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<document id="6C41A244A5D675520B12B52CBA74B104" ID-DOI="10.11646/zootaxa.2462.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5312710" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629676798216" checkinUser="felipe" docAuthor="Hyman, Isabel T. &amp; Ponder, Winston F." docDate="2010" docId="6413F378FFDB6A46F28B750CFBF3F8BF" docLanguage="en" docName="zt02462p148.pdf" docOrigin="Zootaxa 2462 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2462.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Periclocystis ardeni Iredale 1937" docType="treatment" docVersion="3" lastPageNumber="77" masterDocId="982A8B00FF976A0AF21C766BFFEBFF8B" masterDocTitle="A morphological phylogenetic analysis and generic revision of Australian Helicarionidae (Gastropoda: Pulmonata: Stylommatophora), and an assessment of the relationships of the family 2462" masterLastPageNumber="148" masterPageNumber="1" pageNumber="77" updateTime="1699094437759" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="6F78926C9198077951FDE87EC7C51225" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="3F57E75EB4A74A4007A563873576FFFB">
<mods:title id="6A3726880116A8B4706A26D11281055E">A morphological phylogenetic analysis and generic revision of Australian Helicarionidae (Gastropoda: Pulmonata: Stylommatophora), and an assessment of the relationships of the family 2462</mods:title>
</mods:titleInfo>
<mods:name id="6CB8109BE153ADA47DA45020111B144E" type="personal">
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<mods:roleTerm id="B65FF1CC2CD4E1C115CC5AECBFE69392">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F03D9B2860C0D8AB2D6D7C94FA516D4B">Hyman, Isabel T.</mods:namePart>
</mods:name>
<mods:name id="26D655787DD7BE78F1E7F9A6444C3205" type="personal">
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<mods:roleTerm id="72A4881939594F2DCD60FD03FB007E52">Author</mods:roleTerm>
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<mods:namePart id="D19432F465261BE5A6C1F721650EB2D4">Ponder, Winston F.</mods:namePart>
</mods:name>
<mods:typeOfResource id="E3FD738092ABF773973C4443BDC45C4F">text</mods:typeOfResource>
<mods:relatedItem id="F77C886E58FCF77C4959E457FCC3664A" type="host">
<mods:titleInfo id="C59E9B630FB85776CA90C02EABD8F7C7">
<mods:title id="AA680D4BA91B02EC16DDF7A1B4824EB3">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="81E38C02FEB169360A68667024F93845">
<mods:date id="5FB5A8A4E3C858A2D2FB09431783736E">2010</mods:date>
<mods:detail id="EB078BFAD9FB8F74C621F659C3F6FEE5" type="pubDate">
<mods:number id="684AD4D2B56A69417421AFDF08C15064">2010-05-14</mods:number>
</mods:detail>
<mods:detail id="61A4A239D7818F75EB3CB00165D6B726" type="volume">
<mods:number id="6E0A49DBE861DC2861DD3123794AB7BC">2462</mods:number>
</mods:detail>
<mods:detail id="CE5D4117204EAE7BE99B5ED58A9303C1" type="issue">
<mods:number id="76FF4D7443ADC82A60424E7096A6CC15">1</mods:number>
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<mods:start id="433359B4A56CDD984D35AE4E9E533C45">1</mods:start>
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<mods:url id="3140D0E2650392FE5160269265BEBD8F">https://biotaxa.org/Zootaxa/article/view/zootaxa.2462.1.1</mods:url>
</mods:location>
<mods:classification id="5DA988F6D205E25E810E54F5D2B6619B">journal article</mods:classification>
<mods:identifier id="E7E78602E51971C28631015D66F0BDD0" type="DOI">10.11646/zootaxa.2462.1.1</mods:identifier>
<mods:identifier id="74A2D31F04A991A4CC13809B8489D45B" type="ISSN">1175­5334</mods:identifier>
<mods:identifier id="119334BE6458A7A9FE90705C15627514" type="Zenodo-Dep">5312710</mods:identifier>
</mods:mods>
<treatment id="6413F378FFDB6A46F28B750CFBF3F8BF" LSID="urn:lsid:plazi:treatment:6413F378FFDB6A46F28B750CFBF3F8BF" httpUri="http://treatment.plazi.org/id/6413F378FFDB6A46F28B750CFBF3F8BF" lastPageNumber="77" pageId="76" pageNumber="77">
<subSubSection id="A4A011E5FFDB6A46F28B750CFDC7FC0A" box="[151,556,871,897]" pageId="76" pageNumber="77" type="nomenclature">
<paragraph id="EC05426EFFDB6A46F28B750CFDC7FC0A" blockId="76.[151,556,871,932]" box="[151,556,871,897]" pageId="76" pageNumber="77">
<heading id="B74DF502FFDB6A46F28B750CFDC7FC0A" bold="true" box="[151,556,871,897]" fontSize="11" level="1" pageId="76" pageNumber="77" reason="1">
<taxonomicName id="2BBA39EDFFDB6A46F28B750CFDC7FC0A" ID-CoL="6V55Y" authority="Iredale, 1937" authorityName="Iredale" authorityYear="1937" box="[151,556,871,897]" class="Gastropoda" family="Helicarionidae" genus="Periclocystis" kingdom="Animalia" order="Stylommatophora" pageId="76" pageNumber="77" phylum="Mollusca" rank="species" species="ardeni">
<emphasis id="DECE9E7CFFDB6A46F28B750CFDC7FC0A" bold="true" box="[151,556,871,897]" pageId="76" pageNumber="77">
<emphasis id="DECE9E7CFFDB6A46F28B750CFE68FC0A" bold="true" box="[151,387,871,897]" italics="true" pageId="76" pageNumber="77">Periclocystis ardeni</emphasis>
Iredale, 1937
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A4A011E5FFDB6A46F28B75E1FDF4FC2F" box="[151,543,906,932]" pageId="76" pageNumber="77" type="description">
<paragraph id="EC05426EFFDB6A46F28B75E1FDF4FC2F" blockId="76.[151,556,871,932]" box="[151,543,906,932]" pageId="76" pageNumber="77">
<figureCitation id="74815EEBFFDB6A46F28B75E1FEC0FC2F" box="[151,299,906,932]" captionStart="FIGURE 22" captionStartId="59.[151,255,1798,1822]" captionTargetBox="[308,1282,221,1760]" captionTargetId="figure-127@59.[280,1310,194,1786]" captionTargetPageId="59" captionText="FIGURE 22. Penis interiors. A. Nitor circumcincta, AM C356607. B. Nitor subrugata, AM C334537. C. Pravonitor kreffti, AM C425415. D. Expocystis rustica, AM C425540. E. Malandena suturalis, AM C425085. F. Echonitor cyrtochila, SAM D16557. G. Echonitor albumenoidea, SAM D16567. H. Periclocystis ardeni, SAM D16562. I. New Caledonian NC3, MNHN no registration number. J. New Caledonian NC5, MNHN no registration number. K. Caldwellia angularis, AM C205299. L. Caldwellia imperfecta, AM C205300. Scale bars 1 mm (A–L). Abbreviations: ep, epiphallus; pd, penial diverticulum; pp, penial pilaster; ppd, diamond-shaped, ridged pustules; prm, penis retractor muscle; ppu, penial pustules; pv, penial verge." figureDoi="http://doi.org/10.5281/zenodo.5312770" httpUri="https://zenodo.org/record/5312770/files/figure.png" pageId="76" pageNumber="77">Figures 22H</figureCitation>
,
<figureCitation id="74815EEBFFDB6A46F32B75E1FE82FC2F" box="[311,361,906,932]" captionStart="FIGURE 24" captionStartId="68.[151,255,1783,1807]" captionTargetBox="[305,1289,210,1746]" captionTargetId="figure-140@68.[281,1313,194,1768]" captionTargetPageId="68" captionText="FIGURE 24. Reproductive systems. A. Malandena suturalis, AM C425085. B. Westracystis lissus, FMNH 211717. C. Echonitor cyrtochila, SAM D16557. D. Echonitor albumenoidea, SAM D16567. E. Periclocystis ardeni, SAM D16562. F. Tarocystis responsivus, AM C425327. G. New Caledonian species NC3, MNHN no registration number. H. New Caledonian species NC5, MNHN no registration number. Scale bars 1 mm (C, G), 2 mm (A–B, D–F, H). Abbreviations: alb, albumen gland; bc, bursa copulatrix; c, caecum; ep, epiphallus; fl, flagellum; fov, free oviduct; her, hermaphrodite duct; m, muscle attaching penial sheath to epiphallus (only shown in D &amp; G); ov, ovotestis; p, penis; pr, prostate; prm, penis retractor muscle; pt, penial sheath; t, talon; ut, uterus; v, vagina; vd, vas deferens." figureDoi="http://doi.org/10.5281/zenodo.5312774" httpUri="https://zenodo.org/record/5312774/files/figure.png" pageId="76" pageNumber="77">24E</figureCitation>
,
<figureCitation id="74815EEBFFDB6A46F36975E1FE4CFC2F" box="[373,423,906,932]" captionStart="FIGURE 26" captionStartId="71.[151,255,1346,1370]" captionTargetBox="[256,1309,267,1319]" captionTargetId="figure-229@71.[253,1314,250,1328]" captionTargetPageId="71" captionText="FIGURE 26. Shells, dorsal view. A. Westracystis lissus, BMNH 90.12.30.80–4 (S). B. Westracystis lissus, AM C64901 (S of W. tentus). C. Echonitor cyrtochila, BMNH 1922.8.29.16 (H). D. Echonitor albumenoidea, AM C60533 (H). E. Periclocystis ardeni, AM C60541 (H). F. Tarocystis responsivus, AM C104308 (S). G. Conueplecta (Durgellina) calculosa, AM C374873. H. Coneuplecta (Nevelasta) pampini, AM C126763. Scale bars 2 mm (B, G–H), 3 mm (A, C–F). Photographs by M. Allan (F), H. Barlow (B, G–H), D. Beechey (D–E), W. Ponder (A, C). Abbreviations: H, holotype; S, syntype." figureDoi="http://doi.org/10.5281/zenodo.5312780" httpUri="https://zenodo.org/record/5312780/files/figure.png" pageId="76" pageNumber="77">26E</figureCitation>
,
<figureCitation id="74815EEBFFDB6A46F3AF75E1FE0FFC2F" box="[435,484,906,932]" captionStart="FIGURE 27" captionStartId="73.[151,255,1674,1698]" captionTargetBox="[267,1319,605,1658]" captionTargetId="figure-241@73.[267,1320,605,1658]" captionTargetPageId="73" captionText="FIGURE 27. Shells, lateral view. A. Westracystis lissus, BMNH 90.12.30.80–4 (S). B. Westracystis lissus, AM C64901 (S of W. tentus). C. Echonitor cyrtochila, BMNH 1922.8.29.16 (H). D. Echonitor albumenoidea, AM C60533 (H). E. Periclocystis ardeni, AM C60541 (H). F. Tarocystis responsivus, AM C104308 (S). G. Conueplecta (Durgellina) calculosa, AM C374873. H. Coneuplecta (Nevelasta) pampini, AM C126763. Scale bars 2 mm (B, G–H), 3 mm (A, C–F). Photographs by H. Barlow (B, F–H), D. Beechey (D–E), W. Ponder (A, C). Abbreviations: H, holotype; S, syntype." figureDoi="http://doi.org/10.5281/zenodo.5312782" httpUri="https://zenodo.org/record/5312782/files/figure.png" pageId="76" pageNumber="77">27E</figureCitation>
,
<figureCitation id="74815EEBFFDB6A46F3ED75E1FDF4FC2F" box="[497,543,906,932]" captionStart="FIGURE 28" captionStartId="74.[151,255,1666,1690]" captionTargetBox="[266,1319,585,1640]" captionTargetId="figure-316@74.[248,1332,570,1664]" captionTargetPageId="74" captionText="FIGURE 28. Shells, ventral view. A. Westracystis lissus, BMNH90.12.30.80–4 (S). B. Westracystis lissus, AM C64901 (S of W. tentus). C. Echonitor cyrtochila, BMNH 1922.8.29.16 (H). D. Echonitor albumenoidea, AM C60533 (H). E. Periclocystis ardeni, AM C60541 (H). F. Tarocystis responsivus, AM C104308 (S). G. Conueplecta (Durgellina) calculosa, AM C374873. H. Coneuplecta (Nevelasta) pampini, AM C126763. Scale bars 2 mm (B, G–H), 3 mm (A, C–F). Photographs by H. Barlow (B, F–H), D. Beechey (D–E), W. Ponder (A, C). Abbreviations: H, holotype; S, syntype." figureDoi="http://doi.org/10.5281/zenodo.5312786" httpUri="https://zenodo.org/record/5312786/files/figure.png" pageId="76" pageNumber="77">28E</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="A4A011E5FFDB6A46F28B75B1FBBDFC7A" box="[151,1110,986,1009]" pageId="76" pageNumber="77" type="reference_group">
<paragraph id="EC05426EFFDB6A46F28B75B1FBBDFC7A" blockId="76.[151,1110,986,1009]" box="[151,1110,986,1009]" pageId="76" pageNumber="77">
<treatmentCitationGroup id="CCAA6540FFDB6A46F28B75B1FBBDFC7A" box="[151,1110,986,1009]" pageId="76" pageNumber="77">
<treatmentCitation id="6D1B647FFFDB6A46F28B75B1FDDDFC7A" author="Iredale, T." box="[151,566,986,1009]" page="28" pageId="76" pageNumber="77" year="1937">
<taxonomicName id="2BBA39EDFFDB6A46F28B75B1FDDDFC7A" ID-CoL="6V55Y" authority="Iredale, 1937 a: 28" authorityName="Iredale" authorityPageNumber="28" authorityYear="1937" box="[151,566,986,1009]" class="Gastropoda" family="Helicarionidae" genus="Periclocystis" kingdom="Animalia" order="Stylommatophora" pageId="76" pageNumber="77" phylum="Mollusca" rank="species" species="ardeni">
<emphasis id="DECE9E7CFFDB6A46F28B75B1FE87FC7A" box="[151,364,986,1009]" italics="true" pageId="76" pageNumber="77">Periclocystis ardeni</emphasis>
<bibRefCitation id="882B3F9FFFDB6A46F36F75B1FDDDFC7A" author="Iredale, T." box="[371,566,986,1009]" pageId="76" pageNumber="77" pagination="6 - 59" refId="ref75409" refString="Iredale, T. (1937 a) An annotated checklist of the land shells of South and Central Australia. South Australian Naturalist, 18, 6 - 59." type="journal article" year="1937">Iredale, 1937a: 28</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, pl. ii, fig. 13;
<treatmentCitation id="6D1B647FFFDB6A46F0CF75B1FC60FC7A" author="Iredale, T." box="[723,907,986,1009]" page="6" pageId="76" pageNumber="77" year="1937">
<bibRefCitation id="882B3F9FFFDB6A46F0CF75B1FC60FC7A" author="Iredale, T." box="[723,907,986,1009]" pageId="76" pageNumber="77" pagination="287 - 333" refId="ref75440" refString="Iredale, T. (1937 b) A basic list of the land Mollusca of Australia. Australian Zoologist, 8, 287 - 333." type="journal article" year="1937">Iredale, 1937b: 6</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="6D1B647FFFDB6A46F18A75B1FBB9FC7A" author="Smith, B. J." box="[918,1106,986,1009]" page="240" pageId="76" pageNumber="77" year="1992">
<bibRefCitation id="882B3F9FFFDB6A46F18A75B1FBB9FC7A" author="Smith, B. J." box="[918,1106,986,1009]" pageId="76" pageNumber="77" pagination="408" refId="ref77207" refString="Smith, B. J. (1992) Non-marine Mollusca. In: Houston, W. W. K. (Ed.) Zoological Catalogue of Australia. Australian Government Publishing Service, Canberra, pp. 408." type="book chapter" year="1992">Smith, 1992: 240</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="A4A011E5FFDB6A46F28B7249FAEAFB3F" pageId="76" pageNumber="77" type="materials_examined">
<paragraph id="EC05426EFFDB6A46F28B7249FC96FBEF" blockId="76.[151,1437,1058,1844]" pageId="76" pageNumber="77">
<emphasis id="DECE9E7CFFDB6A46F28B7249FE66FBB7" bold="true" box="[151,397,1058,1084]" pageId="76" pageNumber="77">Material examined.</emphasis>
<materialsCitation id="5CD24833FFDB6A46F3BD7249FC96FBEF" accessionNumber="C60541" collectionCode="AM" country="Australia" latitude="-31.116667" location="North Flinders Ranges" longLatPrecision="1218" longitude="138.68333" pageId="76" pageNumber="77" specimenCount="1" stateProvince="South Australia" typeStatus="holotype">
<emphasis id="DECE9E7CFFDB6A46F3BD7249FD04FBB7" bold="true" box="[417,751,1058,1084]" pageId="76" pageNumber="77">
<collectingRegion id="2E7E8C8CFFDB6A46F3BD7249FD80FBB7" box="[417,619,1058,1084]" country="Australia" name="South Australia" pageId="76" pageNumber="77">South Australia</collectingRegion>
,
<collectingCountry id="94AD02FEFFDB6A46F0667249FD04FBB7" box="[634,751,1058,1084]" name="Australia" pageId="76" pageNumber="77">Australia</collectingCountry>
</emphasis>
:
<emphasis id="DECE9E7CFFDB6A46F11D7249FC87FBB0" box="[769,876,1058,1083]" italics="true" pageId="76" pageNumber="77">
<typeStatus id="3301FCCCFFDB6A46F11D7249FC87FBB0" box="[769,876,1058,1083]" pageId="76" pageNumber="77" type="holotype">Holotype</typeStatus>
</emphasis>
:
<collectionCode id="8AABDAABFFDB6A46F1627249FC46FBB7" box="[894,941,1058,1084]" country="Australia" name="Australian Museum" pageId="76" pageNumber="77" type="Museum">AM</collectionCode>
C60541 (shell measured, photographed),
<location id="E96514B5FFDB6A46F28B7221FE48FBEF" LSID="urn:lsid:plazi:treatment:6413F378FFDB6A46F28B750CFBF3F8BF:E96514B5FFDB6A46F28B7221FE48FBEF" box="[151,419,1098,1124]" country="Australia" latitude="-31.116667" longLatPrecision="1218" longitude="138.68333" name="North Flinders Ranges" pageId="76" pageNumber="77" stateProvince="South Australia">North Flinders Ranges</location>
, Blinman (
<geoCoordinate id="898E24A9FFDB6A46F0387221FD93FBEF" box="[548,632,1098,1124]" degrees="31" direction="south" minutes="7" orientation="latitude" pageId="76" pageNumber="77" precision="925" value="-31.116667">31º7' S</geoCoordinate>
,
<geoCoordinate id="898E24A9FFDB6A46F0997221FD12FBEF" box="[645,761,1098,1124]" degrees="138" direction="east" minutes="41" orientation="longitude" pageId="76" pageNumber="77" precision="925" value="138.68333">138º41' E</geoCoordinate>
), pre 1937.
</materialsCitation>
</paragraph>
<paragraph id="EC05426EFFDB6A46F2DF7219FAEAFB3F" blockId="76.[151,1437,1058,1844]" pageId="76" pageNumber="77">
<emphasis id="DECE9E7CFFDB6A46F2DF7219FE60FB07" bold="true" box="[195,395,1138,1164]" pageId="76" pageNumber="77">Other material:</emphasis>
<materialsCitation id="5CD24833FFDB6A46F38A7219FB16FB3F" accessionNumber="D16562" collectingDate="1984-08-18" collectionCode="SAM" collectorName="H. Anderson &amp; A. Gackle &amp; K. Gowlett" location="Parachilna Gorge Rd." municipality="Flinders Ranges" pageId="76" pageNumber="77" specimenCount="1">
<collectionCode id="8AABDAABFFDB6A46F38A7219FE3EFB07" box="[406,469,1138,1164]" country="South Africa" httpUri="http://grbio.org/cool/mhew-9czz" name="South African Museum" pageId="76" pageNumber="77">SAM</collectionCode>
D16562 (
<specimenCount id="FABC89E7FFDB6A46F04D7219FCF9FB07" box="[593,786,1138,1164]" pageId="76" pageNumber="77" type="generic">three specimens</specimenCount>
dissected),
<location id="E96514B5FFDB6A46F1BF7219FB43FB07" LSID="urn:lsid:plazi:treatment:6413F378FFDB6A46F28B750CFBF3F8BF:E96514B5FFDB6A46F1BF7219FB43FB07" box="[931,1192,1138,1164]" municipality="Flinders Ranges" name="Parachilna Gorge Rd." pageId="76" pageNumber="77">Parachilna Gorge Rd.</location>
, approx.
<locationDeviation id="FDB75B4CFFDB6A46F7077219FF14FB3F" location="Blinman" pageId="76" pageNumber="77">
<quantity id="2B42EF8BFFDB6A46F7077219FAB2FB07" box="[1307,1369,1138,1164]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="76" pageNumber="77" unit="km" value="9.0">9 km</quantity>
N of Blinman
</locationDeviation>
,
<collectingMunicipality id="0C61D814FFDB6A46F31672F1FE21FB3F" box="[266,458,1178,1204]" pageId="76" pageNumber="77">Flinders Ranges</collectingMunicipality>
, in rock crevices,
<date id="980464AEFFDB6A46F08272F1FCAAFB3F" box="[670,833,1178,1204]" pageId="76" pageNumber="77" value="1984-08-18">
<collectingDate id="88409D46FFDB6A46F08272F1FCAAFB3F" box="[670,833,1178,1204]" pageId="76" pageNumber="77" value="1984-08-18">18 Aug. 1984</collectingDate>
</date>
,
<collectorName id="414F27B8FFDB6A46F15172F1FC0DFB3F" box="[845,998,1178,1204]" pageId="76" pageNumber="77">H. Anderson</collectorName>
,
<collectorName id="414F27B8FFDB6A46F1EE72F1FB80FB3F" box="[1010,1131,1178,1204]" pageId="76" pageNumber="77">A. Gackle</collectorName>
,
<collectorName id="414F27B8FFDB6A46F66B72F1FB16FB3F" box="[1143,1277,1178,1204]" pageId="76" pageNumber="77">K. Gowlett</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A4A011E5FFDB6A46F2DA72A9FE9CF96F" pageId="76" pageNumber="77" type="description">
<paragraph id="EC05426EFFDB6A46F2DA72A9FBDEFAA7" blockId="76.[151,1437,1058,1844]" pageId="76" pageNumber="77">
<emphasis id="DECE9E7CFFDB6A46F2DA72A9FEB5FB57" bold="true" box="[198,350,1218,1244]" pageId="76" pageNumber="77">Description.</emphasis>
<emphasis id="DECE9E7CFFDB6A46F37072A8FD80FB57" box="[364,619,1219,1244]" italics="true" pageId="76" pageNumber="77">External morphology</emphasis>
: Shell (
<figureCitation id="74815EEBFFDB6A46F0CF72A9FC87FB57" box="[723,876,1218,1244]" captionStart="FIGURE 26" captionStartId="71.[151,255,1346,1370]" captionTargetBox="[256,1309,267,1319]" captionTargetId="figure-229@71.[253,1314,250,1328]" captionTargetPageId="71" captionText="FIGURE 26. Shells, dorsal view. A. Westracystis lissus, BMNH 90.12.30.80–4 (S). B. Westracystis lissus, AM C64901 (S of W. tentus). C. Echonitor cyrtochila, BMNH 1922.8.29.16 (H). D. Echonitor albumenoidea, AM C60533 (H). E. Periclocystis ardeni, AM C60541 (H). F. Tarocystis responsivus, AM C104308 (S). G. Conueplecta (Durgellina) calculosa, AM C374873. H. Coneuplecta (Nevelasta) pampini, AM C126763. Scale bars 2 mm (B, G–H), 3 mm (A, C–F). Photographs by M. Allan (F), H. Barlow (B, G–H), D. Beechey (D–E), W. Ponder (A, C). Abbreviations: H, holotype; S, syntype." figureDoi="http://doi.org/10.5281/zenodo.5312780" httpUri="https://zenodo.org/record/5312780/files/figure.png" pageId="76" pageNumber="77">Figures 26E</figureCitation>
,
<figureCitation id="74815EEBFFDB6A46F16372A9FC5BFB57" box="[895,944,1218,1244]" captionStart="FIGURE 27" captionStartId="73.[151,255,1674,1698]" captionTargetBox="[267,1319,605,1658]" captionTargetId="figure-241@73.[267,1320,605,1658]" captionTargetPageId="73" captionText="FIGURE 27. Shells, lateral view. A. Westracystis lissus, BMNH 90.12.30.80–4 (S). B. Westracystis lissus, AM C64901 (S of W. tentus). C. Echonitor cyrtochila, BMNH 1922.8.29.16 (H). D. Echonitor albumenoidea, AM C60533 (H). E. Periclocystis ardeni, AM C60541 (H). F. Tarocystis responsivus, AM C104308 (S). G. Conueplecta (Durgellina) calculosa, AM C374873. H. Coneuplecta (Nevelasta) pampini, AM C126763. Scale bars 2 mm (B, G–H), 3 mm (A, C–F). Photographs by H. Barlow (B, F–H), D. Beechey (D–E), W. Ponder (A, C). Abbreviations: H, holotype; S, syntype." figureDoi="http://doi.org/10.5281/zenodo.5312782" httpUri="https://zenodo.org/record/5312782/files/figure.png" pageId="76" pageNumber="77">27E</figureCitation>
,
<figureCitation id="74815EEBFFDB6A46F1DF72A9FC18FB57" box="[963,1011,1218,1244]" captionStart="FIGURE 28" captionStartId="74.[151,255,1666,1690]" captionTargetBox="[266,1319,585,1640]" captionTargetId="figure-316@74.[248,1332,570,1664]" captionTargetPageId="74" captionText="FIGURE 28. Shells, ventral view. A. Westracystis lissus, BMNH90.12.30.80–4 (S). B. Westracystis lissus, AM C64901 (S of W. tentus). C. Echonitor cyrtochila, BMNH 1922.8.29.16 (H). D. Echonitor albumenoidea, AM C60533 (H). E. Periclocystis ardeni, AM C60541 (H). F. Tarocystis responsivus, AM C104308 (S). G. Conueplecta (Durgellina) calculosa, AM C374873. H. Coneuplecta (Nevelasta) pampini, AM C126763. Scale bars 2 mm (B, G–H), 3 mm (A, C–F). Photographs by H. Barlow (B, F–H), D. Beechey (D–E), W. Ponder (A, C). Abbreviations: H, holotype; S, syntype." figureDoi="http://doi.org/10.5281/zenodo.5312786" httpUri="https://zenodo.org/record/5312786/files/figure.png" pageId="76" pageNumber="77">28E</figureCitation>
) 4.2–4.5 whorls, light brown, not glossy, spire and apex flattened to slightly raised; otherwise shell shape and sculpture as for genus. Mantle laps and lobes and caudal apparatus as for genus; caudal horn moderately long.
</paragraph>
<paragraph id="EC05426EFFDB6A46F2DF7350FB98FADF" blockId="76.[151,1437,1058,1844]" box="[195,1139,1338,1364]" pageId="76" pageNumber="77">
<emphasis id="DECE9E7CFFDB6A46F2DF7350FE88FADF" box="[195,355,1339,1364]" italics="true" pageId="76" pageNumber="77">Mantle cavity</emphasis>
: As for genus. Mantle with pigmentation of black and white spots.
</paragraph>
<paragraph id="EC05426EFFDB6A46F2DF7308FDDFFAF7" blockId="76.[151,1437,1058,1844]" box="[195,564,1378,1404]" pageId="76" pageNumber="77">
<emphasis id="DECE9E7CFFDB6A46F2DF7308FE6CFAF7" box="[195,391,1379,1404]" italics="true" pageId="76" pageNumber="77">Digestive system</emphasis>
: As for genus.
</paragraph>
<paragraph id="EC05426EFFDB6A46F2DF73E0FE27F9E7" blockId="76.[151,1437,1058,1844]" pageId="76" pageNumber="77">
<emphasis id="DECE9E7CFFDB6A46F2DF73E0FE9BFA2F" box="[195,368,1419,1444]" italics="true" pageId="76" pageNumber="77">Genital system</emphasis>
(
<figureCitation id="74815EEBFFDB6A46F36373E1FDFFFA2F" box="[383,532,1418,1444]" captionStart="FIGURE 22" captionStartId="59.[151,255,1798,1822]" captionTargetBox="[308,1282,221,1760]" captionTargetId="figure-127@59.[280,1310,194,1786]" captionTargetPageId="59" captionText="FIGURE 22. Penis interiors. A. Nitor circumcincta, AM C356607. B. Nitor subrugata, AM C334537. C. Pravonitor kreffti, AM C425415. D. Expocystis rustica, AM C425540. E. Malandena suturalis, AM C425085. F. Echonitor cyrtochila, SAM D16557. G. Echonitor albumenoidea, SAM D16567. H. Periclocystis ardeni, SAM D16562. I. New Caledonian NC3, MNHN no registration number. J. New Caledonian NC5, MNHN no registration number. K. Caldwellia angularis, AM C205299. L. Caldwellia imperfecta, AM C205300. Scale bars 1 mm (A–L). Abbreviations: ep, epiphallus; pd, penial diverticulum; pp, penial pilaster; ppd, diamond-shaped, ridged pustules; prm, penis retractor muscle; ppu, penial pustules; pv, penial verge." figureDoi="http://doi.org/10.5281/zenodo.5312770" httpUri="https://zenodo.org/record/5312770/files/figure.png" pageId="76" pageNumber="77">Figures 22H</figureCitation>
,
<figureCitation id="74815EEBFFDB6A46F03C73E1FDBEFA2F" box="[544,597,1418,1444]" captionStart="FIGURE 24" captionStartId="68.[151,255,1783,1807]" captionTargetBox="[305,1289,210,1746]" captionTargetId="figure-140@68.[281,1313,194,1768]" captionTargetPageId="68" captionText="FIGURE 24. Reproductive systems. A. Malandena suturalis, AM C425085. B. Westracystis lissus, FMNH 211717. C. Echonitor cyrtochila, SAM D16557. D. Echonitor albumenoidea, SAM D16567. E. Periclocystis ardeni, SAM D16562. F. Tarocystis responsivus, AM C425327. G. New Caledonian species NC3, MNHN no registration number. H. New Caledonian species NC5, MNHN no registration number. Scale bars 1 mm (C, G), 2 mm (A–B, D–F, H). Abbreviations: alb, albumen gland; bc, bursa copulatrix; c, caecum; ep, epiphallus; fl, flagellum; fov, free oviduct; her, hermaphrodite duct; m, muscle attaching penial sheath to epiphallus (only shown in D &amp; G); ov, ovotestis; p, penis; pr, prostate; prm, penis retractor muscle; pt, penial sheath; t, talon; ut, uterus; v, vagina; vd, vas deferens." figureDoi="http://doi.org/10.5281/zenodo.5312774" httpUri="https://zenodo.org/record/5312774/files/figure.png" pageId="76" pageNumber="77">24E</figureCitation>
): As for genus. Vagina long. Penis long; epiphallus enters penis through simple pore; penis internally covered in three to four longitudinal rows of flat, ridged, diamond-shaped pustules. Penial sheath extremely thick and muscular, thicker than diameter of penis, enclosing only penis; penis retractor muscle attached to epiphallus. Epiphallus approximately equal to penis length, internally with longitudinal pilasters. Diverticulum at junction of epiphallus and vas deferens in form of very short lime-sac (thinwalled, no axial filament).
</paragraph>
<paragraph id="EC05426EFFDB6A46F2DF7010FE3FF91F" blockId="76.[151,1437,1058,1844]" box="[195,468,1658,1684]" pageId="76" pageNumber="77">
<taxonomicName id="2BBA39EDFFDB6A46F2DF7010FEB8F91F" authority=": Not" authorityName="Not" box="[195,339,1658,1684]" class="Jungermanniopsida" family="Radulaceae" genus="Radula" kingdom="Plantae" order="Porellales" pageId="76" pageNumber="77" phylum="Marchantiophyta" rank="genus">
<emphasis id="DECE9E7CFFDB6A46F2DF7010FEFCF91F" box="[195,279,1659,1684]" italics="true" pageId="76" pageNumber="77">Radula</emphasis>
: Not
</taxonomicName>
examined.
</paragraph>
<paragraph id="EC05426EFFDB6A46F2DA70C9FE9CF96F" blockId="76.[151,1437,1058,1844]" pageId="76" pageNumber="77">
<emphasis id="DECE9E7CFFDB6A46F2DA70C9FE5CF937" bold="true" box="[198,439,1698,1724]" pageId="76" pageNumber="77">Range and habitat.</emphasis>
This species is found throughout the Flinders Ranges,
<collectingRegion id="2E7E8C8CFFDB6A46F65470C9FAE8F937" box="[1096,1283,1698,1724]" country="Australia" name="South Australia" pageId="76" pageNumber="77">South Australia</collectingRegion>
, in rock outcrops and crevices.
</paragraph>
</subSubSection>
<subSubSection id="A4A011E5FFDB6A46F2DA7099FBF3F8BF" pageId="76" pageNumber="77" type="discussion">
<paragraph id="EC05426EFFDB6A46F2DA7099FBF3F8BF" blockId="76.[151,1437,1058,1844]" pageId="76" pageNumber="77">
<emphasis id="DECE9E7CFFDB6A46F2DA7099FED4F887" bold="true" box="[198,319,1778,1804]" pageId="76" pageNumber="77">Remarks.</emphasis>
<taxonomicName id="2BBA39EDFFDB6A46F3547098FDD8F887" authorityName="Iredale" authorityYear="1937" box="[328,563,1779,1804]" class="Gastropoda" family="Helicarionidae" genus="Periclocystis" kingdom="Animalia" order="Stylommatophora" pageId="76" pageNumber="77" phylum="Mollusca" rank="species" species="ardeni">
<emphasis id="DECE9E7CFFDB6A46F3547098FDD8F887" box="[328,563,1779,1804]" italics="true" pageId="76" pageNumber="77">Periclocystis ardeni</emphasis>
</taxonomicName>
can be distinguished from members of
<taxonomicName id="2BBA39EDFFDB6A46F6127098FB69F887" authorityName="Iredale" authorityYear="1937" box="[1038,1154,1779,1804]" class="Gastropoda" family="Helicarionidae" genus="Echonitor" kingdom="Animalia" order="Stylommatophora" pageId="76" pageNumber="77" phylum="Mollusca" rank="genus">
<emphasis id="DECE9E7CFFDB6A46F6127098FB69F887" box="[1038,1154,1779,1804]" italics="true" pageId="76" pageNumber="77">Echonitor</emphasis>
</taxonomicName>
, with which it overlaps in range, by its flatter, non-glossy shell with a rapidly descending last whorl.
</paragraph>
</subSubSection>
</treatment>
</document>
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