<documentid="01FCA65519F3D982C22E1EE8A54346A1"ID-DOI="10.5852/ejt.2024.973.2765"ID-ISSN="2118-9773"ID-Zenodo-Dep="14440264"ID-ZooBank="524B5B20-A190-4023-AC2B-7B48A725930A"IM.bibliography_approvedBy="jonas"IM.illustrations_approvedBy="felipe"IM.materialsCitations_approvedBy="jonas"IM.metadata_approvedBy="jonas"IM.tables_approvedBy="jonas"IM.taxonomicNames_approvedBy="jonas"IM.treatmentCitations_approvedBy="jonas"IM.treatments_approvedBy="jonas"checkinTime="1734081370392"checkinUser="plazi"docAuthor="Herbert, David G."docDate="2024"docId="714FF54E37373217FDCCFB65FDA9FB94"docLanguage="en"docName="EJT.2024.973.1-143.pdf"docOrigin="European Journal of Taxonomy 973"docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2765/12655"docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1"docStyleId="EF2B578F1D15862ADE45B0C07C620911"docStyleName="EJT.2018-.journal_article.type1"docStyleVersion="14"docTitle="Tylorhaphe fritillaria Herbert 2024, gen. et sp. nov."docType="treatment"docVersion="5"lastPageNumber="113"masterDocId="8D768D36375A3267FFF2FFF1FFF6FFA0"masterDocTitle="The Umboniinae (Mollusca: Gastropoda: Trochidae) of New Caledonia, with descriptions of two new genera and eight new species, plus an additional new species from the Bismarck Sea"masterLastPageNumber="143"masterPageNumber="1"pageNumber="110"updateTime="1735266715814"updateUser="jonas"zenodo-license-document="CC-BY-4.0"zenodo-license-figures="CC-BY-4.0"zenodo-license-treatments="UNSPECIFIED">
<mods:titleid="BED5F1CADB15B6F7E5C39E7731BFEABD">The Umboniinae (Mollusca: Gastropoda: Trochidae) of New Caledonia, with descriptions of two new genera and eight new species, plus an additional new species from the Bismarck Sea</mods:title>
<mods:affiliationid="034D404CE9A8E6BB715DB28FF11A7A48">Department of Natural Sciences, National Museum Wales, Cathays Park, Cardiff, CF 10 3 NP, UK. Institute of Systematics, Evolution, Biodiversity (ISYEB), Muséum national d’Histoire naturelle, CNRS, SU, EPHE, UA, CP 51, 57 rue Cuvier, 75005 Paris, France.</mods:affiliation>
<taxonomicNameid="3EE63FDB3737320AFDCCFB65FCC9FB0F"authority="Herbert, 2024"authorityName="Herbert"authorityYear="2024"box="[574,831,1172,1199]"class="Gastropoda"family="Trochidae"genus="Tylorhaphe"kingdom="Animalia"order="Trochida"pageId="109"pageNumber="110"phylum="Mollusca"rank="species"species="fritillaria"status="gen. et sp. nov.">
<taxonomicNameLabelid="D0A125313737320AFCB4FB64FC00FB0F"box="[838,1014,1173,1199]"pageId="109"pageNumber="110"rank="species">gen. et sp. nov.</taxonomicNameLabel>
<figureCitationid="61DD58DD3737320AFD26FB2EFCA9FB59"box="[724,863,1247,1273]"captionStart-0="Fig"captionStart-1="Fig"captionStart-2="Fig"captionStartId-0="110.[189,232,1964,1990]"captionStartId-1="111.[189,232,1035,1061]"captionStartId-2="111.[189,232,1961,1987]"captionTargetBox-0="[209,1379,924,1907]"captionTargetBox-1="[193,1395,681,995]"captionTargetBox-2="[189,1399,1162,1929]"captionTargetId-0="figure-263@110.[189,1399,909,1922]"captionTargetId-1="figure-176@111.[189,1399,678,999]"captionTargetId-2="figure-177@111.[189,1399,1162,1929]"captionTargetPageId-0="110"captionTargetPageId-1="111"captionTargetPageId-2="111"captionText-0="Fig. 60. Tylorhaphe fritillaria gen. et sp. nov. A–D. Holotype, Koumac, Stn 1322, height 2.5 mm, diameter 3.6 mm (MNHN-IM-2000-38871)."captionText-1="Fig. 61. Tylorhaphe fritillaria gen. et sp. nov., Lifou, Stn 1461 (paratype, MNHN-IM-2000-38873). A. Apical view of spire. B. Oblique view of spire, scale bar =0.5 mm. C. Protoconch. Scale bars: A–B =0.5 mm; C =100."captionText-2="Fig. 62. Map of New Caledonia showing the distribution of Tylorhaphe fritillaria gen. et sp. nov. Type locality indicated in pink."figureDoi-0="http://doi.org/10.5281/zenodo.14440443"figureDoi-1="http://doi.org/10.5281/zenodo.14440445"figureDoi-2="http://doi.org/10.5281/zenodo.14440447"httpUri-0="https://zenodo.org/record/14440443/files/figure.png"httpUri-1="https://zenodo.org/record/14440445/files/figure.png"httpUri-2="https://zenodo.org/record/14440447/files/figure.png"pageId="109"pageNumber="110">Figs 60– 62</figureCitation>
<taxonomicNameid="3EE63FDB3737320AFEB8FAA2FDD5FACD"authorityName="Herbert"authorityYear="2024"box="[330,547,1363,1389]"class="Gastropoda"family="Trochidae"genus="Tylorhaphe"kingdom="Animalia"order="Trochida"pageId="109"pageNumber="110"phylum="Mollusca"rank="species"species="alisae"status="gen. et sp. nov.">
<taxonomicNameLabelid="D0A125313737320AFDC0FAA5FD0FFACE"box="[562,761,1364,1390]"pageId="109"pageNumber="110"rank="species">gen. et sp. nov.</taxonomicNameLabel>
, but with a narrower umbilicus, the rim of which conspicuously overhangs the underlying umbilical cavity; rim pleated, but peri-umbilical spiral sculpture lacking; basal colour pattern of numerous white blotches on a pale maroon-brown ground is distinctive.
<collectingRegionid="3B228ABA3737320AFE23F875FD62F83E"box="[465,660,1924,1950]"country="New Caledonia"name="Province des Iles"pageId="109"pageNumber="110">Loyalty Islands</collectingRegion>
<locationid="FC3912833737320AFC1BF875FB6FF83E"LSID="urn:lsid:plazi:treatment:714FF54E37373217FDCCFB65FDA9FB94:FC3912833737320AFC1BF875FB6FF83E"box="[1001,1177,1924,1950]"country="New Caledonia"county="Lifou"latitude="-20.9"longLatPrecision="1266"longitude="167.03334"name="Baie du Santal"pageId="109"pageNumber="110"stateProvince="Loyalty Islands">Baie du Santal</location>
. Small (diameter up to 4.0 mm), trochiform, relatively solid, spire height moderate (H/D = 0.65- 0.75); teleoconch of up to 4.75 whorls; periphery roundly angled, situated close to mid-whorl; suture level with periphery; base relatively narrowly umbilicate. First teleoconch whorl with four fine spiral lirae; two stronger cords develop during second whorl, one at shoulder, the other between this and abapical suture; a third spiral cord arises below adapical suture during third whorl, strengthening and becoming beaded on subsequent whorls; shoulder cord becomes progressively less distinct during fourth whorl and additional spiral lirae develop between cords, particularly between subsutural cord and vestige of shoulder cord; peripheral cord emerges from suture at on last whorl, delineating upper limit of peripheral angulation; whorl profile distinctly concave between peripheral and supraperipheral cords, flatter between supraperipheral and subsutural cords; axial sculpture weak, comprising only indistinct growth-lines, most noticeable in final quarter whorl, microsculpture of axial threads throughout. Base with 3–4 spiral cords below periphery, its mid-region smoother with only collabral growth-lines, sinuous below periphery; peri-umbilical region with strong pleats radiating from umbilical rim, but without spiral sculpture; umbilicus relatively narrow, its rim thickened and roundly angled, crenulated by radiating pleats; sides of umbilicus steep, greatly overhung by umbilical rim, with low spiral cords within; a funicle lacking. Aperture subquadrate; peristome interrupted in parietal region; columella stout, with a low bulge at its base and a shallow notch at end of thickened umbilical rim, at junction of columella and basal lips; outer lip not descendant in material available, shallowly indented below periphery; interior of outer lip smooth, somewhat thickened within.
<taxonomicNameid="3EE63FDB37343209FEDBF85DFDD0F866"authorityName="Herbert"authorityYear="2024"box="[297,550,1964,1990]"class="Gastropoda"family="Trochidae"genus="Tylorhaphe"kingdom="Animalia"order="Trochida"pageId="110"pageNumber="111"phylum="Mollusca"rank="species"species="fritillaria"status="gen. et sp. nov.">
<taxonomicNameLabelid="D0A1253137343209FDC0F85DFD19F866"box="[562,751,1964,1990]"pageId="110"pageNumber="111"rank="species">gen. et sp. nov.</taxonomicNameLabel>
. Pattern variable; ground colour typically whitish, mottled with spots, blotches and lines in shades of brownish-yellow to olive-brown; a spiral band of narrow, alternately white and brownish-yellow axial lines above supraperipheral cord; peripheral cord with small, widely-spaced, maroon-red spots; base with a maroon-brown ground densely spotted with white blotches; umbilical rim and columella whitish. Some specimens more uniformly pale orange above periphery, with pinkish subsutural, shoulder and peripheral cords. Basal ground colour turning rose-pink in older post-mortem specimens.
<taxonomicNameid="3EE63FDB37353208FED5FBFDFDD2FB86"authorityName="Herbert"authorityYear="2024"box="[295,548,1036,1062]"class="Gastropoda"family="Trochidae"genus="Tylorhaphe"kingdom="Animalia"order="Trochida"pageId="111"pageNumber="112"phylum="Mollusca"rank="species"species="fritillaria"status="gen. et sp. nov.">
<taxonomicNameLabelid="D0A1253137353208FDDDFBFDFD1AFB86"box="[559,748,1036,1062]"pageId="111"pageNumber="112"rank="species">gen. et sp. nov.</taxonomicNameLabel>
<taxonomicNameid="3EE63FDB37353208FC71F858FB8AF863"authorityName="Herbert"authorityYear="2024"box="[899,1148,1961,1987]"class="Gastropoda"family="Trochidae"genus="Tylorhaphe"kingdom="Animalia"order="Trochida"pageId="111"pageNumber="112"phylum="Mollusca"rank="species"species="fritillaria"status="gen. et sp. nov.">
<taxonomicNameLabelid="D0A1253137353208FB76F85BFAC3F864"box="[1156,1333,1962,1988]"pageId="111"pageNumber="112"rank="species">gen. et sp. nov.</taxonomicNameLabel>
<figureCitationid="61DD58DD372A3217FEACFEBBFE3CFEC4"box="[350,458,330,356]"captionStart="Fig"captionStartId="111.[189,232,1035,1061]"captionTargetBox="[193,1395,681,995]"captionTargetId="figure-176@111.[189,1399,678,999]"captionTargetPageId="111"captionText="Fig. 61. Tylorhaphe fritillaria gen. et sp. nov., Lifou, Stn 1461 (paratype, MNHN-IM-2000-38873). A. Apical view of spire. B. Oblique view of spire, scale bar =0.5 mm. C. Protoconch. Scale bars: A–B =0.5 mm; C =100."figureDoi="http://doi.org/10.5281/zenodo.14440445"httpUri="https://zenodo.org/record/14440445/files/figure.png"pageId="112"pageNumber="113">Fig. 61C</figureCitation>
). Somewhat worn, but evidently similar to that of
<taxonomicNameid="3EE63FDB372A3217FBECFEBBFAD6FEC4"authorityName="Herbert"authorityYear="2024"box="[1054,1312,330,356]"class="Gastropoda"family="Trochidae"genus="Tylorhaphe"kingdom="Animalia"order="Trochida"pageId="112"pageNumber="113"phylum="Mollusca"rank="species"species="luteopicta"status="gen. et sp. nov.">
<figureCitationid="61DD58DD372A3217FE94FD72FE4AFD3D"box="[358,444,643,669]"captionStart="Fig"captionStartId="111.[189,232,1961,1987]"captionTargetBox="[189,1399,1162,1929]"captionTargetId="figure-177@111.[189,1399,1162,1929]"captionTargetPageId="111"captionText="Fig. 62. Map of New Caledonia showing the distribution of Tylorhaphe fritillaria gen. et sp. nov. Type locality indicated in pink."figureDoi="http://doi.org/10.5281/zenodo.14440447"httpUri="https://zenodo.org/record/14440447/files/figure.png"pageId="112"pageNumber="113">Fig. 62</figureCitation>
<collectingRegionid="3B228ABA372A3217FCC3FD43FC1DFD6C"box="[817,1003,690,716]"country="New Caledonia"name="Province des Iles"pageId="112"pageNumber="113">Loyalty Islands</collectingRegion>
<taxonomicNameid="3EE63FDB372A3217FED9FCD3FDF4FC9C"authorityName="Herbert"authorityYear="2024"box="[299,514,802,828]"class="Gastropoda"family="Trochidae"genus="Tylorhaphe"kingdom="Animalia"order="Trochida"pageId="112"pageNumber="113"phylum="Mollusca"rank="species"species="alisae"status="gen. et sp. nov.">
<taxonomicNameLabelid="D0A12531372A3217FDFCFCD2FD26FC9D"box="[526,720,803,829]"pageId="112"pageNumber="113"rank="species">gen. et sp. nov.</taxonomicNameLabel>
is provisional as data regarding its radula morphology are wanting. However, conchological features, in particular the ontogeny of the spiral sculpture, the strongly beaded subsutural cord, the angular periphery and the morphology of the umbilical region, are consistent with those of
Its small size, narrow umbilicus without peri-umbilical spiral cords, and distinctive basal colour pattern render it easily separable from other species in the genus. The most similar species,
, has a broader umbilicus bordered by distinct spiral cords, and its umbilical rim does not overhang the underlying umbilical cavity to the same extent.
