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<document ID-DOI="http://doi.org/10.5281/zenodo.3792178" ID-GBIF-Dataset="3026c846-a607-4826-b2a9-8f7a531baec5" ID-GBIF-Taxon="163752672" ID-ISSN="1313–2970" ID-Zenodo-Dep="3792178" ID-ZooBank="B32EF8D5-C3C6-4C8C-8845-284125448425" approvalRequired="25" approvalRequired_for_textStreams="25" checkinTime="1587995767728" checkinUser="plazi" docAuthor="Golovatch, Sergei, Geoffroy, Jean-Jacques, Mauriès, Jean-Paul &amp; VandenSpiegel, Didier" docDate="2009" docId="936787D7FF9BFF9F8ABB4688FB92FD03" docLanguage="en" docName="ZK_article_1994.pdf" docOrigin="ZooKeys 12 (12)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@52c7884c" docStyleName="zookeys.2008.journal_article" docTitle="Eutrichodesmus Silvestri 1910" docType="key" docVersion="9" lastPageId="43" lastPageNumber="44" masterDocId="6F5EFFAFFFB3FFB48A374557FFC6FF89" masterDocTitle="Review of the millipede genus Eutrichodesmus Silvestri, 1910 (Diplopoda, Polydesmida, Haplodesmidae), with descriptions of new species" masterLastPageNumber="46" masterPageNumber="1" pageId="40" pageNumber="41" updateTime="1643565021146" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
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<mods:titleInfo>
<mods:title>Review of the millipede genus Eutrichodesmus Silvestri, 1910 (Diplopoda, Polydesmida, Haplodesmidae), with descriptions of new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Golovatch, Sergei</mods:namePart>
<mods:affiliation>Russian Academy of Sciences, Moscow, Russia</mods:affiliation>
</mods:name>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Geoffroy, Jean-Jacques</mods:namePart>
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<mods:affiliation>Muséum national d’Histoire naturelle, Département Ecologie &amp; Gestion de la Biodiversité, UMR 7179 du CNRS, Equipe EVOLTRAIT, Brunoy, France &amp; D 362 Dc 0 - &amp; CC</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Mauriès, Jean-Paul</mods:namePart>
<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
<mods:affiliation>Muséum national d’Histoire naturelle, Département Systématique et Evolution, Section Arthropodes, Paris, France &amp; - &amp; - 42 F &amp; D 01 - 28 Ad- 43 F</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>VandenSpiegel, Didier</mods:namePart>
<mods:affiliation>Musée Royal de l’Afrique centrale, Tervuren, Belgium &amp; A &amp; EE &amp; F- 04802 E 68 Cb</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2009</mods:date>
<mods:detail type="pubDate">
<mods:number>2009-06-18</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>12</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>12</mods:number>
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<mods:start>1</mods:start>
<mods:end>46</mods:end>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.3897/zookeys.12.167</mods:identifier>
<mods:identifier type="GBIF-Dataset">3026c846-a607-4826-b2a9-8f7a531baec5</mods:identifier>
<mods:identifier type="ISSN">1313–2970</mods:identifier>
<mods:identifier type="Zenodo-Dep">576460</mods:identifier>
<mods:identifier type="ZooBank">B32EF8D5-C3C6-4C8C-8845-284125448425</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3792178" ID-GBIF-Taxon="163752672" ID-Zenodo-Dep="3792178" LSID="urn:lsid:plazi:treatment:936787D7FF9BFF9F8ABB4688FB92FD03" httpUri="http://treatment.plazi.org/id/936787D7FF9BFF9F8ABB4688FB92FD03" lastPageId="43" lastPageNumber="44" pageId="40" pageNumber="41">
<subSubSection box="[140,631,991,1018]" pageId="40" pageNumber="41" type="nomenclature">
<paragraph blockId="40.[140,631,991,1018]" box="[140,631,991,1018]" pageId="40" pageNumber="41">
<heading bold="true" box="[140,631,991,1018]" fontSize="11" level="1" pageId="40" pageNumber="41" reason="1">
<emphasis bold="true" box="[140,631,991,1018]" pageId="40" pageNumber="41">
Key to the species of
<taxonomicName authorityName="Silvestri" authorityYear="1910" box="[429,631,991,1018]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[429,631,991,1018]" italics="true" pageId="40" pageNumber="41">Eutrichodesmus</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection lastPageId="43" lastPageNumber="44" pageId="40" pageNumber="41" type="key">
<keyStep pageId="40" pageNumber="41">
<paragraph blockId="40.[140,1108,1047,1636]" pageId="40" pageNumber="41">
<keyLead pageId="40" pageNumber="41">
1 At least some metaterga with an evident mid-dorsal outgrowth or projection (
<figureCitation box="[253,321,1082,1108]" captionStart="Figure 9" captionStartId="12.[140,212,1558,1580]" captionTargetBox="[177,1070,165,1538]" captionTargetId="figure@12.[177,1070,165,1539]" captionTargetPageId="12" captionText="Figure 9. Eutrichodesmus aster sp. n., subadult paratype from Tham Han Cave; A, habitus, lateral view; B, anterior part of body, lateral view; C, midbody segments, lateral view; D, posterior part of body, lateral view; E, metatergal seta, sublateral view; F, anterior part of body, frontodorsal view. – Scale bars: A-D, F, 0.5 mm; E, 0.001 mm." figureDoi="http://doi.org/10.5281/zenodo.3769987" httpUri="https://zenodo.org/record/3769987/files/figure.png" pageId="40" pageNumber="41">Figs 9</figureCitation>
A-D; 20A-D)....................................................................................
<emphasis bold="true" box="[1092,1107,1081,1107]" pageId="40" pageNumber="41">2</emphasis>
</keyLead>
</paragraph>
<paragraph blockId="40.[140,1108,1047,1636]" pageId="40" pageNumber="41">
<keyLead pageId="40" pageNumber="41">
– All metaterga subequal, devoid of an evident mid-dorsal outgrowth or projection (
<figureCitation box="[306,373,1152,1178]" captionStart="Figure 2" captionStartId="4.[140,213,1558,1580]" captionTargetBox="[140,1108,502,1538]" captionTargetId="figure@4.[140,1108,502,1539]" captionTargetPageId="4" captionText="Figure 2. Eutrichodesmus distinctus sp. n., ♁ paratype; A, habitus, lateral view; B, E, anterior part of body, lateral and dorsal views, respectively; C, F, midbody segments, lateral and dorsal views, respectively; D, G, posterior part of body, lateral and dorsal views, respectively. – Scale bars: A, 0.5 mm; B-G, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3769973" httpUri="https://zenodo.org/record/3769973/files/figure.png" pageId="40" pageNumber="41">Figs 2</figureCitation>
A-F, 5A-D; 17A-D)...................................................................
<emphasis bold="true" box="[1092,1107,1152,1178]" pageId="40" pageNumber="41">9</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="40" pageNumber="41">
<paragraph blockId="40.[140,1108,1047,1636]" pageId="40" pageNumber="41">
<keyLead pageId="40" pageNumber="41">
2 Only last 3-8 metaterga in front of telson with a very evident mid-dorsal outgrowth. Body with 20 segments.............................................................
<emphasis bold="true" box="[1092,1107,1222,1248]" pageId="40" pageNumber="41">3</emphasis>
</keyLead>
</paragraph>
<paragraph blockId="40.[140,1108,1047,1636]" pageId="40" pageNumber="41">
<keyLead pageId="40" pageNumber="41">
– Most of metaterga, including some of anterior body portion, with a high, often tuberculated projection (
<figureCitation box="[592,660,1293,1319]" captionStart="Figure 9" captionStartId="12.[140,212,1558,1580]" captionTargetBox="[177,1070,165,1538]" captionTargetId="figure@12.[177,1070,165,1539]" captionTargetPageId="12" captionText="Figure 9. Eutrichodesmus aster sp. n., subadult paratype from Tham Han Cave; A, habitus, lateral view; B, anterior part of body, lateral view; C, midbody segments, lateral view; D, posterior part of body, lateral view; E, metatergal seta, sublateral view; F, anterior part of body, frontodorsal view. – Scale bars: A-D, F, 0.5 mm; E, 0.001 mm." figureDoi="http://doi.org/10.5281/zenodo.3769987" httpUri="https://zenodo.org/record/3769987/files/figure.png" pageId="40" pageNumber="41">Figs 9</figureCitation>
A-D; 20A-D). Body with 19 or 20 segments..........................................................................................................
<emphasis bold="true" box="[1092,1107,1328,1354]" pageId="40" pageNumber="41">5</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="40" pageNumber="41">
<paragraph blockId="40.[140,1108,1047,1636]" pageId="40" pageNumber="41">
<keyLead pageId="40" pageNumber="41">
3 Metaterga 12-19 each with an increasingly evident, subtriangular, mid-dorsal outgrowth. Cave in
<collectingRegion box="[472,626,1399,1425]" country="China" name="Yunnan" pageId="40" pageNumber="41">Yunnan Prov.</collectingRegion>
,
<collectingCountry box="[637,707,1399,1425]" name="China" pageId="40" pageNumber="41">China</collectingCountry>
..........................
<taxonomicName baseAuthorityName="Zhang in Zhang &amp; Wang" baseAuthorityYear="1993" box="[900,1108,1398,1425]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="dorsiangulatus">
<emphasis bold="true" box="[900,1108,1398,1425]" italics="true" pageId="40" pageNumber="41">E. dorsiangulatus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="40.[140,1108,1047,1636]" pageId="40" pageNumber="41">
<keyLead pageId="40" pageNumber="41">
– Only metaterga 16(17)-19 each with an evident, rather rounded, mid-dorsal outgrowth...................................................................................................
<emphasis bold="true" box="[1092,1107,1469,1495]" pageId="40" pageNumber="41">4</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="40" pageNumber="41">
<paragraph blockId="40.[140,1108,1047,1636]" pageId="40" pageNumber="41">
<keyLead pageId="40" pageNumber="41">
4 Paraterga narrower. Gonopod process dp less than one-quarter length of acropodite. North
<collectingCountry box="[409,507,1539,1565]" name="Vietnam" pageId="40" pageNumber="41">Vietnam</collectingCountry>
..................................................
<taxonomicName authorityName="Golovatch, Geoffroy, Mauries &amp; VandenSpiegel" authorityYear="2009" box="[881,1107,1539,1566]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="armatocaudatus">
<emphasis bold="true" box="[881,1107,1539,1566]" italics="true" pageId="40" pageNumber="41">E. armatocaudatus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="40.[140,1108,1047,1636]" pageId="40" pageNumber="41">
<keyLead pageId="40" pageNumber="41">
– Paraterga broader. Gonopod process dp much longer, more than half length of acropodite. Cave in
<collectingRegion box="[503,658,1610,1636]" country="China" name="Yunnan" pageId="40" pageNumber="41">Yunnan Prov.</collectingRegion>
,
<collectingCountry box="[669,739,1610,1636]" name="China" pageId="40" pageNumber="41">China</collectingCountry>
..........................
<taxonomicName baseAuthorityName="Zhang in Zhang &amp; Wang" baseAuthorityYear="1993" box="[931,1107,1609,1636]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="monodentus">
<emphasis bold="true" box="[931,1107,1609,1636]" italics="true" pageId="40" pageNumber="41">E. monodentus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="41" pageNumber="42">
<paragraph blockId="41.[140,1108,166,1636]" pageId="41" pageNumber="42">
<keyLead pageId="41" pageNumber="42">
5 Mid-dorsal projections relatively low, not club-shaped, increasingly evident on metaterga 3–17(18) (
<figureCitation box="[522,623,202,228]" captionStart="Figure 20" captionStartId="25.[140,212,1499,1521]" captionTargetBox="[139,1107,165,1479]" captionTargetId="figure@25.[139,1108,165,1479]" captionTargetPageId="25" captionText="Figure 20. Eutrichodesmus asteroides sp. n., ♀ paratype from Hang Cha Noi Cave; A, habitus, lateral view; B, E, anterior part of body, lateral and frontal views, respectively; C, F, midbody segments, lateral and dorsal views, respectively; D, posterior part of body, lateral view. – Scale bars: A-F, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3769989" httpUri="https://zenodo.org/record/3769989/files/figure.png" pageId="41" pageNumber="42">Fig. 20A</figureCitation>
) ..............................................................
<emphasis bold="true" box="[1092,1107,201,227]" pageId="41" pageNumber="42">6</emphasis>
</keyLead>
</paragraph>
<paragraph blockId="41.[140,1108,166,1636]" pageId="41" pageNumber="42">
<keyLead pageId="41" pageNumber="42">
– Mid-dorsal projections especially prominent, club-shaped, present on metaterga (4)5-19, only slightly lower on metatergum 19...........................
<emphasis bold="true" box="[1092,1107,272,298]" pageId="41" pageNumber="42">7</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="41" pageNumber="42">
<paragraph blockId="41.[140,1108,166,1636]" pageId="41" pageNumber="42">
<keyLead pageId="41" pageNumber="42">
6 Body with 19 segments. Subtriangular mid-dorsal projections evident on metaterga 3-17, only slightly smaller on metatergum 18 (
<figureCitation box="[906,1011,343,369]" captionStart="Figure 20" captionStartId="25.[140,212,1499,1521]" captionTargetBox="[139,1107,165,1479]" captionTargetId="figure@25.[139,1108,165,1479]" captionTargetPageId="25" captionText="Figure 20. Eutrichodesmus asteroides sp. n., ♀ paratype from Hang Cha Noi Cave; A, habitus, lateral view; B, E, anterior part of body, lateral and frontal views, respectively; C, F, midbody segments, lateral and dorsal views, respectively; D, posterior part of body, lateral view. – Scale bars: A-F, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3769989" httpUri="https://zenodo.org/record/3769989/files/figure.png" pageId="41" pageNumber="42">Fig. 20A</figureCitation>
). Gonopod process dp rudimentary (
<figureCitation box="[578,712,378,404]" captionStart="Figure 22" captionStartId="27.[140,212,1558,1580]" captionTargetBox="[139,1107,421,1538]" captionTargetId="figure@27.[139,1108,421,1539]" captionTargetPageId="27" captionText="Figure 22. Eutrichodesmus asteroides sp. n., ♀ (A-D) and ♁ (E-F) paratypes from Hang Cha Noi Cave; A, midbody metaterga, sublateral view; B, C, texture of tegument, dorsal view; D, setae on antennae; E, F, left gonopod, mesal and lateral views, respectively. – Scale bars: A, 0.05 mm; B, 0.1 mm; C, 0.01; D, 0.005 mm; E, F, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3769991" httpUri="https://zenodo.org/record/3769991/files/figure.png" pageId="41" pageNumber="42">Figs 22E, F</figureCitation>
).
<collectingCountry box="[732,830,378,404]" name="Vietnam" pageId="41" pageNumber="42">Vietnam</collectingCountry>
.......
<emphasis bold="true" box="[887,1108,377,404]" pageId="41" pageNumber="42">
<taxonomicName authority="Golovatch &amp; Geoffroy &amp; Mauriès &amp; VandenSpiegel, 2009" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès &amp; VandenSpiegel" authorityYear="2009" box="[887,1035,377,404]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="asteroides" status="sp. n.">
<emphasis bold="true" box="[887,1035,377,404]" italics="true" pageId="41" pageNumber="42">E. asteroides</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1043,1108,378,404]" pageId="41" pageNumber="42" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph blockId="41.[140,1108,166,1636]" pageId="41" pageNumber="42">
<keyLead pageId="41" pageNumber="42">
– Body with 20 segments. Clearly tridentate mid-dorsal projections evident on metaterga 3-18, abruptly smaller on metatergum 19. Gonopod process dp small but evident.
<collectingCountry box="[455,554,483,509]" name="Malaysia" pageId="41" pageNumber="42">Malaysia</collectingCountry>
....................................................
<taxonomicName baseAuthorityName="Sinclair" baseAuthorityYear="1901" box="[940,1107,483,510]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="cavernicola">
<emphasis bold="true" box="[940,1107,483,510]" italics="true" pageId="41" pageNumber="42">E. cavernicola</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="41" pageNumber="42">
<paragraph blockId="41.[140,1108,166,1636]" pageId="41" pageNumber="42">
<keyLead pageId="41" pageNumber="42">
7 Metatergum 4 with a high mid-dorsal projection (
<figureCitation box="[802,889,519,545]" captionStart="Figure 9" captionStartId="12.[140,212,1558,1580]" captionTargetBox="[177,1070,165,1538]" captionTargetId="figure@12.[177,1070,165,1539]" captionTargetPageId="12" captionText="Figure 9. Eutrichodesmus aster sp. n., subadult paratype from Tham Han Cave; A, habitus, lateral view; B, anterior part of body, lateral view; C, midbody segments, lateral view; D, posterior part of body, lateral view; E, metatergal seta, sublateral view; F, anterior part of body, frontodorsal view. – Scale bars: A-D, F, 0.5 mm; E, 0.001 mm." figureDoi="http://doi.org/10.5281/zenodo.3769987" httpUri="https://zenodo.org/record/3769987/files/figure.png" pageId="41" pageNumber="42">Fig. 9A</figureCitation>
). Paraterga strongly declivous and quadrilobate (
<figureCitation box="[564,633,554,580]" captionStart="Figure 9" captionStartId="12.[140,212,1558,1580]" captionTargetBox="[177,1070,165,1538]" captionTargetId="figure@12.[177,1070,165,1539]" captionTargetPageId="12" captionText="Figure 9. Eutrichodesmus aster sp. n., subadult paratype from Tham Han Cave; A, habitus, lateral view; B, anterior part of body, lateral view; C, midbody segments, lateral view; D, posterior part of body, lateral view; E, metatergal seta, sublateral view; F, anterior part of body, frontodorsal view. – Scale bars: A-D, F, 0.5 mm; E, 0.001 mm." figureDoi="http://doi.org/10.5281/zenodo.3769987" httpUri="https://zenodo.org/record/3769987/files/figure.png" pageId="41" pageNumber="42">Figs 9</figureCitation>
A-D, 10E). Gonopods as in
<figureCitation box="[956,1039,554,580]" captionStart="Figure 11" captionStartId="14.[140,212,1622,1644]" captionTargetBox="[194,1057,699,1596]" captionTargetId="figure@14.[186,1062,681,1603]" captionTargetPageId="14" captionText="Figure 11. Eutrichodesmus aster sp. n., ♁ paratype from Tham Han Cave; A, B, left gonopod, mesal and lateral views, respectively. – Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3813113" httpUri="https://zenodo.org/record/3813113/files/figure.png" pageId="41" pageNumber="42">Fig. 11</figureCitation>
.
<collectingCountry name="Vietnam" pageId="41" pageNumber="42">Vietnam</collectingCountry>
........................................................................................
<emphasis bold="true" box="[947,1108,589,615]" pageId="41" pageNumber="42">
<taxonomicName authority="Golovatch &amp; Geoffroy &amp; Mauriès &amp; VandenSpiegel, 2009" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès &amp; VandenSpiegel" authorityYear="2009" box="[947,1036,589,615]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="aster" status="sp. n.">
<emphasis bold="true" box="[947,1036,589,615]" italics="true" pageId="41" pageNumber="42">E. aster</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1043,1108,589,615]" pageId="41" pageNumber="42" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph blockId="41.[140,1108,166,1636]" pageId="41" pageNumber="42">
<keyLead pageId="41" pageNumber="42">
– Mid-dorsal projections absent from metatergum 4. Paraterga less strongly declivous. Gonopods different.
<collectingCountry box="[574,677,659,685]" name="Malaysia" pageId="41" pageNumber="42">Malaysia</collectingCountry>
.........................................................
<emphasis bold="true" box="[1092,1107,659,685]" pageId="41" pageNumber="42">8</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="41" pageNumber="42">
<paragraph blockId="41.[140,1108,166,1636]" pageId="41" pageNumber="42">
<keyLead pageId="41" pageNumber="42">
8 Mid-dorsal projections on metaterga 5 and 6 straight in lateral view............. .................................................................................................
<taxonomicName baseAuthorityName="Hoffman" baseAuthorityYear="1977" box="[960,1107,729,756]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="macclurei">
<emphasis bold="true" box="[960,1107,729,756]" italics="true" pageId="41" pageNumber="42">E. macclurei</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="41.[140,1108,166,1636]" pageId="41" pageNumber="42">
<keyLead pageId="41" pageNumber="42">
– Mid-dorsal projections on metaterga 5 and 6 slightly inclined anteriorly in lateral view................................................................................
<taxonomicName baseAuthorityName="Hoffman" baseAuthorityYear="1977" box="[959,1107,800,827]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="reclinatus">
<emphasis bold="true" box="[959,1107,800,827]" italics="true" pageId="41" pageNumber="42">E. reclinatus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="41" pageNumber="42">
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9 Body with 19 segments. Collum and metaterga virtually smooth, non-tuberculate, beset with long or short trichome (
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A-F, 33C-E). Paraterga too short to allow complete body volvation (
<figureCitation box="[698,801,906,932]" captionStart="Figure 32" captionStartId="38.[140,212,1590,1612]" captionTargetBox="[140,1108,481,1570]" captionTargetId="figure@38.[140,1108,480,1570]" captionTargetPageId="38" captionText="Figure 32. Eutrichodesmus reductus sp. n., ♀ paratype; A, habitus, lateral view; B, anterior part of body, lateral view; C, midbody segments, lateral view; D-F, posterior part of body, lateral, dorsal and caudal views, respectively. – Scale bars: A, 0.5 mm; B-F, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3770009" httpUri="https://zenodo.org/record/3770009/files/figure.png" pageId="41" pageNumber="42">Fig. 32A</figureCitation>
). Gonopod telopodite with a complex, lobiform dp (
<figureCitation box="[526,658,941,967]" captionStart="Figure 33" captionStartId="39.[140,212,1558,1580]" captionTargetBox="[139,1107,420,1538]" captionTargetId="figure@39.[139,1108,420,1539]" captionTargetPageId="39" captionText="Figure 33. Eutrichodesmus reductus sp. n., ♀ (A-E) &amp; ♁ (F-G) paratypes; A, head, ventral view; B, midbody segments, ventral view; C, midbody paratergum with ozopore, dorsal view; D, E, texture of tegument, dorsal view; F, G, left gonopod, mesal and lateral views, respectively. – Scale bars: A, B, 0.1 mm; C, 0.02 mm; D, 0.05 mm, E, 0.01 mm; F, G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3770011" httpUri="https://zenodo.org/record/3770011/files/figure.png" pageId="41" pageNumber="42">Fig. 33F, G</figureCitation>
). Southwestern Pacific......................
<emphasis bold="true" box="[1077,1107,941,967]" pageId="41" pageNumber="42">10</emphasis>
</keyLead>
</paragraph>
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– Body with 20 segments. Collum and metaterga clearly tuberculate, trichome (if present) usually short (
<figureCitation box="[549,650,1011,1038]" captionStart="Figure 25" captionStartId="30.[140,212,1507,1529]" captionTargetBox="[140,1108,165,1487]" captionTargetId="figure@30.[140,1109,165,1488]" captionTargetPageId="30" captionText="Figure 25. Eutrichodesmus griseus sp. n., ♀ paratype; A, D, posterior part of body, dorsal and ventral views, respectively; B, anterior part of body, ventral view; C, midbody segments, ventral view; E, crosssection of a midbody segment, caudal view; F, texture of tegument, dorsal view. – Scale bars: A-E, 0.1 mm; F, 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.3769997" httpUri="https://zenodo.org/record/3769997/files/figure.png" pageId="41" pageNumber="42">Figs 25F</figureCitation>
,
<figureCitation box="[666,715,1012,1038]" captionStart="Figure 29" captionStartId="35.[140,212,1494,1516]" captionTargetBox="[140,1108,165,1474]" captionTargetId="figure@35.[140,1108,165,1474]" captionTargetPageId="35" captionText="Figure 29. Eutrichodesmus multilobatus sp. n., ♀ paratype; A, midbody segments, ventral view; B, posterior part of body, ventral view; C, cross-section of a midbody segment, caudal view; D, part of head with left antenna, subdorsal view; E, texture of tegument, dorsal view; F, metatergal seta. – Scale bars: A, B, E, 0.1 mm; C, 0.2 mm; D, 0.05 mm; F, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.3770005" httpUri="https://zenodo.org/record/3770005/files/figure.png" pageId="41" pageNumber="42">29F</figureCitation>
). Paraterga much broader, mostly reaching level of venter, so volvation usually complete. East and Southeast Asia...........................................................................................................
<emphasis bold="true" box="[1077,1107,1081,1107]" pageId="41" pageNumber="42">11</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="41" pageNumber="42">
<paragraph blockId="41.[140,1108,166,1636]" pageId="41" pageNumber="42">
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10 Collum and metaterga beset with long, 2-segmented, tactile setae. Tip of gonopod telopodite without teeth.
<collectingCountry box="[615,710,1153,1179]" name="Vanuatu" pageId="41" pageNumber="42">Vanuatu</collectingCountry>
..........................
<taxonomicName authorityName="Golovatch, Geoffroy, Mauries &amp; VandenSpiegel" authorityYear="2009" box="[913,1107,1152,1179]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="communicans">
<emphasis bold="true" box="[913,1107,1152,1179]" italics="true" pageId="41" pageNumber="42">E. communicans</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="41.[140,1108,166,1636]" pageId="41" pageNumber="42">
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– Collum and metaterga beset with very short trichome (
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B-E, 33C-E). Tip of gonopod telopodite with a few teeth (
<figureCitation box="[750,886,1223,1249]" captionStart="Figure 33" captionStartId="39.[140,212,1558,1580]" captionTargetBox="[139,1107,420,1538]" captionTargetId="figure@39.[139,1108,420,1539]" captionTargetPageId="39" captionText="Figure 33. Eutrichodesmus reductus sp. n., ♀ (A-E) &amp; ♁ (F-G) paratypes; A, head, ventral view; B, midbody segments, ventral view; C, midbody paratergum with ozopore, dorsal view; D, E, texture of tegument, dorsal view; F, G, left gonopod, mesal and lateral views, respectively. – Scale bars: A, B, 0.1 mm; C, 0.02 mm; D, 0.05 mm, E, 0.01 mm; F, G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3770011" httpUri="https://zenodo.org/record/3770011/files/figure.png" pageId="41" pageNumber="42">Figs 33F, G</figureCitation>
).
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,
<collectingCountry name="Indonesia" pageId="41" pageNumber="42">Indonesia</collectingCountry>
.....................................................................................
<emphasis bold="true" box="[904,1108,1257,1284]" pageId="41" pageNumber="42">
<taxonomicName authority="Golovatch &amp; Geoffroy &amp; Mauriès &amp; VandenSpiegel, 2009" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès &amp; VandenSpiegel" authorityYear="2009" box="[904,1036,1257,1284]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="reductus" status="sp. n.">
<emphasis bold="true" box="[904,1036,1257,1284]" italics="true" pageId="41" pageNumber="42">E. reductus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1043,1108,1258,1284]" pageId="41" pageNumber="42" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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11 Paraterga mostly especially wide and only slightly declivous. Caves in
<collectingRegion country="China" name="Guangxi" pageId="41" pageNumber="42">Guangxi Province</collectingRegion>
,
<collectingCountry box="[358,428,1328,1354]" name="China" pageId="41" pageNumber="42">China</collectingCountry>
........................................................................................
<emphasis bold="true" box="[1077,1107,1328,1354]" pageId="41" pageNumber="42">12</emphasis>
</keyLead>
</paragraph>
<paragraph blockId="41.[140,1108,166,1636]" box="[140,1107,1363,1390]" pageId="41" pageNumber="42">
<keyLead box="[140,1107,1363,1390]" pageId="41" pageNumber="42">
– Paraterga not so wide, strongly declivous (
<figureCitation box="[718,808,1363,1390]" captionStart="Figure 2" captionStartId="4.[140,213,1558,1580]" captionTargetBox="[140,1108,502,1538]" captionTargetId="figure@4.[140,1108,502,1539]" captionTargetPageId="4" captionText="Figure 2. Eutrichodesmus distinctus sp. n., ♁ paratype; A, habitus, lateral view; B, E, anterior part of body, lateral and dorsal views, respectively; C, F, midbody segments, lateral and dorsal views, respectively; D, G, posterior part of body, lateral and dorsal views, respectively. – Scale bars: A, 0.5 mm; B-G, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3769973" httpUri="https://zenodo.org/record/3769973/files/figure.png" pageId="41" pageNumber="42">Figs 2A</figureCitation>
,
<figureCitation box="[819,854,1363,1390]" captionStart="Figure 5" captionStartId="8.[140,212,1498,1520]" captionTargetBox="[140,1108,165,1478]" captionTargetId="figure@8.[140,1108,165,1478]" captionTargetPageId="8" captionText="Figure 5. Eutrichodesmus regularis sp. n., ♁ paratype; A, habitus, lateral view; B, E, anterior part of body, frontodorsal and ventral views, respectively; C, midbody segments, dorsal view; D, F, posterior part of body, dorsal and ventral views, respectively. – Scale bars: A, C, 0.5 mm; B, D-F, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3769979" httpUri="https://zenodo.org/record/3769979/files/figure.png" pageId="41" pageNumber="42">5A</figureCitation>
, 13A, 17A, 28A) ...
<emphasis bold="true" box="[1077,1107,1363,1389]" pageId="41" pageNumber="42">13</emphasis>
</keyLead>
</paragraph>
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<paragraph blockId="41.[140,1108,166,1636]" pageId="41" pageNumber="42">
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12 Collum with a row of conspicuous teeth along fore edge of collum............... ......................................................................................................
<taxonomicName authorityName="Golovatch, Geoffroy, Mauries &amp; VandenSpiegel" authorityYear="2009" box="[999,1108,1433,1460]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="similis">
<emphasis bold="true" box="[999,1108,1433,1460]" italics="true" pageId="41" pageNumber="42">E. similis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="41.[140,1108,166,1636]" box="[140,1108,1468,1496]" pageId="41" pageNumber="42">
<keyLead box="[140,1108,1468,1496]" pageId="41" pageNumber="42">
– Collum with only minute teeth at fore edge of collum......................
<taxonomicName authorityName="Golovatch, Geoffroy, Mauries &amp; VandenSpiegel" authorityYear="2009" box="[1018,1108,1468,1495]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="latus">
<emphasis bold="true" box="[1018,1108,1468,1495]" italics="true" pageId="41" pageNumber="42">E. latus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep lastPageId="42" lastPageNumber="43" pageId="41" pageNumber="42">
<paragraph blockId="41.[140,1108,166,1636]" pageId="41" pageNumber="42">
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13 Most metaterga with two transverse rows of bosses. Epiproct very strongly flattened dorsoventrally, subquadrate, spatuliform, with unincised margins. Gonopod telopodite particularly slender, about twice as long as coxa. Shikoku,
<collectingCountry box="[354,419,1610,1636]" name="Japan" pageId="41" pageNumber="42">Japan</collectingCountry>
..........................................................................
<taxonomicName baseAuthorityName="Murakami" baseAuthorityYear="1966" box="[960,1108,1609,1636]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="peculiaris">
<emphasis bold="true" box="[960,1108,1609,1636]" italics="true" pageId="41" pageNumber="42">E. peculiaris</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
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<keyLead pageId="42" pageNumber="43">
– Most metaterga with three transverse rows of bosses or conical tubercles. Epiproct never so strongly flattened. Gonopod telopodite usually shorter. Southeast Asia....................................................................................................
<emphasis bold="true" box="[1077,1107,237,263]" pageId="42" pageNumber="43">14</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="42" pageNumber="43">
<paragraph blockId="42.[140,1108,167,1636]" pageId="42" pageNumber="43">
<keyLead pageId="42" pageNumber="43">
14 Paraterga mostly set off laterally at base by a clear sulcus. Metaterga with three mixostictic rows of evident, conical tubercles, each surmounted by a long, 2-segmented seta. Gonopod telopodite with a particularly small, dentiform dp.
<collectingCountry box="[290,390,378,404]" name="Thailand" pageId="42" pageNumber="43">Thailand</collectingCountry>
..............................................................................
<taxonomicName baseAuthorityName="Hoffman" baseAuthorityYear="1982" box="[967,1107,377,404]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="gremialis">
<emphasis bold="true" box="[967,1107,377,404]" italics="true" pageId="42" pageNumber="43">E. gremialis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="42.[140,1108,167,1636]" pageId="42" pageNumber="43">
<keyLead pageId="42" pageNumber="43">
– Paraterga less markedly (if at all) set off laterally at base (
<figureCitation box="[898,990,413,439]" captionStart="Figure 2" captionStartId="4.[140,213,1558,1580]" captionTargetBox="[140,1108,502,1538]" captionTargetId="figure@4.[140,1108,502,1539]" captionTargetPageId="4" captionText="Figure 2. Eutrichodesmus distinctus sp. n., ♁ paratype; A, habitus, lateral view; B, E, anterior part of body, lateral and dorsal views, respectively; C, F, midbody segments, lateral and dorsal views, respectively; D, G, posterior part of body, lateral and dorsal views, respectively. – Scale bars: A, 0.5 mm; B-G, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3769973" httpUri="https://zenodo.org/record/3769973/files/figure.png" pageId="42" pageNumber="43">Figs 2A</figureCitation>
,
<figureCitation box="[1003,1038,413,440]" captionStart="Figure 5" captionStartId="8.[140,212,1498,1520]" captionTargetBox="[140,1108,165,1478]" captionTargetId="figure@8.[140,1108,165,1478]" captionTargetPageId="8" captionText="Figure 5. Eutrichodesmus regularis sp. n., ♁ paratype; A, habitus, lateral view; B, E, anterior part of body, frontodorsal and ventral views, respectively; C, midbody segments, dorsal view; D, F, posterior part of body, dorsal and ventral views, respectively. – Scale bars: A, C, 0.5 mm; B, D-F, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3769979" httpUri="https://zenodo.org/record/3769979/files/figure.png" pageId="42" pageNumber="43">5A</figureCitation>
, 13A, 17A, 28A), largely continuing general outline of metaterga or nearly so (Figs 18D, 29C). Metatergal trichome very short or missing. Gonopod telopodite different....................................................................................................
<emphasis bold="true" box="[1077,1107,518,544]" pageId="42" pageNumber="43">15</emphasis>
</keyLead>
</paragraph>
</keyStep>
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15 Paraterga relatively short, volvation apparently imperfect (
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). Body small, up to
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long.
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..........................................................
<emphasis bold="true" box="[1077,1107,589,615]" pageId="42" pageNumber="43">16</emphasis>
</keyLead>
</paragraph>
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– Paraterga broad enough to allow complete volvation (Figs 17A, 28A). Body usually larger.............................................................................................
<emphasis bold="true" box="[1077,1107,659,685]" pageId="42" pageNumber="43">17</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="42" pageNumber="43">
<paragraph blockId="42.[140,1108,167,1636]" box="[140,1108,694,721]" pageId="42" pageNumber="43">
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16 Entire body pallid, up to 5.0 mm long. Gonopods simple .............
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<emphasis bold="true" box="[995,1108,694,721]" italics="true" pageId="42" pageNumber="43">E. basalis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
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<keyLead pageId="42" pageNumber="43">
– Vertex, collum, metaterga and much of telson grey, this strongly contrasting with pallid venter and legs (
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). Body 6.0-
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long. Gonopods more elaborate (
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C-E).........................................................
<emphasis bold="true" box="[924,1108,800,827]" pageId="42" pageNumber="43">
<taxonomicName authority="Golovatch &amp; Geoffroy &amp; Mauriès &amp; VandenSpiegel, 2009" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès &amp; VandenSpiegel" authorityYear="2009" box="[924,1035,800,827]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="griseus" status="sp. n.">
<emphasis bold="true" box="[924,1035,800,827]" italics="true" pageId="42" pageNumber="43">E. griseus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1043,1108,800,826]" pageId="42" pageNumber="43" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
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</paragraph>
</keyStep>
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17 Pattern of metatergal tuberculation isostictic (
<figureCitation box="[780,851,835,862]" captionStart="Figure 5" captionStartId="8.[140,212,1498,1520]" captionTargetBox="[140,1108,165,1478]" captionTargetId="figure@8.[140,1108,165,1478]" captionTargetPageId="8" captionText="Figure 5. Eutrichodesmus regularis sp. n., ♁ paratype; A, habitus, lateral view; B, E, anterior part of body, frontodorsal and ventral views, respectively; C, midbody segments, dorsal view; D, F, posterior part of body, dorsal and ventral views, respectively. – Scale bars: A, C, 0.5 mm; B, D-F, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3769979" httpUri="https://zenodo.org/record/3769979/files/figure.png" pageId="42" pageNumber="43">Figs 5</figureCitation>
A-D). Seminal groove terminating on a very evident, pilose-spiculate pulvillus near base of a relatively shortened, lamellar acropodite (
<figureCitation box="[670,789,906,932]" captionStart="Figure 6" captionStartId="9.[140,212,1558,1580]" captionTargetBox="[140,1108,478,1538]" captionTargetId="figure@9.[140,1108,478,1538]" captionTargetPageId="9" captionText="Figure 6. Eutrichodesmus regularis sp. n., ♁ paratype; A, texture of tegument, dorsal view; B, metatergal tubercle, dorsal view; C, metatergal seta, dorsal view; D, cross-section of a midbody segment, caudal view; E, midbody legs, caudal view; F, segments 7 and 8, ventral view; G, left gonopod in situ, caudoventral view. – Scale bars: A, B, 0.05 mm; C, 0.01 mm; D, F, 0.2 mm; E, G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3769981" httpUri="https://zenodo.org/record/3769981/files/figure.png" pageId="42" pageNumber="43">Figs 6F, G</figureCitation>
,
<figureCitation box="[802,815,906,932]" captionStart="Figure 7" captionStartId="10.[140,212,1622,1644]" captionTargetBox="[140,1108,308,1602]" captionTargetId="figure@10.[140,1108,308,1603]" captionTargetPageId="10" captionText="Figure 7. Eutrichodesmus regularis sp. n., ♁ paratypes; A-F, gonopods, frontoventral, submesal, mesal, mesal, mesal and lateral views, respectively. – Scale bars: A-C, 0.1 mm; D, 0.05 mm; E, F, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3769983" httpUri="https://zenodo.org/record/3769983/files/figure.png" pageId="42" pageNumber="43">7</figureCitation>
A-F)....
<emphasis bold="true" box="[900,1108,905,932]" pageId="42" pageNumber="43">
<taxonomicName authority="Golovatch &amp; Geoffroy &amp; Mauriès &amp; VandenSpiegel, 2009" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès &amp; VandenSpiegel" authorityYear="2009" box="[900,1036,905,932]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="regularis" status="sp. n.">
<emphasis bold="true" box="[900,1036,905,932]" italics="true" pageId="42" pageNumber="43">E. regularis</emphasis>
</taxonomicName>
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</emphasis>
</keyLead>
</paragraph>
<paragraph blockId="42.[140,1108,167,1636]" pageId="42" pageNumber="43">
<keyLead pageId="42" pageNumber="43">
– Metatergal tuberculation mixostictic (Figs 13A-F, 24A). Seminal groove terminating distally to subapically, mostly without a pulvillus (seldom with a hairpad) (Figs 18G, H, 30) .......................................................................
<emphasis bold="true" box="[1077,1107,1011,1037]" pageId="42" pageNumber="43">18</emphasis>
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</paragraph>
</keyStep>
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<paragraph blockId="42.[140,1108,167,1636]" box="[140,1107,1046,1073]" pageId="42" pageNumber="43">
<keyLead box="[140,1107,1046,1073]" pageId="42" pageNumber="43">
18 Distofemoral process of gonotelopodite bare, subunciform, pointed.........
<emphasis bold="true" box="[1077,1107,1046,1072]" pageId="42" pageNumber="43">19</emphasis>
</keyLead>
</paragraph>
<paragraph blockId="42.[140,1108,167,1636]" pageId="42" pageNumber="43">
<keyLead pageId="42" pageNumber="43">
– Distofemoral process of gonotelopodite nearly always conspicuously papillate (Figs 18G, H, 30), seldom redumentary (Figs 15A-F), never pointed .......
<emphasis bold="true" box="[1077,1107,1117,1143]" pageId="42" pageNumber="43">20</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="42" pageNumber="43">
<paragraph blockId="42.[140,1108,167,1636]" box="[140,1108,1152,1179]" pageId="42" pageNumber="43">
<keyLead box="[140,1108,1152,1179]" pageId="42" pageNumber="43">
19 Body ca 8.0 mm long. Gonopod tip bifid.
<collectingCountry box="[727,825,1152,1178]" name="Vietnam" pageId="42" pageNumber="43">Vietnam</collectingCountry>
..................
<taxonomicName authorityName="Silvestri" authorityYear="1910" box="[962,1108,1152,1179]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="demangei">
<emphasis bold="true" box="[962,1108,1152,1179]" italics="true" pageId="42" pageNumber="43">E. demangei</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="42.[140,1108,167,1636]" pageId="42" pageNumber="43">
<keyLead pageId="42" pageNumber="43">
– Body ca 14.0 mm long. Gonopod tip simple, unciform. Cave in
<collectingRegion country="China" name="Yunnan" pageId="42" pageNumber="43">Yunnan Prov.</collectingRegion>
,
<collectingCountry box="[317,387,1223,1249]" name="China" pageId="42" pageNumber="43">China</collectingCountry>
............................................................................
<taxonomicName authorityName="Zhang in Zhang &amp; Wang" authorityYear="1993" box="[943,1107,1222,1249]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="arcicollaris">
<emphasis bold="true" box="[943,1107,1222,1249]" italics="true" pageId="42" pageNumber="43">E. arcicollaris</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="42" pageNumber="43">
<paragraph blockId="42.[140,1108,167,1636]" pageId="42" pageNumber="43">
<keyLead pageId="42" pageNumber="43">
20 Paraterga mostly bilobate laterally and unilobate caudolaterally (Fig. 17A). Very small:
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long and
<quantity box="[640,792,1293,1320]" metricMagnitude="-4" metricUnit="m" metricValue="6.75" metricValueMax="7.0" metricValueMin="6.5" pageId="42" pageNumber="43" unit="mm" value="0.675" valueMax="0.7" valueMin="0.65">0.65-0.7 mm</quantity>
wide. Gonopods as in Figs 18G, H.
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.........................................................
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<taxonomicName authority="Golovatch &amp; Geoffroy &amp; Mauriès &amp; VandenSpiegel, 2009" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès &amp; VandenSpiegel" authorityYear="2009" box="[875,1035,1328,1355]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="curticornis" status="sp. n.">
<emphasis bold="true" box="[875,1035,1328,1355]" italics="true" pageId="42" pageNumber="43">E. curticornis</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1043,1108,1328,1354]" pageId="42" pageNumber="43" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
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<keyLead pageId="42" pageNumber="43">
– Paraterga mostly 3- or 4-lobate laterally (Figs 13A-D). Length ≥ 6.0 mm, width ≥
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. Gonopods different .......................................................
<emphasis bold="true" box="[1077,1107,1398,1424]" pageId="42" pageNumber="43">21</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="42" pageNumber="43">
<paragraph blockId="42.[140,1108,167,1636]" pageId="42" pageNumber="43">
<keyLead pageId="42" pageNumber="43">
21 Metaterga 18 and 19 not elevated dorsally. Caudolateral lobulations mostly deeply incised. Distofemoral process of gonopod bipartite. Caves in
<collectingRegion country="China" name="Guizhou" pageId="42" pageNumber="43">Guizhou Prov.</collectingRegion>
,
<collectingCountry box="[317,387,1504,1530]" name="China" pageId="42" pageNumber="43">China</collectingCountry>
...................................................................................
<taxonomicName authorityName="Golovatch, Geoffroy, Mauries &amp; VandenSpiegel" authorityYear="2009" box="[995,1107,1504,1531]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="incisus">
<emphasis bold="true" box="[995,1107,1504,1531]" italics="true" pageId="42" pageNumber="43">E. incisus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="42.[140,1108,167,1636]" pageId="42" pageNumber="43">
<keyLead pageId="42" pageNumber="43">
– Metaterga 18 and 19 slightly elevated dorsally (
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A-F). Caudolateral lobulations not incised, tuberculiform. Distofemoral process of gonopod unipartite (
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).........................................................................................
<emphasis bold="true" box="[1077,1107,1609,1635]" pageId="42" pageNumber="43">22</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="43" pageNumber="44">
<paragraph blockId="43.[140,1108,166,651]" pageId="43" pageNumber="44">
<keyLead pageId="43" pageNumber="44">
22 Body 6.0-
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long and
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wide. Ozopores opening on top of porosteles representing penultimate of five lateral lobulations of respective paraterga, with caudolateral lobulations missing. Gonocoxae with only few macrosetae. Gonopods as in
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.
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..................
<emphasis bold="true" box="[852,1108,272,299]" pageId="43" pageNumber="44">
<taxonomicName authority="Golovatch &amp; Geoffroy &amp; Mauriès &amp; VandenSpiegel, 2009" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès &amp; VandenSpiegel" authorityYear="2009" box="[852,1035,272,299]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="multilobatus" status="sp. n.">
<emphasis bold="true" box="[852,1035,272,299]" italics="true" pageId="43" pageNumber="44">E. multilobatus</emphasis>
</taxonomicName>
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</emphasis>
</keyLead>
</paragraph>
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– Body ≥ 8.0 mm long and
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wide. Ozopores opening dorsally on caudalmost of 3-4 lateral lobulations of respective paraterga, caudolateral lobulations present. Gonocoxae with abundant macrosetae. Gonopods different.... .................................................................................................................
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</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="43" pageNumber="44">
<paragraph blockId="43.[140,1108,166,651]" pageId="43" pageNumber="44">
<keyLead pageId="43" pageNumber="44">
23 Body 8.0-
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long and
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wide. Paraterga mostly 3-lobate (
<figureCitation box="[253,321,483,510]" captionStart="Figure 2" captionStartId="4.[140,213,1558,1580]" captionTargetBox="[140,1108,502,1538]" captionTargetId="figure@4.[140,1108,502,1539]" captionTargetPageId="4" captionText="Figure 2. Eutrichodesmus distinctus sp. n., ♁ paratype; A, habitus, lateral view; B, E, anterior part of body, lateral and dorsal views, respectively; C, F, midbody segments, lateral and dorsal views, respectively; D, G, posterior part of body, lateral and dorsal views, respectively. – Scale bars: A, 0.5 mm; B-G, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3769973" httpUri="https://zenodo.org/record/3769973/files/figure.png" pageId="43" pageNumber="44">Figs 2</figureCitation>
A-G). Gonopods as in
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E-H. Cave in
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,
<collectingCountry box="[983,1055,483,509]" name="China" pageId="43" pageNumber="44">China</collectingCountry>
....... .......................................................................................
<emphasis bold="true" box="[889,1108,518,545]" pageId="43" pageNumber="44">
<taxonomicName authority="Golovatch &amp; Geoffroy &amp; Mauriès &amp; VandenSpiegel, 2009" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès &amp; VandenSpiegel" authorityYear="2009" box="[889,1036,518,545]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="distinctus" status="sp. n.">
<emphasis bold="true" box="[889,1036,518,545]" italics="true" pageId="43" pageNumber="44">E. distinctus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1043,1108,519,545]" pageId="43" pageNumber="44" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph blockId="43.[140,1108,166,651]" pageId="43" pageNumber="44">
<keyLead pageId="43" pageNumber="44">
– Body
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long and
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wide. Paraterga mostly 4-lobate (Figs 13A-D). Gonopods as in Figs 15A-F. Caves in
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,
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... ........................................................................................
<emphasis bold="true" box="[895,1108,624,651]" pageId="43" pageNumber="44">
<taxonomicName authority="Golovatch &amp; Geoffroy &amp; Mauriès &amp; VandenSpiegel, 2009" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès &amp; VandenSpiegel" authorityYear="2009" box="[895,1036,624,651]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="filisetiger" status="sp. n.">
<emphasis bold="true" box="[895,1036,624,651]" italics="true" pageId="43" pageNumber="44">E. filisetiger</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1043,1108,624,650]" pageId="43" pageNumber="44" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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