treatments-xml/data/86/20/98/862098811AD37CD4171CB6D7174F1C33.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.905.39592" ID-GBIF-Dataset="ee5a0817-7cfc-4c21-ba7d-3beabb06cccd" ID-PMC="PMC7004242" ID-Pensoft-Pub="1313-2970-905-1" ID-Pensoft-UUID="4C7E0CB428DF56BDB15B7AC8D6707961" ID-PubMed="32063726" ID-ZooBank="F4C766E4633A41039FFDE952718F41FB" ModsDocID="1313-2970-905-1" checkinTime="1579539902851" checkinUser="pensoft" docAuthor="Salata, Sebastian &amp; Fisher, Brian L." docDate="2020" docId="862098811AD37CD4171CB6D7174F1C33" docLanguage="en" docName="ZooKeys 905: 1-235" docOrigin="ZooKeys 905" docSource="http://dx.doi.org/10.3897/zookeys.905.39592" docTitle="Pheidole boribora Salata &amp; Fisher 2020, sp. nov." docType="treatment" docUuid="EDD6BF4F-581F-4246-B931-502AB27EC1BB" docUuidSource="ZooBank" docVersion="5" id="4C7E0CB428DF56BDB15B7AC8D6707961" lastPageNumber="1" masterDocId="4C7E0CB428DF56BDB15B7AC8D6707961" masterDocTitle="Pheidole Westwood, 1839 (Hymenoptera, Formicidae) of Madagascar - an introduction and a taxonomic revision of eleven species groups" masterLastPageNumber="235" masterPageNumber="1" pageNumber="1" updateTime="1668168337901" updateUser="ExternalLinkService">
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<mods:title>Pheidole Westwood, 1839 (Hymenoptera, Formicidae) of Madagascar - an introduction and a taxonomic revision of eleven species groups</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Salata, Sebastian</mods:namePart>
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<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Fisher, Brian L.</mods:namePart>
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<mods:date>2020</mods:date>
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<mods:number>905</mods:number>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.905.39592</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-905-1</mods:identifier>
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<treatment ID-GBIF-Taxon="161777020" LSID="urn:lsid:zoobank.org:act:EDD6BF4F-581F-4246-B931-502AB27EC1BB" httpUri="http://treatment.plazi.org/id/862098811AD37CD4171CB6D7174F1C33" lastPageNumber="1" pageId="0" pageNumber="1">
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<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="http://zoobank.org/EDD6BF4F-581F-4246-B931-502AB27EC1BB" authority="Salata &amp; Fisher, 2020" authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole boribora" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="boribora" status="sp. nov.">Pheidole boribora</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 73" captionStartId="F73" captionText="Figure 73. Pheidole boribora sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0456014) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0456013)." figureDoi="10.3897/zookeys.905.39592.figure73" httpUri="https://binary.pensoft.net/fig/373120" pageId="0" pageNumber="1">Figs 73A-F</figureCitation>
<figureCitation captionStart="Figure 84" captionStartId="F84" captionText="Figure 84. Hypostomal teeth. Pheidole aelloea sp. nov. (A). P. ala sp. nov. (B). P. andapa sp. nov. (C). P. ankerana sp. nov. (D). P. annemariae Forel (E). P. avaratra sp. nov. (F). P. bemarahaensis sp. nov. (G). P. bemarivoensis sp. nov. (H). P. binara sp. nov. (I). P. boribora sp. nov. (J). P. brevipilosa sp. nov. (K). P. curvistriata sp. nov. (L). P. diakritos sp. nov. (M). P. ehazoara sp. nov. (N). P. ensifera Forel (O). P. ferruginea sp. nov. (P). P. fisaka sp. nov. (R). P. fitarata sp. nov. (S). P. glabra sp. nov. (T). P. goavana sp. nov. (U). P. lamperos sp. nov. (W). P. longipilosa sp. nov. (X). P. longispinosa Forel (Y). P. lutea sp. nov. (Z)." figureDoi="10.3897/zookeys.905.39592.figure84" httpUri="https://binary.pensoft.net/fig/373131" pageId="0" pageNumber="1">, 84J</figureCitation>
<figureCitation captionStart="Figure 86" captionStartId="F86" captionText="Figure 86. Distribution. Pheidole aelloea sp. nov. (A). P. ala sp. nov. (B). P. andapa sp. nov. (C). P. ankerana sp. nov. (D). P. annemariae Forel (E). P. avaratra sp. nov. (F). P. bemarahaensis sp. nov. (G). P. bemarivoensis sp. nov. (H). P. binara sp. nov. (I). P. boribora sp. nov. (J). P. brevipilosa sp. nov. (K). P. curvistriata sp. nov. (L). P. diakritos sp. nov. (M). P. ehazoara sp. nov. (N). P. ensifera Forel (O). P. ferruginea sp. nov. (P). P. fisaka sp. nov. (R). P. fitarata sp. nov. (S). P. glabra sp. nov. (T). P. goavana sp. nov. (U)." figureDoi="10.3897/zookeys.905.39592.figure86" httpUri="https://binary.pensoft.net/fig/373133" pageId="0" pageNumber="1">, 86J</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type material">
<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
.
</emphasis>
Madagascar. •1 major worker; Toliara; Parc National 
<normalizedToken originalValue="d’Andohahela">d'Andohahela</normalizedToken>
, Col du Sedro, 3.8 km 113°ESE Mahamavo, 37.6 km 341°NNW Tolagnaro; 
<geoCoordinate degrees="24.76389" direction="south" orientation="latitude" precision="1" value="-24.76389">-24.76389</geoCoordinate>
, 
<geoCoordinate degrees="46.75167" direction="east" orientation="longitude" precision="1" value="46.75167">46.75167</geoCoordinate>
; alt. 900 m; 21 Jan 2002; B.L. Fisher et al. leg.; BLF05130, CASENT0456013 (CASC). 
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
.
</emphasis>
Madagascar. •4w.; same data as for holotype; CASENT0456014-CASENT0456016, CASENT0872187-CASENT0872190 (CASC).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Major workers</emphasis>
.
</emphasis>
Head in full-face view oval, relatively as long as wide, anterior and posterior of eyes moderately convex; sides of the head with moderately dense, long, erect pilosity; genae smooth and shiny; inner hypostomal teeth distinct, distinct, low, thick, and triangular, with rounded apex; outer hypostomal teeth approximately the same size as inner hypostomal teeth, thick, dentate, with rounded tips; propodeal spines small, triangular; first gastral tergite finely shagreened; body reddish brown. 
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Minor workers</emphasis>
.
</emphasis>
Head foveolate; promesonotum low, short, slightly convex, with relatively steep posterior declivity; propodeal spines small; mesosoma foveolate, foveolate, mesonotum, anepisternum, katepisternum and lateral surfaces of propodeum smooth; body yellow.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Major workers.</emphasis>
Measurements (
<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
= 1): HL: 1.5; HW: 1.4; SL: 0.66; EL: 0.17; WL: 1.07; PSL: 0.17; MTL: 0.66; PNW: 0.58; PTW: 0.19; PPW: 0.41; CI: 93.4; SI: 47.2; PSLI: 11.2; PPI: 45.5; PNI: 41.6; MTI: 46.9. 
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Head</emphasis>
.
</emphasis>
In full-face view oval, anterior and posterior of eyes moderately convex (Fig. 
<figureCitation captionStart="Figure 73" captionStartId="F73" captionText="Figure 73. Pheidole boribora sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0456014) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0456013)." figureDoi="10.3897/zookeys.905.39592.figure73" httpUri="https://binary.pensoft.net/fig/373120" pageId="0" pageNumber="1">73B</figureCitation>
). In lateral view sub-rectangular; ventral and dorsal faces finely convex; dorsal face finely depressed posteriorly, forming shallow transverse depression between frons and occipital lobes; inner hypostomal teeth invisible. Sides of the head with moderately dense, long, erect pilosity; whole head with moderately dense, short, suberect to erect pilosity. Antennal scrobes indistinct and not delimited by carinulae. Occipital lobes shiny, with thick, sparse, irregular rugae, interspaces smooth to finely foveolate; frons with thick, moderately dense, longitudinal rugae, interspaces smooth to finely foveolate; lateral sides of head foveolate, with additional thin, sparse, longitudinal rugae; malar area with dense and thin longitudinal rugulae; genae smooth and shiny. Clypeus shiny and smooth, with thin, longitudinal rugulae on the lateral sides; median notch present, narrow, and moderately deep; median longitudinal carina absent; lateral longitudinal carinae absent. Scape, when laid back, reaching the midlength of head; pilosity decumbent to erect (Fig. 
<figureCitation captionStart="Figure 73" captionStartId="F73" captionText="Figure 73. Pheidole boribora sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0456014) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0456013)." figureDoi="10.3897/zookeys.905.39592.figure73" httpUri="https://binary.pensoft.net/fig/373120" pageId="0" pageNumber="1">73B, D</figureCitation>
). Inner hypostomal teeth distinct, low, thick, and triangular, with rounded apex; outer hypostomal teeth approximately the same size as inner hypostomal teeth, thick, dentate, with rounded tips (Fig. 
<figureCitation captionStart="Figure 84" captionStartId="F84" captionText="Figure 84. Hypostomal teeth. Pheidole aelloea sp. nov. (A). P. ala sp. nov. (B). P. andapa sp. nov. (C). P. ankerana sp. nov. (D). P. annemariae Forel (E). P. avaratra sp. nov. (F). P. bemarahaensis sp. nov. (G). P. bemarivoensis sp. nov. (H). P. binara sp. nov. (I). P. boribora sp. nov. (J). P. brevipilosa sp. nov. (K). P. curvistriata sp. nov. (L). P. diakritos sp. nov. (M). P. ehazoara sp. nov. (N). P. ensifera Forel (O). P. ferruginea sp. nov. (P). P. fisaka sp. nov. (R). P. fitarata sp. nov. (S). P. glabra sp. nov. (T). P. goavana sp. nov. (U). P. lamperos sp. nov. (W). P. longipilosa sp. nov. (X). P. longispinosa Forel (Y). P. lutea sp. nov. (Z)." figureDoi="10.3897/zookeys.905.39592.figure84" httpUri="https://binary.pensoft.net/fig/373131" pageId="0" pageNumber="1">84J</figureCitation>
). 
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
.
</emphasis>
In lateral view, promesonotum short, angular, and relatively low, posterior mesonotum steep, with moderately large, tubercle-like projections; promesonotal groove absent; metanotal groove absent or indistinct; propodeal spines small, triangular, with base wide, apex rounded; humeral area with small and flat tubercles (Fig. 
<figureCitation captionStart="Figure 73" captionStartId="F73" captionText="Figure 73. Pheidole boribora sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0456014) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0456013)." figureDoi="10.3897/zookeys.905.39592.figure73" httpUri="https://binary.pensoft.net/fig/373120" pageId="0" pageNumber="1">73D</figureCitation>
). Surface shiny, with very sparse, transverse to irregular thin rugae, pronotum and propodeum with additional indistinct and sparse foveolae. Pilosity moderately dense, long, and erect (Fig. 
<figureCitation captionStart="Figure 73" captionStartId="F73" captionText="Figure 73. Pheidole boribora sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0456014) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0456013)." figureDoi="10.3897/zookeys.905.39592.figure73" httpUri="https://binary.pensoft.net/fig/373120" pageId="0" pageNumber="1">73D, F</figureCitation>
). 
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
.
</emphasis>
Shiny, finely shagreened to smooth; peduncle short, with small, rounded, horizontal lobes on its basal part; node moderately high and triangular, with convex apex, in rear view node relatively straight; pilosity moderately sparse and erect (Fig. 
<figureCitation captionStart="Figure 73" captionStartId="F73" captionText="Figure 73. Pheidole boribora sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0456014) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0456013)." figureDoi="10.3897/zookeys.905.39592.figure73" httpUri="https://binary.pensoft.net/fig/373120" pageId="0" pageNumber="1">73D, F</figureCitation>
). 
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
.
</emphasis>
Shiny and finely shagreened; short and rounded; in dorsal view sides with short, acute, triangular projections; pilosity long, moderately dense, and erect (Fig. 
<figureCitation captionStart="Figure 73" captionStartId="F73" captionText="Figure 73. Pheidole boribora sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0456014) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0456013)." figureDoi="10.3897/zookeys.905.39592.figure73" httpUri="https://binary.pensoft.net/fig/373120" pageId="0" pageNumber="1">73D, F</figureCitation>
). 
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
.
</emphasis>
First gastral tergite shiny and finely shagreened; pilosity dense, long and erect (Fig. 
<figureCitation captionStart="Figure 73" captionStartId="F73" captionText="Figure 73. Pheidole boribora sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0456014) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0456013)." figureDoi="10.3897/zookeys.905.39592.figure73" httpUri="https://binary.pensoft.net/fig/373120" pageId="0" pageNumber="1">73D, F</figureCitation>
). 
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Colour</emphasis>
.
</emphasis>
Head reddish brown; mesosoma and gaster yellowish brown to brown; legs dark yellow (Fig. 
<figureCitation captionStart="Figure 73" captionStartId="F73" captionText="Figure 73. Pheidole boribora sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0456014) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0456013)." figureDoi="10.3897/zookeys.905.39592.figure73" httpUri="https://binary.pensoft.net/fig/373120" pageId="0" pageNumber="1">73D, F</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.905.39592.figure73" httpUri="https://binary.pensoft.net/fig/373120" pageId="0" pageNumber="1" start="Figure 73" startId="F73">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 73.</emphasis>
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole boribora" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="boribora">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole boribora</emphasis>
</taxonomicName>
sp. nov., full-face view (
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
), profile (
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
), and dorsal view (
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
) of paratype minor worker (CASENT0456014) and full-face view (
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
), profile (
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
), and dorsal view (
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
) of holotype major worker (CASENT0456013).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
Measurements (
<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
= 3): HL: 0.55-0.58 (0.56); HW: 0.49-0.5 (0.49); SL: 0.57-0.58 (0.57); EL: 0.11-0.11 (0.11); WL: 0.68-0.76 (0.74); PSL: 0.07-0.1 (0.08); MTL: 0.42-0.46 (0.43); PNW: 0.31-0.32 (0.31); PTW: 0.06-0.08 (0.07); PPW: 0.12-0.14 (0.13); CI: 85.8-89.4 (87.5); SI: 116.2-116.8 (116.4); PSLI: 12.0-17.0 (14.6); PPI: 46.0-59.8 (53.3); PNI: 62.9-64.2 (63.5); MTI: 85.0-91.2 (88.0). 
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Head</emphasis>
.
</emphasis>
Occipital margin straight or indistinctly concave; occipital carina indistinct, weakly developed (Fig. 
<figureCitation captionStart="Figure 73" captionStartId="F73" captionText="Figure 73. Pheidole boribora sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0456014) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0456013)." figureDoi="10.3897/zookeys.905.39592.figure73" httpUri="https://binary.pensoft.net/fig/373120" pageId="0" pageNumber="1">73A</figureCitation>
). Pilosity moderately dense, long, erect. Whole head foveolate. Clypeus with median longitudinal carina absent; two lateral longitudinal carinae absent. Scape, when laid back, surpassing the posterior head margin by two-fifths of its length; pilosity erect (Fig. 
<figureCitation captionStart="Figure 73" captionStartId="F73" captionText="Figure 73. Pheidole boribora sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0456014) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0456013)." figureDoi="10.3897/zookeys.905.39592.figure73" httpUri="https://binary.pensoft.net/fig/373120" pageId="0" pageNumber="1">73A, C</figureCitation>
). 
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
.
</emphasis>
In lateral view, promesonotum low, short, slightly convex, with relatively steep posterior declivity; promesonotal groove absent; metanotal groove absent; propodeal spines small, triangular, apex acute (Fig. 
<figureCitation captionStart="Figure 73" captionStartId="F73" captionText="Figure 73. Pheidole boribora sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0456014) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0456013)." figureDoi="10.3897/zookeys.905.39592.figure73" httpUri="https://binary.pensoft.net/fig/373120" pageId="0" pageNumber="1">73C</figureCitation>
). Sculpture foveolate; katepisternum, anepisternum and mesonotum smooth. Pilosity sparse, long, and erect (Fig. 
<figureCitation captionStart="Figure 73" captionStartId="F73" captionText="Figure 73. Pheidole boribora sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0456014) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0456013)." figureDoi="10.3897/zookeys.905.39592.figure73" httpUri="https://binary.pensoft.net/fig/373120" pageId="0" pageNumber="1">73C, E</figureCitation>
). 
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
.
</emphasis>
Peduncle short and thin with ventral face slightly convex; node low, triangular, and small; with few moderately long, erect setae (Fig. 
<figureCitation captionStart="Figure 73" captionStartId="F73" captionText="Figure 73. Pheidole boribora sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0456014) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0456013)." figureDoi="10.3897/zookeys.905.39592.figure73" httpUri="https://binary.pensoft.net/fig/373120" pageId="0" pageNumber="1">73C, E</figureCitation>
). 
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
.
</emphasis>
Short, low, and convex; with few moderately long, erect setae (Fig. 
<figureCitation captionStart="Figure 73" captionStartId="F73" captionText="Figure 73. Pheidole boribora sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0456014) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0456013)." figureDoi="10.3897/zookeys.905.39592.figure73" httpUri="https://binary.pensoft.net/fig/373120" pageId="0" pageNumber="1">73C, E</figureCitation>
). 
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
.
</emphasis>
With moderately sparse, erect pilosity (Fig. 
<figureCitation captionStart="Figure 73" captionStartId="F73" captionText="Figure 73. Pheidole boribora sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0456014) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0456013)." figureDoi="10.3897/zookeys.905.39592.figure73" httpUri="https://binary.pensoft.net/fig/373120" pageId="0" pageNumber="1">73C, E</figureCitation>
). 
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Colour</emphasis>
.
</emphasis>
Unicolourous, yellow (Fig. 
<figureCitation captionStart="Figure 73" captionStartId="F73" captionText="Figure 73. Pheidole boribora sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0456014) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0456013)." figureDoi="10.3897/zookeys.905.39592.figure73" httpUri="https://binary.pensoft.net/fig/373120" pageId="0" pageNumber="1">73C, E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">Malagasy for oval, in reference to the head shape of major workers.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="biology_ecology">
<paragraph pageId="0" pageNumber="1">Biology.</paragraph>
<paragraph pageId="0" pageNumber="1">The species was collected at 900 m in elevation, in montane rainforest. Nest was located in a rotten log.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="comments">
<paragraph pageId="0" pageNumber="1">Comments.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Major workers</emphasis>
.
</emphasis>
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole boribora" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="boribora">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole boribora</emphasis>
</taxonomicName>
sp. nov. is most similar to 
<taxonomicName lsidName="P. miramila" pageId="0" pageNumber="1" rank="species" species="miramila">
<emphasis italics="true" pageId="0" pageNumber="1">P. miramila</emphasis>
</taxonomicName>
sp. nov. and differs from it by presence of foveolae on head, dense and long pilosity of sides of head, dentate inner hypostomal teeth, and small propodeal spines. 
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Minor workers</emphasis>
.
</emphasis>
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole boribora" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="boribora">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole boribora</emphasis>
</taxonomicName>
sp. nov. is most similar to 
<taxonomicName lsidName="P. petax" pageId="0" pageNumber="1" rank="species" species="petax">
<emphasis italics="true" pageId="0" pageNumber="1">P. petax</emphasis>
</taxonomicName>
and differs from it by smooth mesonotum, anepisternum, and lateral surfaces of propodeum.
</paragraph>
</subSubSection>
</treatment>
</document>