>
Spermathecal;
<emphasis id="82B04C65DE5AFFE0FF1D7292DB39A8FC" bold="true" box="[255,280,1939,1963]" pageId="2" pageNumber="595">D.</emphasis>
Sternite VIII;
<emphasis id="82B04C65DE5AFFE0FE4B7292DB9EA8FC" bold="true" box="[425,447,1939,1963]" pageId="2" pageNumber="595">E.</emphasis>
Tergite VIII;
<emphasis id="82B04C65DE5AFFE0FDAA7292D87AA8FC" bold="true" box="[584,603,1939,1963]" pageId="2" pageNumber="595">F.</emphasis>
Paramere;
<emphasis id="82B04C65DE5AFFE0FD2F7293D8C6A8FC" bold="true" box="[717,743,1938,1963]" pageId="2" pageNumber="595">G.</emphasis>
Male segments IX–X, dorsal view;
<emphasis id="82B04C65DE5AFFE0FBB27292DE4BA8FC" bold="true" box="[1104,1130,1939,1963]" pageId="2" pageNumber="595">H.</emphasis>
Same, ventral view;
<emphasis id="82B04C65DE5AFFE0FADE7292DF6DA8FC" bold="true" box="[1340,1356,1939,1963]" pageId="2" pageNumber="595">I.</emphasis>
Female segments IX–X, dorsal view;
<emphasis id="82B04C65DE5AFFE0FE2472B6DBF8A898" bold="true" box="[454,473,1975,1999]" pageId="2" pageNumber="595">J.</emphasis>
Same, ventral view;
<emphasis id="82B04C65DE5AFFE0FD5372B6D8EAA898" bold="true" box="[689,715,1975,1999]" pageId="2" pageNumber="595">K.</emphasis>
Media lobe of aedegus, ventral view;
<emphasis id="82B04C65DE5AFFE0FBB372B6DE49A898" bold="true" box="[1105,1128,1975,1999]" pageId="2" pageNumber="595">L.</emphasis>
Same, lateral view;
<emphasis id="82B04C65DE5AFFE0FADB72B6DF76A898" bold="true" box="[1337,1367,1975,1999]" pageId="2" pageNumber="595">M.</emphasis>
Same, dorsal view.
</paragraph>
</caption>
<paragraph id="B07B9077DE5BFFE1FF257598D804AF80" blockId="3.[151,1436,153,431]" pageId="3" pageNumber="596">
Media lobe of aedeagus broad and short (
<figureCitation id="28FF8CF2DE5BFFE1FD787598D8C3AFE4" box="[666,738,153,179]" captionStart="FIGURE 2" captionStartId="2.[151,250,1902,1927]" captionTargetBox="[160,1424,841,1876]" captionTargetId="figure@2.[160,1427,813,1878]" captionTargetPageId="2" captionText="FIGURE 2. Diagnostic features of Orphnebius lilizheni sp. nov. A. Head and pronotum, in dorsal view; B. Antenna; C. Spermathecal; D. Sternite VIII; E. Tergite VIII; F. Paramere; G. Male segments IX–X, dorsal view; H. Same, ventral view; I. Female segments IX–X, dorsal view; J. Same, ventral view; K. Media lobe of aedegus, ventral view; L. Same, lateral view; M. Same, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3891271" httpUri="https://zenodo.org/record/3891271/files/figure.png" pageId="3" pageNumber="596">Figs 2</figureCitation>
K–M). Paramere small in relation to median lobe, apical truncate, with two short setae (
<figureCitation id="28FF8CF2DE5BFFE1FE2075BCD839AF80" box="[450,536,189,215]" captionStart="FIGURE 2" captionStartId="2.[151,250,1902,1927]" captionTargetBox="[160,1424,841,1876]" captionTargetId="figure@2.[160,1427,813,1878]" captionTargetPageId="2" captionText="FIGURE 2. Diagnostic features of Orphnebius lilizheni sp. nov. A. Head and pronotum, in dorsal view; B. Antenna; C. Spermathecal; D. Sternite VIII; E. Tergite VIII; F. Paramere; G. Male segments IX–X, dorsal view; H. Same, ventral view; I. Female segments IX–X, dorsal view; J. Same, ventral view; K. Media lobe of aedegus, ventral view; L. Same, lateral view; M. Same, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3891271" httpUri="https://zenodo.org/record/3891271/files/figure.png" pageId="3" pageNumber="596">Fig. 2F</figureCitation>
).
</paragraph>
<paragraph id="B07B9077DE5BFFE1FF2575E0DAEBAE14" blockId="3.[151,1436,153,431]" pageId="3" pageNumber="596">
Measurements. AnL
<quantity id="773C3D92DE5BFFE1FE5375E0D876AFAC" box="[433,599,225,251]" metricMagnitude="-3" metricUnit="m" metricValue="1.91" metricValueMax="1.94" metricValueMin="1.88" pageId="3" pageNumber="596" unit="mm" value="1.91" valueMax="1.94" valueMin="1.88">1.88–1.94 mm</quantity>
, BL
<quantity id="773C3D92DE5BFFE1FD6C75E1D912AFAC" box="[654,819,224,251]" metricMagnitude="-3" metricUnit="m" metricValue="4.99" metricValueMax="5.04" metricValueMin="4.94" pageId="3" pageNumber="596" unit="mm" value="4.99" valueMax="5.04" valueMin="4.94">4.94–5.04 mm</quantity>
, HL
<quantity id="773C3D92DE5BFFE1FC8975E0DE30AFAC" box="[875,1041,225,251]" metricMagnitude="-4" metricUnit="m" metricValue="8.25" metricValueMax="8.4" metricValueMin="8.1" pageId="3" pageNumber="596" unit="mm" value="0.825" valueMax="0.84" valueMin="0.81">0.81–0.84 mm</quantity>
, HW 0.99–1.00 mm, PL
<quantity id="773C3D92DE5BFFE1FACE75E0DAE4AE48" metricMagnitude="-4" metricUnit="m" metricValue="7.9" metricValueMax="8.0" metricValueMin="7.8" pageId="3" pageNumber="596" unit="mm" value="0.79" valueMax="0.8" valueMin="0.78">0.78–0.80 mm</quantity>
, PW
<quantity id="773C3D92DE5BFFE1FEE17404DB88AE48" box="[259,425,261,287]" metricMagnitude="-3" metricUnit="m" metricValue="1.1549999999999998" metricValueMax="1.17" metricValueMin="1.14" pageId="3" pageNumber="596" unit="mm" value="1.1549999999999998" valueMax="1.17" valueMin="1.14">1.14–1.17 mm</quantity>
, EL
<quantity id="773C3D92DE5BFFE1FE3D7404D8A4AE48" box="[479,645,261,287]" metricMagnitude="-4" metricUnit="m" metricValue="7.65" metricValueMax="7.7" metricValueMin="7.6" pageId="3" pageNumber="596" unit="mm" value="0.765" valueMax="0.77" valueMin="0.76">0.76–0.77 mm</quantity>
, EW
<quantity id="773C3D92DE5BFFE1FD277404D94AAE48" box="[709,875,261,287]" metricMagnitude="-3" metricUnit="m" metricValue="1.9049999999999998" metricValueMax="1.93" metricValueMin="1.88" pageId="3" pageNumber="596" unit="mm" value="1.9049999999999998" valueMax="1.93" valueMin="1.88">1.88–1.93 mm</quantity>
, AL
<quantity id="773C3D92DE5BFFE1FC417405DE68AE48" box="[931,1097,260,287]" metricMagnitude="-3" metricUnit="m" metricValue="2.605" metricValueMax="2.63" metricValueMin="2.58" pageId="3" pageNumber="596" unit="mm" value="2.605" valueMax="2.63" valueMin="2.58">2.58–2.63 mm</quantity>
, AW
<quantity id="773C3D92DE5BFFE1FB6A7404DF0FAE48" box="[1160,1326,261,287]" metricMagnitude="-3" metricUnit="m" metricValue="1.855" metricValueMax="1.88" metricValueMin="1.83" pageId="3" pageNumber="596" unit="mm" value="1.855" valueMax="1.88" valueMin="1.83">1.83–1.88 mm</quantity>
, Adl
<quantity id="773C3D92DE5BFFE1FA897404DAE7AE14" metricMagnitude="-3" metricUnit="m" metricValue="1.29" pageId="3" pageNumber="596" unit="mm" value="1.29">1.29 mm</quantity>
.
</paragraph>
<paragraph id="B07B9077DE5BFFE1FF25744CDBA5AEF8" blockId="3.[151,1436,153,431]" pageId="3" pageNumber="596">
Female (
<figureCitation id="28FF8CF2DE5BFFE1FEC8744CDBA6AE30" box="[298,391,333,359]" captionStart="FIGURE 1" captionStartId="1.[151,250,2017,2042]" captionTargetBox="[186,1398,819,1981]" captionTargetId="figure@1.[176,1412,813,1992]" captionTargetPageId="1" captionText="FIGURE 1. Habitus of Orphnebius lilizheni sp. nov. A. Male; B. Female." figureDoi="http://doi.org/10.5281/zenodo.3891269" httpUri="https://zenodo.org/record/3891269/files/figure.png" pageId="3" pageNumber="596">Figs 1B</figureCitation>
,
<figureCitation id="28FF8CF2DE5BFFE1FE74744CDB98AE30" box="[406,441,333,359]" captionStart="FIGURE 2" captionStartId="2.[151,250,1902,1927]" captionTargetBox="[160,1424,841,1876]" captionTargetId="figure@2.[160,1427,813,1878]" captionTargetPageId="2" captionText="FIGURE 2. Diagnostic features of Orphnebius lilizheni sp. nov. A. Head and pronotum, in dorsal view; B. Antenna; C. Spermathecal; D. Sternite VIII; E. Tergite VIII; F. Paramere; G. Male segments IX–X, dorsal view; H. Same, ventral view; I. Female segments IX–X, dorsal view; J. Same, ventral view; K. Media lobe of aedegus, ventral view; L. Same, lateral view; M. Same, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3891271" httpUri="https://zenodo.org/record/3891271/files/figure.png" pageId="3" pageNumber="596">2C</figureCitation>
, I–J) generally similar to male, usually slightly larger. Segments IX–X (
<figureCitation id="28FF8CF2DE5BFFE1FB17744CDF1FAE30" box="[1269,1342,333,359]" captionStart="FIGURE 2" captionStartId="2.[151,250,1902,1927]" captionTargetBox="[160,1424,841,1876]" captionTargetId="figure@2.[160,1427,813,1878]" captionTargetPageId="2" captionText="FIGURE 2. Diagnostic features of Orphnebius lilizheni sp. nov. A. Head and pronotum, in dorsal view; B. Antenna; C. Spermathecal; D. Sternite VIII; E. Tergite VIII; F. Paramere; G. Male segments IX–X, dorsal view; H. Same, ventral view; I. Female segments IX–X, dorsal view; J. Same, ventral view; K. Media lobe of aedegus, ventral view; L. Same, lateral view; M. Same, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3891271" httpUri="https://zenodo.org/record/3891271/files/figure.png" pageId="3" pageNumber="596">Figs 2</figureCitation>
I–J) distinctly modified, with long and dense setae; spermathecal (
<figureCitation id="28FF8CF2DE5BFFE1FCD67470D9AAAEDC" box="[820,907,369,395]" captionStart="FIGURE 2" captionStartId="2.[151,250,1902,1927]" captionTargetBox="[160,1424,841,1876]" captionTargetId="figure@2.[160,1427,813,1878]" captionTargetPageId="2" captionText="FIGURE 2. Diagnostic features of Orphnebius lilizheni sp. nov. A. Head and pronotum, in dorsal view; B. Antenna; C. Spermathecal; D. Sternite VIII; E. Tergite VIII; F. Paramere; G. Male segments IX–X, dorsal view; H. Same, ventral view; I. Female segments IX–X, dorsal view; J. Same, ventral view; K. Media lobe of aedegus, ventral view; L. Same, lateral view; M. Same, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3891271" httpUri="https://zenodo.org/record/3891271/files/figure.png" pageId="3" pageNumber="596">Fig. 2C</figureCitation>
) duct simply bent, proximally straight, neither twisted nor undulate.
</paragraph>
<caption id="E4BBC0FFDE5BFFE1FF757294D901A886" ID-DOI="http://doi.org/10.5281/zenodo.3891273" ID-Zenodo-Dep="3891273" httpUri="https://zenodo.org/record/3891273/files/figure.png" pageId="3" pageNumber="596" startId="3.[151,250,1941,1966]" targetBox="[151,1436,453,1916]" targetPageId="3">
<paragraph id="B07B9077DE5BFFE1FF757294D901A886" blockId="3.[151,1436,1941,2002]" pageId="3" pageNumber="596">
<emphasis id="82B04C65DE5BFFE1FF757294DB13A8F9" bold="true" box="[151,306,1941,1966]" pageId="3" pageNumber="596">FIGURE 3. A.</emphasis>
Larva of
<taxonomicName id="77C4EBF4DE5BFFE1FE777294D844A8FA" authority="Jiang &amp; Li &amp; Wang, 2020" authorityName="Jiang &amp; Li &amp; Wang" authorityYear="2020" box="[405,613,1941,1965]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="596" phylum="Arthropoda" rank="species" species="lilizheni" status="sp. nov.">
<emphasis id="82B04C65DE5BFFE1FE777294D844A8FA" box="[405,613,1941,1965]" italics="true" pageId="3" pageNumber="596">Orphnebius lilizheni</emphasis>
</taxonomicName>
<emphasis id="82B04C65DE5BFFE1FD897297D89DA8F9" bold="true" box="[619,700,1942,1966]" pageId="3" pageNumber="596">
<taxonomicNameLabel id="9983F11EDE5BFFE1FD897297D89DA8F9" box="[619,700,1942,1966]" pageId="3" pageNumber="596" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, dorsal view;
<emphasis id="82B04C65DE5BFFE1FCAE7297D942A8F9" bold="true" box="[844,867,1942,1966]" pageId="3" pageNumber="596">B.</emphasis>
Same, lateral view;
<emphasis id="82B04C65DE5BFFE1FBD17294DE6AA8F9" bold="true" box="[1075,1099,1941,1966]" pageId="3" pageNumber="596">C.</emphasis>
Same, ventral view;
<emphasis id="82B04C65DE5BFFE1FAC07297DF1BA8F9" bold="true" box="[1314,1338,1942,1966]" pageId="3" pageNumber="596">D.</emphasis>
<taxonomicName id="77C4EBF4DE5BFFE1FAA37297DFBDA8FA" authorityName="Latreille" authorityYear="1804" box="[1345,1436,1942,1965]" class="Insecta" family="Formicidae" genus="Myrmica" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="596" phylum="Arthropoda" rank="genus">
<emphasis id="82B04C65DE5BFFE1FAA37297DFBDA8FA" box="[1345,1436,1942,1965]" italics="true" pageId="3" pageNumber="596">Myrmica</emphasis>
</taxonomicName>
sp., dorsal view;
<emphasis id="82B04C65DE5BFFE1FEA672BBDB7BA885" bold="true" box="[324,346,1978,2002]" pageId="3" pageNumber="596">E.</emphasis>
Same, lateral view;
<emphasis id="82B04C65DE5BFFE1FDC872BBD81CA885" bold="true" box="[554,573,1978,2002]" pageId="3" pageNumber="596">F.</emphasis>
Larva of
<taxonomicName id="77C4EBF4DE5BFFE1FD4372BBD8DDA886" authorityName="Latreille" authorityYear="1804" box="[673,764,1978,2001]" class="Insecta" family="Formicidae" genus="Myrmica" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="596" phylum="Arthropoda" rank="genus">
<emphasis id="82B04C65DE5BFFE1FD4372BBD8DDA886" box="[673,764,1978,2001]" italics="true" pageId="3" pageNumber="596">Myrmica</emphasis>
</taxonomicName>
sp.
</paragraph>
</caption>
<paragraph id="B07B9077DE5CFFE6FF257598DF02AF80" blockId="4.[151,1436,152,287]" pageId="4" pageNumber="597">
Measurements. AnL
<quantity id="773C3D92DE5CFFE6FE537599D876AFE4" box="[433,599,152,179]" metricMagnitude="-3" metricUnit="m" metricValue="1.9249999999999998" metricValueMax="1.95" metricValueMin="1.9" pageId="4" pageNumber="597" unit="mm" value="1.9249999999999998" valueMax="1.95" valueMin="1.9">1.90–1.95 mm</quantity>
, BL 5.00–
<quantity id="773C3D92DE5CFFE6FD2F7599D912AFE4" box="[717,819,152,179]" metricMagnitude="-3" metricUnit="m" metricValue="5.17" pageId="4" pageNumber="597" unit="mm" value="5.17">5.17 mm</quantity>
, HL
<quantity id="773C3D92DE5CFFE6FC897598DE30AFE4" box="[875,1041,153,179]" metricMagnitude="-4" metricUnit="m" metricValue="8.25" metricValueMax="8.4" metricValueMin="8.1" pageId="4" pageNumber="597" unit="mm" value="0.825" valueMax="0.84" valueMin="0.81">0.81–0.84 mm</quantity>
, HW 1.00–
<quantity id="773C3D92DE5CFFE6FB717598DED8AFE4" box="[1171,1273,153,179]" metricMagnitude="-3" metricUnit="m" metricValue="1.03" pageId="4" pageNumber="597" unit="mm" value="1.03">1.03 mm</quantity>
, PL
<quantity id="773C3D92DE5CFFE6FACE7598DAE4AF80" metricMagnitude="-4" metricUnit="m" metricValue="8.1" metricValueMax="8.2" metricValueMin="8.0" pageId="4" pageNumber="597" unit="mm" value="0.81" valueMax="0.82" valueMin="0.8">0.80–0.82 mm</quantity>
, PW
<quantity id="773C3D92DE5CFFE6FEE375BCDB87AF80" box="[257,422,189,215]" metricMagnitude="-3" metricUnit="m" metricValue="1.1749999999999998" metricValueMax="1.19" metricValueMin="1.16" pageId="4" pageNumber="597" unit="mm" value="1.1749999999999998" valueMax="1.19" valueMin="1.16">1.16–1.19 mm</quantity>
, EL
<quantity id="773C3D92DE5CFFE6FE3875BCD85EAF80" box="[474,639,189,215]" metricMagnitude="-4" metricUnit="m" metricValue="7.75" metricValueMax="7.9" metricValueMin="7.6" pageId="4" pageNumber="597" unit="mm" value="0.775" valueMax="0.79" valueMin="0.76">0.76–0.79 mm</quantity>
, EW
<quantity id="773C3D92DE5CFFE6FD5F75BDD943AF80" box="[701,866,188,215]" metricMagnitude="-3" metricUnit="m" metricValue="1.915" metricValueMax="1.95" metricValueMin="1.88" pageId="4" pageNumber="597" unit="mm" value="1.915" valueMax="1.95" valueMin="1.88">1.88–1.95 mm</quantity>
, AL
<quantity id="773C3D92DE5CFFE6FC7575BCDE1DAF80" box="[919,1084,189,215]" metricMagnitude="-3" metricUnit="m" metricValue="2.6799999999999997" metricValueMax="2.73" metricValueMin="2.63" pageId="4" pageNumber="597" unit="mm" value="2.6799999999999997" valueMax="2.73" valueMin="2.63">2.63–2.73 mm</quantity>
, AW
<quantity id="773C3D92DE5CFFE6FB9B75BCDF3EAF80" box="[1145,1311,189,215]" metricMagnitude="-3" metricUnit="m" metricValue="1.88" metricValueMax="1.93" metricValueMin="1.83" pageId="4" pageNumber="597" unit="mm" value="1.88" valueMax="1.93" valueMin="1.83">1.83–1.93 mm</quantity>
.
</paragraph>
<paragraph id="B07B9077DE5CFFE6FF2575E0D800AE48" blockId="4.[151,1436,152,287]" pageId="4" pageNumber="597">
Larvae (
<figureCitation id="28FF8CF2DE5CFFE6FEC175E0DB46AFAC" box="[291,359,225,251]" captionStart="FIGURE 3" captionStartId="3.[151,250,1941,1966]" captionTargetBox="[151,1436,453,1916]" captionTargetId="figure@3.[151,1436,453,1916]" captionTargetPageId="3" captionText="FIGURE 3. A. Larva of Orphnebius lilizheni sp. nov., dorsal view; B. Same, lateral view; C. Same, ventral view; D. Myrmica sp., dorsal view; E. Same, lateral view; F. Larva of Myrmica sp." figureDoi="http://doi.org/10.5281/zenodo.3891273" httpUri="https://zenodo.org/record/3891273/files/figure.png" pageId="4" pageNumber="597">Figs 3</figureCitation>
A–C, third instar), ivory-white, fat, soft and much weakly osteogenated, more or less resembling the larvae of
<taxonomicName id="77C4EBF4DE5CFFE6FEC87407DBAEAE48" authorityName="Latreille" authorityYear="1804" box="[298,399,262,287]" class="Insecta" family="Formicidae" genus="Myrmica" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="597" phylum="Arthropoda" rank="genus">
<emphasis id="82B04C65DE5CFFE6FEC87407DBAEAE48" box="[298,399,262,287]" italics="true" pageId="4" pageNumber="597">Myrmica</emphasis>
</taxonomicName>
sp (
<figureCitation id="28FF8CF2DE5CFFE6FE5C7404D835AE48" box="[446,532,261,287]" captionStart="FIGURE 3" captionStartId="3.[151,250,1941,1966]" captionTargetBox="[151,1436,453,1916]" captionTargetId="figure@3.[151,1436,453,1916]" captionTargetPageId="3" captionText="FIGURE 3. A. Larva of Orphnebius lilizheni sp. nov., dorsal view; B. Same, lateral view; C. Same, ventral view; D. Myrmica sp., dorsal view; E. Same, lateral view; F. Larva of Myrmica sp." figureDoi="http://doi.org/10.5281/zenodo.3891273" httpUri="https://zenodo.org/record/3891273/files/figure.png" pageId="4" pageNumber="597">Fig. 3F</figureCitation>
).
</paragraph>
<caption id="E4BBC0FFDE5CFFE6FF7573C7D94DA870" ID-DOI="http://doi.org/10.5281/zenodo.3891275" ID-Zenodo-Dep="3891275" httpUri="https://zenodo.org/record/3891275/files/figure.png" pageId="4" pageNumber="597" startId="4.[151,250,1734,1759]" targetBox="[151,1436,309,1709]" targetPageId="4">
<paragraph id="B07B9077DE5CFFE6FF7573C7D94DA870" blockId="4.[151,1437,1734,1831]" pageId="4" pageNumber="597">
<emphasis id="82B04C65DE5CFFE6FF7573C7DB35A988" bold="true" box="[151,276,1734,1759]" pageId="4" pageNumber="597">FIGURE 4.</emphasis>
Habitat of
<taxonomicName id="77C4EBF4DE5CFFE6FE6973C7D87DA989" authority="Jiang &amp; Li &amp; Wang, 2020" authorityName="Jiang &amp; Li &amp; Wang" authorityYear="2020" box="[395,604,1734,1758]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="597" phylum="Arthropoda" rank="species" species="lilizheni" status="sp. nov.">
<emphasis id="82B04C65DE5CFFE6FE6973C7D87DA989" box="[395,604,1734,1758]" italics="true" pageId="4" pageNumber="597">Orphnebius lilizheni</emphasis>
</taxonomicName>
<emphasis id="82B04C65DE5CFFE6FD8173C6D8F7A988" bold="true" box="[611,726,1734,1759]" pageId="4" pageNumber="597">
<taxonomicNameLabel id="9983F11EDE5CFFE6FD8173C6D894A988" box="[611,693,1735,1759]" pageId="4" pageNumber="597" rank="species">sp. nov.</taxonomicNameLabel>
A.
</emphasis>
Adults and larvae in a nest of
<taxonomicName id="77C4EBF4DE5CFFE6FBF673C6DE4EA989" authorityName="Latreille" authorityYear="1804" box="[1044,1135,1735,1758]" class="Insecta" family="Formicidae" genus="Myrmica" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="597" phylum="Arthropoda" rank="genus">
<emphasis id="82B04C65DE5CFFE6FBF673C6DE4EA989" box="[1044,1135,1735,1758]" italics="true" pageId="4" pageNumber="597">Myrmica</emphasis>
</taxonomicName>
sp.;
<emphasis id="82B04C65DE5CFFE6FB4173C6DE9BA988" bold="true" box="[1187,1210,1735,1759]" pageId="4" pageNumber="597">B.</emphasis>
A
<taxonomicName id="77C4EBF4DE5CFFE6FB3873C6DF14A989" authorityName="Latreille" authorityYear="1804" box="[1242,1333,1735,1758]" class="Insecta" family="Formicidae" genus="Myrmica" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="597" phylum="Arthropoda" rank="genus">
<emphasis id="82B04C65DE5CFFE6FB3873C6DF14A989" box="[1242,1333,1735,1758]" italics="true" pageId="4" pageNumber="597">Myrmica</emphasis>
</taxonomicName>
sp. ant in nest;
<emphasis id="82B04C65DE5CFFE6FF2C73EBDAC7A854" bold="true" box="[206,230,1770,1795]" pageId="4" pageNumber="597">C.</emphasis>
General environment of the type locality: Baotianman (Xizang, China). Laa—Larva of the host ant; La—Larva of
<taxonomicName id="77C4EBF4DE5CFFE6FA9A73EADB6AA871" authority="Jiang &amp; Li &amp; Wang, 2020" authorityName="Jiang &amp; Li &amp; Wang" authorityYear="2020" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="597" phylum="Arthropoda" rank="species" species="lilizheni" status="sp. nov.">
<emphasis id="82B04C65DE5CFFE6FA9A73EADB6AA871" italics="true" pageId="4" pageNumber="597">Orphnebius lilizheni</emphasis>
</taxonomicName>
<emphasis id="82B04C65DE5CFFE6FEB3720EDB80A870" bold="true" box="[337,417,1807,1831]" pageId="4" pageNumber="597">
<taxonomicNameLabel id="9983F11EDE5CFFE6FEB3720EDB80A870" box="[337,417,1807,1831]" pageId="4" pageNumber="597" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; ad—Adults of
<taxonomicName id="77C4EBF4DE5CFFE6FDA4720FD937A871" authority="Jiang &amp; Li &amp; Wang, 2020" authorityName="Jiang &amp; Li &amp; Wang" authorityYear="2020" box="[582,790,1806,1830]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="597" phylum="Arthropoda" rank="species" species="lilizheni" status="sp. nov.">
<emphasis id="82B04C65DE5CFFE6FDA4720FD937A871" box="[582,790,1806,1830]" italics="true" pageId="4" pageNumber="597">Orphnebius lilizheni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9983F11EDE5CFFE6FCFE720ED94DA870" box="[796,876,1807,1831]" pageId="4" pageNumber="597" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="B07B9077DE5CFFE7FF257255DEACAFAC" blockId="4.[151,1436,1876,2011]" lastBlockId="5.[151,1436,153,431]" lastPageId="5" lastPageNumber="598" pageId="4" pageNumber="597">
<emphasis id="82B04C65DE5CFFE6FF257255DB95A838" bold="true" box="[199,436,1876,1903]" pageId="4" pageNumber="597">Comparative Notes.</emphasis>
The new species is similar to
<taxonomicName id="77C4EBF4DE5CFFE6FD1C7254DE8EA838" authority="Assing, 2006" authorityName="Assing" authorityYear="2006" box="[766,1199,1877,1903]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="597" phylum="Arthropoda" rank="species" species="longistriatus">
<emphasis id="82B04C65DE5CFFE6FD1C7254DE37A838" box="[766,1046,1877,1903]" italics="true" pageId="4" pageNumber="597">Orphnebius longistriatus</emphasis>
Assing, 2006
</taxonomicName>
and
<taxonomicName id="77C4EBF4DE5CFFE6FB017254DB21A8C4" authority="Assing, 2006" authorityName="Assing" authorityYear="2006" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="597" phylum="Arthropoda" rank="species" species="hamatus">
<emphasis id="82B04C65DE5CFFE6FB017254DF42A838" box="[1251,1379,1877,1903]" italics="true" pageId="4" pageNumber="597">O. hamatus</emphasis>
Assing,
<quantity id="773C3D92DE5CFFE6FF2A7278DB21A8C4" box="[200,256,1913,1939]" metricMagnitude="1" metricUnit="m" metricValue="5.0952399999999995" pageId="4" pageNumber="597" unit="in" value="2006.0">2006</quantity>
</taxonomicName>
in general appearance. But
<taxonomicName id="77C4EBF4DE5CFFE6FDD37278D88DA8C4" authority="Jiang &amp; Li &amp; Wang, 2020" authorityName="Jiang &amp; Li &amp; Wang" authorityYear="2020" box="[561,684,1913,1939]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="597" phylum="Arthropoda" rank="species" species="lilizheni" status="sp. nov.">
<emphasis id="82B04C65DE5CFFE6FDD37278D88DA8C4" box="[561,684,1913,1939]" italics="true" pageId="4" pageNumber="597">O. lilizheni</emphasis>
</taxonomicName>
<emphasis id="82B04C65DE5CFFE6FD507278D92BA8C4" bold="true" box="[690,778,1913,1939]" pageId="4" pageNumber="597">
<taxonomicNameLabel id="9983F11EDE5CFFE6FD507278D92BA8C4" box="[690,778,1913,1939]" pageId="4" pageNumber="597" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has concolorous abdominal segments, while the abdominal segments III–V, VI and VII–VIII are respectively blackish, brown and reddish in
<taxonomicName id="77C4EBF4DE5CFFE6FBC4729CDEF7A8E0" authorityName="Assing" authorityYear="2006" box="[1062,1238,1949,1975]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="597" phylum="Arthropoda" rank="species" species="longistriatus">
<emphasis id="82B04C65DE5CFFE6FBC4729CDEF7A8E0" box="[1062,1238,1949,1975]" italics="true" pageId="4" pageNumber="597">O. longistriatus</emphasis>
</taxonomicName>
. The new species can be distinguished from
<taxonomicName id="77C4EBF4DE5CFFE6FE5D72C0D861A88C" authorityName="Assing" authorityYear="2006" box="[447,576,1985,2011]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="597" phylum="Arthropoda" rank="species" species="hamatus">
<emphasis id="82B04C65DE5CFFE6FE5D72C0D861A88C" box="[447,576,1985,2011]" italics="true" pageId="4" pageNumber="597">O. hamatus</emphasis>
</taxonomicName>
by the different form of median lobe of aedeagus.
<taxonomicName id="77C4EBF4DE5CFFE6FB9772C0DED7A88C" authorityName="Assing" authorityYear="2006" box="[1141,1270,1985,2011]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="597" phylum="Arthropoda" rank="species" species="hamatus">
<emphasis id="82B04C65DE5CFFE6FB9772C0DED7A88C" box="[1141,1270,1985,2011]" italics="true" pageId="4" pageNumber="597">O. hamatus</emphasis>
</taxonomicName>
possess a very large median lobe, and the ventral process is short, broad and apically curved, while the median lobe of the new species is simple and short, and the ventral process of the new species is more strongly curved. The paramere of
<taxonomicName id="77C4EBF4DE5DFFE7FA6375BCDAD7AFAC" authorityName="Assing" authorityYear="2006" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="598" phylum="Arthropoda" rank="species" species="hamatus">
<emphasis id="82B04C65DE5DFFE7FA6375BCDAD7AFAC" italics="true" pageId="5" pageNumber="598">O. hamatus</emphasis>
</taxonomicName>
has four long and two short setae while the new species with only two long setae.
</paragraph>
</subSubSection>
<subSubSection id="F8DEC3FCDE5DFFE7FF257405DACAAE30" pageId="5" pageNumber="598" type="distribution">
<paragraph id="B07B9077DE5DFFE7FF257405D833AE48" blockId="5.[151,1436,153,431]" box="[199,530,260,287]" pageId="5" pageNumber="598">
<emphasis id="82B04C65DE5DFFE7FF257405DB40AE48" bold="true" box="[199,353,260,287]" pageId="5" pageNumber="598">Distribution.</emphasis>
<collectingCountry id="C8D3D0E7DE5DFFE7FE857404DB8FAE48" box="[359,430,261,287]" name="China" pageId="5" pageNumber="598">China</collectingCountry>
:
<collectingRegion id="72005E95DE5DFFE7FE5B7404D82CAE48" box="[441,525,261,287]" country="China" name="Xizang" pageId="5" pageNumber="598">Xizang</collectingRegion>
.
</paragraph>
<paragraph id="B07B9077DE5DFFE7FF257429DACAAE30" blockId="5.[151,1436,153,431]" pageId="5" pageNumber="598">
<emphasis id="82B04C65DE5DFFE7FF257429DBACAE14" bold="true" box="[199,397,296,323]" pageId="5" pageNumber="598">Biological notes.</emphasis>
The adults and larvae of this new species were collected in a nest of
<taxonomicName id="77C4EBF4DE5DFFE7FB2A742BDF0CAE14" authorityName="Latreille" authorityYear="1804" box="[1224,1325,298,323]" class="Insecta" family="Formicidae" genus="Myrmica" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="598" phylum="Arthropoda" rank="genus">
<emphasis id="82B04C65DE5DFFE7FB2A742BDF0CAE14" box="[1224,1325,298,323]" italics="true" pageId="5" pageNumber="598">Myrmica</emphasis>
</taxonomicName>
sp. (
<figureCitation id="28FF8CF2DE5DFFE7FA887428DA87AE30" captionStart="FIGURE 4" captionStartId="4.[151,250,1734,1759]" captionTargetBox="[151,1436,309,1709]" captionTargetId="figure@4.[151,1436,309,1709]" captionTargetPageId="4" captionText="FIGURE 4. Habitat of Orphnebius lilizheni sp. nov. A. Adults and larvae in a nest of Myrmica sp.; B. A Myrmica sp. ant in nest; C. General environment of the type locality: Baotianman (Xizang, China). Laa—Larva of the host ant; La—Larva of Orphnebius lilizheni sp. nov.; ad—Adults of Orphnebius lilizheni sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3891275" httpUri="https://zenodo.org/record/3891275/files/figure.png" pageId="5" pageNumber="598">Figs 4</figureCitation>
A–B).
</paragraph>
</subSubSection>
<subSubSection id="F8DEC3FCDE5DFFE7FF257470DEE2AEF8" pageId="5" pageNumber="598" type="etymology">
<paragraph id="B07B9077DE5DFFE7FF257470DEE2AEF8" blockId="5.[151,1436,153,431]" pageId="5" pageNumber="598">
<emphasis id="82B04C65DE5DFFE7FF257470DB6FAEDC" bold="true" box="[199,334,369,395]" pageId="5" pageNumber="598">Etymology.</emphasis>
This species is named in honor of Prof. Li-Zhen Li (
<collectingRegion id="72005E95DE5DFFE7FC407470DE2DAEDC" box="[930,1036,369,395]" country="China" name="Shanghai" pageId="5" pageNumber="598">Shanghai</collectingRegion>
Normal University,
<collectingRegion id="72005E95DE5DFFE7FB167470DF7EAEDC" box="[1268,1375,369,395]" country="China" name="Shanghai" pageId="5" pageNumber="598">Shanghai</collectingRegion>
,
<collectingCountry id="C8D3D0E7DE5DFFE7FA887470DA94AEF8" name="China" pageId="5" pageNumber="598">China</collectingCountry>
), a famous Chinese insect taxonomist, who works on the Staphylinid beetles for a long time.
</paragraph>
</subSubSection>
</treatment>
</document>
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