269 lines
31 KiB
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269 lines
31 KiB
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<document ID-DOI="http://doi.org/10.5281/zenodo.6214791" ID-GBIF-Dataset="3026c846-a607-4826-b2a9-8f7a531baec5" ID-GBIF-Taxon="163971737" ID-ISSN="1313–2970" ID-Zenodo-Dep="6214791" ID-ZooBank="B32EF8D5-C3C6-4C8C-8845-284125448425" LSID="urn:lsid:zoobank.org:act:1726A727-8FC2-4BF8-807D-8DFC1C54380C" approvalRequired="25" approvalRequired_for_textStreams="25" checkinTime="1587995767728" checkinUser="plazi" docAuthor="Golovatch, Sergei, Geoffroy, Jean-Jacques, Mauriès, Jean-Paul & VandenSpiegel, Didier" docDate="2009" docId="936787D7FF97FF938ABB405BFCAAFEF8" docLanguage="en" docName="ZK_article_1994.pdf" docOrigin="ZooKeys 12 (12)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@52c7884c" docStyleName="zookeys.2008.journal_article" docTitle="Eutrichodesmus reductus Golovatch, Geoffroy, Mauries & VandenSpiegel 2009, sp. n." docType="treatment" docUuid="1726A727-8FC2-4BF8-807D-8DFC1C54380C" docUuidSource="ZooBank" docVersion="5" lastPageId="39" lastPageNumber="40" masterDocId="6F5EFFAFFFB3FFB48A374557FFC6FF89" masterDocTitle="Review of the millipede genus Eutrichodesmus Silvestri, 1910 (Diplopoda, Polydesmida, Haplodesmidae), with descriptions of new species" masterLastPageNumber="46" masterPageNumber="1" pageId="36" pageNumber="37" updateTime="1645494461179" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Review of the millipede genus Eutrichodesmus Silvestri, 1910 (Diplopoda, Polydesmida, Haplodesmidae), with descriptions of new species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Golovatch, Sergei</mods:namePart>
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<mods:affiliation>Russian Academy of Sciences, Moscow, Russia</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Geoffroy, Jean-Jacques</mods:namePart>
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<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
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<mods:affiliation>Muséum national d’Histoire naturelle, Département Ecologie & Gestion de la Biodiversité, UMR 7179 du CNRS, Equipe EVOLTRAIT, Brunoy, France & D 362 Dc 0 - & CC</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Mauriès, Jean-Paul</mods:namePart>
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<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
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<mods:affiliation>Muséum national d’Histoire naturelle, Département Systématique et Evolution, Section Arthropodes, Paris, France & - & - 42 F & D 01 - 28 Ad- 43 F</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>VandenSpiegel, Didier</mods:namePart>
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<mods:affiliation>Musée Royal de l’Afrique centrale, Tervuren, Belgium & A & EE & F- 04802 E 68 Cb</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2009</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2009-06-18</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>12</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>12</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>46</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.3897/zookeys.12.167</mods:identifier>
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<mods:identifier type="GBIF-Dataset">3026c846-a607-4826-b2a9-8f7a531baec5</mods:identifier>
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<mods:identifier type="ISSN">1313–2970</mods:identifier>
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<mods:identifier type="Zenodo-Dep">576460</mods:identifier>
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<mods:identifier type="ZooBank">B32EF8D5-C3C6-4C8C-8845-284125448425</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6214791" ID-GBIF-Taxon="163971737" ID-Zenodo-Dep="6214791" LSID="urn:lsid:zoobank.org:act:1726A727-8FC2-4BF8-807D-8DFC1C54380C" httpUri="http://treatment.plazi.org/id/936787D7FF97FF938ABB405BFCAAFEF8" lastPageId="39" lastPageNumber="40" pageId="36" pageNumber="37">
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<subSubSection box="[140,1086,1292,1319]" pageId="36" pageNumber="37" type="nomenclature">
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<paragraph blockId="36.[140,1086,1292,1390]" box="[140,1086,1292,1319]" pageId="36" pageNumber="37">
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<heading box="[140,1086,1292,1319]" level="2" pageId="36" pageNumber="37">
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<emphasis bold="true" box="[140,1086,1292,1319]" pageId="36" pageNumber="37">
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<taxonomicName authority="Golovatch, Geoffroy, Mauries & VandenSpiegel" authorityName="Golovatch, Geoffroy, Mauries & VandenSpiegel" authorityYear="2009" box="[140,1009,1292,1319]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="reductus" status="sp. n.">
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<emphasis bold="true" box="[140,434,1292,1319]" italics="true" pageId="36" pageNumber="37">
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<emphasis box="[140,328,1292,1319]" inLineHeading="true" pageId="36" pageNumber="37" reason="1">Eutrichodesmus</emphasis>
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reductus
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</emphasis>
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Golovatch, Geoffroy, Mauriès & VandenSpiegel
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</taxonomicName>
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,
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<taxonomicNameLabel box="[1021,1086,1293,1319]" pageId="36" pageNumber="37" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="36" pageNumber="37" type="description">
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<paragraph blockId="36.[140,1086,1292,1390]" box="[140,977,1328,1355]" pageId="36" pageNumber="37">
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<uri box="[140,977,1328,1355]" pageId="36" pageNumber="37">
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urn:lsid:zoobank.org:act:
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<uuid box="[425,977,1328,1355]" pageId="36" pageNumber="37">1726A727-8FC2-4BF8-807D-8DFC1C54380C</uuid>
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</uri>
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</paragraph>
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<paragraph blockId="36.[140,1086,1292,1390]" box="[140,260,1363,1390]" pageId="36" pageNumber="37">
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Figs
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<figureCitation box="[192,260,1364,1390]" captionStart-0="Figure 31" captionStart-1="Figure 32" captionStart-2="Figure 33" captionStartId-0="37.[140,212,902,924]" captionStartId-1="38.[140,212,1590,1612]" captionStartId-2="39.[140,212,1558,1580]" captionTargetBox-0="[140,1108,165,882]" captionTargetBox-1="[140,1108,481,1570]" captionTargetBox-2="[139,1107,420,1538]" captionTargetId-0="figure@37.[140,1108,165,883]" captionTargetId-1="figure@38.[140,1108,480,1570]" captionTargetId-2="figure@39.[139,1108,420,1539]" captionTargetPageId-0="37" captionTargetPageId-1="38" captionTargetPageId-2="39" captionText-0="Figure 31. Eutrichodesmus reductus sp. n., ♁ and ♀ paratypes; habitus, lateral view. (Photographed not to scale by L. Deharveng)." captionText-1="Figure 32. Eutrichodesmus reductus sp. n., ♀ paratype; A, habitus, lateral view; B, anterior part of body, lateral view; C, midbody segments, lateral view; D-F, posterior part of body, lateral, dorsal and caudal views, respectively. – Scale bars: A, 0.5 mm; B-F, 0.1 mm." captionText-2="Figure 33. Eutrichodesmus reductus sp. n., ♀ (A-E) & ♁ (F-G) paratypes; A, head, ventral view; B, midbody segments, ventral view; C, midbody paratergum with ozopore, dorsal view; D, E, texture of tegument, dorsal view; F, G, left gonopod, mesal and lateral views, respectively. – Scale bars: A, B, 0.1 mm; C, 0.02 mm; D, 0.05 mm, E, 0.01 mm; F, G, 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3813134" figureDoi-1="http://doi.org/10.5281/zenodo.3770009" figureDoi-2="http://doi.org/10.5281/zenodo.3770011" httpUri-0="https://zenodo.org/record/3813134/files/figure.png" httpUri-1="https://zenodo.org/record/3770009/files/figure.png" httpUri-2="https://zenodo.org/record/3770011/files/figure.png" pageId="36" pageNumber="37">31-33</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="36" pageNumber="37" type="materials_examined">
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<paragraph blockId="36.[140,1108,1433,1672]" pageId="36" pageNumber="37">
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<emphasis bold="true" box="[140,312,1433,1460]" pageId="36" pageNumber="37">
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<emphasis box="[140,198,1434,1460]" inLineHeading="true" pageId="36" pageNumber="37" reason="1">Type</emphasis>
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material:
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</emphasis>
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<materialsCitation ID-GBIF-Occurrence="2610648312" collectingDate="1990-08-18" collectorName="A. Bedos & L. Deharveng" country="Indonesia" county="Maros" location="Gua Saripa Cave" municipality="Samanggi" pageId="36" pageNumber="37" specimenCount="1" specimenCount-female="1" stateProvince="Sulawesi Selatan" typeStatus="holotype">
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<collectingCountry box="[319,432,1434,1460]" name="Indonesia" pageId="36" pageNumber="37">Indonesia</collectingCountry>
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,
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<collectingRegion box="[444,631,1434,1460]" country="Indonesia" name="Sulawesi Selatan" pageId="36" pageNumber="37">Sulawesi Selatan</collectingRegion>
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, kab.
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<collectingCounty box="[699,774,1434,1460]" pageId="36" pageNumber="37">Maros</collectingCounty>
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:
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<collectingMunicipality box="[785,898,1434,1460]" pageId="36" pageNumber="37">Samanggi</collectingMunicipality>
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,
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<location LSID="urn:lsid:plazi:treatment:936787D7FF97FF938ABB405BFCAAFEF8:1E11601AFF97FF9089B940CDFB89FA3D" box="[910,1103,1434,1460]" country="Indonesia" county="Maros" municipality="Samanggi" name="Gua Saripa Cave" pageId="36" pageNumber="37" stateProvince="Sulawesi Selatan">Gua Saripa Cave</location>
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, hand collected,
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<date box="[319,474,1469,1495]" pageId="36" pageNumber="37" value="1990-08-18">
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<collectingDate box="[319,474,1469,1495]" pageId="36" pageNumber="37" value="1990-08-18">18.VIII.1990</collectingDate>
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</date>
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, leg.
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<collectorName box="[533,634,1469,1495]" pageId="36" pageNumber="37">A. Bedos</collectorName>
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and
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<collectorName box="[691,847,1469,1496]" pageId="36" pageNumber="37">L. Deharveng</collectorName>
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(SULS-214),
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<typeStatus box="[1008,1107,1469,1495]" pageId="36" pageNumber="37">holotype</typeStatus>
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♁ (MZB),
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<typeStatus box="[266,376,1505,1531]" pageId="36" pageNumber="37">paratypes</typeStatus>
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: 1 ♁ (MNHN JC 325),
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<specimenCount box="[657,694,1504,1531]" pageId="36" pageNumber="37" type="female">1 ♀</specimenCount>
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(SEM)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="36" pageNumber="37" type="etymology">
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<paragraph blockId="36.[140,1108,1433,1672]" pageId="36" pageNumber="37">
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<emphasis bold="true" box="[185,263,1539,1565]" pageId="36" pageNumber="37">
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<emphasis box="[185,258,1539,1565]" inLineHeading="true" pageId="36" pageNumber="37" reason="1">Name</emphasis>
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:
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</emphasis>
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To emphasize the strongly underdeveloped paraterga 2 and lack of metatergal tuberculation.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37" type="diagnosis">
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<paragraph blockId="36.[140,1108,1433,1672]" pageId="36" pageNumber="37">
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<emphasis bold="true" box="[185,312,1609,1635]" pageId="36" pageNumber="37">
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<emphasis box="[185,307,1609,1635]" inLineHeading="true" pageId="36" pageNumber="37" reason="1">Diagnosis</emphasis>
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:
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</emphasis>
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Differs from congeners except
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<taxonomicName authority="Golovatch, Geoffroy, Mauries & VandenSpiegel, 2009" authorityName="Golovatch, Geoffroy, Mauries & VandenSpiegel" authorityYear="2009" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="communicans">
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<emphasis box="[677,858,1611,1636]" italics="true" pageId="36" pageNumber="37">E. communicans</emphasis>
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<bibRefCitation author="Golovatch SI & Geoffroy J-J & Mauries J-P & VandenSpiegel D" pageId="36" pageNumber="37" pagination="1 - 53" refId="ref12534" refString="Golovatch SI, Geoffroy J-J, Mauries J-P, VandenSpiegel D (2009) Review of the millipede family Haplodesmidae, with descriptions of some new or poorly-known species (Diplopoda: Polydesmida). In: Golovatch SI, Mesibov R (Eds) Advances in the Systematics of Diplopoda I. ZooKeys 7: 1 - 53." type="journal article" year="2009">Golovatch, Geoffroy, Mauriès & VandenSpiegel, 2009</bibRefCitation>
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</taxonomicName>
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by the strongly underdeveloped paraterga 2, coupled
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</paragraph>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.3813134" ID-Zenodo-Dep="3813134" httpUri="https://zenodo.org/record/3813134/files/figure.png" pageId="37" pageNumber="38" startId="37.[140,212,902,924]" targetBox="[140,1108,165,882]" targetPageId="37">
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<paragraph blockId="37.[140,1108,902,956]" pageId="37" pageNumber="38">
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<emphasis bold="true" box="[140,251,902,924]" pageId="37" pageNumber="38">
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<figureCitation box="[140,245,902,924]" captionStart="Figure 3" captionStartId="5.[140,212,1442,1464]" captionTargetBox="[140,1108,165,1422]" captionTargetId="figure@5.[140,1108,165,1423]" captionTargetPageId="5" captionText="Figure 3. Eutrichodesmus distinctus sp. n., ♁ paratypes; A, midbody segments, ventral view; B, telson, ventral view; C, anterior part of body, ventral view; D, texture of tegument, dorsal view; E, both gonopods in situ, ventral view; F-H, gonopod, ventral, mesal and lateral views, respectively. – Scale bars: A, 0.5 mm; B, E, G, 0.1 mm; C, H, 0.2 mm; D, 0.02 mm; F, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3769975" httpUri="https://zenodo.org/record/3769975/files/figure.png" pageId="37" pageNumber="38">Figure 31</figureCitation>
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.
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</emphasis>
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<taxonomicName authority="Golovatch & Geoffroy & Mauriès & VandenSpiegel, 2009" authorityName="Golovatch & Geoffroy & Mauriès & VandenSpiegel" authorityYear="2009" box="[258,475,902,923]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="reductus" status="sp. n.">
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<emphasis box="[258,475,902,923]" italics="true" pageId="37" pageNumber="38">Eutrichodesmus reductus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[482,534,903,924]" pageId="37" pageNumber="38" rank="species">sp. n.</taxonomicNameLabel>
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, ♁ and
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<specimenCount box="[611,623,903,924]" pageId="37" pageNumber="38" type="female">♀</specimenCount>
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<typeStatus box="[635,724,903,924]" pageId="37" pageNumber="38">paratypes</typeStatus>
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; habitus, lateral view. (Photographed not to scale by L. Deharveng).
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</paragraph>
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</caption>
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<paragraph blockId="37.[140,1108,1011,1672]" pageId="37" pageNumber="38">
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with 19 body segments and the absence of metatergal tuberculation; from
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<taxonomicName authorityName="Golovatch, Geoffroy, Mauries & VandenSpiegel" authorityYear="2009" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="communicans">
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<emphasis italics="true" pageId="37" pageNumber="38">E. communicans</emphasis>
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</taxonomicName>
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by a very short tergal trichome, from it and other congeners in a few minor details of gonopod structure (in particular, the shape of the telopodite).
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="39" lastPageNumber="40" pageId="37" pageNumber="38" type="description">
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<paragraph blockId="37.[140,1108,1011,1672]" pageId="37" pageNumber="38">
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<emphasis bold="true" box="[185,334,1117,1143]" pageId="37" pageNumber="38">
|
|||
|
|
<emphasis box="[185,328,1117,1143]" inLineHeading="true" pageId="37" pageNumber="38" reason="1">Description</emphasis>
|
|||
|
|
:
|
|||
|
|
</emphasis>
|
|||
|
|
Length of adults of both sexes ca 4.0-
|
|||
|
|
<quantity box="[768,863,1117,1144]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="37" pageNumber="38" unit="mm" value="4.2">4.2 mm</quantity>
|
|||
|
|
, width
|
|||
|
|
<quantity box="[949,1103,1117,1144]" metricMagnitude="-4" metricUnit="m" metricValue="4.75" metricValueMax="5.0" metricValueMin="4.5" pageId="37" pageNumber="38" unit="mm" value="0.475" valueMax="0.5" valueMin="0.45">0.45-0.5 mm</quantity>
|
|||
|
|
, body broadest at midbody segments.
|
|||
|
|
<typeStatus box="[570,676,1152,1178]" pageId="37" pageNumber="38">Holotype</typeStatus>
|
|||
|
|
ca 4.0 mm long and
|
|||
|
|
<quantity box="[931,1038,1152,1179]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="37" pageNumber="38" unit="mm" value="0.45">0.45 mm</quantity>
|
|||
|
|
wide. Coloration uniformly pallid, shown pinkish because of a photographic artifact (Fig. 31).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph blockId="37.[140,1108,1011,1672]" lastBlockId="38.[140,1108,167,440]" lastPageId="38" lastPageNumber="39" pageId="37" pageNumber="38">
|
|||
|
|
Adults with 19 segments, body subcylindrical (Figs 31, 32A), volvation apparently incomplete due to insufficiently wide and low paraterga. Head (
|
|||
|
|
<figureCitation box="[844,942,1258,1284]" captionStart="Figure 33" captionStartId="39.[140,212,1558,1580]" captionTargetBox="[139,1107,420,1538]" captionTargetId="figure@39.[139,1108,420,1539]" captionTargetPageId="39" captionText="Figure 33. Eutrichodesmus reductus sp. n., ♀ (A-E) & ♁ (F-G) paratypes; A, head, ventral view; B, midbody segments, ventral view; C, midbody paratergum with ozopore, dorsal view; D, E, texture of tegument, dorsal view; F, G, left gonopod, mesal and lateral views, respectively. – Scale bars: A, B, 0.1 mm; C, 0.02 mm; D, 0.05 mm, E, 0.01 mm; F, G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3770011" httpUri="https://zenodo.org/record/3770011/files/figure.png" pageId="37" pageNumber="38">Fig. 33A</figureCitation>
|
|||
|
|
) slightly transverse, with a poorly separated pair of very low, paramedian tubercles above antennal sockets; antennae relatively short and clavate, antennomere 6 longest; tegument (
|
|||
|
|
<figureCitation captionStart="Figure 33" captionStartId="39.[140,212,1558,1580]" captionTargetBox="[139,1107,420,1538]" captionTargetId="figure@39.[139,1108,420,1539]" captionTargetPageId="39" captionText="Figure 33. Eutrichodesmus reductus sp. n., ♀ (A-E) & ♁ (F-G) paratypes; A, head, ventral view; B, midbody segments, ventral view; C, midbody paratergum with ozopore, dorsal view; D, E, texture of tegument, dorsal view; F, G, left gonopod, mesal and lateral views, respectively. – Scale bars: A, B, 0.1 mm; C, 0.02 mm; D, 0.05 mm, E, 0.01 mm; F, G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3770011" httpUri="https://zenodo.org/record/3770011/files/figure.png" pageId="37" pageNumber="38">Figs 33</figureCitation>
|
|||
|
|
C-E) and many other characters (
|
|||
|
|
<figureCitation box="[544,642,1363,1390]" captionStart="Figure 32" captionStartId="38.[140,212,1590,1612]" captionTargetBox="[140,1108,481,1570]" captionTargetId="figure@38.[140,1108,480,1570]" captionTargetPageId="38" captionText="Figure 32. Eutrichodesmus reductus sp. n., ♀ paratype; A, habitus, lateral view; B, anterior part of body, lateral view; C, midbody segments, lateral view; D-F, posterior part of body, lateral, dorsal and caudal views, respectively. – Scale bars: A, 0.5 mm; B-F, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3770009" httpUri="https://zenodo.org/record/3770009/files/figure.png" pageId="37" pageNumber="38">Figs 32F</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation box="[654,700,1364,1390]" captionStart="Figure 33" captionStartId="39.[140,212,1558,1580]" captionTargetBox="[139,1107,420,1538]" captionTargetId="figure@39.[139,1108,420,1539]" captionTargetPageId="39" captionText="Figure 33. Eutrichodesmus reductus sp. n., ♀ (A-E) & ♁ (F-G) paratypes; A, head, ventral view; B, midbody segments, ventral view; C, midbody paratergum with ozopore, dorsal view; D, E, texture of tegument, dorsal view; F, G, left gonopod, mesal and lateral views, respectively. – Scale bars: A, B, 0.1 mm; C, 0.02 mm; D, 0.05 mm, E, 0.01 mm; F, G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3770011" httpUri="https://zenodo.org/record/3770011/files/figure.png" pageId="37" pageNumber="38">33B</figureCitation>
|
|||
|
|
) much as in
|
|||
|
|
<taxonomicName authority="Golovatch & Geoffroy & Mauriès & VandenSpiegel, 2009" authorityName="Golovatch & Geoffroy & Mauriès & VandenSpiegel" authorityYear="2009" box="[850,980,1363,1389]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="distinctus" status="sp. n.">
|
|||
|
|
<emphasis box="[850,980,1363,1389]" italics="true" pageId="37" pageNumber="38">E. distinctus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel box="[989,1051,1364,1390]" pageId="37" pageNumber="38" rank="species">sp. n.</taxonomicNameLabel>
|
|||
|
|
; collum and following metaterga devoid of tuberculation, beset with numerous, irregularly arranged, extremely short setae (
|
|||
|
|
<figureCitation box="[507,587,1434,1460]" captionStart="Figure 32" captionStartId="38.[140,212,1590,1612]" captionTargetBox="[140,1108,481,1570]" captionTargetId="figure@38.[140,1108,480,1570]" captionTargetPageId="38" captionText="Figure 32. Eutrichodesmus reductus sp. n., ♀ paratype; A, habitus, lateral view; B, anterior part of body, lateral view; C, midbody segments, lateral view; D-F, posterior part of body, lateral, dorsal and caudal views, respectively. – Scale bars: A, 0.5 mm; B-F, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3770009" httpUri="https://zenodo.org/record/3770009/files/figure.png" pageId="37" pageNumber="38">Fig. 32</figureCitation>
|
|||
|
|
B-F, 33C-E). Paraterga directed ventrolaterad, slightly interrupting contour of convex dorsum, short, not reaching level of venter (
|
|||
|
|
<figureCitation captionStart="Figure 32" captionStartId="38.[140,212,1590,1612]" captionTargetBox="[140,1108,481,1570]" captionTargetId="figure@38.[140,1108,480,1570]" captionTargetPageId="38" captionText="Figure 32. Eutrichodesmus reductus sp. n., ♀ paratype; A, habitus, lateral view; B, anterior part of body, lateral view; C, midbody segments, lateral view; D-F, posterior part of body, lateral, dorsal and caudal views, respectively. – Scale bars: A, 0.5 mm; B-F, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3770009" httpUri="https://zenodo.org/record/3770009/files/figure.png" pageId="37" pageNumber="38">Figs 32</figureCitation>
|
|||
|
|
A-D); paraterga 2 (
|
|||
|
|
<figureCitation box="[386,486,1504,1530]" captionStart="Figure 32" captionStartId="38.[140,212,1590,1612]" captionTargetBox="[140,1108,481,1570]" captionTargetId="figure@38.[140,1108,480,1570]" captionTargetPageId="38" captionText="Figure 32. Eutrichodesmus reductus sp. n., ♀ paratype; A, habitus, lateral view; B, anterior part of body, lateral view; C, midbody segments, lateral view; D-F, posterior part of body, lateral, dorsal and caudal views, respectively. – Scale bars: A, 0.5 mm; B-F, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3770009" httpUri="https://zenodo.org/record/3770009/files/figure.png" pageId="37" pageNumber="38">Fig. 32B</figureCitation>
|
|||
|
|
) only a little enlarged compared to following ones, indistinctly lobulate both anterolaterally and laterally, with only a single noticeable lobe forming a schism ledge, both schism and hyposchism very small; paraterga 3 and 4 not narrower than others, overlap typical already from paraterga 4, not paraterga 5 as in preceding congeners. Paraterga broadly rounded caudally, very indistinctly lobulate laterally and with a single evident lobulation caudolaterally (
|
|||
|
|
<figureCitation box="[703,784,167,193]" captionStart="Figure 32" captionStartId="38.[140,212,1590,1612]" captionTargetBox="[140,1108,481,1570]" captionTargetId="figure@38.[140,1108,480,1570]" captionTargetPageId="38" captionText="Figure 32. Eutrichodesmus reductus sp. n., ♀ paratype; A, habitus, lateral view; B, anterior part of body, lateral view; C, midbody segments, lateral view; D-F, posterior part of body, lateral, dorsal and caudal views, respectively. – Scale bars: A, 0.5 mm; B-F, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3770009" httpUri="https://zenodo.org/record/3770009/files/figure.png" pageId="38" pageNumber="39">Figs 32</figureCitation>
|
|||
|
|
C-E, 33C). Limbus distinctly denticulate, almost hidden by nearby abundant microvilli (
|
|||
|
|
<figureCitation box="[801,940,202,228]" captionStart="Figure 33" captionStartId="39.[140,212,1558,1580]" captionTargetBox="[139,1107,420,1538]" captionTargetId="figure@39.[139,1108,420,1539]" captionTargetPageId="39" captionText="Figure 33. Eutrichodesmus reductus sp. n., ♀ (A-E) & ♁ (F-G) paratypes; A, head, ventral view; B, midbody segments, ventral view; C, midbody paratergum with ozopore, dorsal view; D, E, texture of tegument, dorsal view; F, G, left gonopod, mesal and lateral views, respectively. – Scale bars: A, B, 0.1 mm; C, 0.02 mm; D, 0.05 mm, E, 0.01 mm; F, G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3770011" httpUri="https://zenodo.org/record/3770011/files/figure.png" pageId="38" pageNumber="39">Fig. 33D, E</figureCitation>
|
|||
|
|
). Pore formula normal, ozopores located dorsally near base of caudal corner of paraterga (
|
|||
|
|
<figureCitation box="[953,1055,237,263]" captionStart="Figure 33" captionStartId="39.[140,212,1558,1580]" captionTargetBox="[139,1107,420,1538]" captionTargetId="figure@39.[139,1108,420,1539]" captionTargetPageId="39" captionText="Figure 33. Eutrichodesmus reductus sp. n., ♀ (A-E) & ♁ (F-G) paratypes; A, head, ventral view; B, midbody segments, ventral view; C, midbody paratergum with ozopore, dorsal view; D, E, texture of tegument, dorsal view; F, G, left gonopod, mesal and lateral views, respectively. – Scale bars: A, B, 0.1 mm; C, 0.02 mm; D, 0.05 mm, E, 0.01 mm; F, G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3770011" httpUri="https://zenodo.org/record/3770011/files/figure.png" pageId="38" pageNumber="39">Fig. 33C</figureCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph blockId="38.[140,1108,167,440]" box="[185,1107,272,299]" pageId="38" pageNumber="39">
|
|||
|
|
Legs relatively long and slender, evidently surpassing edge of paraterga (
|
|||
|
|
<figureCitation box="[992,1094,272,298]" captionStart="Figure 33" captionStartId="39.[140,212,1558,1580]" captionTargetBox="[139,1107,420,1538]" captionTargetId="figure@39.[139,1108,420,1539]" captionTargetPageId="39" captionText="Figure 33. Eutrichodesmus reductus sp. n., ♀ (A-E) & ♁ (F-G) paratypes; A, head, ventral view; B, midbody segments, ventral view; C, midbody paratergum with ozopore, dorsal view; D, E, texture of tegument, dorsal view; F, G, left gonopod, mesal and lateral views, respectively. – Scale bars: A, B, 0.1 mm; C, 0.02 mm; D, 0.05 mm, E, 0.01 mm; F, G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3770011" httpUri="https://zenodo.org/record/3770011/files/figure.png" pageId="38" pageNumber="39">Fig. 33B</figureCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph blockId="38.[140,1108,167,440]" lastBlockId="39.[140,1108,167,369]" lastPageId="39" lastPageNumber="40" pageId="38" pageNumber="39">
|
|||
|
|
Gonopods (
|
|||
|
|
<figureCitation box="[319,448,307,334]" captionStart="Figure 33" captionStartId="39.[140,212,1558,1580]" captionTargetBox="[139,1107,420,1538]" captionTargetId="figure@39.[139,1108,420,1539]" captionTargetPageId="39" captionText="Figure 33. Eutrichodesmus reductus sp. n., ♀ (A-E) & ♁ (F-G) paratypes; A, head, ventral view; B, midbody segments, ventral view; C, midbody paratergum with ozopore, dorsal view; D, E, texture of tegument, dorsal view; F, G, left gonopod, mesal and lateral views, respectively. – Scale bars: A, B, 0.1 mm; C, 0.02 mm; D, 0.05 mm, E, 0.01 mm; F, G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3770011" httpUri="https://zenodo.org/record/3770011/files/figure.png" pageId="38" pageNumber="39">Fig. 33F, G</figureCitation>
|
|||
|
|
) relatively complex. Coxae abundantly micropapillate, but only with a few macrosetae. Telopodite elongate, only slightly arcuate, with a large, papillate, peculiar, distofemoral outgrowth (dp) in distal one-third and a slender but short solenomere bearing a few small setae (but no pad!) subapically at base of a very complex tip represented by two erect teeth, a longer uncus and a group of minute outgrowths.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<caption ID-DOI="http://doi.org/10.5281/zenodo.3770009" ID-Zenodo-Dep="3770009" httpUri="https://zenodo.org/record/3770009/files/figure.png" pageId="38" pageNumber="39" startId="38.[140,212,1590,1612]" targetBox="[140,1108,481,1570]" targetPageId="38">
|
|||
|
|
<paragraph blockId="38.[140,1108,1590,1676]" pageId="38" pageNumber="39">
|
|||
|
|
<emphasis bold="true" box="[140,251,1590,1612]" pageId="38" pageNumber="39">Figure 32.</emphasis>
|
|||
|
|
<taxonomicName authority="Golovatch & Geoffroy & Mauriès & VandenSpiegel, 2009" authorityName="Golovatch & Geoffroy & Mauriès & VandenSpiegel" authorityYear="2009" box="[257,472,1590,1611]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="reductus" status="sp. n.">
|
|||
|
|
<emphasis box="[257,472,1590,1611]" italics="true" pageId="38" pageNumber="39">Eutrichodesmus reductus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel box="[479,530,1591,1612]" pageId="38" pageNumber="39" rank="species">sp. n.</taxonomicNameLabel>
|
|||
|
|
, ♀ paratype;
|
|||
|
|
<emphasis bold="true" box="[655,674,1590,1612]" pageId="38" pageNumber="39">A</emphasis>
|
|||
|
|
, habitus, lateral view;
|
|||
|
|
<emphasis bold="true" box="[882,899,1590,1612]" pageId="38" pageNumber="39">B</emphasis>
|
|||
|
|
, anterior part of body, lateral view;
|
|||
|
|
<emphasis bold="true" box="[257,276,1622,1644]" pageId="38" pageNumber="39">C</emphasis>
|
|||
|
|
, midbody segments, lateral view;
|
|||
|
|
<emphasis bold="true" box="[600,642,1622,1644]" pageId="38" pageNumber="39">D-F</emphasis>
|
|||
|
|
, posterior part of body, lateral, dorsal and caudal views, respectively. – Scale bars: A, 0.5 mm; B-F, 0.1 mm.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<subSubSection pageId="39" pageNumber="40" type="discussion">
|
|||
|
|
<paragraph blockId="39.[140,1108,167,369]" pageId="39" pageNumber="40">
|
|||
|
|
<emphasis bold="true" box="[185,298,237,263]" pageId="39" pageNumber="40">
|
|||
|
|
<emphasis box="[185,293,237,263]" inLineHeading="true" pageId="39" pageNumber="40" reason="1">Remarks</emphasis>
|
|||
|
|
:
|
|||
|
|
</emphasis>
|
|||
|
|
This small-bodied, pallid, non-volvatory (= “haplodesmid”, see
|
|||
|
|
<bibRefCitation author="Golovatch SI & Geoffroy J-J & Mauries J-P & VandenSpiegel D" pageId="39" pageNumber="40" pagination="1 - 53" refId="ref12534" refString="Golovatch SI, Geoffroy J-J, Mauries J-P, VandenSpiegel D (2009) Review of the millipede family Haplodesmidae, with descriptions of some new or poorly-known species (Diplopoda: Polydesmida). In: Golovatch SI, Mesibov R (Eds) Advances in the Systematics of Diplopoda I. ZooKeys 7: 1 - 53." type="journal article" year="2009">Golovatch et al. 2009</bibRefCitation>
|
|||
|
|
) species seems to be especially close, morphologically as well as geographically, to
|
|||
|
|
<taxonomicName authorityName="Golovatch, Geoffroy, Mauries & VandenSpiegel" authorityYear="2009" box="[306,485,308,333]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="communicans">
|
|||
|
|
<emphasis box="[306,485,308,333]" italics="true" pageId="39" pageNumber="40">E. communicans</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
from
|
|||
|
|
<collectingCountry box="[556,655,308,334]" name="Vanuatu" pageId="39" pageNumber="40">Vanuatu</collectingCountry>
|
|||
|
|
, Melanesia, southwestern Pacific (
|
|||
|
|
<bibRefCitation author="Golovatch SI & Geoffroy J-J & Mauries J-P & VandenSpiegel D" pageId="39" pageNumber="40" pagination="1 - 53" refId="ref12534" refString="Golovatch SI, Geoffroy J-J, Mauries J-P, VandenSpiegel D (2009) Review of the millipede family Haplodesmidae, with descriptions of some new or poorly-known species (Diplopoda: Polydesmida). In: Golovatch SI, Mesibov R (Eds) Advances in the Systematics of Diplopoda I. ZooKeys 7: 1 - 53." type="journal article" year="2009">Golovatch et al. 2009</bibRefCitation>
|
|||
|
|
), possibly representing still one more troglobite.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|