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123 lines
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<document ID-ISSN="0003-0090" approvalRequired="248" approvalRequired_for_originalDoi="1" approvalRequired_for_taxonomicNames="67" approvalRequired_for_textStreams="120" approvalRequired_for_treatments="60" checkinTime="1630255760165" checkinUser="felipe" docAuthor="Conrad, J. L." docDate="2008" docId="BF23879DD12EFFC4FD5BA8434E75D4CF" docLanguage="en" docName="B310.pdf" docOrigin="Bulletin of the American Museum of Natural History 2008 (310)" docStyle="DocumentStyle:2B34016C1EBFE58B07368F9517618743.1:BulAmeMusNatHis.2000-2010.journal_article.0cover.type1" docStyleId="2B34016C1EBFE58B07368F9517618743" docStyleName="BulAmeMusNatHis.2000-2010.journal_article.0cover.type1" docStyleVersion="1" docTitle="Mosasauridae Gervais 1853" docType="treatment" docVersion="3" lastPageNumber="127" masterDocId="431AFFE5D153FFBAFFA0AE674F56D039" masterDocTitle="Phylogeny And Systematics Of Squamata (Reptilia) Based On Morphology" masterLastPageNumber="182" masterPageNumber="1" pageNumber="126" updateTime="1630877461833" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Phylogeny And Systematics Of Squamata (Reptilia) Based On Morphology</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Conrad, J. L.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Bulletin of the American Museum of Natural History</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2008</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2008-06-03</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>2008</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>310</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>182</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="ISSN">0003-0090</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.5459370" ID-Zenodo-Dep="5459370" LSID="urn:lsid:plazi:treatment:BF23879DD12EFFC4FD5BA8434E75D4CF" httpUri="http://treatment.plazi.org/id/BF23879DD12EFFC4FD5BA8434E75D4CF" lastPageId="126" lastPageNumber="127" pageId="125" pageNumber="126">
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<subSubSection pageId="125" pageNumber="126" type="nomenclature">
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<paragraph blockId="125.[763,1097,1572,1624]" box="[763,1097,1572,1594]" pageId="125" pageNumber="126">
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<heading box="[763,1097,1572,1594]" centered="true" fontSize="9" level="2" pageId="125" pageNumber="126" reason="2">
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<taxonomicName authority="Gervais, 1853" authorityName="Gervais" authorityYear="1853" box="[763,1097,1572,1594]" class="Reptilia" family="Mosasauridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="125" pageNumber="126" phylum="Chordata" rank="family">
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Mosasauridae
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<bibRefCitation author="Gervais, P." box="[936,1097,1572,1594]" pageId="125" pageNumber="126" pagination="470 - 474" refId="ref89805" refString="Gervais, P. 1853. Observations relatives aux reptiles fossiles de France. Comptes Rendus de l'Academie des Sciences de Paris 36: 374 - 377, 470 - 474." type="journal article" year="1853">Gervais, 1853</bibRefCitation>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph blockId="125.[763,1097,1572,1624]" box="[839,1021,1602,1624]" pageId="125" pageNumber="126">
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<heading box="[839,1021,1602,1624]" centered="true" fontSize="10" level="2" pageId="125" pageNumber="126" reason="4">(figs. 54F, 55F)</heading>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="125" pageNumber="126" type="description">
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<paragraph blockId="125.[667,1193,1647,1759]" pageId="125" pageNumber="126">
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DEFINITION:
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<taxonomicName baseAuthorityName="Marsh" baseAuthorityYear="1869" box="[870,1185,1648,1671]" class="Reptilia" family="Mosasauridae" genus="Halisaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="125" pageNumber="126" phylum="Chordata" rank="species" species="platyspondylus">
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<emphasis box="[870,1185,1648,1671]" italics="true" pageId="125" pageNumber="126">Halisaurus platyspondylus</emphasis>
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</taxonomicName>
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,
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<taxonomicName box="[667,897,1677,1700]" class="Actinopterygii" family="Belonidae" genus="Tylosaurus" kingdom="Animalia" order="Beloniformes" pageId="125" pageNumber="126" phylum="Chordata" rank="species" species="proriger">
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<emphasis box="[667,897,1677,1700]" italics="true" pageId="125" pageNumber="126">Tylosaurus proriger</emphasis>
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</taxonomicName>
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,
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<taxonomicName baseAuthorityName="Mulder" baseAuthorityYear="1999" box="[919,1185,1677,1700]" class="Reptilia" family="Mosasauridae" genus="Mosasaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="125" pageNumber="126" phylum="Chordata" rank="species" species="hoffmanni">
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<emphasis box="[919,1185,1677,1700]" italics="true" pageId="125" pageNumber="126">Mosasaurus hoffmanni</emphasis>
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</taxonomicName>
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, and all descendants of their last common ancestor.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="126" pageNumber="127" type="diagnosis">
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<paragraph blockId="126.[128,654,211,1270]" pageId="126" pageNumber="127">DIAGNOSIS: Mosasaurids are diagnosed by two unambiguous synapomorphies: 247(1) absence of a functional sacrum and 275(1) presence of a flattened ‘‘hourglass-shaped’’ humerus.</paragraph>
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</subSubSection>
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<subSubSection pageId="126" pageNumber="127" type="discussion">
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<paragraph blockId="126.[128,654,211,1270]" pageId="126" pageNumber="127">
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COMMENTS: The definition of
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<taxonomicName authorityName="Gervais" authorityYear="1853" class="Reptilia" family="Mosasauridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="126" pageNumber="127" phylum="Chordata" rank="family">Mosasauridae</taxonomicName>
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implemented here follows that of Bell (1997) who divided mosasaurids into
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<taxonomicName authorityName="Marsh" authorityYear="1869" class="Reptilia" family="Mosasauridae" genus="Halisaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="126" pageNumber="127" phylum="Chordata" rank="genus">
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<emphasis italics="true" pageId="126" pageNumber="127">Halisaurus</emphasis>
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</taxonomicName>
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and Natantia. The sister-group relationship of
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<taxonomicName authorityName="Marsh" authorityYear="1869" box="[273,396,479,502]" class="Reptilia" family="Mosasauridae" genus="Halisaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="126" pageNumber="127" phylum="Chordata" rank="genus">
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<emphasis box="[273,396,479,502]" italics="true" pageId="126" pageNumber="127">Halisaurus</emphasis>
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</taxonomicName>
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and all other mosasaurids has been recovered in several analyses (
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<bibRefCitation author="DeBraga, M. & R. L. Carroll" box="[135,461,539,561]" pageId="126" pageNumber="127" pagination="245 - 322" refId="ref87038" refString="DeBraga, M., and R. L. Carroll. 1993. The origin of mosasaurs as a model of macroevolutionary patternsand processes. Evolutionary Biology 27: 245 - 322." type="journal article" year="1993">DeBraga and Carroll, 1993</bibRefCitation>
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;
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<bibRefCitation author="Caldwell, M. W." box="[474,650,539,561]" pageId="126" pageNumber="127" pagination="407 - 436" refId="ref85009" refString="Caldwell, M. W. 1996. Ontogeny and phylogeny of the mesopodial skeleton in mosasauroid reptiles. Zoological Journal of the Linnean Society 116: 407 - 436." type="journal article" year="1996">Caldwell, 1996</bibRefCitation>
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; Bardet and Suberbiola, 2001; Bardet et al., 2003, 2005), but has been questioned recently (
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<bibRefCitation author="Polcyn, M. J. & G. L. Bell, Jr." box="[135,422,627,649]" pageId="126" pageNumber="127" pagination="321 - 333" refId="ref97390" refString="Polcyn, M. J., and G. L. Bell, Jr. 2005. Russellosaurus coheni n. gen., n. sp.: a 92 million-yearold mosasaur from Texas (USA), and the definition of the parafamily Russellosaurina. Netherlands Journal of Geosciences 84: 321 - 333." type="journal article" year="2005">Bell and Polcyn, 2005</bibRefCitation>
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;
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<bibRefCitation author="Polcyn, M. J. & G. L. Bell, Jr." pageId="126" pageNumber="127" pagination="321 - 333" refId="ref97390" refString="Polcyn, M. J., and G. L. Bell, Jr. 2005. Russellosaurus coheni n. gen., n. sp.: a 92 million-yearold mosasaur from Texas (USA), and the definition of the parafamily Russellosaurina. Netherlands Journal of Geosciences 84: 321 - 333." type="journal article" year="2005">Polcyn and Bell, 2005</bibRefCitation>
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). This problem will require further examination given the recent description of new taxa such as
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<taxonomicName authority="(Polcyn and Bell, 2005)" baseAuthorityName="Polcyn and Bell" baseAuthorityYear="2005" class="Reptilia" family="Mosasauridae" genus="Russellosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="126" pageNumber="127" phylum="Chordata" rank="species" species="coheni">
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<emphasis box="[392,653,715,738]" italics="true" pageId="126" pageNumber="127">Russellosaurus coheni</emphasis>
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(
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<bibRefCitation author="Polcyn, M. J. & G. L. Bell, Jr." box="[135,427,745,768]" pageId="126" pageNumber="127" pagination="321 - 333" refId="ref97390" refString="Polcyn, M. J., and G. L. Bell, Jr. 2005. Russellosaurus coheni n. gen., n. sp.: a 92 million-yearold mosasaur from Texas (USA), and the definition of the parafamily Russellosaurina. Netherlands Journal of Geosciences 84: 321 - 333." type="journal article" year="2005">Polcyn and Bell, 2005</bibRefCitation>
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)
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</taxonomicName>
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and
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<taxonomicName authority="(Bell and Polcyn, 2005)" baseAuthorityName="Bell and Polcyn" baseAuthorityYear="2005" class="Reptilia" family="Mosasauridae" genus="Dallasaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="126" pageNumber="127" phylum="Chordata" rank="species" species="turneri">
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<emphasis italics="true" pageId="126" pageNumber="127">Dallasaurus turneri</emphasis>
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(
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<bibRefCitation author="Polcyn, M. J. & G. L. Bell, Jr." box="[223,495,775,797]" pageId="126" pageNumber="127" pagination="321 - 333" refId="ref97390" refString="Polcyn, M. J., and G. L. Bell, Jr. 2005. Russellosaurus coheni n. gen., n. sp.: a 92 million-yearold mosasaur from Texas (USA), and the definition of the parafamily Russellosaurina. Netherlands Journal of Geosciences 84: 321 - 333." type="journal article" year="2005">Bell and Polcyn, 2005</bibRefCitation>
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)
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</taxonomicName>
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. Note, however, that although some existing phylogenetic studies of mosasauroids have sampled densely for species than the current analysis (Bell, 1997;
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<bibRefCitation author="Polcyn, M. J. & G. L. Bell, Jr." box="[279,556,893,915]" pageId="126" pageNumber="127" pagination="321 - 333" refId="ref97390" refString="Polcyn, M. J., and G. L. Bell, Jr. 2005. Russellosaurus coheni n. gen., n. sp.: a 92 million-yearold mosasaur from Texas (USA), and the definition of the parafamily Russellosaurina. Netherlands Journal of Geosciences 84: 321 - 333." type="journal article" year="2005">Bell and Polcyn, 2005</bibRefCitation>
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;
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<bibRefCitation author="Polcyn, M. J. & G. L. Bell, Jr." pageId="126" pageNumber="127" pagination="321 - 333" refId="ref97390" refString="Polcyn, M. J., and G. L. Bell, Jr. 2005. Russellosaurus coheni n. gen., n. sp.: a 92 million-yearold mosasaur from Texas (USA), and the definition of the parafamily Russellosaurina. Netherlands Journal of Geosciences 84: 321 - 333." type="journal article" year="2005">Polcyn and Bell, 2005</bibRefCitation>
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), they have limited their taxonomic sampling to mainly North American and European taxa. The current analysis benefits from more complete taxonomic sampling from all over the world (e.g.,
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<taxonomicName box="[201,509,1070,1093]" class="Reptilia" family="Mosasauridae" genus="Lakumasaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="126" pageNumber="127" phylum="Chordata" rank="species" species="antarcticus">
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<emphasis box="[201,509,1070,1093]" italics="true" pageId="126" pageNumber="127">Lakumasaurus antarcticus</emphasis>
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</taxonomicName>
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from
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<collectingCountry name="Antarctica" pageId="126" pageNumber="127">Antarctica</collectingCountry>
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and
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<taxonomicName box="[279,582,1101,1122]" class="Reptilia" family="Mosasauridae" genus="Goronyosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="126" pageNumber="127" phylum="Chordata" rank="species" species="nigeriensis">
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<emphasis box="[279,582,1101,1122]" italics="true" pageId="126" pageNumber="127">Goronyosaurus nigeriensis</emphasis>
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</taxonomicName>
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from Africa), but does not include many of the American species represented in earlier analyses. A more definitive phylogenetic and taxonomic analysis should include all of these data.
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</paragraph>
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</subSubSection>
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</treatment>
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</document>
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