409 lines
36 KiB
XML
409 lines
36 KiB
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<document ID-DOI="http://dx.doi.org/10.3897/BDJ.4.e7775" ID-PMC="PMC4822071" ID-Pensoft-Pub="1314-2828-4-7775" ID-PubMed="27099555" ModsDocAuthor="" ModsDocDate="2016" ModsDocID="1314-2828-4-e7775" ModsDocOrigin="Biodiversity Data Journal 4" ModsDocTitle="Redescription of the little-known grasshopper Willemsella (Acrididae, Hemiacridinae) from Peninsular Malaysia" checkinTime="1457426927455" checkinUser="pensoft" docAuthor="Tan, Ming Kai & Kamaruddin, Khairul N" docDate="2016" docId="AAD56D1AB5A498996C8F9AA08018CD07" docLanguage="en" docName="BiodivDatJour 4: e7775" docOrigin="Biodiversity Data Journal 4" docSource="http://dx.doi.org/10.3897/BDJ.4.e7775" docTitle="Willemsella bicolor Miller 1934" docType="treatment" docVersion="3" lastPageNumber="7775" masterDocId="FFBAFFD9FF834866DD72FFB5E20AFFB3" masterDocTitle="Redescription of the little-known grasshopper Willemsella (Acrididae, Hemiacridinae) from Peninsular Malaysia" masterLastPageNumber="7775" masterPageNumber="7775" pageNumber="7775" updateTime="1668123551845" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Redescription of the little-known grasshopper Willemsella (Acrididae, Hemiacridinae) from Peninsular Malaysia</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Tan, Ming Kai</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kamaruddin, Khairul N</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Biodiversity Data Journal</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2016</mods:date>
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<mods:detail type="volume">
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<mods:number>4</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>7775</mods:start>
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<mods:end>7775</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/BDJ.4.e7775</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/BDJ.4.e7775</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-2828-4-7775</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:AAD56D1AB5A498996C8F9AA08018CD07" httpUri="http://treatment.plazi.org/id/AAD56D1AB5A498996C8F9AA08018CD07" lastPageNumber="7775" pageId="0" pageNumber="7775">
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<subSubSection pageId="0" pageNumber="7775" type="nomenclature">
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<paragraph pageId="0" pageNumber="7775">
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<taxonomicName authority="Miller, 1934" authorityName="Miller" authorityYear="1934" class="Insecta" family="Acrididae" genus="Willemsella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Willemsella bicolor" order="Orthoptera" pageId="0" pageNumber="7775" phylum="Arthropoda" rank="species" species="bicolor">Willemsella bicolor Miller, 1934</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="7775" type="reference_group">
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<paragraph pageId="0" pageNumber="7775">
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<taxonomicName class="Insecta" family="Acrididae" genus="Willemsella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Willemsella bicolor" order="Orthoptera" pageId="0" pageNumber="7775" phylum="Arthropoda" rank="species" species="bicolor">Willemsella bicolor</taxonomicName>
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The genus should be placed more correctly under
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<taxonomicName lsidName="" pageId="0" pageNumber="7775" rank="subfamily" subfamily="Hemiacridinae">Hemiacridinae</taxonomicName>
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, rather than
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<taxonomicName lsidName="" pageId="0" pageNumber="7775" rank="subfamily" subfamily="Catantopinae">Catantopinae</taxonomicName>
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. In
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<taxonomicName class="Insecta" family="Acrididae" genus="Willemsella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Willemsella" order="Orthoptera" pageId="0" pageNumber="7775" phylum="Arthropoda" rank="genus">Willemsella</taxonomicName>
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, the basal and apical penis valves are separated, diagnostic for species in
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<taxonomicName lsidName="" pageId="0" pageNumber="7775" rank="subfamily" subfamily="Hemiacridinae">Hemiacridinae</taxonomicName>
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. In
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<taxonomicName lsidName="" pageId="0" pageNumber="7775" rank="subfamily" subfamily="Catantopinae">Catantopinae</taxonomicName>
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, the basal and penis valves are connected. The genus also exhibits a series of regular transverse veinlets near the radial area of tegmen, similar to those found in
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<taxonomicName lsidName="" pageId="0" pageNumber="7775" rank="subfamily" subfamily="Hemiacridinae">Hemiacridinae</taxonomicName>
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and usually absent in
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<taxonomicName lsidName="" pageId="0" pageNumber="7775" rank="subfamily" subfamily="Catantopinae">Catantopinae</taxonomicName>
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.
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<taxonomicName class="Insecta" family="Acrididae" genus="Willemsella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Willemsella" order="Orthoptera" pageId="0" pageNumber="7775" phylum="Arthropoda" rank="genus">Willemsella</taxonomicName>
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also shares characters of
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<taxonomicName lsidName="" pageId="0" pageNumber="7775" rank="subfamily" subfamily="Catantopinae">Catantopinae</taxonomicName>
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: presence of prosternal spine, rounded mesosternal lobes and absence of intercalary veins in the medial area of tegmen.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="7775" type="materials_examined">
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<paragraph pageId="0" pageNumber="7775">Materials</paragraph>
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<paragraph pageId="0" pageNumber="7775">
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<materialsCitation collectingDate="06/18/2015" collectingMethod="active collecting" collectionCode="LAR" collectorName="Ming Kai Tan et al." country="Peninsular Malaysia" latitude="4.86184" location="Bukit Larut (Maxwell Hill)" longitude="100.79276" pageId="0" pageNumber="7775" specimenCode="LAR. 15.53" specimenCount="1" specimenCount-male="1" typeStatus="Other material">
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Type status:
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<typeStatus pageId="0" pageNumber="7775">Other material</typeStatus>
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. Occurrence: catalogNumber:
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<specimenCode pageId="0" pageNumber="7775">LAR.15.53</specimenCode>
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; recordedBy:
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<collectorName pageId="0" pageNumber="7775">Ming Kai Tan et al.</collectorName>
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; individualCount:
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<specimenCount pageId="0" pageNumber="7775">1</specimenCount>
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; sex:
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<specimenType pageId="0" pageNumber="7775">male</specimenType>
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; lifeStage:
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<specimenType pageId="0" pageNumber="7775">adult</specimenType>
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; disposition: ZRC; Taxon: scientificName: Willemsellabicolor; nameAccordingTo: Miller; namePublishedIn: 1934; kingdom: Animalia; phylum: Arthropod; class: Insecta; order: Orthoptera; family: Acrididae; genus: Willemsella; specificEpithet: bicolor; Location: country:
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<collectingCountry pageId="0" pageNumber="7775">Peninsular Malaysia</collectingCountry>
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; stateProvince: Perak; locality:
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<location LSID="urn:lsid:plazi:treatment:AAD56D1AB5A498996C8F9AA08018CD07:7095F8799A5C1ACEED2E7C83F8A63598" country="Peninsular Malaysia" latitude="4.86184" longitude="100.79276" name="Bukit Larut (Maxwell Hill)" pageId="0" pageNumber="7775">Bukit Larut (Maxwell Hill)</location>
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; verbatimElevation: 1056 m; decimalLatitude:
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<geoCoordinate orientation="latitude" pageId="0" pageNumber="7775" value="4.86184">4.86184</geoCoordinate>
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; decimalLongitude:
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<geoCoordinate orientation="longitude" pageId="0" pageNumber="7775" value="100.79276">100.79276</geoCoordinate>
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; Identification: identifiedBy: M. K. Tan; dateIdentified: 2015; Event: samplingProtocol:
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<collectingMethod pageId="0" pageNumber="7775">active collecting</collectingMethod>
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; eventDate:
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<collectingDate pageId="0" pageNumber="7775" value="06/18/2015">06/18/2015</collectingDate>
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; eventTime: 1053-1201
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</materialsCitation>
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</paragraph>
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<paragraph pageId="0" pageNumber="7775">
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<materialsCitation collectingDate="09/19/2015" collectingMethod="active collecting" collectionCode="LAR" collectorName="Ming Kai Tan et al." country="Peninsular Malaysia" latitude="4.86626" location="Bukit Larut (Maxwell Hill)" longitude="100.78865" pageId="0" pageNumber="7775" specimenCode="LAR. 15.189" specimenCount="1" specimenCount-male="1" typeStatus="Other material">
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Type status:
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<typeStatus pageId="0" pageNumber="7775">Other material</typeStatus>
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. Occurrence: catalogNumber:
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<specimenCode pageId="0" pageNumber="7775">LAR.15.189</specimenCode>
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; recordedBy:
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<collectorName pageId="0" pageNumber="7775">Ming Kai Tan et al.</collectorName>
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; individualCount:
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<specimenCount pageId="0" pageNumber="7775">1</specimenCount>
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; sex:
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<specimenType pageId="0" pageNumber="7775">male</specimenType>
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; lifeStage:
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<specimenType pageId="0" pageNumber="7775">adult</specimenType>
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; disposition: ZRC; Taxon: scientificName: Willemsellabicolor; nameAccordingTo: Miller; namePublishedIn: 1934; kingdom: Animalia; phylum: Arthropod; class: Insecta; order: Orthoptera; family: Acrididae; genus: Willemsella; specificEpithet: bicolor; Location: country:
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<collectingCountry pageId="0" pageNumber="7775">Peninsular Malaysia</collectingCountry>
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; stateProvince: Perak; locality:
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<location LSID="urn:lsid:plazi:treatment:AAD56D1AB5A498996C8F9AA08018CD07:1D9FAF9B75C576C1A45EAE3624949CB6" country="Peninsular Malaysia" latitude="4.86626" longitude="100.78865" name="Bukit Larut (Maxwell Hill)" pageId="0" pageNumber="7775">Bukit Larut (Maxwell Hill)</location>
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; verbatimElevation: 920.4 m; decimalLatitude:
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<geoCoordinate orientation="latitude" pageId="0" pageNumber="7775" value="4.86626">4.86626</geoCoordinate>
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; decimalLongitude:
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<geoCoordinate orientation="longitude" pageId="0" pageNumber="7775" value="100.78865">100.78865</geoCoordinate>
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; Identification: identifiedBy: M. K. Tan; dateIdentified: 2015; Event: samplingProtocol:
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<collectingMethod pageId="0" pageNumber="7775">active collecting</collectingMethod>
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; eventDate:
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<collectingDate pageId="0" pageNumber="7775" value="09/19/2015">09/19/2015</collectingDate>
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; eventTime: 1925-2049
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</materialsCitation>
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</paragraph>
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<paragraph pageId="0" pageNumber="7775">
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<materialsCitation collectingDate="09/20/2015" collectingMethod="active collecting" collectionCode="LAR" collectorName="Ming Kai Tan et al." country="Peninsular Malaysia" latitude="4.86289" location="Bukit Larut (Maxwell Hill)" longitude="100.79163" pageId="0" pageNumber="7775" specimenCode="LAR. 15.199" specimenCount="1" specimenCount-male="1" typeStatus="Other material">
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Type status:
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<typeStatus pageId="0" pageNumber="7775">Other material</typeStatus>
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. Occurrence: catalogNumber:
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<specimenCode pageId="0" pageNumber="7775">LAR.15.199</specimenCode>
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; recordedBy:
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<collectorName pageId="0" pageNumber="7775">Ming Kai Tan et al.</collectorName>
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; individualCount:
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<specimenCount pageId="0" pageNumber="7775">1</specimenCount>
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; sex:
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<specimenType pageId="0" pageNumber="7775">male</specimenType>
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; lifeStage:
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<specimenType pageId="0" pageNumber="7775">adult</specimenType>
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; disposition: ZRC; Taxon: scientificName: Willemsellabicolor; nameAccordingTo: Miller; namePublishedIn: 1934; kingdom: Animalia; phylum: Arthropod; class: Insecta; order: Orthoptera; family: Acrididae; genus: Willemsella; specificEpithet: bicolor; Location: country:
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<collectingCountry pageId="0" pageNumber="7775">Peninsular Malaysia</collectingCountry>
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; stateProvince: Perak; locality:
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<location LSID="urn:lsid:plazi:treatment:AAD56D1AB5A498996C8F9AA08018CD07:873BCE8CCDD33E9E4472BAF04CB11300" country="Peninsular Malaysia" latitude="4.86289" longitude="100.79163" name="Bukit Larut (Maxwell Hill)" pageId="0" pageNumber="7775">Bukit Larut (Maxwell Hill)</location>
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; verbatimElevation: 1001 m; decimalLatitude:
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<geoCoordinate orientation="latitude" pageId="0" pageNumber="7775" value="4.86289">4.86289</geoCoordinate>
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; decimalLongitude:
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<geoCoordinate orientation="longitude" pageId="0" pageNumber="7775" value="100.79163">100.79163</geoCoordinate>
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; Identification: identifiedBy: M. K. Tan; dateIdentified: 2015; Event: samplingProtocol:
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<collectingMethod pageId="0" pageNumber="7775">active collecting</collectingMethod>
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; eventDate:
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<collectingDate pageId="0" pageNumber="7775" value="09/20/2015">09/20/2015</collectingDate>
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; eventTime: 1043-1236
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</materialsCitation>
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</paragraph>
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<paragraph pageId="0" pageNumber="7775">
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<materialsCitation collectingDate="09/20/2015" collectingMethod="active collecting" collectionCode="LAR" collectorName="Ming Kai Tan et al." country="Peninsular Malaysia" latitude="4.86167" location="Bukit Larut (Maxwell Hill)" longitude="100.79434" pageId="0" pageNumber="7775" specimenCode="LAR. 15.203" specimenCount="1" specimenCount-male="1" typeStatus="Other material">
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Type status:
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<typeStatus pageId="0" pageNumber="7775">Other material</typeStatus>
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. Occurrence: catalogNumber:
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<specimenCode pageId="0" pageNumber="7775">LAR.15.203</specimenCode>
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; recordedBy:
|
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<collectorName pageId="0" pageNumber="7775">Ming Kai Tan et al.</collectorName>
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; individualCount:
|
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<specimenCount pageId="0" pageNumber="7775">1</specimenCount>
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; sex:
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<specimenType pageId="0" pageNumber="7775">male</specimenType>
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; lifeStage:
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<specimenType pageId="0" pageNumber="7775">adult</specimenType>
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; disposition: ZRC; Taxon: scientificName: Willemsellabicolor; nameAccordingTo: Miller; namePublishedIn: 1934; kingdom: Animalia; phylum: Arthropod; class: Insecta; order: Orthoptera; family: Acrididae; genus: Willemsella; specificEpithet: bicolor; Location: country:
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<collectingCountry pageId="0" pageNumber="7775">Peninsular Malaysia</collectingCountry>
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; stateProvince: Perak; locality:
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<location LSID="urn:lsid:plazi:treatment:AAD56D1AB5A498996C8F9AA08018CD07:2A99BF1CD3B35F1B7A4FEAFBECBD4CCE" country="Peninsular Malaysia" latitude="4.86167" longitude="100.79434" name="Bukit Larut (Maxwell Hill)" pageId="0" pageNumber="7775">Bukit Larut (Maxwell Hill)</location>
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|||
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; verbatimElevation: 1097 m; decimalLatitude:
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|||
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<geoCoordinate orientation="latitude" pageId="0" pageNumber="7775" value="4.86167">4.86167</geoCoordinate>
|
|||
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; decimalLongitude:
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<geoCoordinate orientation="longitude" pageId="0" pageNumber="7775" value="100.79434">100.79434</geoCoordinate>
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; Identification: identifiedBy: M. K. Tan; dateIdentified: 2015; Event: samplingProtocol:
|
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<collectingMethod pageId="0" pageNumber="7775">active collecting</collectingMethod>
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; eventDate:
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<collectingDate pageId="0" pageNumber="7775" value="09/20/2015">09/20/2015</collectingDate>
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; eventTime: 1928-2055
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</materialsCitation>
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</paragraph>
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<paragraph pageId="0" pageNumber="7775">
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<materialsCitation collectingDate="09/20/2015" collectingMethod="active collecting" collectionCode="LAR" collectorName="Ming Kai Tan et al." country="Peninsular Malaysia" latitude="4.86167" location="Bukit Larut (Maxwell Hill)" longitude="100.79434" pageId="0" pageNumber="7775" specimenCode="LAR. 15.210" specimenCount="1" specimenCount-female="1" typeStatus="Other material">
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Type status:
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<typeStatus pageId="0" pageNumber="7775">Other material</typeStatus>
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. Occurrence: catalogNumber:
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<specimenCode pageId="0" pageNumber="7775">LAR.15.210</specimenCode>
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; recordedBy:
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<collectorName pageId="0" pageNumber="7775">Ming Kai Tan et al.</collectorName>
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; individualCount:
|
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<specimenCount pageId="0" pageNumber="7775">1</specimenCount>
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; sex:
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<specimenType pageId="0" pageNumber="7775">female</specimenType>
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; lifeStage:
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<specimenType pageId="0" pageNumber="7775">adult</specimenType>
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; disposition: ZRC; Taxon: scientificName: Willemsellabicolor; nameAccordingTo: Miller; namePublishedIn: 1934; kingdom: Animalia; phylum: Arthropod; class: Insecta; order: Orthoptera; family: Acrididae; genus: Willemsella; specificEpithet: bicolor; Location: country:
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<collectingCountry pageId="0" pageNumber="7775">Peninsular Malaysia</collectingCountry>
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; stateProvince: Perak; locality:
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|
|
<location LSID="urn:lsid:plazi:treatment:AAD56D1AB5A498996C8F9AA08018CD07:1581CDA9852AB408DE537C654FBDD0A9" country="Peninsular Malaysia" latitude="4.86167" longitude="100.79434" name="Bukit Larut (Maxwell Hill)" pageId="0" pageNumber="7775">Bukit Larut (Maxwell Hill)</location>
|
|||
|
|
; verbatimElevation: 1097 m; decimalLatitude:
|
|||
|
|
<geoCoordinate orientation="latitude" pageId="0" pageNumber="7775" value="4.86167">4.86167</geoCoordinate>
|
|||
|
|
; decimalLongitude:
|
|||
|
|
<geoCoordinate orientation="longitude" pageId="0" pageNumber="7775" value="100.79434">100.79434</geoCoordinate>
|
|||
|
|
; Identification: identifiedBy: M. K. Tan; dateIdentified: 2015; Event: samplingProtocol:
|
|||
|
|
<collectingMethod pageId="0" pageNumber="7775">active collecting</collectingMethod>
|
|||
|
|
; eventDate:
|
|||
|
|
<collectingDate pageId="0" pageNumber="7775" value="09/20/2015">09/20/2015</collectingDate>
|
|||
|
|
; eventTime: 1928-2055
|
|||
|
|
</materialsCitation>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph pageId="0" pageNumber="7775">
|
|||
|
|
<materialsCitation collectingDate="09/20/2015" collectingMethod="active collecting" collectionCode="LAR" collectorName="Ming Kai Tan et al." country="Peninsular Malaysia" latitude="4.86191" location="Bukit Larut (Maxwell Hill)" longitude="100.79305" pageId="0" pageNumber="7775" specimenCode="LAR. 15.211" specimenCount="1" specimenCount-female="1" typeStatus="Other material">
|
|||
|
|
Type status:
|
|||
|
|
<typeStatus pageId="0" pageNumber="7775">Other material</typeStatus>
|
|||
|
|
. Occurrence: catalogNumber:
|
|||
|
|
<specimenCode pageId="0" pageNumber="7775">LAR.15.211</specimenCode>
|
|||
|
|
; recordedBy:
|
|||
|
|
<collectorName pageId="0" pageNumber="7775">Ming Kai Tan et al.</collectorName>
|
|||
|
|
; individualCount:
|
|||
|
|
<specimenCount pageId="0" pageNumber="7775">1</specimenCount>
|
|||
|
|
; sex:
|
|||
|
|
<specimenType pageId="0" pageNumber="7775">female</specimenType>
|
|||
|
|
; lifeStage:
|
|||
|
|
<specimenType pageId="0" pageNumber="7775">adult</specimenType>
|
|||
|
|
; disposition: ZRC; Taxon: scientificName: Willemsellabicolor; nameAccordingTo: Miller; namePublishedIn: 1934; kingdom: Animalia; phylum: Arthropod; class: Insecta; order: Orthoptera; family: Acrididae; genus: Willemsella; specificEpithet: bicolor; Location: country:
|
|||
|
|
<collectingCountry pageId="0" pageNumber="7775">Peninsular Malaysia</collectingCountry>
|
|||
|
|
; stateProvince: Perak; locality:
|
|||
|
|
<location LSID="urn:lsid:plazi:treatment:AAD56D1AB5A498996C8F9AA08018CD07:8B8E2C0D35F5253A1D511ADA5AFA2A08" country="Peninsular Malaysia" latitude="4.86191" longitude="100.79305" name="Bukit Larut (Maxwell Hill)" pageId="0" pageNumber="7775">Bukit Larut (Maxwell Hill)</location>
|
|||
|
|
; verbatimElevation: 1052 m; decimalLatitude:
|
|||
|
|
<geoCoordinate orientation="latitude" pageId="0" pageNumber="7775" value="4.86191">4.86191</geoCoordinate>
|
|||
|
|
; decimalLongitude:
|
|||
|
|
<geoCoordinate orientation="longitude" pageId="0" pageNumber="7775" value="100.79305">100.79305</geoCoordinate>
|
|||
|
|
; Identification: identifiedBy: M. K. Tan; dateIdentified: 2015; Event: samplingProtocol:
|
|||
|
|
<collectingMethod pageId="0" pageNumber="7775">active collecting</collectingMethod>
|
|||
|
|
; eventDate:
|
|||
|
|
<collectingDate pageId="0" pageNumber="7775" value="09/20/2015">09/20/2015</collectingDate>
|
|||
|
|
; eventTime: 2055-2136
|
|||
|
|
</materialsCitation>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph pageId="0" pageNumber="7775">
|
|||
|
|
<materialsCitation collectingDate="12/27/2012" collectingMethod="active collecting" collectionCode="FRA" collectorName="Ming Kai Tan et al." country="Peninsular Malaysia" latitude="3.71944" location="Bukit Fraser" longitude="101.72936" pageId="0" pageNumber="7775" specimenCode="FRA. 12.88" specimenCount="1" specimenCount-male="1" typeStatus="Other material">
|
|||
|
|
Type status:
|
|||
|
|
<typeStatus pageId="0" pageNumber="7775">Other material</typeStatus>
|
|||
|
|
. Occurrence: catalogNumber:
|
|||
|
|
<specimenCode pageId="0" pageNumber="7775">FRA.12.88</specimenCode>
|
|||
|
|
; recordedBy:
|
|||
|
|
<collectorName pageId="0" pageNumber="7775">Ming Kai Tan et al.</collectorName>
|
|||
|
|
; individualCount:
|
|||
|
|
<specimenCount pageId="0" pageNumber="7775">1</specimenCount>
|
|||
|
|
; sex:
|
|||
|
|
<specimenType pageId="0" pageNumber="7775">male</specimenType>
|
|||
|
|
; lifeStage:
|
|||
|
|
<specimenType pageId="0" pageNumber="7775">adult</specimenType>
|
|||
|
|
; disposition: ZRC; Taxon: scientificName: Willemsellabicolor; nameAccordingTo: Miller; namePublishedIn: 1934; kingdom: Animalia; phylum: Arthropod; class: Insecta; order: Orthoptera; family: Acrididae; genus: Willemsella; specificEpithet: bicolor; Location: country:
|
|||
|
|
<collectingCountry pageId="0" pageNumber="7775">Peninsular Malaysia</collectingCountry>
|
|||
|
|
; stateProvince: Pahang; locality:
|
|||
|
|
<location LSID="urn:lsid:plazi:treatment:AAD56D1AB5A498996C8F9AA08018CD07:1CECABB29EEC48E3BF884F2C534469ED" country="Peninsular Malaysia" latitude="3.71944" longitude="101.72936" name="Bukit Fraser" pageId="0" pageNumber="7775">Bukit Fraser</location>
|
|||
|
|
; verbatimElevation: 1214 m; decimalLatitude:
|
|||
|
|
<geoCoordinate orientation="latitude" pageId="0" pageNumber="7775" value="3.71944">3.71944</geoCoordinate>
|
|||
|
|
; decimalLongitude:
|
|||
|
|
<geoCoordinate orientation="longitude" pageId="0" pageNumber="7775" value="101.72936">101.72936</geoCoordinate>
|
|||
|
|
; Identification: identifiedBy: M. K. Tan; dateIdentified: 2015; Event: samplingProtocol:
|
|||
|
|
<collectingMethod pageId="0" pageNumber="7775">active collecting</collectingMethod>
|
|||
|
|
; eventDate:
|
|||
|
|
<collectingDate pageId="0" pageNumber="7775" value="12/27/2012">12/27/2012</collectingDate>
|
|||
|
|
; eventTime: 1338
|
|||
|
|
</materialsCitation>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph pageId="0" pageNumber="7775">
|
|||
|
|
<materialsCitation collectingDate="12/27/2012" collectingMethod="active collecting" collectionCode="FRA" collectorName="Ming Kai Tan et al." country="Peninsular Malaysia" latitude="3.71952" location="Bukit Fraser" longitude="101.72937" pageId="0" pageNumber="7775" specimenCode="FRA. 12.90" specimenCount="1" specimenCount-male="1" typeStatus="Other material">
|
|||
|
|
Type status:
|
|||
|
|
<typeStatus pageId="0" pageNumber="7775">Other material</typeStatus>
|
|||
|
|
. Occurrence: catalogNumber:
|
|||
|
|
<specimenCode pageId="0" pageNumber="7775">FRA.12.90</specimenCode>
|
|||
|
|
; recordedBy:
|
|||
|
|
<collectorName pageId="0" pageNumber="7775">Ming Kai Tan et al.</collectorName>
|
|||
|
|
; individualCount:
|
|||
|
|
<specimenCount pageId="0" pageNumber="7775">1</specimenCount>
|
|||
|
|
; sex:
|
|||
|
|
<specimenType pageId="0" pageNumber="7775">male</specimenType>
|
|||
|
|
; lifeStage:
|
|||
|
|
<specimenType pageId="0" pageNumber="7775">adult</specimenType>
|
|||
|
|
; disposition: ZRC; Taxon: scientificName: Willemsellabicolor; nameAccordingTo: Miller; namePublishedIn: 1934; kingdom: Animalia; phylum: Arthropod; class: Insecta; order: Orthoptera; family: Acrididae; genus: Willemsella; specificEpithet: bicolor; Location: country:
|
|||
|
|
<collectingCountry pageId="0" pageNumber="7775">Peninsular Malaysia</collectingCountry>
|
|||
|
|
; stateProvince: Pahang; locality:
|
|||
|
|
<location LSID="urn:lsid:plazi:treatment:AAD56D1AB5A498996C8F9AA08018CD07:95976DC949EEBA43D4A305735377046B" country="Peninsular Malaysia" latitude="3.71952" longitude="101.72937" name="Bukit Fraser" pageId="0" pageNumber="7775">Bukit Fraser</location>
|
|||
|
|
; verbatimElevation: 1203 m; decimalLatitude:
|
|||
|
|
<geoCoordinate orientation="latitude" pageId="0" pageNumber="7775" value="3.71952">3.71952</geoCoordinate>
|
|||
|
|
; decimalLongitude:
|
|||
|
|
<geoCoordinate orientation="longitude" pageId="0" pageNumber="7775" value="101.72937">101.72937</geoCoordinate>
|
|||
|
|
; Identification: identifiedBy: M. K. Tan; dateIdentified: 2015; Event: samplingProtocol:
|
|||
|
|
<collectingMethod pageId="0" pageNumber="7775">active collecting</collectingMethod>
|
|||
|
|
; eventDate:
|
|||
|
|
<collectingDate pageId="0" pageNumber="7775" value="12/27/2012">12/27/2012</collectingDate>
|
|||
|
|
; eventTime: 1346
|
|||
|
|
</materialsCitation>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph pageId="0" pageNumber="7775">
|
|||
|
|
<materialsCitation collectingDate="12/29/2012" collectingMethod="active collecting" collectionCode="FRA" collectorName="Ming Kai Tan et al." country="Peninsular Malaysia" latitude="3.71943" location="Bukit Fraser" longitude="101.72935" pageId="0" pageNumber="7775" specimenCode="FRA. 12.112" specimenCount="1" specimenCount-male="1" typeStatus="Other material">
|
|||
|
|
Type status:
|
|||
|
|
<typeStatus pageId="0" pageNumber="7775">Other material</typeStatus>
|
|||
|
|
. Occurrence: catalogNumber:
|
|||
|
|
<specimenCode pageId="0" pageNumber="7775">FRA.12.112</specimenCode>
|
|||
|
|
; recordedBy:
|
|||
|
|
<collectorName pageId="0" pageNumber="7775">Ming Kai Tan et al.</collectorName>
|
|||
|
|
; individualCount:
|
|||
|
|
<specimenCount pageId="0" pageNumber="7775">1</specimenCount>
|
|||
|
|
; sex:
|
|||
|
|
<specimenType pageId="0" pageNumber="7775">male</specimenType>
|
|||
|
|
; lifeStage:
|
|||
|
|
<specimenType pageId="0" pageNumber="7775">adult</specimenType>
|
|||
|
|
; disposition: ZRC; Taxon: scientificName: Willemsellabicolor; nameAccordingTo: Miller; namePublishedIn: 1934; kingdom: Animalia; phylum: Arthropod; class: Insecta; order: Orthoptera; family: Acrididae; genus: Willemsella; specificEpithet: bicolor; Location: country:
|
|||
|
|
<collectingCountry pageId="0" pageNumber="7775">Peninsular Malaysia</collectingCountry>
|
|||
|
|
; stateProvince: Pahang; locality:
|
|||
|
|
<location LSID="urn:lsid:plazi:treatment:AAD56D1AB5A498996C8F9AA08018CD07:6C2308A4718A93C30BBD88D46B465A0F" country="Peninsular Malaysia" latitude="3.71943" longitude="101.72935" name="Bukit Fraser" pageId="0" pageNumber="7775">Bukit Fraser</location>
|
|||
|
|
; verbatimElevation: 1209 m; decimalLatitude:
|
|||
|
|
<geoCoordinate orientation="latitude" pageId="0" pageNumber="7775" value="3.71943">3.71943</geoCoordinate>
|
|||
|
|
; decimalLongitude:
|
|||
|
|
<geoCoordinate orientation="longitude" pageId="0" pageNumber="7775" value="101.72935">101.72935</geoCoordinate>
|
|||
|
|
; Identification: identifiedBy: M. K. Tan; dateIdentified: 2015; Event: samplingProtocol:
|
|||
|
|
<collectingMethod pageId="0" pageNumber="7775">active collecting</collectingMethod>
|
|||
|
|
; eventDate:
|
|||
|
|
<collectingDate pageId="0" pageNumber="7775" value="12/29/2012">12/29/2012</collectingDate>
|
|||
|
|
; eventTime: 2005
|
|||
|
|
</materialsCitation>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph pageId="0" pageNumber="7775">
|
|||
|
|
<materialsCitation collectingDate="05/19/2013" collectingMethod="active collecting" collectionCode="FRA" collectorName="Ming Kai Tan et al." country="Peninsular Malaysia" latitude="3.71836" location="Bukit Fraser" longitude="101.72987" pageId="0" pageNumber="7775" specimenCode="FRA. 13.35" specimenCount="1" specimenCount-female="1" typeStatus="Other material">
|
|||
|
|
Type status:
|
|||
|
|
<typeStatus pageId="0" pageNumber="7775">Other material</typeStatus>
|
|||
|
|
. Occurrence: catalogNumber:
|
|||
|
|
<specimenCode pageId="0" pageNumber="7775">FRA.13.35</specimenCode>
|
|||
|
|
; recordedBy:
|
|||
|
|
<collectorName pageId="0" pageNumber="7775">Ming Kai Tan et al.</collectorName>
|
|||
|
|
; individualCount:
|
|||
|
|
<specimenCount pageId="0" pageNumber="7775">1</specimenCount>
|
|||
|
|
; sex:
|
|||
|
|
<specimenType pageId="0" pageNumber="7775">female</specimenType>
|
|||
|
|
; lifeStage:
|
|||
|
|
<specimenType pageId="0" pageNumber="7775">adult</specimenType>
|
|||
|
|
; disposition: ZRC; Taxon: scientificName: Willemsellabicolor; nameAccordingTo: Miller; namePublishedIn: 1934; kingdom: Animalia; phylum: Arthropod; class: Insecta; order: Orthoptera; family: Acrididae; genus: Willemsella; specificEpithet: bicolor; Location: country:
|
|||
|
|
<collectingCountry pageId="0" pageNumber="7775">Peninsular Malaysia</collectingCountry>
|
|||
|
|
; stateProvince: Pahang; locality:
|
|||
|
|
<location LSID="urn:lsid:plazi:treatment:AAD56D1AB5A498996C8F9AA08018CD07:9705E135AB73BB74FE6B93F59D4B7C29" country="Peninsular Malaysia" latitude="3.71836" longitude="101.72987" name="Bukit Fraser" pageId="0" pageNumber="7775">Bukit Fraser</location>
|
|||
|
|
; verbatimElevation: 1227 m; decimalLatitude:
|
|||
|
|
<geoCoordinate orientation="latitude" pageId="0" pageNumber="7775" value="3.71836">3.71836</geoCoordinate>
|
|||
|
|
; decimalLongitude:
|
|||
|
|
<geoCoordinate orientation="longitude" pageId="0" pageNumber="7775" value="101.72987">101.72987</geoCoordinate>
|
|||
|
|
; Identification: identifiedBy: M. K. Tan; dateIdentified: 2015; Event: samplingProtocol:
|
|||
|
|
<collectingMethod pageId="0" pageNumber="7775">active collecting</collectingMethod>
|
|||
|
|
; eventDate:
|
|||
|
|
<collectingDate pageId="0" pageNumber="7775" value="05/19/2013">05/19/2013</collectingDate>
|
|||
|
|
; eventTime: 1409
|
|||
|
|
</materialsCitation>
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection pageId="0" pageNumber="7775" type="description">
|
|||
|
|
<paragraph pageId="0" pageNumber="7775">Description</paragraph>
|
|||
|
|
<paragraph pageId="0" pageNumber="7775">Adult habitus as shown in Fig. 1. Head about as long as pronotum (Fig. 2a, b, d, e). Frontal ridge narrow, parallel-sided dorsal and ventral of median ocellus; ridge more deep but narrower in males than females (Fig. 2c, f). Fastigium of vertex broadly rounded to subtruncated (Fig. 2b, e). Vertex between eyes average about 4.3 times in males (n=3, min=3.5, max=5.0) and 2.0 times (n=2, min=2.0, max=2.0) in females as broad as frontal ridge between antennae. Eyes large. Antennae about 20-segmented, flattened; reaching and surpassing posterior margin of pronotum in males, and reaching middle but not surpassing posterior margin of pronotum in females. Pronotum long, crossed by three transverse furrows (Fig. 2b, e); prozona 1.7 times in male (n=3, min=1.5, max=2.1) and 2.1 times in female (n=2, min=1.9, max=2.4) as long as metazona; anterior margin of prozona truncated to feebly convex; posterior margin of metazona broadly rounded; lateral carinae absent; median carina indistinct, more distinct along metazona. Prosternal spine distinct, flattened and triangular basally, narrowing into a long and subacute apex. Mesosternal lophi with internal margin broadly rounded; mesosternal interspace slightly less broad than lobe. Metasternal lophi with internal margin broadly rounded, separated feebly by metasternal interspace. Tegmen truncated with rounded apex, overlapping each other in rest position, elongated and reaching base of epiproct in males and base of 8th tergite in females. Radial area of tegmen with a series of regular transverse veinlets (Fig. 2a, d). Hind wings concealed under tegmen. Hind femora 4.7 times (n=5, min=4.6, max=4.8) as long as maximal width of these femora; dorsal carina smooth; knee with dorsal external lobe subtruncated to angular. Hind femora 1.1 times (n=5, min=1.1, max=1.1) longer than hind tibiae. Hind tibiae setose, with 8 outer dorsal subapical spines and 8 inner dorsal subapical spines; with 2 outer and 2 inner apical dorsal spines. Hind tarsi about a third of hind tibiae; third segment (without claws) longer than the two basal segments together; arolium large, almost reaching apex of claws.</paragraph>
|
|||
|
|
<paragraph pageId="0" pageNumber="7775">
|
|||
|
|
Male. Epiproct broadly tongue-shaped with an apical lobe; median sulcus shallow, broad and feeble, only in the basal half bordered by broad parallel carinae; apical lobe triangular with obtuse apex (Fig. 3a). Cercus in dorsal view compressed and gently curved externally; apex obtusely rounded. Subgenital plate conical, with obtuse apex (Fig. 3a). Male phallic complex as in Fig. 3b, c. Epiphallus bridge (B), very broad and stout, slightly curved along apical margin. Anchora (A) articulated, extending in less than 90
|
|||
|
|
<normalizedToken originalValue="º-angle">°-angle</normalizedToken>
|
|||
|
|
to epiphallus bridge, apex obtuse from dorsal view; membrane between lophi sparsely granulated. Anterior projection (An) of epiphallus swollen and somewhat sparsely granular, forming at less than 90
|
|||
|
|
<normalizedToken originalValue="º-angle">°-angle</normalizedToken>
|
|||
|
|
to the anchora. Lophi (L) stout and transverse, forming highly sclerotized lateral angles; internal angle bulbous and granular, external angle slightly flattened and obtuse, less sclerotized and granular. Ectophallic membrane thickened to form a dorsal and paired ventral ectophallic sclerites; dorsal ectophallic sclerite (Dsv) more membraneous, bridge-shaped with paired cranial appendages covering the cingulum dorsally and laterally; ventral appendages ventral ectophallic sclerites (Esv) rounded, sclerotized and granular. Apodemes (Apd) robust, weakly sclerotized, extending to basal end. Arch of cingulum (Ac) sclerotized, more so internally; finely granulated externally. Basal and apical penis valves separated (Fig. 3c). Basal valves of penis (Bp) broadening basally, more sclerotized internally; apical valves (Ap) truncated and sclerotized, slightly surpassing arch of cingulum.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph pageId="0" pageNumber="7775">
|
|||
|
|
Female. Tenth abdominal tergite with posterior margin rounded-triangularly excised, without lateral lobe (Fig. 3d).
|
|||
|
|
<normalizedToken originalValue="Supra–anal">Supra-anal</normalizedToken>
|
|||
|
|
plate elongated, separated into two parts by a middle transverse carina, apical half broadly tongue-shaped with obtuse apex (Fig. 3d). Transverse carina of supra-anal plate continuing laterally. Cercus triangular with pointed apex. Subgenital plate rectangular slightly longer than broad; posterior margin with large median angular projection and without lateral excisions (Fig. 3f). Ovipositor with long setae, short, hook-like; dorsal valves smooth with apex gently curved dorsad; ventral valves with apex bent ventrad (Fig. 3e).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph pageId="0" pageNumber="7775">Colouration (Fig. 1). Male generally brightly green, female brown to olive brown. Head in male mostly green with some patches of yellow beneath eyes at gena and frons; in females, brown dorsally and anteriorly with some dark olive green patch, dark olive green laterally. Scapus and pedicel black, sometimes with white ring apically. Antennal segments bright red. Disc of pronotum green in male and brown in female; lateral lobes with dorsal third dark green in male or dark olive brown in female, basally with a bright yellow spot extending from the anterior to the posterior margins of lateral lobe. Tegmen green in male and brown in female. Fore and mid femora and tibiae green, with knees black, tarsi dark in male; brown in female. Hind femur generally green in male and brown in female; dorso-basal area yellow, ventro-internal bright red; hind knee black. Hind tibia dark blue. Abdominal tergite green in male or brown in female with dark and pale yellow spots. Thoracic and abdominal sternites bright red.</paragraph>
|
|||
|
|
<paragraph pageId="0" pageNumber="7775">Measurement of alcohol-preserved dry-pinned specimens, in mm (mean in bracket). Male (n=3) BL: 18.4-19.9 (19.1), VW: 0.5-0.7 (0.6), FRW: 0.1-0.2 (0.1), PL: 3.4-4.2 (3.8), PZL: 2.3-2.6 (2.4), MZL: 1.1-1.6 (1.4), PW: 3.0-3.4 (3.2), TL: 7.9-8.7 (8.4), HFL: 12.7-13.6 (13.1), HFW: 2.7-2.9 (2.8), HTL: 11.5-12.5 (12.0); female (n=2) BL: 21.9-22.4 (22.2), VW: 1.0, FRW: 0.5, PL: 5.2-5.5 (5.4), PZL: 3.4-3.9 (3.7), MZL: 1.6-1.8 (1.7), PW: 4.1-4.6 (4.4), TL: 9.9-10.2 (10.1), HFL: 15.4-15.9 (15.7), HFW: 3.2-3.4 (3.3), HTL: 14.0-14.2 (14.1).</paragraph>
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</subSubSection>
|
|||
|
|
<subSubSection pageId="0" pageNumber="7775" type="diagnosis">
|
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<paragraph pageId="0" pageNumber="7775">Diagnosis</paragraph>
|
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|
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<paragraph pageId="0" pageNumber="7775">
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Prosternal spine distinct, flattened and triangular basally, narrowing into a long and subacute apex. Radial area of tegmen with a series of regular transverse veinlets. Anchora (A) articulated, extending in less than 90
|
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|
|
<normalizedToken originalValue="º-angle">°-angle</normalizedToken>
|
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|
|
to epiphallus bridge, apex obtuse from dorsal view; membrane between lophi sparsely granulated. Anterior projection (An) of epiphallus swollen and somewhat sparsely granular, forming at less than 90
|
|||
|
|
<normalizedToken originalValue="º-angle">°-angle</normalizedToken>
|
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|
|
to the anchora. Basal and apical penis valves separated. Antennal segments bright red. Lateral lobes with a basal bright yellow spot extending from the anterior to the posterior margins of lateral lobe.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection pageId="0" pageNumber="7775" type="distribution">
|
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|
|
<paragraph pageId="0" pageNumber="7775">Distribution</paragraph>
|
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|
|
<paragraph pageId="0" pageNumber="7775">
|
|||
|
|
Malay Peninsula: Bukit Larut, Perak; Bukit Fraser, Pahang; Dusan Tua, Selangor (
|
|||
|
|
<bibRefCitation author="Miller, N. C. E." journalOrPublisher="Journal of the Federation of Malay State Museum" pageId="0" pageNumber="7775" pagination=": 526" title="Notes on Malayan Acrididae and description of some new genera and species" volume="17" year="1934">Miller 1934</bibRefCitation>
|
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|
).
|
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|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection pageId="0" pageNumber="7775" type="ecology">
|
|||
|
|
<paragraph pageId="0" pageNumber="7775">Ecology</paragraph>
|
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|
|
<paragraph pageId="0" pageNumber="7775">Little is known about the ecology of this grasshopper species. they are fairly common among tall grasses in Bukit Fraser and Bukit Larut, suggesting that they may prefer high elevations.</paragraph>
|
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|
|
</subSubSection>
|
|||
|
|
<subSubSection pageId="0" pageNumber="7775" type="conservation">
|
|||
|
|
<paragraph pageId="0" pageNumber="7775">Conservation</paragraph>
|
|||
|
|
<paragraph pageId="0" pageNumber="7775">So far endemic to Peninsular Malaysia.</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|