treatments-xml/data/C4/18/0E/C4180E85D7E15724905623B0878186A7.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.909.46838" ID-GBIF-Dataset="5cc495f4-4a41-41a0-b362-0e6d8c629b52" ID-PMC="PMC7015954" ID-Pensoft-Pub="1313-2970-909-79" ID-Pensoft-UUID="0AEF1785B8655964A36D859FBA7D8755" ID-PubMed="32089636" ID-ZooBank="22B7FF16D0A240F9B20EF7C6E0AF1842" ModsDocID="1313-2970-909-79" checkinTime="1580983798284" checkinUser="pensoft" docAuthor="Vujic, Ante, Likov, Laura, Radenkovic, Snezana, Tubic, Natasa Kocis, Djan, Mihajla, Sebic, Anja, Perez-Banon, Celeste, Barkalov, Anatolij, Hayat, Ruestem, Rojo, Santos, Andric, Andrijana &amp; Stahls, Gunilla" docDate="2020" docId="C4180E85D7E15724905623B0878186A7" docLanguage="en" docName="ZooKeys 909: 79-158" docOrigin="ZooKeys 909" docSource="http://dx.doi.org/10.3897/zookeys.909.46838" docTitle="Merodon medium Vujic, Likov &amp; Radenkovic 2020, sp. nov." docType="treatment" docUuid="E384B35E-E377-49B6-90CE-DEBDCBCEDDCB" docUuidSource="ZooBank" docVersion="4" id="0AEF1785B8655964A36D859FBA7D8755" lastPageNumber="79" masterDocId="0AEF1785B8655964A36D859FBA7D8755" masterDocTitle="Revision of the Merodon serrulatus group (Diptera, Syrphidae)" masterLastPageNumber="158" masterPageNumber="79" pageNumber="79" updateTime="1668168397753" updateUser="ExternalLinkService">
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<mods:title>Revision of the Merodon serrulatus group (Diptera, Syrphidae)</mods:title>
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<mods:namePart>Vujic, Ante</mods:namePart>
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<mods:namePart>Likov, Laura</mods:namePart>
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<mods:namePart>Radenkovic, Snezana</mods:namePart>
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<mods:namePart>Tubic, Natasa Kocis</mods:namePart>
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<mods:namePart>Djan, Mihajla</mods:namePart>
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<mods:namePart>Sebic, Anja</mods:namePart>
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<mods:namePart>Perez-Banon, Celeste</mods:namePart>
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<mods:namePart>Barkalov, Anatolij</mods:namePart>
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<mods:namePart>Hayat, Ruestem</mods:namePart>
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<mods:namePart>Rojo, Santos</mods:namePart>
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<mods:namePart>Andric, Andrijana</mods:namePart>
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<mods:namePart>Stahls, Gunilla</mods:namePart>
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<treatment ID-GBIF-Taxon="162047731" LSID="urn:lsid:zoobank.org:act:E384B35E-E377-49B6-90CE-DEBDCBCEDDCB" httpUri="http://treatment.plazi.org/id/C4180E85D7E15724905623B0878186A7" lastPageNumber="79" pageId="0" pageNumber="79">
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<paragraph pageId="0" pageNumber="79">
<taxonomicName LSID="C4180E85-D7E1-5724-9056-23B0878186A7" authority="Vujić, Likov &amp; Radenković" authorityName="Vujic, Likov &amp; Radenkovic" authorityYear="2020" class="Insecta" family="Syrphidae" genus="Merodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Merodon medium" order="Diptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="medium" status="sp. nov.">
Merodon medium 
<normalizedToken originalValue="Vujić">Vujic</normalizedToken>
, Likov &amp; 
<normalizedToken originalValue="Radenković">Radenkovic</normalizedToken>
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<taxonomicNameLabel pageId="0" pageNumber="79">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Male genitalia, surstylus, lateral view. A Merodon serrulatus, Spain B Merodon serrulatus, France C Merodon serrulatus, Greece (Pindos) D Merodon serrulatus, Greece (Olympos) E Merodon serrulatus, Greece (Peloponnese) F Merodon serrulatus, Montenegro G Merodon serrulatus, Russia H Merodon medium sp. nov. G, H margin of anterior surstyle lobe marked with arrow. Abbreviation: bp-basolateral protrusion. Scale bar: 0.2 mm." figureDoi="10.3897/zookeys.909.46838.figure6" httpUri="https://binary.pensoft.net/fig/377367" pageId="0" pageNumber="79">Figs 6H</figureCitation>
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Antenna. A Merodon hirsutus, outer side, lateral view, male B Merodon hirsutus, inner side, lateral view, male C Merodon hirsutus, dorsal view, male D Merodon hirsutus, outer side, lateral view, female E Merodon kawamurae, outer side, lateral view, male F Merodon kawamurae, inner side, lateral view, male G Merodon kawamurae, dorsal view, male H Merodon kawamurae, outer side, lateral view, female I Merodon medium sp. nov., outer side, lateral view, male J Merodon medium sp. nov., outer side, lateral view, female. Scale bar: 1 mm." figureDoi="10.3897/zookeys.909.46838.figure19" httpUri="https://binary.pensoft.net/fig/377380" pageId="0" pageNumber="79">, 19I, J</figureCitation>
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. A, B metaleg, C, D metatrochanter, metafemur and metatibia, lateral view. A Merodon kawamurae, male B Merodon kawamurae, female C Merodon medium sp. nov., male D Merodon medium sp. nov., female. Scale bar: 2 mm." figureDoi="10.3897/zookeys.909.46838.figure22" httpUri="https://binary.pensoft.net/fig/377383" pageId="0" pageNumber="79">, 22C, D</figureCitation>
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Merodon medium sp. nov., abdomen, dorsal view. A male B female. Scale bar: 2 mm." figureDoi="10.3897/zookeys.909.46838.figure24" httpUri="https://binary.pensoft.net/fig/377385" pageId="0" pageNumber="79">, 24A, B</figureCitation>
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Male genitalia. A Merodon medium sp. nov., epandrium, lateral view B Merodon medium sp. nov., epandrium, ventral view C Merodon medium sp. nov., hypandrium, lateral view D Merodon nigrocapillatus sp. nov., epandrium, lateral view E Merodon nigrocapillatus sp. nov., epandrium, ventral view F Merodon nigrocapillatus sp. nov., hypandrium, lateral view. Abbreviations: al-anterior surstyle lobe, bp-basolateral protrusion, l-lingula; arrow marks the hook-like extension in A. Scale bar: 0.2 mm." figureDoi="10.3897/zookeys.909.46838.figure25" httpUri="https://binary.pensoft.net/fig/377386" pageId="0" pageNumber="79">, 25A-C</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="diagnosis">
<paragraph pageId="0" pageNumber="79">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="79">
Large species (10.3-13 mm) with wide dark brown abdomen and yellow-orange maculae on lateral sides of tergum 2 (Fig. 
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Merodon medium sp. nov., abdomen, dorsal view. A male B female. Scale bar: 2 mm." figureDoi="10.3897/zookeys.909.46838.figure24" httpUri="https://binary.pensoft.net/fig/377385" pageId="0" pageNumber="79">24A, B</figureCitation>
); basoflagellomere elongated, ca. 2.5 times longer than broad (Fig. 
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Antenna. A Merodon hirsutus, outer side, lateral view, male B Merodon hirsutus, inner side, lateral view, male C Merodon hirsutus, dorsal view, male D Merodon hirsutus, outer side, lateral view, female E Merodon kawamurae, outer side, lateral view, male F Merodon kawamurae, inner side, lateral view, male G Merodon kawamurae, dorsal view, male H Merodon kawamurae, outer side, lateral view, female I Merodon medium sp. nov., outer side, lateral view, male J Merodon medium sp. nov., outer side, lateral view, female. Scale bar: 1 mm." figureDoi="10.3897/zookeys.909.46838.figure19" httpUri="https://binary.pensoft.net/fig/377380" pageId="0" pageNumber="79">19 I, J</figureCitation>
,); metafemur incrassate (Fig. 
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. A, B metaleg, C, D metatrochanter, metafemur and metatibia, lateral view. A Merodon kawamurae, male B Merodon kawamurae, female C Merodon medium sp. nov., male D Merodon medium sp. nov., female. Scale bar: 2 mm." figureDoi="10.3897/zookeys.909.46838.figure22" httpUri="https://binary.pensoft.net/fig/377383" pageId="0" pageNumber="79">22C, D</figureCitation>
); terga 2-4 with conspicuous microtrichose fasciae (Fig. 
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Merodon medium sp. nov., abdomen, dorsal view. A male B female. Scale bar: 2 mm." figureDoi="10.3897/zookeys.909.46838.figure24" httpUri="https://binary.pensoft.net/fig/377385" pageId="0" pageNumber="79">24A, B</figureCitation>
). Similar to some populations of 
<taxonomicName baseAuthorityName="Wiedemann in Meigen" baseAuthorityYear="1822" class="Insecta" family="Syrphidae" genus="Merodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Merodon serrulatus" order="Diptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="serrulatus">
<emphasis italics="true" pageId="0" pageNumber="79">Merodon serrulatus</emphasis>
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, but clearly differs in shape of abdomen: relation between maximum width of tergum 2 and its medial length is 3.3 in male and 3.5 in female of 
<taxonomicName lsidName="M. medium" pageId="0" pageNumber="79" rank="species" species="medium">
<emphasis italics="true" pageId="0" pageNumber="79">M. medium</emphasis>
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sp. nov. compared with 2.3 in 
<taxonomicName lsidName="M. serrulatus" pageId="0" pageNumber="79" rank="species" species="serrulatus">
<emphasis italics="true" pageId="0" pageNumber="79">M. serrulatus</emphasis>
</taxonomicName>
male and 2.7 in female; male genitalia: anterior surstyle lobe with concave margin in 
<taxonomicName lsidName="M. medium" pageId="0" pageNumber="79" rank="species" species="medium">
<emphasis italics="true" pageId="0" pageNumber="79">M. medium</emphasis>
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sp. nov. (Fig. 
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Male genitalia, surstylus, lateral view. A Merodon serrulatus, Spain B Merodon serrulatus, France C Merodon serrulatus, Greece (Pindos) D Merodon serrulatus, Greece (Olympos) E Merodon serrulatus, Greece (Peloponnese) F Merodon serrulatus, Montenegro G Merodon serrulatus, Russia H Merodon medium sp. nov. G, H margin of anterior surstyle lobe marked with arrow. Abbreviation: bp-basolateral protrusion. Scale bar: 0.2 mm." figureDoi="10.3897/zookeys.909.46838.figure6" httpUri="https://binary.pensoft.net/fig/377367" pageId="0" pageNumber="79">6H</figureCitation>
: marked with arrow), convex in 
<taxonomicName lsidName="M. serrulatus" pageId="0" pageNumber="79" rank="species" species="serrulatus">
<emphasis italics="true" pageId="0" pageNumber="79">M. serrulatus</emphasis>
</taxonomicName>
(Fig. 
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Male genitalia, surstylus, lateral view. A Merodon serrulatus, Spain B Merodon serrulatus, France C Merodon serrulatus, Greece (Pindos) D Merodon serrulatus, Greece (Olympos) E Merodon serrulatus, Greece (Peloponnese) F Merodon serrulatus, Montenegro G Merodon serrulatus, Russia H Merodon medium sp. nov. G, H margin of anterior surstyle lobe marked with arrow. Abbreviation: bp-basolateral protrusion. Scale bar: 0.2 mm." figureDoi="10.3897/zookeys.909.46838.figure6" httpUri="https://binary.pensoft.net/fig/377367" pageId="0" pageNumber="79">6G</figureCitation>
: marked with arrow); apical microtrichose area of anterior surstyle lobe 2.5 times broader than long in 
<taxonomicName lsidName="M. medium" pageId="0" pageNumber="79" rank="species" species="medium">
<emphasis italics="true" pageId="0" pageNumber="79">M. medium</emphasis>
</taxonomicName>
sp. nov. (Fig. 
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Male genitalia, surstylus, lateral view. A Merodon serrulatus, Spain B Merodon serrulatus, France C Merodon serrulatus, Greece (Pindos) D Merodon serrulatus, Greece (Olympos) E Merodon serrulatus, Greece (Peloponnese) F Merodon serrulatus, Montenegro G Merodon serrulatus, Russia H Merodon medium sp. nov. G, H margin of anterior surstyle lobe marked with arrow. Abbreviation: bp-basolateral protrusion. Scale bar: 0.2 mm." figureDoi="10.3897/zookeys.909.46838.figure6" httpUri="https://binary.pensoft.net/fig/377367" pageId="0" pageNumber="79">6G</figureCitation>
: al), less than one time in 
<taxonomicName lsidName="M. serrulatus" pageId="0" pageNumber="79" rank="species" species="serrulatus">
<emphasis italics="true" pageId="0" pageNumber="79">M. serrulatus</emphasis>
</taxonomicName>
(Fig. 
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Male genitalia, surstylus, lateral view. A Merodon serrulatus, Spain B Merodon serrulatus, France C Merodon serrulatus, Greece (Pindos) D Merodon serrulatus, Greece (Olympos) E Merodon serrulatus, Greece (Peloponnese) F Merodon serrulatus, Montenegro G Merodon serrulatus, Russia H Merodon medium sp. nov. G, H margin of anterior surstyle lobe marked with arrow. Abbreviation: bp-basolateral protrusion. Scale bar: 0.2 mm." figureDoi="10.3897/zookeys.909.46838.figure6" httpUri="https://binary.pensoft.net/fig/377367" pageId="0" pageNumber="79">6G</figureCitation>
: al); molecular data and distribution (
<taxonomicName lsidName="M. medium" pageId="0" pageNumber="79" rank="species" species="medium">
<emphasis italics="true" pageId="0" pageNumber="79">M. medium</emphasis>
</taxonomicName>
sp. nov. is an endemic to the island of Crete in Greece).
</paragraph>
<caption doi="10.3897/zookeys.909.46838.figure24" httpUri="https://binary.pensoft.net/fig/377385" pageId="0" pageNumber="79" start="Figure 24" startId="F24">
<paragraph pageId="0" pageNumber="79">
<emphasis bold="true" pageId="0" pageNumber="79">Figure 24.</emphasis>
<taxonomicName authorityName="Vujic, Likov &amp; Radenkovic" authorityYear="2020" class="Insecta" family="Syrphidae" genus="Merodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Merodon medium" order="Diptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="medium">
<emphasis italics="true" pageId="0" pageNumber="79">Merodon medium</emphasis>
</taxonomicName>
sp. nov., abdomen, dorsal view. 
<emphasis bold="true" pageId="0" pageNumber="79">A</emphasis>
male 
<emphasis bold="true" pageId="0" pageNumber="79">B</emphasis>
female. Scale bar: 2 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="description">
<paragraph pageId="0" pageNumber="79">Description.</paragraph>
<paragraph pageId="0" pageNumber="79">
<emphasis bold="true" pageId="0" pageNumber="79">Male.</emphasis>
Head. Antennae black to dark brown; basoflagellomere elongated ca. 2.2 times as long as wide, and ca. 2.5 times as long as pedicel, concave dorsally with acute apex; dorsolateral fossette narrow; arista dark and thickened at basal one third, covered with dense microtrichia, ca. 1.5 times as long as basoflagellomere (Fig. 
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Antenna. A Merodon hirsutus, outer side, lateral view, male B Merodon hirsutus, inner side, lateral view, male C Merodon hirsutus, dorsal view, male D Merodon hirsutus, outer side, lateral view, female E Merodon kawamurae, outer side, lateral view, male F Merodon kawamurae, inner side, lateral view, male G Merodon kawamurae, dorsal view, male H Merodon kawamurae, outer side, lateral view, female I Merodon medium sp. nov., outer side, lateral view, male J Merodon medium sp. nov., outer side, lateral view, female. Scale bar: 1 mm." figureDoi="10.3897/zookeys.909.46838.figure19" httpUri="https://binary.pensoft.net/fig/377380" pageId="0" pageNumber="79">19I</figureCitation>
); face and frons black with gray microtrichia, face covered with dense whitish gray, and frons with yellowish gray pile; oral margin microtrichose with shiny lateral areas; lunule shiny black, bare; vertex shiny black, except microtrichose area in front of anterior ocellus; vertex isosceles, with long, pale whitish yellow pile, mixed with few black pile on the ocellar triangle; ocellar triangle equilateral; eyes covered with dense pile; occiput with gray-yellow pile, ventrally covered with a dense, gray microtrichia; eye contiguity 10-12 facets long; vertical triangle: eye contiguity: frons = 1.2: 1: 2.
</paragraph>
<paragraph pageId="0" pageNumber="79">
Thorax. Scutum and scutellum black with bronze luster, covered with dense, erect, yellow pile, except sides of scutum at wing basis with patch of short black pile and fascia of black pile between wing basis; scutum with two or more microtrichose vittae, anteriorly connected and posteriorly reaching the scutellum; scutum dull; posterodorsal part of anterior anepisternum, posterior anepisternum (except anteroventral angle), anterior anepimeron, dorsomedial anepimeron, and posterodorsal and anteroventral parts of katepisternum with long, pale yellow pile and grayish microtrichia; wings entirely covered with microtrichia; wing veins brown; calypteres and halteres yellowish; legs mostly black, except brown tarsi ventrally in some specimens; pile on legs pale yellow, except few black pile in apical fifth of metafemur in some specimens; metafemur incrassate, ca. three times longer than wide; pile on postero- and anteroventral surface short, except few sparse pile approximately the same length as pile on dorsal surface (Fig. 
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. A, B metaleg, C, D metatrochanter, metafemur and metatibia, lateral view. A Merodon kawamurae, male B Merodon kawamurae, female C Merodon medium sp. nov., male D Merodon medium sp. nov., female. Scale bar: 2 mm." figureDoi="10.3897/zookeys.909.46838.figure22" httpUri="https://binary.pensoft.net/fig/377383" pageId="0" pageNumber="79">22C</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="79">
Abdomen. Broad, tapering, 1.2 times longer than mesonotum; terga dark, except for a pair of yellow-orange, triangular, lateral maculae on tergum 2; terga 2-4 each with a pair of white microtrichose, oblique fasciae (on tergum 2 more triangular); pile on terga all yellow (Fig. 
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Merodon medium sp. nov., abdomen, dorsal view. A male B female. Scale bar: 2 mm." figureDoi="10.3897/zookeys.909.46838.figure24" httpUri="https://binary.pensoft.net/fig/377385" pageId="0" pageNumber="79">24A</figureCitation>
); sterna dark brown, covered with long whitish yellow pile.
</paragraph>
<paragraph pageId="0" pageNumber="79">
Male genitalia. Apical part of anterior surstyle lobe rhomboid shape, 1.5 times longer than wide, covered with dense, short pile (Fig. 
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Male genitalia. A Merodon medium sp. nov., epandrium, lateral view B Merodon medium sp. nov., epandrium, ventral view C Merodon medium sp. nov., hypandrium, lateral view D Merodon nigrocapillatus sp. nov., epandrium, lateral view E Merodon nigrocapillatus sp. nov., epandrium, ventral view F Merodon nigrocapillatus sp. nov., hypandrium, lateral view. Abbreviations: al-anterior surstyle lobe, bp-basolateral protrusion, l-lingula; arrow marks the hook-like extension in A. Scale bar: 0.2 mm." figureDoi="10.3897/zookeys.909.46838.figure25" httpUri="https://binary.pensoft.net/fig/377386" pageId="0" pageNumber="79">25A</figureCitation>
: al) and with concave margin (Fig. 
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Male genitalia, surstylus, lateral view. A Merodon serrulatus, Spain B Merodon serrulatus, France C Merodon serrulatus, Greece (Pindos) D Merodon serrulatus, Greece (Olympos) E Merodon serrulatus, Greece (Peloponnese) F Merodon serrulatus, Montenegro G Merodon serrulatus, Russia H Merodon medium sp. nov. G, H margin of anterior surstyle lobe marked with arrow. Abbreviation: bp-basolateral protrusion. Scale bar: 0.2 mm." figureDoi="10.3897/zookeys.909.46838.figure6" httpUri="https://binary.pensoft.net/fig/377367" pageId="0" pageNumber="79">6H</figureCitation>
: marked with arrow); posterior surstyle lobe oval with basolateral protrusion (lateral hump) (Fig. 
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Male genitalia. A Merodon medium sp. nov., epandrium, lateral view B Merodon medium sp. nov., epandrium, ventral view C Merodon medium sp. nov., hypandrium, lateral view D Merodon nigrocapillatus sp. nov., epandrium, lateral view E Merodon nigrocapillatus sp. nov., epandrium, ventral view F Merodon nigrocapillatus sp. nov., hypandrium, lateral view. Abbreviations: al-anterior surstyle lobe, bp-basolateral protrusion, l-lingula; arrow marks the hook-like extension in A. Scale bar: 0.2 mm." figureDoi="10.3897/zookeys.909.46838.figure25" httpUri="https://binary.pensoft.net/fig/377386" pageId="0" pageNumber="79">25B</figureCitation>
: bp) and basal hook-like extension (Fig. 
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Male genitalia. A Merodon medium sp. nov., epandrium, lateral view B Merodon medium sp. nov., epandrium, ventral view C Merodon medium sp. nov., hypandrium, lateral view D Merodon nigrocapillatus sp. nov., epandrium, lateral view E Merodon nigrocapillatus sp. nov., epandrium, ventral view F Merodon nigrocapillatus sp. nov., hypandrium, lateral view. Abbreviations: al-anterior surstyle lobe, bp-basolateral protrusion, l-lingula; arrow marks the hook-like extension in A. Scale bar: 0.2 mm." figureDoi="10.3897/zookeys.909.46838.figure25" httpUri="https://binary.pensoft.net/fig/377386" pageId="0" pageNumber="79">25A</figureCitation>
: marked with arrow); hypandrium sickle-shaped, without lateral projections; lingula 
<taxonomicName authorityName="Vujic, Likov &amp; Radenkovic" authorityYear="2020" class="Insecta" family="Syrphidae" genus="Merodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="medium" order="Diptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="medium">medium</taxonomicName>
sized (Fig. 
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Male genitalia. A Merodon medium sp. nov., epandrium, lateral view B Merodon medium sp. nov., epandrium, ventral view C Merodon medium sp. nov., hypandrium, lateral view D Merodon nigrocapillatus sp. nov., epandrium, lateral view E Merodon nigrocapillatus sp. nov., epandrium, ventral view F Merodon nigrocapillatus sp. nov., hypandrium, lateral view. Abbreviations: al-anterior surstyle lobe, bp-basolateral protrusion, l-lingula; arrow marks the hook-like extension in A. Scale bar: 0.2 mm." figureDoi="10.3897/zookeys.909.46838.figure25" httpUri="https://binary.pensoft.net/fig/377386" pageId="0" pageNumber="79">25C</figureCitation>
: l).
</paragraph>
<caption doi="10.3897/zookeys.909.46838.figure25" httpUri="https://binary.pensoft.net/fig/377386" pageId="0" pageNumber="79" start="Figure 25" startId="F25">
<paragraph pageId="0" pageNumber="79">
<emphasis bold="true" pageId="0" pageNumber="79">Figure 25.</emphasis>
Male genitalia. 
<emphasis bold="true" pageId="0" pageNumber="79">A</emphasis>
<taxonomicName authorityName="Vujic, Likov &amp; Radenkovic" authorityYear="2020" class="Insecta" family="Syrphidae" genus="Merodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Merodon medium" order="Diptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="medium">
<emphasis italics="true" pageId="0" pageNumber="79">Merodon medium</emphasis>
</taxonomicName>
sp. nov., epandrium, lateral view 
<emphasis bold="true" pageId="0" pageNumber="79">B</emphasis>
<taxonomicName authorityName="Vujic, Likov &amp; Radenkovic" authorityYear="2020" class="Insecta" family="Syrphidae" genus="Merodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Merodon medium" order="Diptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="medium">
<emphasis italics="true" pageId="0" pageNumber="79">Merodon medium</emphasis>
</taxonomicName>
sp. nov., epandrium, ventral view 
<emphasis bold="true" pageId="0" pageNumber="79">C</emphasis>
<taxonomicName authorityName="Vujic, Likov &amp; Radenkovic" authorityYear="2020" class="Insecta" family="Syrphidae" genus="Merodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Merodon medium" order="Diptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="medium">
<emphasis italics="true" pageId="0" pageNumber="79">Merodon medium</emphasis>
</taxonomicName>
sp. nov., hypandrium, lateral view 
<emphasis bold="true" pageId="0" pageNumber="79">D</emphasis>
<taxonomicName authorityName="Vujic, Likov &amp; Radenkovic" authorityYear="2020" class="Insecta" family="Syrphidae" genus="Merodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Merodon nigrocapillatus" order="Diptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="nigrocapillatus">
<emphasis italics="true" pageId="0" pageNumber="79">Merodon nigrocapillatus</emphasis>
</taxonomicName>
sp. nov., epandrium, lateral view 
<emphasis bold="true" pageId="0" pageNumber="79">E</emphasis>
<taxonomicName authorityName="Vujic, Likov &amp; Radenkovic" authorityYear="2020" class="Insecta" family="Syrphidae" genus="Merodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Merodon nigrocapillatus" order="Diptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="nigrocapillatus">
<emphasis italics="true" pageId="0" pageNumber="79">Merodon nigrocapillatus</emphasis>
</taxonomicName>
sp. nov., epandrium, ventral view 
<emphasis bold="true" pageId="0" pageNumber="79">F</emphasis>
<taxonomicName authorityName="Vujic, Likov &amp; Radenkovic" authorityYear="2020" class="Insecta" family="Syrphidae" genus="Merodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Merodon nigrocapillatus" order="Diptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="nigrocapillatus">
<emphasis italics="true" pageId="0" pageNumber="79">Merodon nigrocapillatus</emphasis>
</taxonomicName>
sp. nov., hypandrium, lateral view. Abbreviations: al-anterior surstyle lobe, bp-basolateral protrusion, l-lingula; arrow marks the hook-like extension in 
<emphasis bold="true" pageId="0" pageNumber="79">A</emphasis>
. Scale bar: 0.2 mm.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="79">
<emphasis bold="true" pageId="0" pageNumber="79">Female.</emphasis>
Similar to the male except for normal sexual dimorphism and for the following characteristics: antennae with rounded tip, basoflagellomere ca. two times longer than wide, (Fig. 
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Antenna. A Merodon hirsutus, outer side, lateral view, male B Merodon hirsutus, inner side, lateral view, male C Merodon hirsutus, dorsal view, male D Merodon hirsutus, outer side, lateral view, female E Merodon kawamurae, outer side, lateral view, male F Merodon kawamurae, inner side, lateral view, male G Merodon kawamurae, dorsal view, male H Merodon kawamurae, outer side, lateral view, female I Merodon medium sp. nov., outer side, lateral view, male J Merodon medium sp. nov., outer side, lateral view, female. Scale bar: 1 mm." figureDoi="10.3897/zookeys.909.46838.figure19" httpUri="https://binary.pensoft.net/fig/377380" pageId="0" pageNumber="79">19J</figureCitation>
); frons with broad microtrichose vittae along eye margins; frons covered with pilosity of variable color, from mostly gray-yellow until predominately black pile; ocellar triangle covered with black pile; metafemur incrassate, pile on postero- and anteroventral surface short (Fig. 
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. A, B metaleg, C, D metatrochanter, metafemur and metatibia, lateral view. A Merodon kawamurae, male B Merodon kawamurae, female C Merodon medium sp. nov., male D Merodon medium sp. nov., female. Scale bar: 2 mm." figureDoi="10.3897/zookeys.909.46838.figure22" httpUri="https://binary.pensoft.net/fig/377383" pageId="0" pageNumber="79">22D</figureCitation>
); terga pale yellow pilose at lateral sides, anterior two thirds of tergum 2 and all terga 4 and 5; terga 2 and 3 medially with short adpressed black pile; microtrichose fasciae on terga 3 and 4 broad (Fig. 
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Merodon medium sp. nov., abdomen, dorsal view. A male B female. Scale bar: 2 mm." figureDoi="10.3897/zookeys.909.46838.figure24" httpUri="https://binary.pensoft.net/fig/377385" pageId="0" pageNumber="79">24B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="etymology">
<paragraph pageId="0" pageNumber="79">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="79">
Medium (middle, center) refers to the 
<normalizedToken originalValue="species’">species'</normalizedToken>
distribution, being the only taxon of the group found on Crete, in the middle of Mediterranean Sea.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="distribution">
<paragraph pageId="0" pageNumber="79">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="79">
<taxonomicName authorityName="Vujic, Likov &amp; Radenkovic" authorityYear="2020" class="Insecta" family="Syrphidae" genus="Merodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Merodon medium" order="Diptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="medium">
<emphasis italics="true" pageId="0" pageNumber="79">Merodon medium</emphasis>
</taxonomicName>
sp. nov. is endemic to the Greek island of Crete (Fig. 
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Distribution map of Merodon serrulatus species group." figureDoi="10.3897/zookeys.909.46838.figure7" httpUri="https://binary.pensoft.net/fig/377368" pageId="0" pageNumber="79">7</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="ecology">
<paragraph pageId="0" pageNumber="79">Ecology.</paragraph>
<paragraph pageId="0" pageNumber="79">
Preferred environment: forest/open ground; evergreen oak forest, dry 
<taxonomicName class="Pinopsida" family="Pinaceae" genus="Pinus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pinus" order="Pinales" pageId="0" pageNumber="79" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="79">Pinus</emphasis>
</taxonomicName>
forest; scrub with 
<taxonomicName class="Magnoliopsida" family="Anacardiaceae" genus="Pistacia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pistacia lentiscus" order="Sapindales" pageId="0" pageNumber="79" phylum="Tracheophyta" rank="species" species="lentiscus">
<emphasis italics="true" pageId="0" pageNumber="79">Pistacia lentiscus</emphasis>
</taxonomicName>
L.; well-vegetated, unimproved grassland. Flowers visited: 
<taxonomicName class="Liliopsida" family="Asparagaceae" genus="Ornithogalum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ornithogalum" order="Asparagales" pageId="0" pageNumber="79" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="79">Ornithogalum</emphasis>
</taxonomicName>
spp., 
<taxonomicName class="Magnoliopsida" family="Rosaceae" genus="Potentilla" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Potentilla" order="Rosales" pageId="0" pageNumber="79" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="79">Potentilla</emphasis>
</taxonomicName>
spp. and 
<taxonomicName class="Insecta" family="Eulophidae" genus="Thymus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Thymus" order="Hymenoptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="79">Thymus</emphasis>
</taxonomicName>
spp. Flight period: May.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="type material">
<paragraph pageId="0" pageNumber="79">Type material.</paragraph>
<paragraph pageId="0" pageNumber="79">
<emphasis bold="true" pageId="0" pageNumber="79">Holotype</emphasis>
. Greece • ♂; Crete, Chania, Omalos plain; 
<geoCoordinate degrees="35" direction="north" minutes="19" orientation="latitude" precision="15" seconds="21" value="35.3225">35°19'21&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="23" direction="east" minutes="55" orientation="longitude" precision="15" seconds="50" value="23.930555">23°55'50&quot;E</geoCoordinate>
; 28 May 2014; A. 
<normalizedToken originalValue="Vujić">Vujic</normalizedToken>
leg.; FSUNS 06729. Original label: &quot;HOLOTYPE of 
<emphasis italics="true" pageId="0" pageNumber="79">Merodon</emphasis>
/ 
<emphasis italics="true" pageId="0" pageNumber="79">medium</emphasis>
<normalizedToken originalValue="Vujić">Vujic</normalizedToken>
, Likov et / 
<normalizedToken originalValue="Radenković">Radenkovic</normalizedToken>
sp.n. 2019&quot; [red label], &quot;Greece, Crete, Chania, / Omalos plain / 28.05.2014. 35.322593 / 23.930496 Leg. 
<normalizedToken originalValue="Vujić”">Vujic&quot;</normalizedToken>
, 
<normalizedToken originalValue="“AU298”">&quot;AU298&quot;</normalizedToken>
, 
<normalizedToken originalValue="“06729”">&quot;06729&quot;</normalizedToken>
(See Supplementary file 3: Figure 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Merodon serrulatus antenna. A outer side, lateral view, male (Spain) B inner side, lateral view, male (Spain) C dorsal view, male (Spain) D outer side, lateral view, female (Spain) E inner side, lateral view, female (Spain) F dorsal view, female (Spain) G outer side, lateral view, male (Greece) H inner side, lateral view, male (Greece) I dorsal view, male (Greece) J outer side, lateral view, male (Russia). Abbreviation: f-fossette. Scale bar: 1 mm." figureDoi="10.3897/zookeys.909.46838.figure3" httpUri="https://binary.pensoft.net/fig/377364" pageId="0" pageNumber="79">3</figureCitation>
). 
<emphasis bold="true" pageId="0" pageNumber="79">Paratypes</emphasis>
. Greece • 1 ♀; Crete, Chania, Imbors; 
<geoCoordinate degrees="35" direction="north" minutes="15" orientation="latitude" precision="15" seconds="08" value="35.252224">35°15'08&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="24" direction="east" minutes="10" orientation="longitude" precision="15" seconds="28" value="24.174444">24°10'28&quot;E</geoCoordinate>
; 27 May 2014; A. 
<normalizedToken originalValue="Vujić">Vujic</normalizedToken>
leg.; FSUNS 06706 • 1 ♂; Crete, Chania, Omalos plain; 
<geoCoordinate degrees="35" direction="north" minutes="19" orientation="latitude" precision="15" seconds="06" value="35.318333">35°19'06&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="23" direction="east" minutes="54" orientation="longitude" precision="15" seconds="51" value="23.914165">23°54'51&quot;E</geoCoordinate>
; 28 May 2014; A. 
<normalizedToken originalValue="Vujić">Vujic</normalizedToken>
leg.; FSUNS 06723 • 1 ♂; Crete, Chania, Omalos plain; 
<geoCoordinate degrees="35" direction="north" minutes="19" orientation="latitude" precision="15" seconds="21" value="35.3225">35°19'21&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="23" direction="east" minutes="55" orientation="longitude" precision="15" seconds="50" value="23.930555">23°55'50&quot;E</geoCoordinate>
; 28 May 2014; A. 
<normalizedToken originalValue="Vujić">Vujic</normalizedToken>
leg.; FSUNS 06731 • 1 ♀, Crete, Chania, Mescla, 
<geoCoordinate degrees="35" direction="north" minutes="24" orientation="latitude" precision="15" seconds="05" value="35.40139">35°24'05&quot;N</geoCoordinate>
<geoCoordinate degrees="23" direction="east" minutes="56" orientation="longitude" precision="15" seconds="26" value="23.940556">23°56'26&quot;E</geoCoordinate>
; 28 May 2014; A. 
<normalizedToken originalValue="Vujić">Vujic</normalizedToken>
leg.; FSUNS 06718.
</paragraph>
</subSubSection>
</treatment>
</document>