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<document id="9BAD05EE7EA0F15B348666F770872E4B" ID-CLB-Dataset="80413" ID-DOI="10.11646/zootaxa.5155.4.3" ID-GBIF-Dataset="37214782-4329-4fd3-9e82-3ffcfbadd12c" ID-ISSN="1175-5326" ID-Zenodo-Dep="6691435" ID-ZooBank="DA3FFBB6-8CB7-48AE-9F37-1E62CDB91211" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1655955024723" checkinUser="plazi" docAuthor="Gnezdilov, Vladimir M." docDate="2022" docId="99442B4A6003CE78359A891FFAB2D3EE" docLanguage="en" docName="zootaxa.5155.4.3.pdf" docOrigin="Zootaxa 5155 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lethierium Dlabola 1980" docType="treatment" docVersion="7" lastPageNumber="518" masterDocId="657D53326002CE79350D8B3DFF91D43C" masterDocTitle="Revision of the genus Lethierium Dlabola, 1980 (Hemiptera: Auchenorrhyncha Issidae), with description of two new species from Central Morocco" masterLastPageNumber="538" masterPageNumber="517" pageNumber="518" updateTime="1699349937175" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="907C048650610C88A9DED5D216DEB6C9">Revision of the genus Lethierium Dlabola, 1980 (Hemiptera: Auchenorrhyncha Issidae), with description of two new species from Central Morocco</mods:title>
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<mods:namePart id="920FA217FBAD1D81CA119CCE6845EA21">Gnezdilov, Vladimir M.</mods:namePart>
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<treatment id="99442B4A6003CE78359A891FFAB2D3EE" ID-DOI="http://doi.org/10.5281/zenodo.6691296" ID-GBIF-Taxon="196193191" ID-Zenodo-Dep="6691296" LSID="urn:lsid:plazi:treatment:99442B4A6003CE78359A891FFAB2D3EE" httpUri="http://treatment.plazi.org/id/99442B4A6003CE78359A891FFAB2D3EE" lastPageNumber="518" pageId="1" pageNumber="518">
<subSubSection id="59F7C9D76003CE78359A891FFDB9D600" box="[151,552,546,573]" pageId="1" pageNumber="518" type="nomenclature">
<paragraph id="11529A5C6003CE78359A891FFDB9D600" blockId="1.[151,552,546,573]" box="[151,552,546,573]" pageId="1" pageNumber="518">
<heading id="4A1A2D306003CE78359A891FFDB9D600" bold="true" box="[151,552,546,573]" fontSize="11" level="1" pageId="1" pageNumber="518" reason="1">
<emphasis id="2399464E6003CE78359A891FFDB9D600" bold="true" box="[151,552,546,573]" pageId="1" pageNumber="518">
Genus
<taxonomicName id="D6EDE1DF6003CE7835E3891FFDB9D600" ID-CoL="5DPD" authority="Dlabola, 1980" authorityName="Dlabola" authorityYear="1980" box="[238,552,546,573]" class="Insecta" family="Issidae" genus="Lethierium" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="518" phylum="Arthropoda" rank="genus">
<emphasis id="2399464E6003CE7835E3891FFEE4D601" bold="true" box="[238,373,546,573]" italics="true" pageId="1" pageNumber="518">Lethierium</emphasis>
<bibRefCitation id="757CE7AD6003CE783470891EFDB9D600" author="Dlabola, J." box="[381,552,546,573]" pageId="1" pageNumber="518" pagination="173 - 248" refId="ref8741" refString="Dlabola, J. (1980) Tribus-Einteilung, neue Gattungen und Arten der Subf. Issinae in der eremischen Zone (Homoptera, Auchenorrhyncha). Acta Musei Nationalis Pragae, 36 B (4), 173 - 248." type="journal article" year="1980">Dlabola, 1980</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="59F7C9D76003CE78359A895AFE46D643" box="[151,471,615,639]" pageId="1" pageNumber="518" type="reference_group">
<paragraph id="11529A5C6003CE78359A895AFE46D643" blockId="1.[151,936,615,670]" box="[151,471,615,639]" pageId="1" pageNumber="518">
<treatmentCitationGroup id="31FDBD726003CE78359A895AFE46D643" box="[151,471,615,639]" pageId="1" pageNumber="518">
<treatmentCitation id="904CBC4D6003CE78359A895AFE42D643" author="Dlabola, J." box="[151,467,615,639]" page="235" pageId="1" pageNumber="518" year="1980">
<taxonomicName id="D6EDE1DF6003CE78359A895AFE42D643" ID-CoL="5DPD" authority="Dlabola, 1980: 235" authorityName="Dlabola" authorityPageNumber="235" authorityYear="1980" box="[151,467,615,639]" class="Insecta" family="Issidae" genus="Lethierium" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="518" phylum="Arthropoda" rank="genus">
<emphasis id="2399464E6003CE78359A895AFE97D643" box="[151,262,615,639]" italics="true" pageId="1" pageNumber="518">Lethierium</emphasis>
<bibRefCitation id="757CE7AD6003CE783400895AFE42D643" author="Dlabola, J." box="[269,467,615,639]" pageId="1" pageNumber="518" pagination="173 - 248" refId="ref8741" refString="Dlabola, J. (1980) Tribus-Einteilung, neue Gattungen und Arten der Subf. Issinae in der eremischen Zone (Homoptera, Auchenorrhyncha). Acta Musei Nationalis Pragae, 36 B (4), 173 - 248." type="journal article" year="1980">Dlabola, 1980: 235</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="59F7C9D76003CE78359A89BBFC39D6A2" box="[151,936,646,670]" pageId="1" pageNumber="518" type="materials_examined">
<paragraph id="11529A5C6003CE78359A89BBFC39D6A2" blockId="1.[151,936,615,670]" box="[151,936,646,670]" pageId="1" pageNumber="518">
<typeStatus id="CE5624FE6003CE78359A89BBFF58D6A2" box="[151,201,646,670]" pageId="1" pageNumber="518">Type</typeStatus>
species:
<taxonomicName id="D6EDE1DF6003CE78342B89BAFD3DD6A2" authority="Lethierry, 1878" authorityName="Lethierry" authorityYear="1878" box="[294,684,646,670]" class="Insecta" family="Issidae" genus="Hysteropterum" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="518" phylum="Arthropoda" rank="species" species="reiberi">
<emphasis id="2399464E6003CE78342B89BAFD99D6A2" box="[294,520,646,670]" italics="true" pageId="1" pageNumber="518">Hysteropterum reiberi</emphasis>
<bibRefCitation id="757CE7AD6003CE78370389BBFD3DD6A2" author="Lethierry, M. L." box="[526,684,646,670]" pageId="1" pageNumber="518" pagination="25 - 31" refId="ref9505" refString="Lethierry, M. L. (1878) Homopteres nouveaux d'Europe et des contrees voisines. Pt. 2. Annales de la Societe Entomologique de Belgique, 21, 25 - 31." type="journal article" year="1878">Lethierry, 1878</bibRefCitation>
</taxonomicName>
, by original designation.
</paragraph>
</subSubSection>
<subSubSection id="59F7C9D76003CE78359A89F1FAB2D3EE" pageId="1" pageNumber="518" type="diagnosis">
<paragraph id="11529A5C6003CE78359A89F1FB7CD713" blockId="1.[151,1437,716,2002]" pageId="1" pageNumber="518">
<emphasis id="2399464E6003CE78359A89F1FE83D6DA" bold="true" box="[151,274,716,742]" pageId="1" pageNumber="518">Diagnosis.</emphasis>
Metope wide, often smooth and without carinae. Coryphe transverse. Forewings wide, with wide hypocostal plate. Hind wings rudimentary. Abdominal sternites IV–VI in imago each with two large sensory pits medially. Phallobase strongly curved, its proximal part (basement) with a pair of lobe-shaped processes.
</paragraph>
<paragraph id="11529A5C6003CE7835CA8805FE3AD11B" blockId="1.[151,1437,716,2002]" pageId="1" pageNumber="518">
<emphasis id="2399464E6003CE7835CA8805FD84D76E" bold="true" box="[199,533,824,850]" pageId="1" pageNumber="518">Supplementary description.</emphasis>
Eyes large, 0.5 as wide as coryphe (in dorsal view). Coryphe transverse, with lateral margins weakly diverging anteriorly (
<figureCitation id="89D686D96003CE78379F8861FD4BD74A" box="[658,730,860,886]" captionStart="FIGURES 1–3" captionStartId="3.[151,264,1230,1254]" captionTargetBox="[152,1417,201,1225]" captionTargetId="figure-334@3.[151,1435,179,1239]" captionTargetPageId="3" captionText="FIGURES 1–3. Lethierium reiberi (Lethierry, 1878), male, Mékalis. 1—dorsal view; 2—lateral view; 3—frontal view. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.6691441" httpUri="https://zenodo.org/record/6691441/files/figure.png" pageId="1" pageNumber="518">Figs 1</figureCitation>
,
<figureCitation id="89D686D96003CE7837E48861FD68D74A" box="[745,761,860,886]" captionStart="FIGURES 6–8" captionStartId="5.[151,264,1738,1762]" captionTargetBox="[182,1420,678,1715]" captionTargetId="figure-251@5.[151,1435,669,1717]" captionTargetPageId="5" captionText="FIGURES 6–8. Lethierium juniperi (Bergevin, 1915), female, Aïn Kerma. 6—dorsal view; 7—lateral view; 8—frontal view. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.6691453" httpUri="https://zenodo.org/record/6691453/files/figure.png" pageId="1" pageNumber="518">6</figureCitation>
,
<figureCitation id="89D686D96003CE78360A8861FC86D74A" box="[775,791,860,886]" captionStart="FIGURES 9–11" captionStartId="6.[151,264,1757,1781]" captionTargetBox="[170,1420,788,1682]" captionTargetId="figure-298@6.[151,1436,776,1701]" captionTargetPageId="6" captionText="FIGURES 9–11. Lethierium cedricolum (Dlabola, 1989), male, Timhadit. 9—dorsal view; 10—lateral view; 11—frontal view. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.6691457" httpUri="https://zenodo.org/record/6691457/files/figure.png" pageId="1" pageNumber="518">9</figureCitation>
,
<figureCitation id="89D686D96003CE7836288861FCD2D74A" box="[805,835,860,886]" captionStart="FIGURES 12–15" captionStartId="7.[151,264,1287,1311]" captionTargetBox="[161,1406,181,1236]" captionTargetId="figure-15@7.[151,1436,181,1266]" captionTargetPageId="7" captionText="FIGURES 12–15. Lethierium grandilobum sp. nov., 12–14—holotype, 15—female paratype, Marrakech. 12—dorsal view; 13, 15—lateral view; 14—frontal view.Abbreviation: kp — knee prominence. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.6691461" httpUri="https://zenodo.org/record/6691461/files/figure.png" pageId="1" pageNumber="518">12</figureCitation>
,
<figureCitation id="89D686D96003CE78365D8861FCFFD74A" box="[848,878,860,886]" captionStart="FIGURES 16–19" captionStartId="8.[151,264,1521,1545]" captionTargetBox="[151,1433,273,1498]" captionTargetId="figure-86@8.[151,1435,273,1500]" captionTargetPageId="8" captionText="FIGURES 16–19. Lethierium carinatum sp. nov., female paratype, Bou Naceur. 16—dorsal view; 17—lateral view; 18—frontal view; 19—caudal view. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.6691465" httpUri="https://zenodo.org/record/6691465/files/figure.png" pageId="1" pageNumber="518">16</figureCitation>
,
<figureCitation id="89D686D96003CE7836718861FC0BD74A" box="[892,922,860,886]" captionStart="FIGURES 20–22" captionStartId="9.[151,264,1302,1326]" captionTargetBox="[156,1425,159,1271]" captionTargetId="figure-314@9.[151,1436,151,1278]" captionTargetPageId="9" captionText="FIGURES 20–22. Lethierium reiberi (Lethierry, 1878), male, Aïn Sefra. 20—face; 21—head and pronotum, dorsal view; 22—hind wing. Out of scale." figureDoi="http://doi.org/10.5281/zenodo.6691469" httpUri="https://zenodo.org/record/6691469/files/figure.png" pageId="1" pageNumber="518">21</figureCitation>
,
<figureCitation id="89D686D96003CE7836AA8861FC54D74A" box="[935,965,860,886]" captionStart="FIGURES 30–31" captionStartId="11.[151,264,1950,1974]" captionTargetBox="[167,1422,166,1900]" captionTargetId="figure-37@11.[151,1436,151,1922]" captionTargetPageId="11" captionText="FIGURES 30–31. Lethierium juniperi (Bergevin, 1915), male, Aïn Kerma. 30 – face; 31 – head and pronotum, dorsal view. Out of scale." figureDoi="http://doi.org/10.5281/zenodo.6691481" httpUri="https://zenodo.org/record/6691481/files/figure.png" pageId="1" pageNumber="518">31</figureCitation>
,
<figureCitation id="89D686D96003CE7836DE8861FC63D74A" box="[979,1010,860,886]" captionStart="FIGURES 39–40" captionStartId="13.[151,264,1900,1924]" captionTargetBox="[205,1382,209,1864]" captionTargetId="figure-15@13.[195,1393,199,1876]" captionTargetPageId="13" captionText="FIGURES 39–40. Lethierium cedricolum (Dlabola, 1989), male, Timhadit. 39—face; 40—head and pronotum, dorsal view. Out of scale." figureDoi="http://doi.org/10.5281/zenodo.6691489" httpUri="https://zenodo.org/record/6691489/files/figure.png" pageId="1" pageNumber="518">40</figureCitation>
,
<figureCitation id="89D686D96003CE7836F38861FB8FD74B" box="[1022,1054,860,887]" captionStart="FIGURES 53–55" captionStartId="16.[151,264,1566,1590]" captionTargetBox="[161,1429,191,1534]" captionTargetId="figure-28@16.[151,1436,181,1542]" captionTargetPageId="16" captionText="FIGURES 53–55. Lethierium carinatum sp. nov., female paratype, Bou Naceur. 53—face; 54—eye, scapus, pedicel, and paranotal lobe, frontal view; 55—head and pronotum, dorsal view. Out of scale." figureDoi="http://doi.org/10.5281/zenodo.6691503" httpUri="https://zenodo.org/record/6691503/files/figure.png" pageId="1" pageNumber="518">55</figureCitation>
). Metope wide, often smooth and without carinae (
<figureCitation id="89D686D96003CE78345B88BDFE31D7A6" box="[342,416,896,922]" captionStart="FIGURES 1–3" captionStartId="3.[151,264,1230,1254]" captionTargetBox="[152,1417,201,1225]" captionTargetId="figure-334@3.[151,1435,179,1239]" captionTargetPageId="3" captionText="FIGURES 1–3. Lethierium reiberi (Lethierry, 1878), male, Mékalis. 1—dorsal view; 2—lateral view; 3—frontal view. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.6691441" httpUri="https://zenodo.org/record/6691441/files/figure.png" pageId="1" pageNumber="518">Figs 3</figureCitation>
,
<figureCitation id="89D686D96003CE7834A388BDFE2CD7A6" box="[430,445,896,922]" captionStart="FIGURES 6–8" captionStartId="5.[151,264,1738,1762]" captionTargetBox="[182,1420,678,1715]" captionTargetId="figure-251@5.[151,1435,669,1717]" captionTargetPageId="5" captionText="FIGURES 6–8. Lethierium juniperi (Bergevin, 1915), female, Aïn Kerma. 6—dorsal view; 7—lateral view; 8—frontal view. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.6691453" httpUri="https://zenodo.org/record/6691453/files/figure.png" pageId="1" pageNumber="518">8</figureCitation>
,
<figureCitation id="89D686D96003CE7834C188BDFE78D7A6" box="[460,489,896,922]" captionStart="FIGURES 9–11" captionStartId="6.[151,264,1757,1781]" captionTargetBox="[170,1420,788,1682]" captionTargetId="figure-298@6.[151,1436,776,1701]" captionTargetPageId="6" captionText="FIGURES 9–11. Lethierium cedricolum (Dlabola, 1989), male, Timhadit. 9—dorsal view; 10—lateral view; 11—frontal view. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.6691457" httpUri="https://zenodo.org/record/6691457/files/figure.png" pageId="1" pageNumber="518">11</figureCitation>
,
<figureCitation id="89D686D96003CE7834FB88BDFD85D7A6" box="[502,532,896,922]" captionStart="FIGURES 12–15" captionStartId="7.[151,264,1287,1311]" captionTargetBox="[161,1406,181,1236]" captionTargetId="figure-15@7.[151,1436,181,1266]" captionTargetPageId="7" captionText="FIGURES 12–15. Lethierium grandilobum sp. nov., 12–14—holotype, 15—female paratype, Marrakech. 12—dorsal view; 13, 15—lateral view; 14—frontal view.Abbreviation: kp — knee prominence. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.6691461" httpUri="https://zenodo.org/record/6691461/files/figure.png" pageId="1" pageNumber="518">14</figureCitation>
,
<figureCitation id="89D686D96003CE78372F88BDFDD0D7A6" box="[546,577,896,922]" captionStart="FIGURES 20–22" captionStartId="9.[151,264,1302,1326]" captionTargetBox="[156,1425,159,1271]" captionTargetId="figure-314@9.[151,1436,151,1278]" captionTargetPageId="9" captionText="FIGURES 20–22. Lethierium reiberi (Lethierry, 1878), male, Aïn Sefra. 20—face; 21—head and pronotum, dorsal view; 22—hind wing. Out of scale." figureDoi="http://doi.org/10.5281/zenodo.6691469" httpUri="https://zenodo.org/record/6691469/files/figure.png" pageId="1" pageNumber="518">20</figureCitation>
,
<figureCitation id="89D686D96003CE78374088BDFDFAD7A6" box="[589,619,896,922]" captionStart="FIGURES 30–31" captionStartId="11.[151,264,1950,1974]" captionTargetBox="[167,1422,166,1900]" captionTargetId="figure-37@11.[151,1436,151,1922]" captionTargetPageId="11" captionText="FIGURES 30–31. Lethierium juniperi (Bergevin, 1915), male, Aïn Kerma. 30 – face; 31 – head and pronotum, dorsal view. Out of scale." figureDoi="http://doi.org/10.5281/zenodo.6691481" httpUri="https://zenodo.org/record/6691481/files/figure.png" pageId="1" pageNumber="518">30</figureCitation>
,
<figureCitation id="89D686D96003CE78377488BDFD08D7A6" box="[633,665,896,922]" captionStart="FIGURES 39–40" captionStartId="13.[151,264,1900,1924]" captionTargetBox="[205,1382,209,1864]" captionTargetId="figure-15@13.[195,1393,199,1876]" captionTargetPageId="13" captionText="FIGURES 39–40. Lethierium cedricolum (Dlabola, 1989), male, Timhadit. 39—face; 40—head and pronotum, dorsal view. Out of scale." figureDoi="http://doi.org/10.5281/zenodo.6691489" httpUri="https://zenodo.org/record/6691489/files/figure.png" pageId="1" pageNumber="518">39</figureCitation>
). Coryphe and metope joint at obtuse angle (in lateral view). Ocelli absent. Postclypeus smooth, without carinae. Rostrum reaching hind coxae; 3
<superScript id="E69837146003CE7836F98899FB93D78E" attach="left" box="[1012,1026,932,946]" fontSize="6" pageId="1" pageNumber="518">rd</superScript>
segment narrowing apically, slightly shorter than 2
<superScript id="E69837146003CE78343C88F5FED0D7EA" attach="left" box="[305,321,968,982]" fontSize="6" pageId="1" pageNumber="518">nd</superScript>
one. Pedicel globular. Pronotum with strongly convex anterior margin and nearly straight posterior margin. Paradiscal fields of pronotum wide behind the eyes. Paranotal lobes of pronotum wide, without carine, with slightly bent lower margin (
<figureCitation id="89D686D96003CE7837098F2DFDC9D017" box="[516,600,1040,1067]" captionStart="FIGURES 53–55" captionStartId="16.[151,264,1566,1590]" captionTargetBox="[161,1429,191,1534]" captionTargetId="figure-28@16.[151,1436,181,1542]" captionTargetPageId="16" captionText="FIGURES 53–55. Lethierium carinatum sp. nov., female paratype, Bou Naceur. 53—face; 54—eye, scapus, pedicel, and paranotal lobe, frontal view; 55—head and pronotum, dorsal view. Out of scale." figureDoi="http://doi.org/10.5281/zenodo.6691503" httpUri="https://zenodo.org/record/6691503/files/figure.png" pageId="1" pageNumber="518">Fig. 54</figureCitation>
). Forewings wide, extend beyond the abdomen apex, with weakly obtusely angulate costal margin and distinct membrane (
<figureCitation id="89D686D96003CE7837A88F09FD7CD072" box="[677,749,1076,1102]" captionStart="FIGURES 1–3" captionStartId="3.[151,264,1230,1254]" captionTargetBox="[152,1417,201,1225]" captionTargetId="figure-334@3.[151,1435,179,1239]" captionTargetPageId="3" captionText="FIGURES 1–3. Lethierium reiberi (Lethierry, 1878), male, Mékalis. 1—dorsal view; 2—lateral view; 3—frontal view. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.6691441" httpUri="https://zenodo.org/record/6691441/files/figure.png" pageId="1" pageNumber="518">Figs 2</figureCitation>
,
<figureCitation id="89D686D96003CE7837F58F08FC99D073" box="[760,776,1077,1103]" captionStart="FIGURES 6–8" captionStartId="5.[151,264,1738,1762]" captionTargetBox="[182,1420,678,1715]" captionTargetId="figure-251@5.[151,1435,669,1717]" captionTargetPageId="5" captionText="FIGURES 6–8. Lethierium juniperi (Bergevin, 1915), female, Aïn Kerma. 6—dorsal view; 7—lateral view; 8—frontal view. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.6691453" httpUri="https://zenodo.org/record/6691453/files/figure.png" pageId="1" pageNumber="518">7</figureCitation>
,
<figureCitation id="89D686D96003CE78361E8F09FCA0D072" box="[787,817,1076,1102]" captionStart="FIGURES 9–11" captionStartId="6.[151,264,1757,1781]" captionTargetBox="[170,1420,788,1682]" captionTargetId="figure-298@6.[151,1436,776,1701]" captionTargetPageId="6" captionText="FIGURES 9–11. Lethierium cedricolum (Dlabola, 1989), male, Timhadit. 9—dorsal view; 10—lateral view; 11—frontal view. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.6691457" httpUri="https://zenodo.org/record/6691457/files/figure.png" pageId="1" pageNumber="518">10</figureCitation>
,
<figureCitation id="89D686D96003CE7836318F09FCCBD072" box="[828,858,1076,1102]" captionStart="FIGURES 12–15" captionStartId="7.[151,264,1287,1311]" captionTargetBox="[161,1406,181,1236]" captionTargetId="figure-15@7.[151,1436,181,1266]" captionTargetPageId="7" captionText="FIGURES 12–15. Lethierium grandilobum sp. nov., 12–14—holotype, 15—female paratype, Marrakech. 12—dorsal view; 13, 15—lateral view; 14—frontal view.Abbreviation: kp — knee prominence. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.6691461" httpUri="https://zenodo.org/record/6691461/files/figure.png" pageId="1" pageNumber="518">13</figureCitation>
,
<figureCitation id="89D686D96003CE78366B8F09FC14D072" box="[870,901,1076,1102]" captionStart="FIGURES 16–19" captionStartId="8.[151,264,1521,1545]" captionTargetBox="[151,1433,273,1498]" captionTargetId="figure-86@8.[151,1435,273,1500]" captionTargetPageId="8" captionText="FIGURES 16–19. Lethierium carinatum sp. nov., female paratype, Bou Naceur. 16—dorsal view; 17—lateral view; 18—frontal view; 19—caudal view. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.6691465" httpUri="https://zenodo.org/record/6691465/files/figure.png" pageId="1" pageNumber="518">17</figureCitation>
). Forewings often with distinct shoulder-shaped prominence (or knee prominence) well recognized as bulge at first furcation of radius (
<figureCitation id="89D686D96003CE7831528F65FB20D04E" box="[1119,1201,1112,1138]" captionStart="FIGURES 12–15" captionStartId="7.[151,264,1287,1311]" captionTargetBox="[161,1406,181,1236]" captionTargetId="figure-15@7.[151,1436,181,1266]" captionTargetPageId="7" captionText="FIGURES 12–15. Lethierium grandilobum sp. nov., 12–14—holotype, 15—female paratype, Marrakech. 12—dorsal view; 13, 15—lateral view; 14—frontal view.Abbreviation: kp — knee prominence. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.6691461" httpUri="https://zenodo.org/record/6691461/files/figure.png" pageId="1" pageNumber="518">Fig. 12</figureCitation>
,
<emphasis id="2399464E6003CE7831B18F65FB47D04E" box="[1212,1238,1112,1138]" italics="true" pageId="1" pageNumber="518">kp</emphasis>
). Forewings with wide precostal area and wide hypocostal plate, basal cell small. Clavus long, ⅔ of wing length. Hind wings rudimentary, nearly reaching abdomen basement (
<figureCitation id="89D686D96003CE7837998F9DFD77D086" box="[660,742,1184,1210]" captionStart="FIGURES 20–22" captionStartId="9.[151,264,1302,1326]" captionTargetBox="[156,1425,159,1271]" captionTargetId="figure-314@9.[151,1436,151,1278]" captionTargetPageId="9" captionText="FIGURES 20–22. Lethierium reiberi (Lethierry, 1878), male, Aïn Sefra. 20—face; 21—head and pronotum, dorsal view; 22—hind wing. Out of scale." figureDoi="http://doi.org/10.5281/zenodo.6691469" httpUri="https://zenodo.org/record/6691469/files/figure.png" pageId="1" pageNumber="518">Fig. 22</figureCitation>
), matte. Hind tibia with 1–2 lateral spines. First metatarsomere short and wide, with two latero-apical and 6–7 intermediate spines in continuous row. Second metatarsomere equal in length to first one, but narrower, with only two latero-apical spines. Ventral surface of first and second metatarsomeres with long setae.
</paragraph>
<paragraph id="11529A5C6003CE7835CA8E0DFE50D1C2" blockId="1.[151,1437,716,2002]" pageId="1" pageNumber="518">
Abdominal sternites IV–VI of imago each with two large sensory pits medially (
<figureCitation id="89D686D96003CE7831408E0DFB23D177" box="[1101,1202,1328,1355]" captionStart="FIGURES 4–5" captionStartId="4.[151,264,1590,1614]" captionTargetBox="[204,1363,237,1519]" captionTargetId="figure-46@4.[151,1435,237,1552]" captionTargetPageId="4" captionText="FIGURES 4–5. Lethierium spp. 4—L. reiberi (Lethierry, 1878), male (Mékalis), ventral view (scale bar—1 mm); 5—L. juniperi (Bergevin, 1915), female (Aïn Kerma), sternite VI, ventral view (out of scale)." figureDoi="http://doi.org/10.5281/zenodo.6691449" httpUri="https://zenodo.org/record/6691449/files/figure.png" pageId="1" pageNumber="518">Figs 4, 5</figureCitation>
). Each pit comprises of wide circle depression of the cuticle with a sensory hair in its centre arising from dark brown socket, laying nearly horizontally and directed to the rim of the depression at acute angle to the midline of the sternite. Besides these large sensory pits the sternites IV–VI have also several trichobothria, from three to seven on each side along its posterior margin, which are represented by long sensory hair (longer than surrounding setae) arising from dark brown socket vertically from the cuticle.
</paragraph>
<paragraph id="11529A5C6003CE7835CA8D35FB7CD2C7" blockId="1.[151,1437,716,2002]" pageId="1" pageNumber="518">
Male pygofer elongate vertically, with weakly obtuse or nearly right upper angle and nearly straight hind margins (
<figureCitation id="89D686D96003CE7835DA8D11FEA0D27A" box="[215,305,1580,1606]" captionStart="FIGURES 23–29" captionStartId="10.[151,264,1599,1623]" captionTargetBox="[182,1405,185,1566]" captionTargetId="figure-28@10.[171,1415,181,1575]" captionTargetPageId="10" captionText="FIGURES 23–29. Lethierium reiberi (Lethierry, 1878), Aïn Sefra, male genitalia. 23—pygofer, lateral view; 24—anal tube, dorsal view; 25—anal tube, lateral view; 26—penis and connective, lateral view; 27—penis, ventral view; 28—style, dorsal view; 29—style, lateral view. Out of scale." figureDoi="http://doi.org/10.5281/zenodo.6691475" httpUri="https://zenodo.org/record/6691475/files/figure.png" pageId="1" pageNumber="518">Figs 23</figureCitation>
,
<figureCitation id="89D686D96003CE7834328D11FECCD27A" box="[319,349,1580,1606]" captionStart="FIGURES 32–38" captionStartId="12.[151,264,1482,1506]" captionTargetBox="[174,1429,254,1409]" captionTargetId="figure-250@12.[151,1436,224,1437]" captionTargetPageId="12" captionText="FIGURES 32–38. Lethierium juniperi (Bergevin, 1915), Aïn Kerma, male genitalia. 32 – pygofer, lateral view; 33 – penis, ven- tral view; 34 – penis and connective, lateral view; 35 – style, lateral view; 36 – style, dorsal view; 37 – anal tube, dorsal view; 38 – anal tube, lateral view. Out of scale." figureDoi="http://doi.org/10.5281/zenodo.6691485" httpUri="https://zenodo.org/record/6691485/files/figure.png" pageId="1" pageNumber="518">32</figureCitation>
,
<figureCitation id="89D686D96003CE7834618D11FE1BD27A" box="[364,394,1580,1606]" captionStart="FIGURES 41–46" captionStartId="14.[151,264,1513,1537]" captionTargetBox="[158,1430,188,1454]" captionTargetId="figure-28@14.[151,1436,181,1462]" captionTargetPageId="14" captionText="FIGURES 41–46. Lethierium cedricolum (Dlabola, 1989), Timhadit, male genitalia. 41—anal tube and pygofer, lateral view; 42—anal tube, dorsal view; 43—style, lateral view; 44—style, dorsal view; 45—penis, ventral view; 46—penis and connective, lateral view. Out of scale." figureDoi="http://doi.org/10.5281/zenodo.6691491" httpUri="https://zenodo.org/record/6691491/files/figure.png" pageId="1" pageNumber="518">41</figureCitation>
,
<figureCitation id="89D686D96003CE7834958D11FE27D27A" box="[408,438,1580,1606]" captionStart="FIGURES 47–52" captionStartId="15.[151,264,1887,1911]" captionTargetBox="[166,1423,188,1854]" captionTargetId="figure-15@15.[151,1436,181,1863]" captionTargetPageId="15" captionText="FIGURES 47–52. Lethierium grandilobum sp. nov., holotype, male genitalia. 47—anal tube, pygofer, and penis, lateral view; 48—anal tube, dorsal view; 49—penis and connective, lateral view; 50—penis and connective, ventral view; 51—style, lateral view; 52—style, dorsal view. Out of scale." figureDoi="http://doi.org/10.5281/zenodo.6691493" httpUri="https://zenodo.org/record/6691493/files/figure.png" pageId="1" pageNumber="518">47</figureCitation>
,
<figureCitation id="89D686D96003CE7834C88D11FE74D27B" box="[453,485,1580,1607]" captionStart="FIGURES 56–63" captionStartId="17.[151,264,1697,1721]" captionTargetBox="[161,1425,532,1642]" captionTargetId="figure-160@17.[151,1436,524,1655]" captionTargetPageId="17" captionText="FIGURES 56–63. Lethierium carinatum sp. nov., holotype, male genitalia. 56—pygofer, lateral view; 57—anal tube, lateral view; 58—anal tube, dorsal view; 59—penis and connective, lateral view; 60—connective, caudal view; 61—penis, ventral view; 62—style, lateral view; 63—style, dorsal view. Out of scale." figureDoi="http://doi.org/10.5281/zenodo.6691509" httpUri="https://zenodo.org/record/6691509/files/figure.png" pageId="1" pageNumber="518">56</figureCitation>
). Lateral margins of anal tube turned downwards (25, 38, 57). Phallobase strongly curved, with a weakly sclerotized sac dorso-apically, its basement with a pair of lobe-shaped processes (
<figureCitation id="89D686D96003CE7830328D6DFA06D256" box="[1343,1431,1616,1642]" captionStart="FIGURES 23–29" captionStartId="10.[151,264,1599,1623]" captionTargetBox="[182,1405,185,1566]" captionTargetId="figure-28@10.[171,1415,181,1575]" captionTargetPageId="10" captionText="FIGURES 23–29. Lethierium reiberi (Lethierry, 1878), Aïn Sefra, male genitalia. 23—pygofer, lateral view; 24—anal tube, dorsal view; 25—anal tube, lateral view; 26—penis and connective, lateral view; 27—penis, ventral view; 28—style, dorsal view; 29—style, lateral view. Out of scale." figureDoi="http://doi.org/10.5281/zenodo.6691475" httpUri="https://zenodo.org/record/6691475/files/figure.png" pageId="1" pageNumber="518">Figs 26</figureCitation>
,
<figureCitation id="89D686D96003CE78359A8D49FF24D2B2" box="[151,181,1652,1678]" captionStart="FIGURES 32–38" captionStartId="12.[151,264,1482,1506]" captionTargetBox="[174,1429,254,1409]" captionTargetId="figure-250@12.[151,1436,224,1437]" captionTargetPageId="12" captionText="FIGURES 32–38. Lethierium juniperi (Bergevin, 1915), Aïn Kerma, male genitalia. 32 – pygofer, lateral view; 33 – penis, ven- tral view; 34 – penis and connective, lateral view; 35 – style, lateral view; 36 – style, dorsal view; 37 – anal tube, dorsal view; 38 – anal tube, lateral view. Out of scale." figureDoi="http://doi.org/10.5281/zenodo.6691485" httpUri="https://zenodo.org/record/6691485/files/figure.png" pageId="1" pageNumber="518">34</figureCitation>
,
<figureCitation id="89D686D96003CE7835CF8D49FF70D2B2" box="[194,225,1652,1678]" captionStart="FIGURES 41–46" captionStartId="14.[151,264,1513,1537]" captionTargetBox="[158,1430,188,1454]" captionTargetId="figure-28@14.[151,1436,181,1462]" captionTargetPageId="14" captionText="FIGURES 41–46. Lethierium cedricolum (Dlabola, 1989), Timhadit, male genitalia. 41—anal tube and pygofer, lateral view; 42—anal tube, dorsal view; 43—style, lateral view; 44—style, dorsal view; 45—penis, ventral view; 46—penis and connective, lateral view. Out of scale." figureDoi="http://doi.org/10.5281/zenodo.6691491" httpUri="https://zenodo.org/record/6691491/files/figure.png" pageId="1" pageNumber="518">46</figureCitation>
,
<figureCitation id="89D686D96003CE7835E08D49FE9DD2B2" box="[237,268,1652,1678]" captionStart="FIGURES 47–52" captionStartId="15.[151,264,1887,1911]" captionTargetBox="[166,1423,188,1854]" captionTargetId="figure-15@15.[151,1436,181,1863]" captionTargetPageId="15" captionText="FIGURES 47–52. Lethierium grandilobum sp. nov., holotype, male genitalia. 47—anal tube, pygofer, and penis, lateral view; 48—anal tube, dorsal view; 49—penis and connective, lateral view; 50—penis and connective, ventral view; 51—style, lateral view; 52—style, dorsal view. Out of scale." figureDoi="http://doi.org/10.5281/zenodo.6691493" httpUri="https://zenodo.org/record/6691493/files/figure.png" pageId="1" pageNumber="518">49</figureCitation>
,
<figureCitation id="89D686D96003CE7834158D49FEA9D2B3" box="[280,312,1652,1679]" captionStart="FIGURES 56–63" captionStartId="17.[151,264,1697,1721]" captionTargetBox="[161,1425,532,1642]" captionTargetId="figure-160@17.[151,1436,524,1655]" captionTargetPageId="17" captionText="FIGURES 56–63. Lethierium carinatum sp. nov., holotype, male genitalia. 56—pygofer, lateral view; 57—anal tube, lateral view; 58—anal tube, dorsal view; 59—penis and connective, lateral view; 60—connective, caudal view; 61—penis, ventral view; 62—style, lateral view; 63—style, dorsal view. Out of scale." figureDoi="http://doi.org/10.5281/zenodo.6691509" httpUri="https://zenodo.org/record/6691509/files/figure.png" pageId="1" pageNumber="518">59</figureCitation>
). Dorsolateral phallobase lobes narrowing apically, without processes. Ventral phallobase lobe long and wide, narrowing apically (
<figureCitation id="89D686D96003CE7834E08DA5FDD5D28E" box="[493,580,1688,1714]" captionStart="FIGURES 23–29" captionStartId="10.[151,264,1599,1623]" captionTargetBox="[182,1405,185,1566]" captionTargetId="figure-28@10.[171,1415,181,1575]" captionTargetPageId="10" captionText="FIGURES 23–29. Lethierium reiberi (Lethierry, 1878), Aïn Sefra, male genitalia. 23—pygofer, lateral view; 24—anal tube, dorsal view; 25—anal tube, lateral view; 26—penis and connective, lateral view; 27—penis, ventral view; 28—style, dorsal view; 29—style, lateral view. Out of scale." figureDoi="http://doi.org/10.5281/zenodo.6691475" httpUri="https://zenodo.org/record/6691475/files/figure.png" pageId="1" pageNumber="518">Figs 27</figureCitation>
,
<figureCitation id="89D686D96003CE78375D8DA5FDFFD28E" box="[592,622,1688,1714]" captionStart="FIGURES 32–38" captionStartId="12.[151,264,1482,1506]" captionTargetBox="[174,1429,254,1409]" captionTargetId="figure-250@12.[151,1436,224,1437]" captionTargetPageId="12" captionText="FIGURES 32–38. Lethierium juniperi (Bergevin, 1915), Aïn Kerma, male genitalia. 32 – pygofer, lateral view; 33 – penis, ven- tral view; 34 – penis and connective, lateral view; 35 – style, lateral view; 36 – style, dorsal view; 37 – anal tube, dorsal view; 38 – anal tube, lateral view. Out of scale." figureDoi="http://doi.org/10.5281/zenodo.6691485" httpUri="https://zenodo.org/record/6691485/files/figure.png" pageId="1" pageNumber="518">33</figureCitation>
,
<figureCitation id="89D686D96003CE7837778DA5FD09D28F" box="[634,664,1688,1715]" captionStart="FIGURES 41–46" captionStartId="14.[151,264,1513,1537]" captionTargetBox="[158,1430,188,1454]" captionTargetId="figure-28@14.[151,1436,181,1462]" captionTargetPageId="14" captionText="FIGURES 41–46. Lethierium cedricolum (Dlabola, 1989), Timhadit, male genitalia. 41—anal tube and pygofer, lateral view; 42—anal tube, dorsal view; 43—style, lateral view; 44—style, dorsal view; 45—penis, ventral view; 46—penis and connective, lateral view. Out of scale." figureDoi="http://doi.org/10.5281/zenodo.6691491" httpUri="https://zenodo.org/record/6691491/files/figure.png" pageId="1" pageNumber="518">45</figureCitation>
,
<figureCitation id="89D686D96003CE7837A98DA5FD53D28F" box="[676,706,1688,1715]" captionStart="FIGURES 47–52" captionStartId="15.[151,264,1887,1911]" captionTargetBox="[166,1423,188,1854]" captionTargetId="figure-15@15.[151,1436,181,1863]" captionTargetPageId="15" captionText="FIGURES 47–52. Lethierium grandilobum sp. nov., holotype, male genitalia. 47—anal tube, pygofer, and penis, lateral view; 48—anal tube, dorsal view; 49—penis and connective, lateral view; 50—penis and connective, ventral view; 51—style, lateral view; 52—style, dorsal view. Out of scale." figureDoi="http://doi.org/10.5281/zenodo.6691493" httpUri="https://zenodo.org/record/6691493/files/figure.png" pageId="1" pageNumber="518">50</figureCitation>
,
<figureCitation id="89D686D96003CE7837C38DA5FD7CD28F" box="[718,749,1688,1715]" captionStart="FIGURES 56–63" captionStartId="17.[151,264,1697,1721]" captionTargetBox="[161,1425,532,1642]" captionTargetId="figure-160@17.[151,1436,524,1655]" captionTargetPageId="17" captionText="FIGURES 56–63. Lethierium carinatum sp. nov., holotype, male genitalia. 56—pygofer, lateral view; 57—anal tube, lateral view; 58—anal tube, dorsal view; 59—penis and connective, lateral view; 60—connective, caudal view; 61—penis, ventral view; 62—style, lateral view; 63—style, dorsal view. Out of scale." figureDoi="http://doi.org/10.5281/zenodo.6691509" httpUri="https://zenodo.org/record/6691509/files/figure.png" pageId="1" pageNumber="518">61</figureCitation>
). Apical aedeagal processes not visible above the phallobase, narrowing apically. Aedeagus with a pair of ventral hooks, often strongly curved. Style with well developed apical and lateral teeth (
<figureCitation id="89D686D96003CE7834548DDDFE21D2C6" box="[345,432,1760,1786]" captionStart="FIGURES 23–29" captionStartId="10.[151,264,1599,1623]" captionTargetBox="[182,1405,185,1566]" captionTargetId="figure-28@10.[171,1415,181,1575]" captionTargetPageId="10" captionText="FIGURES 23–29. Lethierium reiberi (Lethierry, 1878), Aïn Sefra, male genitalia. 23—pygofer, lateral view; 24—anal tube, dorsal view; 25—anal tube, lateral view; 26—penis and connective, lateral view; 27—penis, ventral view; 28—style, dorsal view; 29—style, lateral view. Out of scale." figureDoi="http://doi.org/10.5281/zenodo.6691475" httpUri="https://zenodo.org/record/6691475/files/figure.png" pageId="1" pageNumber="518">Figs 29</figureCitation>
,
<figureCitation id="89D686D96003CE7834B18DDDFE4BD2C7" box="[444,474,1760,1787]" captionStart="FIGURES 32–38" captionStartId="12.[151,264,1482,1506]" captionTargetBox="[174,1429,254,1409]" captionTargetId="figure-250@12.[151,1436,224,1437]" captionTargetPageId="12" captionText="FIGURES 32–38. Lethierium juniperi (Bergevin, 1915), Aïn Kerma, male genitalia. 32 – pygofer, lateral view; 33 – penis, ven- tral view; 34 – penis and connective, lateral view; 35 – style, lateral view; 36 – style, dorsal view; 37 – anal tube, dorsal view; 38 – anal tube, lateral view. Out of scale." figureDoi="http://doi.org/10.5281/zenodo.6691485" httpUri="https://zenodo.org/record/6691485/files/figure.png" pageId="1" pageNumber="518">35</figureCitation>
,
<figureCitation id="89D686D96003CE7834EB8DDDFD95D2C6" box="[486,516,1760,1786]" captionStart="FIGURES 41–46" captionStartId="14.[151,264,1513,1537]" captionTargetBox="[158,1430,188,1454]" captionTargetId="figure-28@14.[151,1436,181,1462]" captionTargetPageId="14" captionText="FIGURES 41–46. Lethierium cedricolum (Dlabola, 1989), Timhadit, male genitalia. 41—anal tube and pygofer, lateral view; 42—anal tube, dorsal view; 43—style, lateral view; 44—style, dorsal view; 45—penis, ventral view; 46—penis and connective, lateral view. Out of scale." figureDoi="http://doi.org/10.5281/zenodo.6691491" httpUri="https://zenodo.org/record/6691491/files/figure.png" pageId="1" pageNumber="518">43</figureCitation>
,
<figureCitation id="89D686D96003CE78371D8DDDFDBFD2C7" box="[528,558,1760,1787]" captionStart="FIGURES 47–52" captionStartId="15.[151,264,1887,1911]" captionTargetBox="[166,1423,188,1854]" captionTargetId="figure-15@15.[151,1436,181,1863]" captionTargetPageId="15" captionText="FIGURES 47–52. Lethierium grandilobum sp. nov., holotype, male genitalia. 47—anal tube, pygofer, and penis, lateral view; 48—anal tube, dorsal view; 49—penis and connective, lateral view; 50—penis and connective, ventral view; 51—style, lateral view; 52—style, dorsal view. Out of scale." figureDoi="http://doi.org/10.5281/zenodo.6691493" httpUri="https://zenodo.org/record/6691493/files/figure.png" pageId="1" pageNumber="518">51</figureCitation>
,
<figureCitation id="89D686D96003CE7837378DDDFDCBD2C7" box="[570,602,1760,1787]" captionStart="FIGURES 56–63" captionStartId="17.[151,264,1697,1721]" captionTargetBox="[161,1425,532,1642]" captionTargetId="figure-160@17.[151,1436,524,1655]" captionTargetPageId="17" captionText="FIGURES 56–63. Lethierium carinatum sp. nov., holotype, male genitalia. 56—pygofer, lateral view; 57—anal tube, lateral view; 58—anal tube, dorsal view; 59—penis and connective, lateral view; 60—connective, caudal view; 61—penis, ventral view; 62—style, lateral view; 63—style, dorsal view. Out of scale." figureDoi="http://doi.org/10.5281/zenodo.6691509" httpUri="https://zenodo.org/record/6691509/files/figure.png" pageId="1" pageNumber="518">62</figureCitation>
). Connective with a large cup (
<figureCitation id="89D686D96003CE7836B88DDDFB9DD2C6" box="[949,1036,1760,1786]" captionStart="FIGURES 23–29" captionStartId="10.[151,264,1599,1623]" captionTargetBox="[182,1405,185,1566]" captionTargetId="figure-28@10.[171,1415,181,1575]" captionTargetPageId="10" captionText="FIGURES 23–29. Lethierium reiberi (Lethierry, 1878), Aïn Sefra, male genitalia. 23—pygofer, lateral view; 24—anal tube, dorsal view; 25—anal tube, lateral view; 26—penis and connective, lateral view; 27—penis, ventral view; 28—style, dorsal view; 29—style, lateral view. Out of scale." figureDoi="http://doi.org/10.5281/zenodo.6691475" httpUri="https://zenodo.org/record/6691475/files/figure.png" pageId="1" pageNumber="518">Figs 26</figureCitation>
,
<figureCitation id="89D686D96003CE7831158DDDFBA7D2C6" box="[1048,1078,1760,1786]" captionStart="FIGURES 32–38" captionStartId="12.[151,264,1482,1506]" captionTargetBox="[174,1429,254,1409]" captionTargetId="figure-250@12.[151,1436,224,1437]" captionTargetPageId="12" captionText="FIGURES 32–38. Lethierium juniperi (Bergevin, 1915), Aïn Kerma, male genitalia. 32 – pygofer, lateral view; 33 – penis, ven- tral view; 34 – penis and connective, lateral view; 35 – style, lateral view; 36 – style, dorsal view; 37 – anal tube, dorsal view; 38 – anal tube, lateral view. Out of scale." figureDoi="http://doi.org/10.5281/zenodo.6691485" httpUri="https://zenodo.org/record/6691485/files/figure.png" pageId="1" pageNumber="518">34</figureCitation>
,
<figureCitation id="89D686D96003CE78314F8DDDFBF0D2C6" box="[1090,1121,1760,1786]" captionStart="FIGURES 41–46" captionStartId="14.[151,264,1513,1537]" captionTargetBox="[158,1430,188,1454]" captionTargetId="figure-28@14.[151,1436,181,1462]" captionTargetPageId="14" captionText="FIGURES 41–46. Lethierium cedricolum (Dlabola, 1989), Timhadit, male genitalia. 41—anal tube and pygofer, lateral view; 42—anal tube, dorsal view; 43—style, lateral view; 44—style, dorsal view; 45—penis, ventral view; 46—penis and connective, lateral view. Out of scale." figureDoi="http://doi.org/10.5281/zenodo.6691491" httpUri="https://zenodo.org/record/6691491/files/figure.png" pageId="1" pageNumber="518">46</figureCitation>
,
<figureCitation id="89D686D96003CE7831618DDDFB1AD2C6" box="[1132,1163,1760,1786]" captionStart="FIGURES 47–52" captionStartId="15.[151,264,1887,1911]" captionTargetBox="[166,1423,188,1854]" captionTargetId="figure-15@15.[151,1436,181,1863]" captionTargetPageId="15" captionText="FIGURES 47–52. Lethierium grandilobum sp. nov., holotype, male genitalia. 47—anal tube, pygofer, and penis, lateral view; 48—anal tube, dorsal view; 49—penis and connective, lateral view; 50—penis and connective, ventral view; 51—style, lateral view; 52—style, dorsal view. Out of scale." figureDoi="http://doi.org/10.5281/zenodo.6691493" httpUri="https://zenodo.org/record/6691493/files/figure.png" pageId="1" pageNumber="518">49</figureCitation>
,
<figureCitation id="89D686D96003CE78319B8DDDFB71D2C7" box="[1174,1248,1760,1787]" captionStart="FIGURES 56–63" captionStartId="17.[151,264,1697,1721]" captionTargetBox="[161,1425,532,1642]" captionTargetId="figure-160@17.[151,1436,524,1655]" captionTargetPageId="17" captionText="FIGURES 56–63. Lethierium carinatum sp. nov., holotype, male genitalia. 56—pygofer, lateral view; 57—anal tube, lateral view; 58—anal tube, dorsal view; 59—penis and connective, lateral view; 60—connective, caudal view; 61—penis, ventral view; 62—style, lateral view; 63—style, dorsal view. Out of scale." figureDoi="http://doi.org/10.5281/zenodo.6691509" httpUri="https://zenodo.org/record/6691509/files/figure.png" pageId="1" pageNumber="518">59, 60</figureCitation>
).
</paragraph>
<paragraph id="11529A5C6003CE7835CA8C39FAB2D3EE" blockId="1.[151,1437,716,2002]" pageId="1" pageNumber="518">
Hind margin of female sternite VII widely concave (in ventral view), with labial median process (in dorsal view) (
<figureCitation id="89D686D96003CE7835EB8C15FE03D37E" box="[230,402,1832,1858]" captionStart="FIGURES 64–70" captionStartId="18.[151,264,1215,1239]" captionTargetBox="[160,1427,187,1184]" captionTargetId="figure-28@18.[151,1436,181,1192]" captionTargetPageId="18" captionText="FIGURES 64–70. Lethierium spp., female. 64—L. cedricolum (Dlabola, 1989), Ras el ma, sternite VII, dorsal view; 65—same, ventral view; 66—L. reiberi (Lethierry, 1878), Mékalis, sternite VII, dorsal view; 67—L. juniperi (Bergevin, 1915), Aïn Kerma, sternite VII, dorsal view; 68—same, ventral view; 69—L. carinatum sp. nov., paratype, Bou Naceur, anal tube, dorsal view; 70—L. carinatum sp. nov., paratype, Bou Naceur, sternite VII, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.6691513" httpUri="https://zenodo.org/record/6691513/files/figure.png" pageId="1" pageNumber="518">Figs 64–68, 70</figureCitation>
). Gonoplacs convex, without carinae (
<figureCitation id="89D686D96003CE7836338C15FC04D37E" box="[830,917,1832,1858]" captionStart="FIGURES 16–19" captionStartId="8.[151,264,1521,1545]" captionTargetBox="[151,1433,273,1498]" captionTargetId="figure-86@8.[151,1435,273,1500]" captionTargetPageId="8" captionText="FIGURES 16–19. Lethierium carinatum sp. nov., female paratype, Bou Naceur. 16—dorsal view; 17—lateral view; 18—frontal view; 19—caudal view. Scale bar—1 mm." figureDoi="http://doi.org/10.5281/zenodo.6691465" httpUri="https://zenodo.org/record/6691465/files/figure.png" pageId="1" pageNumber="518">Figs 19</figureCitation>
,
<figureCitation id="89D686D96003CE7836AC8C15FC51D37E" box="[929,960,1832,1858]" captionStart="FIGURES 71–77" captionStartId="19.[151,264,1708,1732]" captionTargetBox="[157,1428,189,1675]" captionTargetId="figure-15@19.[151,1436,181,1685]" captionTargetPageId="19" captionText="FIGURES 71–77. Lethierium spp., ovipositor structure, 71–75—L. juniperi (Bergevin, 1915), Aïn Kerma, 76–77—L. cedricolum (Dlabola, 1989), Ras el ma. 71—anal tube and gonoplac, dorsal view; 72, 76—gonapophyses IX and gonospiculum bridge, lateral view; 73—same, ventral view; 74—gonapophysis VIII; 75—apex of endogonocoxal lobe; 77—basement of gonapophyses IX and gonospiculum bridge, caudal view. Out of scale." figureDoi="http://doi.org/10.5281/zenodo.6691517" httpUri="https://zenodo.org/record/6691517/files/figure.png" pageId="1" pageNumber="518">71</figureCitation>
). Gonocoxa VIII with weakly lobe-shaped hind margin (
<figureCitation id="89D686D96003CE7834228C71FE15D35A" box="[303,388,1868,1894]" captionStart="FIGURES 71–77" captionStartId="19.[151,264,1708,1732]" captionTargetBox="[157,1428,189,1675]" captionTargetId="figure-15@19.[151,1436,181,1685]" captionTargetPageId="19" captionText="FIGURES 71–77. Lethierium spp., ovipositor structure, 71–75—L. juniperi (Bergevin, 1915), Aïn Kerma, 76–77—L. cedricolum (Dlabola, 1989), Ras el ma. 71—anal tube and gonoplac, dorsal view; 72, 76—gonapophyses IX and gonospiculum bridge, lateral view; 73—same, ventral view; 74—gonapophysis VIII; 75—apex of endogonocoxal lobe; 77—basement of gonapophyses IX and gonospiculum bridge, caudal view. Out of scale." figureDoi="http://doi.org/10.5281/zenodo.6691517" httpUri="https://zenodo.org/record/6691517/files/figure.png" pageId="1" pageNumber="518">Fig. 74</figureCitation>
). Anterior connective lamina of gonapophysis VIII wide, with three teeth in apical group and 3–4 teeth in lateral group. Endogonocoxal process slightly furcating apically (
<figureCitation id="89D686D96003CE7836F98C4DFBD6D3B7" box="[1012,1095,1904,1931]" captionStart="FIGURES 71–77" captionStartId="19.[151,264,1708,1732]" captionTargetBox="[157,1428,189,1675]" captionTargetId="figure-15@19.[151,1436,181,1685]" captionTargetPageId="19" captionText="FIGURES 71–77. Lethierium spp., ovipositor structure, 71–75—L. juniperi (Bergevin, 1915), Aïn Kerma, 76–77—L. cedricolum (Dlabola, 1989), Ras el ma. 71—anal tube and gonoplac, dorsal view; 72, 76—gonapophyses IX and gonospiculum bridge, lateral view; 73—same, ventral view; 74—gonapophysis VIII; 75—apex of endogonocoxal lobe; 77—basement of gonapophyses IX and gonospiculum bridge, caudal view. Out of scale." figureDoi="http://doi.org/10.5281/zenodo.6691517" httpUri="https://zenodo.org/record/6691517/files/figure.png" pageId="1" pageNumber="518">Fig. 75</figureCitation>
). Posterior connective laminae of gonapophyses IX vertically elongate, with its distal parts curved at obtuse angle (
<figureCitation id="89D686D96003CE7831588CA9FAA3D393" box="[1109,1330,1940,1967]" captionStart="FIGURES 71–77" captionStartId="19.[151,264,1708,1732]" captionTargetBox="[157,1428,189,1675]" captionTargetId="figure-15@19.[151,1436,181,1685]" captionTargetPageId="19" captionText="FIGURES 71–77. Lethierium spp., ovipositor structure, 71–75—L. juniperi (Bergevin, 1915), Aïn Kerma, 76–77—L. cedricolum (Dlabola, 1989), Ras el ma. 71—anal tube and gonoplac, dorsal view; 72, 76—gonapophyses IX and gonospiculum bridge, lateral view; 73—same, ventral view; 74—gonapophysis VIII; 75—apex of endogonocoxal lobe; 77—basement of gonapophyses IX and gonospiculum bridge, caudal view. Out of scale." figureDoi="http://doi.org/10.5281/zenodo.6691517" httpUri="https://zenodo.org/record/6691517/files/figure.png" pageId="1" pageNumber="518">Figs 72, 73, 76, 77</figureCitation>
). Median field of the laminae two-lobed (
<figureCitation id="89D686D96003CE7834F78C85FDDFD3EE" box="[506,590,1976,2002]" captionStart="FIGURES 71–77" captionStartId="19.[151,264,1708,1732]" captionTargetBox="[157,1428,189,1675]" captionTargetId="figure-15@19.[151,1436,181,1685]" captionTargetPageId="19" captionText="FIGURES 71–77. Lethierium spp., ovipositor structure, 71–75—L. juniperi (Bergevin, 1915), Aïn Kerma, 76–77—L. cedricolum (Dlabola, 1989), Ras el ma. 71—anal tube and gonoplac, dorsal view; 72, 76—gonapophyses IX and gonospiculum bridge, lateral view; 73—same, ventral view; 74—gonapophysis VIII; 75—apex of endogonocoxal lobe; 77—basement of gonapophyses IX and gonospiculum bridge, caudal view. Out of scale." figureDoi="http://doi.org/10.5281/zenodo.6691517" httpUri="https://zenodo.org/record/6691517/files/figure.png" pageId="1" pageNumber="518">Fig. 73</figureCitation>
). Lateral fields of the laminae short. Gonospiculum bridge large.
</paragraph>
</subSubSection>
</treatment>
</document>
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