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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1008.56412" ID-GBIF-Dataset="1543f73a-77cf-483f-8050-92c440d7aa60" ID-PMC="PMC7790805" ID-Pensoft-Pub="1313-2970-1008-139" ID-Pensoft-UUID="82E8DE7E05CD55D5AA7AE498E4F70660" ID-PubMed="33505191" ID-ZooBank="0D4C047F1B794FE59ED4D93C62BCF6A0" ModsDocID="1313-2970-1008-139" checkinTime="1609463494510" checkinUser="pensoft" docAuthor="Lyu, Jing-Cai, Dai, Liang-Liang, Wei, Ping-Fan, He, Yan-Hong, Yuan, Zhi-Yong, Shi, Wen-Li, Zhou, Sheng-Lun, Ran, Si-Yu, Kuang, Zhong-Fan, Guo, Xuan, Wei, Gang &amp; Yuan, Guo" docDate="2020" docId="1A5B45F4AC3B50D489426873D63DE751" docLanguage="en" docName="ZooKeys 1008: 139-157" docOrigin="ZooKeys 1008" docPubDate="2020-12-31" docSource="http://dx.doi.org/10.3897/zookeys.1008.56412" docTitle="Leptobrachella bashaensis Lyu &amp; Dai &amp; Wei &amp; He &amp; Yuan &amp; Shi &amp; Zhou &amp; Ran &amp; Kuang &amp; Guo &amp; Wei &amp; Yuan 2020, sp. nov." docType="treatment" docUuid="6257CDCB-B124-4649-BE10-9A4874C67F20" docUuidSource="ZooBank" docVersion="4" id="82E8DE7E05CD55D5AA7AE498E4F70660" lastPageNumber="139" masterDocId="82E8DE7E05CD55D5AA7AE498E4F70660" masterDocTitle="A new species of the genus Leptobrachella Smith, 1925 (Anura, Megophryidae) from Guizhou, China" masterLastPageNumber="157" masterPageNumber="139" pageNumber="139" updateTime="1668149666256" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A new species of the genus Leptobrachella Smith, 1925 (Anura, Megophryidae) from Guizhou, China</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lyu, Jing-Cai</mods:namePart>
<mods:affiliation>Guizhou Institute of Biology, Guizhou Academy of Sciences, Guiyang, 550002, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dai, Liang-Liang</mods:namePart>
<mods:affiliation>Guizhou Institute of Biology, Guizhou Academy of Sciences, Guiyang, 550002, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wei, Ping-Fan</mods:namePart>
<mods:affiliation>Key Laboratory for Conserving Wildlife with Small Populations in Yunnan, Southwest Forestry University, Kunming 650224, Yunnan, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>He, Yan-Hong</mods:namePart>
<mods:affiliation>Key Laboratory for Conserving Wildlife with Small Populations in Yunnan, Southwest Forestry University, Kunming 650224, Yunnan, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Yuan, Zhi-Yong</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-5991-3021</mods:nameIdentifier>
<mods:affiliation>Key Laboratory for Conserving Wildlife with Small Populations in Yunnan, Southwest Forestry University, Kunming 650224, Yunnan, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Shi, Wen-Li</mods:namePart>
<mods:affiliation>Forestry Bureau of Congjiang County, Congjiang, 557400, Guizhou, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zhou, Sheng-Lun</mods:namePart>
<mods:affiliation>Mt. Foding National Nature Reserve Bureau, shiqian, 555100, Guizhou, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ran, Si-Yu</mods:namePart>
<mods:affiliation>Mt. Foding National Nature Reserve Bureau, shiqian, 555100, Guizhou, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kuang, Zhong-Fan</mods:namePart>
<mods:affiliation>Guizhou Institute of Biology, Guizhou Academy of Sciences, Guiyang, 550002, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Guo, Xuan</mods:namePart>
<mods:affiliation>Guizhou Institute of Biology, Guizhou Academy of Sciences, Guiyang, 550002, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wei, Gang</mods:namePart>
<mods:affiliation>Biodiversity Conservation Key Laboratory, Guiyang College, Guiyang, 550002, China</mods:affiliation>
<mods:nameIdentifier type="email">wg198553@126.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Yuan, Guo</mods:namePart>
<mods:affiliation>Guizhou Institute of Biology, Guizhou Academy of Sciences, Guiyang, 550002, China</mods:affiliation>
<mods:nameIdentifier type="email">875932286@qq.com</mods:nameIdentifier>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2020</mods:date>
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<mods:number>2020-12-31</mods:number>
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<mods:number>1008</mods:number>
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<mods:start>139</mods:start>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1008.56412</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1008.56412</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1008-139</mods:identifier>
<mods:identifier type="ZooBank">0D4C047F1B794FE59ED4D93C62BCF6A0</mods:identifier>
<mods:identifier type="Pensoft-UUID">82E8DE7E05CD55D5AA7AE498E4F70660</mods:identifier>
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<treatment ID-GBIF-Taxon="176352567" LSID="urn:lsid:zoobank.org:act:6257CDCB-B124-4649-BE10-9A4874C67F20" httpUri="http://treatment.plazi.org/id/1A5B45F4AC3B50D489426873D63DE751" lastPageNumber="139" pageId="0" pageNumber="139">
<subSubSection pageId="0" pageNumber="139" type="nomenclature">
<paragraph pageId="0" pageNumber="139">
<taxonomicName LSID="http://zoobank.org/6257CDCB-B124-4649-BE10-9A4874C67F20" authority="Lyu &amp; Dai &amp; Wei &amp; He &amp; Yuan &amp; Shi &amp; Zhou &amp; Ran &amp; Kuang &amp; Guo &amp; Wei &amp; Yuan, 2020" authorityName="Lyu &amp; Dai &amp; Wei &amp; He &amp; Yuan &amp; Shi &amp; Zhou &amp; Ran &amp; Kuang &amp; Guo &amp; Wei &amp; Yuan" authorityYear="2020" class="Amphibia" family="Megophryidae" genus="Leptobrachella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptobrachella bashaensis" order="Anura" pageId="0" pageNumber="139" phylum="Chordata" rank="species" species="bashaensis" status="sp. nov.">Leptobrachella bashaensis</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="139">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Holotype of Leptobrachella bashaensis sp. nov. (GIB 196403) in life A frontolateral view B lateral view C iris coloration D W-shaped marking E dorsal view of thighs F posterioventral view of thighs G ventral view H throat view." figureDoi="10.3897/zookeys.1008.56412.figure3" httpUri="https://binary.pensoft.net/fig/492940" pageId="0" pageNumber="139">Figs 3</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Holotype of Leptobrachella bashaensis sp. nov. (GIB 196403) in life A, B ventral and dorsal view of the hand C, D dorsal and ventral views of the foot." figureDoi="10.3897/zookeys.1008.56412.figure4" httpUri="https://binary.pensoft.net/fig/492941" pageId="0" pageNumber="139">, 4</figureCitation>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Holotype of Leptobrachella bashaensis sp. nov. (GIB 196403) in preservation A dorsal view B ventral view C volar view of hand D plantar view of foot." figureDoi="10.3897/zookeys.1008.56412.figure5" httpUri="https://binary.pensoft.net/fig/492942" pageId="0" pageNumber="139">, 5</figureCitation>
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Tadpole of Leptobrachella bashaensis sp. nov. in preservative A dorsal view B ventral view C dorsal view D labial tooth." figureDoi="10.3897/zookeys.1008.56412.figure6" httpUri="https://binary.pensoft.net/fig/492943" pageId="0" pageNumber="139">, 6</figureCitation>
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Paratypes of Leptobrachella bashaensis sp. nov. in life (GIB 196404)." figureDoi="10.3897/zookeys.1008.56412.figure7" httpUri="https://binary.pensoft.net/fig/492944" pageId="0" pageNumber="139">, 7</figureCitation>
;
<figureCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. List of voucher specimens and GenBank accession numbers for all DNA sequences of the mitochondrial 16 S rRNA gene fragments of Leptobrachella included in this study." pageId="0" pageNumber="139">Tables 1</figureCitation>
,
<figureCitation captionStart="Table 2" captionStartId="T2" captionText="Table 2. Pairwise genetic divergence between Leptobrachella species of monophyletic clades A based on uncorrected p - distance at a 16 S rRNA fragment." pageId="0" pageNumber="139">2</figureCitation>
,
<figureCitation captionStart="Table 3" captionStartId="T3" captionText="Table 3. Morphometric measurements (in mm) of Leptobrachella bashaensis sp. nov. Abbreviations are defined in the text." pageId="0" pageNumber="139">3</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="139" type="type material">
<paragraph pageId="0" pageNumber="139">Type material.</paragraph>
<paragraph pageId="0" pageNumber="139">
<emphasis bold="true" pageId="0" pageNumber="139">
<emphasis italics="true" pageId="0" pageNumber="139">Holotype</emphasis>
.
</emphasis>
GIB196403, adult male, from Basha Nature Reserve, Congjiang, Guizhou Province, China (
<geoCoordinate degrees="25.8534" direction="north" orientation="latitude" precision="5" value="25.8534">25.8534°N</geoCoordinate>
,
<geoCoordinate degrees="108.7356" direction="east" orientation="longitude" precision="5" value="108.7356">108.7356°E</geoCoordinate>
, elevation 900 m. a.s.l.; Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Collection locality (Basha, red star) of Leptobrachella bashaensis sp. nov. specimens from Basha, Guizhou, China used in this study." figureDoi="10.3897/zookeys.1008.56412.figure1" httpUri="https://binary.pensoft.net/fig/492938" pageId="0" pageNumber="139">1</figureCitation>
), collected by Jingcai Lyu and Liangliang Dai on 4 June 2019.
<emphasis bold="true" pageId="0" pageNumber="139">
<emphasis italics="true" pageId="0" pageNumber="139">Paratypes</emphasis>
.
</emphasis>
GIB196401-02, GIB196404, GIB196406-07 (five adult males) collected from the holotype locality by Jingcai Lyu on 4 June 2019.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="139" type="diagnosis">
<paragraph pageId="0" pageNumber="139">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="139">
The new species is assigned to the genus
<taxonomicName authorityName="Smith" authorityYear="1925" class="Amphibia" family="Megophryidae" genus="Leptobrachella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptobrachella" order="Anura" pageId="0" pageNumber="139" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="139">Leptobrachella</emphasis>
</taxonomicName>
on the basis of the following characters: small size, rounded fingertips, presence of an elevated inner palmar tubercle not continuous to the thumb, presence of supra-axillary, femoral and ventrolateral glands, vomerine teeth absent, tubercles on eyelids, and pale vertical bar present on anterior tip of snout (
<bibRefCitation DOI="https://doi.org/10.3406/linly.1980.10444" author="Dubois, A" journalOrPublisher="Bulletin Mensuel de la Societe Linneenne de Lyon" pageId="0" pageNumber="139" pagination="469 - 482" refId="B6" refString="Dubois, A, 1980. Notes sur la systematique et la repartition des amphibiens anoures de Chine et des regions avoisinantes. IV. Classification generique et subgenerique des Pelobatidae Megophryinae. Bulletin Mensuel de la Societe Linneenne de Lyon 49: 469 - 482, DOI: https://doi.org/10.3406/linly.1980.10444" title="Notes sur la systematique et la repartition des amphibiens anoures de Chine et des regions avoisinantes. IV. Classification generique et subgenerique des Pelobatidae Megophryinae." url="https://doi.org/10.3406/linly.1980.10444" volume="49" year="1980">Dubois 1980</bibRefCitation>
,
<bibRefCitation author="Dubois, A" journalOrPublisher="Alytes" pageId="0" pageNumber="139" pagination="147 - 153" refId="B7" refString="Dubois, A, 1983. Note preliminaire sur le genre Leptobrachella Dubois, 1980 (Amphibiens, Anoures), avec diagnose d'uneespeceouvelle du Vietnam. Alytes 2: 147 - 153" title="Note preliminaire sur le genre Leptobrachella Dubois, 1980 (Amphibiens, Anoures), avec diagnose d'uneespeceouvelle du Vietnam." volume="2" year="1983">1983</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.3147.1.1" author="Ohler, A" journalOrPublisher="Zootaxa" pageId="0" pageNumber="139" pagination="1 - 83" refId="B21" refString="Ohler, A, Wollenberg, KC, Grosjean, S, Hendrix, R, Vences, M, Ziegler, T, Dubois, A, 2011. Sorting out Lalos: description of new species and additional taxonomic data on megophryid frogs from northern Indochina (genus Leptobrachella, Megophryidae, Anura). Zootaxa 3147: 1 - 83, DOI: https://doi.org/10.11646/zootaxa.3147.1.1" title="Sorting out Lalos: description of new species and additional taxonomic data on megophryid frogs from northern Indochina (genus Leptobrachella, Megophryidae, Anura)." url="https://doi.org/10.11646/zootaxa.3147.1.1" volume="3147" year="2011">Ohler et al. 2011</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.2198.1.5" author="Rowley, JJL" journalOrPublisher="Zootaxa" pageId="0" pageNumber="139" pagination="51 - 60" refId="B24" refString="Rowley, JJL, Cao, TT, 2009. A new species of Leptobrachella (Anura: Megophryidae) from central Vietnam. Zootaxa 2198: 51 - 60, DOI: https://doi.org/10.11646/zootaxa.2198.1.5" title="A new species of Leptobrachella (Anura: Megophryidae) from central Vietnam." url="https://doi.org/10.11646/zootaxa.2198.1.5" volume="2198" year="2009">Rowley and Cao 2009</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.3737.4.5" author="Rowley, JJL" journalOrPublisher="Zootaxa" pageId="0" pageNumber="139" pagination="415 - 428" refId="B25" refString="Rowley, JJL, Dau, QV, Nguyen, TT, 2013. A new species of Leptobrachella (Anura: Megophryidae) from the highest mountain in Indochina. Zootaxa 3737 (4): 415 - 428, DOI: https://doi.org/10.11646/zootaxa.3737.4.5" title="A new species of Leptobrachella (Anura: Megophryidae) from the highest mountain in Indochina." url="https://doi.org/10.11646/zootaxa.3737.4.5" volume="3737" year="2013">Rowley et al. 2013</bibRefCitation>
).
<taxonomicName authorityName="Lyu &amp; Dai &amp; Wei &amp; He &amp; Yuan &amp; Shi &amp; Zhou &amp; Ran &amp; Kuang &amp; Guo &amp; Wei &amp; Yuan" authorityYear="2020" class="Amphibia" family="Megophryidae" genus="Leptobrachella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptobrachella bashaensis" order="Anura" pageId="0" pageNumber="139" phylum="Chordata" rank="species" species="bashaensis">
<emphasis italics="true" pageId="0" pageNumber="139">Leptobrachella bashaensis</emphasis>
</taxonomicName>
sp. nov. is distinguished from its congeners by a combination of the following morphological characters: (1) small size (SVL 22.9-25.6 mm in six adult males and 27.1 mm in one adult female), (2) head longer than wide, (3) externally distinct tympanum, (4) dorsal skin slightly shagreened with small tubercles and irregular brown stripes, (5) distinct dark spots on the flank, (6) creamy-white chest and off-white belly with irregular black spots, (7) grey-pinkish to dark brownish-violet ventral skin of limbs with numerous whitish speckles, (8) distinct ventrolateral glands, forming a distinct white line, (9) finger webbing and fringes absent, (10) toe webbing rudimentary and lateral fringes narrow, (11) longitudinal ridges under toes and not interrupted at the articulations, (12) a distinctly bicolored iris, typically bright orange in upper half, fading to silver in lower half.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="139" type="description">
<paragraph pageId="0" pageNumber="139">Description of the holotype.</paragraph>
<paragraph pageId="0" pageNumber="139">
Adult male. SVL 24.0 mm; head slightly longer than wide (HDL/HDW = 1.10), rectangular in dorsal view (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Holotype of Leptobrachella bashaensis sp. nov. (GIB 196403) in life A frontolateral view B lateral view C iris coloration D W-shaped marking E dorsal view of thighs F posterioventral view of thighs G ventral view H throat view." figureDoi="10.3897/zookeys.1008.56412.figure3" httpUri="https://binary.pensoft.net/fig/492940" pageId="0" pageNumber="139">3A, B</figureCitation>
); snout rounded in both ventral view and lateral view, protruding slightly beyond lower jaw (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Holotype of Leptobrachella bashaensis sp. nov. (GIB 196403) in life A frontolateral view B lateral view C iris coloration D W-shaped marking E dorsal view of thighs F posterioventral view of thighs G ventral view H throat view." figureDoi="10.3897/zookeys.1008.56412.figure3" httpUri="https://binary.pensoft.net/fig/492940" pageId="0" pageNumber="139">3D</figureCitation>
); nostril oval, located closer to tip of snout than to eye; loreal region oblique; canthus rostralis distinct; eye large (EYE/HDL = 0.35), diameter slightly smaller than snout length (EYE/SNT = 0.88), notably protuberant in both dorsal and lateral view; pupil vertical; tympanum distinct, rounded, diameter smaller than that of eye (TMP/EYE = 0.60) (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Holotype of Leptobrachella bashaensis sp. nov. (GIB 196403) in life A frontolateral view B lateral view C iris coloration D W-shaped marking E dorsal view of thighs F posterioventral view of thighs G ventral view H throat view." figureDoi="10.3897/zookeys.1008.56412.figure3" httpUri="https://binary.pensoft.net/fig/492940" pageId="0" pageNumber="139">3C</figureCitation>
); vomerine teeth absent; vocal sac openings slit-like, located posterolateral on floor of mouth; tongue long and moderately wide, with a shallow notch at posterior tip; supratympanic ridge distinct, running from eye towards axillary with raised tubercles. Forelimb long and slender, fingers long and slender, without webbing and lateral fringes; relative length of fingers II&lt;I&lt;IV&lt;III; tips of fingers rounded and slightly swollen; nuptial pad absent; subarticular tubercles absent in fingers; inner metacarpal tubercle large and rounded, separated from laterally compressed and much smaller outer metacarpal tubercle (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Holotype of Leptobrachella bashaensis sp. nov. (GIB 196403) in life A, B ventral and dorsal view of the hand C, D dorsal and ventral views of the foot." figureDoi="10.3897/zookeys.1008.56412.figure4" httpUri="https://binary.pensoft.net/fig/492941" pageId="0" pageNumber="139">4A, B</figureCitation>
). Hindlimb moderately long; tibia half of snout-vent length (TIB/SVL = 0.50); tibiotarsal articulation of adpressed limb reaching snout, well beyond anterior margin of eye, but not beyond snout tip; relative toe lengths I&lt;II&lt;V&lt;III&lt;IV; toe tips rounded and thickened; greatly reduced basal webbing present between all five toes; narrow lateral fringes present only on II and III toes (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Holotype of Leptobrachella bashaensis sp. nov. (GIB 196403) in life A, B ventral and dorsal view of the hand C, D dorsal and ventral views of the foot." figureDoi="10.3897/zookeys.1008.56412.figure4" httpUri="https://binary.pensoft.net/fig/492941" pageId="0" pageNumber="139">4C, D</figureCitation>
); subarticular tubercles hardly discernible under toes II and III; dorsal skin slightly rough with small tubercles and irregular pustules; ventral skin smooth; oval supra-axillary gland present at forelimb base on ventral surface of axillary region (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Holotype of Leptobrachella bashaensis sp. nov. (GIB 196403) in life A frontolateral view B lateral view C iris coloration D W-shaped marking E dorsal view of thighs F posterioventral view of thighs G ventral view H throat view." figureDoi="10.3897/zookeys.1008.56412.figure3" httpUri="https://binary.pensoft.net/fig/492940" pageId="0" pageNumber="139">3F</figureCitation>
); oval femoral glands distinct on posteroventral surface of thigh, closer to knee than to vent; ventrolateral glands forming a distinct line on flanks.
</paragraph>
<caption doi="10.3897/zookeys.1008.56412.figure3" httpUri="https://binary.pensoft.net/fig/492940" pageId="0" pageNumber="139" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="139">
<emphasis bold="true" pageId="0" pageNumber="139">Figure 3.</emphasis>
Holotype of
<taxonomicName authorityName="Lyu &amp; Dai &amp; Wei &amp; He &amp; Yuan &amp; Shi &amp; Zhou &amp; Ran &amp; Kuang &amp; Guo &amp; Wei &amp; Yuan" authorityYear="2020" class="Amphibia" family="Megophryidae" genus="Leptobrachella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptobrachella bashaensis" order="Anura" pageId="0" pageNumber="139" phylum="Chordata" rank="species" species="bashaensis">
<emphasis italics="true" pageId="0" pageNumber="139">Leptobrachella bashaensis</emphasis>
</taxonomicName>
sp. nov. (GIB196403) in life
<emphasis bold="true" pageId="0" pageNumber="139">A</emphasis>
frontolateral view
<emphasis bold="true" pageId="0" pageNumber="139">B</emphasis>
lateral view
<emphasis bold="true" pageId="0" pageNumber="139">C</emphasis>
iris coloration
<emphasis bold="true" pageId="0" pageNumber="139">D</emphasis>
W-shaped marking
<emphasis bold="true" pageId="0" pageNumber="139">E</emphasis>
dorsal view of thighs
<emphasis bold="true" pageId="0" pageNumber="139">F</emphasis>
posterioventral view of thighs
<emphasis bold="true" pageId="0" pageNumber="139">G</emphasis>
ventral view
<emphasis bold="true" pageId="0" pageNumber="139">H</emphasis>
throat view.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1008.56412.figure4" httpUri="https://binary.pensoft.net/fig/492941" pageId="0" pageNumber="139" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="139">
<emphasis bold="true" pageId="0" pageNumber="139">Figure 4.</emphasis>
Holotype of
<taxonomicName authorityName="Lyu &amp; Dai &amp; Wei &amp; He &amp; Yuan &amp; Shi &amp; Zhou &amp; Ran &amp; Kuang &amp; Guo &amp; Wei &amp; Yuan" authorityYear="2020" class="Amphibia" family="Megophryidae" genus="Leptobrachella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptobrachella bashaensis" order="Anura" pageId="0" pageNumber="139" phylum="Chordata" rank="species" species="bashaensis">
<emphasis italics="true" pageId="0" pageNumber="139">Leptobrachella bashaensis</emphasis>
</taxonomicName>
sp. nov. (GIB196403) in life
<emphasis bold="true" pageId="0" pageNumber="139">A, B</emphasis>
ventral and dorsal view of the hand
<emphasis bold="true" pageId="0" pageNumber="139">C, D</emphasis>
dorsal and ventral views of the foot.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="139" type="coloration of holotype in life">
<paragraph pageId="0" pageNumber="139">Coloration of holotype in life.</paragraph>
<paragraph pageId="0" pageNumber="139">
Dorsal surface of head and trunk brown with small, dark brown, irregularly shaped markings; large reverse-triangle dark brown marking between eyes, connected to the W-shaped mark between axillae (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Holotype of Leptobrachella bashaensis sp. nov. (GIB 196403) in life A frontolateral view B lateral view C iris coloration D W-shaped marking E dorsal view of thighs F posterioventral view of thighs G ventral view H throat view." figureDoi="10.3897/zookeys.1008.56412.figure3" httpUri="https://binary.pensoft.net/fig/492940" pageId="0" pageNumber="139">3</figureCitation>
); supratympanic ridge present; elbow to upper arm reddish in color; fine, distinct reddish tubercles scattered on upper eyelids, dorsal surfaces of head, body, and limbs. Ventral surface of throat and belly off-white, with faint spots; chest creamy white, with irregular black spots (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Holotype of Leptobrachella bashaensis sp. nov. (GIB 196403) in life A frontolateral view B lateral view C iris coloration D W-shaped marking E dorsal view of thighs F posterioventral view of thighs G ventral view H throat view." figureDoi="10.3897/zookeys.1008.56412.figure3" httpUri="https://binary.pensoft.net/fig/492940" pageId="0" pageNumber="139">3G, H</figureCitation>
); ventral surfaces of lip and limbs covered with irregular white speckles; ventral surface of thighs grey-brown with white spots. Supra-axillary gland, ventrolateral glands, and femoral glands white. Iris bicolored: bright orange in upper half, silver in lower half, with black reticulations throughout. Fine, faint transverse dark brown bars on dorsal surface of fingers and toes, lower arms, tarsus, thighs, and tibia (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Holotype of Leptobrachella bashaensis sp. nov. (GIB 196403) in life A, B ventral and dorsal view of the hand C, D dorsal and ventral views of the foot." figureDoi="10.3897/zookeys.1008.56412.figure4" httpUri="https://binary.pensoft.net/fig/492941" pageId="0" pageNumber="139">4</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="139" type="coloration of holotype in preservation">
<paragraph pageId="0" pageNumber="139">Coloration of holotype in preservation.</paragraph>
<paragraph pageId="0" pageNumber="139">
In preservative (75% ethanol), dorsal surface dark grey-brown, and ventral and lateral white glands not evident (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Holotype of Leptobrachella bashaensis sp. nov. (GIB 196403) in preservation A dorsal view B ventral view C volar view of hand D plantar view of foot." figureDoi="10.3897/zookeys.1008.56412.figure5" httpUri="https://binary.pensoft.net/fig/492942" pageId="0" pageNumber="139">5</figureCitation>
); ventral surface of throat, chest, belly, and interior portions of arms fade to creamy white, ventral surface of thighs dark brown with white spots; bars and blotches on dorsum and limbs are dark brown and less apparent. The color of the tympanum fades to brown.
</paragraph>
<caption doi="10.3897/zookeys.1008.56412.figure5" httpUri="https://binary.pensoft.net/fig/492942" pageId="0" pageNumber="139" start="Figure 5" startId="F5">
<paragraph pageId="0" pageNumber="139">
<emphasis bold="true" pageId="0" pageNumber="139">Figure 5.</emphasis>
Holotype of
<taxonomicName authorityName="Lyu &amp; Dai &amp; Wei &amp; He &amp; Yuan &amp; Shi &amp; Zhou &amp; Ran &amp; Kuang &amp; Guo &amp; Wei &amp; Yuan" authorityYear="2020" class="Amphibia" family="Megophryidae" genus="Leptobrachella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptobrachella bashaensis" order="Anura" pageId="0" pageNumber="139" phylum="Chordata" rank="species" species="bashaensis">
<emphasis italics="true" pageId="0" pageNumber="139">Leptobrachella bashaensis</emphasis>
</taxonomicName>
sp. nov. (GIB196403) in preservation
<emphasis bold="true" pageId="0" pageNumber="139">A</emphasis>
dorsal view
<emphasis bold="true" pageId="0" pageNumber="139">B</emphasis>
ventral view
<emphasis bold="true" pageId="0" pageNumber="139">C</emphasis>
volar view of hand
<emphasis bold="true" pageId="0" pageNumber="139">D</emphasis>
plantar view of foot.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="139" type="tadpoles">
<paragraph pageId="0" pageNumber="139">Tadpoles.</paragraph>
<paragraph pageId="0" pageNumber="139">
Brown, narrow; BH 4.4 mm; BL 14.1 mm; BW 6.2; ED 2.0 mm; TH 3.8; SS 4.9 mm; TMW 3.4 mm; TL 29.3 mm; TOL 43.8 mm. TAL/TOL = 0.67, with a I:3+3/1+1:II labial tooth row formula (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Tadpole of Leptobrachella bashaensis sp. nov. in preservative A dorsal view B ventral view C dorsal view D labial tooth." figureDoi="10.3897/zookeys.1008.56412.figure6" httpUri="https://binary.pensoft.net/fig/492943" pageId="0" pageNumber="139">6</figureCitation>
). In life, dorsal surface of head dark brown with small, brown, irregularly shaped spot; air sac-shaped bulges on both sides of the body; upper lip and lower lip nearly round shape. Tadpoles were collected in the field from a stream surveyed on 16 October 2017 in Basha Nature Reserve by Lyu.
</paragraph>
<caption doi="10.3897/zookeys.1008.56412.figure6" httpUri="https://binary.pensoft.net/fig/492943" pageId="0" pageNumber="139" start="Figure 6" startId="F6">
<paragraph pageId="0" pageNumber="139">
<emphasis bold="true" pageId="0" pageNumber="139">Figure 6.</emphasis>
Tadpole of
<taxonomicName authorityName="Lyu &amp; Dai &amp; Wei &amp; He &amp; Yuan &amp; Shi &amp; Zhou &amp; Ran &amp; Kuang &amp; Guo &amp; Wei &amp; Yuan" authorityYear="2020" class="Amphibia" family="Megophryidae" genus="Leptobrachella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptobrachella bashaensis" order="Anura" pageId="0" pageNumber="139" phylum="Chordata" rank="species" species="bashaensis">
<emphasis italics="true" pageId="0" pageNumber="139">Leptobrachella bashaensis</emphasis>
</taxonomicName>
sp. nov. in preservative
<emphasis bold="true" pageId="0" pageNumber="139">A</emphasis>
dorsal view
<emphasis bold="true" pageId="0" pageNumber="139">B</emphasis>
ventral view
<emphasis bold="true" pageId="0" pageNumber="139">C</emphasis>
dorsal view
<emphasis bold="true" pageId="0" pageNumber="139">D</emphasis>
labial tooth.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="139" type="etymology">
<paragraph pageId="0" pageNumber="139">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="139">
The specific epithet, &quot;
<taxonomicName authorityName="Lyu &amp; Dai &amp; Wei &amp; He &amp; Yuan &amp; Shi &amp; Zhou &amp; Ran &amp; Kuang &amp; Guo &amp; Wei &amp; Yuan" authorityYear="2020" class="Amphibia" family="Megophryidae" genus="Leptobrachella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptobrachella bashaensis" order="Anura" pageId="0" pageNumber="139" phylum="Chordata" rank="species" species="bashaensis">
<emphasis italics="true" pageId="0" pageNumber="139">Leptobrachella bashaensis</emphasis>
</taxonomicName>
&quot;/Basha Zhang Tu Chan (岜沙掌突蟾 in Chinese), refers to the location where the specimens were collected, Basha Nature Reserve of Guizhou Province, China.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="139" type="morphological variation">
<paragraph pageId="0" pageNumber="139">Morphological variation.</paragraph>
<paragraph pageId="0" pageNumber="139">
Variation in size and body proportions of the type series are shown in Table
<tableCitation captionStart="Table 3" captionStartId="T3" captionText="Table 3. Morphometric measurements (in mm) of Leptobrachella bashaensis sp. nov. Abbreviations are defined in the text." httpUri="http://table.plazi.org/id/7C5052AC3F573E98A1AB66AC680B10D0" pageId="0" pageNumber="139" tableUuid="7C5052AC3F573E98A1AB66AC680B10D0">3</tableCitation>
. Representative photographs of paratypes in life are shown in Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Paratypes of Leptobrachella bashaensis sp. nov. in life (GIB 196404)." figureDoi="10.3897/zookeys.1008.56412.figure7" httpUri="https://binary.pensoft.net/fig/492944" pageId="0" pageNumber="139">7</figureCitation>
. In life, the dorsal surface of the head and trunk varies from olive-brown to reddish; the ventral surface of the lower jaw among these individuals varies from cream-yellow, pink, to grey; the ventral surface of chest and belly is grey-pink to white. There is variation among individuals in the shape of tubercles, pustules, black ventrolateral blotches, and dark stripes, bars on the dorsum or dorsal surface of limbs. The W-shaped marking between axillae is distinct in all individuals.
</paragraph>
<caption doi="10.3897/zookeys.1008.56412.figure7" httpUri="https://binary.pensoft.net/fig/492944" pageId="0" pageNumber="139" start="Figure 7" startId="F7">
<paragraph pageId="0" pageNumber="139">
<emphasis bold="true" pageId="0" pageNumber="139">Figure 7.</emphasis>
Paratypes of
<taxonomicName authorityName="Lyu &amp; Dai &amp; Wei &amp; He &amp; Yuan &amp; Shi &amp; Zhou &amp; Ran &amp; Kuang &amp; Guo &amp; Wei &amp; Yuan" authorityYear="2020" class="Amphibia" family="Megophryidae" genus="Leptobrachella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptobrachella bashaensis" order="Anura" pageId="0" pageNumber="139" phylum="Chordata" rank="species" species="bashaensis">
<emphasis italics="true" pageId="0" pageNumber="139">Leptobrachella bashaensis</emphasis>
</taxonomicName>
sp. nov. in life (GIB196404).
</paragraph>
</caption>
<caption ID-Table-UUID="7C5052AC3F573E98A1AB66AC680B10D0" httpUri="http://table.plazi.org/id/7C5052AC3F573E98A1AB66AC680B10D0" pageId="0" pageNumber="139" start="Table 3" startId="T3">
<paragraph pageId="0" pageNumber="139">
<emphasis bold="true" pageId="0" pageNumber="139">Table 3.</emphasis>
Morphometric measurements (in mm) of
<taxonomicName authorityName="Lyu &amp; Dai &amp; Wei &amp; He &amp; Yuan &amp; Shi &amp; Zhou &amp; Ran &amp; Kuang &amp; Guo &amp; Wei &amp; Yuan" authorityYear="2020" class="Amphibia" family="Megophryidae" genus="Leptobrachella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptobrachella bashaensis" order="Anura" pageId="0" pageNumber="139" phylum="Chordata" rank="species" species="bashaensis">
<emphasis italics="true" pageId="0" pageNumber="139">Leptobrachella bashaensis</emphasis>
</taxonomicName>
sp. nov. Abbreviations are defined in the text.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="139">
<table pageId="0" pageNumber="139">
<tr pageId="0" pageNumber="139">
<th colspan="1" pageId="0" pageNumber="139" rowspan="1">Voucher number</th>
<th colspan="1" pageId="0" pageNumber="139" rowspan="1">GIB196401</th>
<th colspan="1" pageId="0" pageNumber="139" rowspan="1">GIB196402</th>
<th colspan="1" pageId="0" pageNumber="139" rowspan="1">GIB196403</th>
<th colspan="1" pageId="0" pageNumber="139" rowspan="1">GIB196404</th>
<th colspan="1" pageId="0" pageNumber="139" rowspan="1">GIB196406</th>
<th colspan="1" pageId="0" pageNumber="139" rowspan="1">GIB196407</th>
<th colspan="1" pageId="0" pageNumber="139" rowspan="1">GIB196405</th>
</tr>
<tr pageId="0" pageNumber="139">
<th colspan="1" pageId="0" pageNumber="139" rowspan="1">Sex</th>
<th colspan="1" pageId="0" pageNumber="139" rowspan="1">Male</th>
<th colspan="1" pageId="0" pageNumber="139" rowspan="1">Male</th>
<th colspan="1" pageId="0" pageNumber="139" rowspan="1">Male</th>
<th colspan="1" pageId="0" pageNumber="139" rowspan="1">Male</th>
<th colspan="1" pageId="0" pageNumber="139" rowspan="1">Male</th>
<th colspan="1" pageId="0" pageNumber="139" rowspan="1">Male</th>
<th colspan="1" pageId="0" pageNumber="139" rowspan="1">Female</th>
</tr>
<tr pageId="0" pageNumber="139">
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">SVL</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">22.9</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">23.0</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">24.0</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">24.6</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">25.6</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">24.2</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">27.1</td>
</tr>
<tr pageId="0" pageNumber="139">
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">HDL</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">8.4</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">8.3</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">9.4</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">8.7</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">9.4</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">9.1</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">9.6</td>
</tr>
<tr pageId="0" pageNumber="139">
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">HDW</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">8.0</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">7.5</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">8.7</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">8.1</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">8.2</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">8.0</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">8.2</td>
</tr>
<tr pageId="0" pageNumber="139">
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">SNT</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">3.3</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">3.4</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">3.5</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">3.5</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">4.0</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">3.8</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">3.8</td>
</tr>
<tr pageId="0" pageNumber="139">
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">IND</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">2.5</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">2.4</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">2.4</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">2.7</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">2.7</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">2.5</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">2.9</td>
</tr>
<tr pageId="0" pageNumber="139">
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">IOD</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">3.2</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">3.5</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">3.3</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">3.3</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">3.3</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">3.2</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">3.3</td>
</tr>
<tr pageId="0" pageNumber="139">
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">UEW</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">2.3</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">2.1</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">2.2</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">2.2</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">2.3</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">2.5</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">2.3</td>
</tr>
<tr pageId="0" pageNumber="139">
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">EYE</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">2.8</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">2.7</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">2.9</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">3.1</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">3.5</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">3.3</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">3.6</td>
</tr>
<tr pageId="0" pageNumber="139">
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">NEL</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">1.5</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">1.6</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">1.8</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">2.0</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">1.9</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">1.8</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">2.0</td>
</tr>
<tr pageId="0" pageNumber="139">
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">NSL</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">1.9</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">1.8</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">1.7</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">1.7</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">1.9</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">1.6</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">1.9</td>
</tr>
<tr pageId="0" pageNumber="139">
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">TEY</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">0.7</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">0.9</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">1.1</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">0.9</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">1.0</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">1.0</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">1.1</td>
</tr>
<tr pageId="0" pageNumber="139">
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">ML</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">6.2</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">5.6</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">6.1</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">6.0</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">6.0</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">6.3</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">6.3</td>
</tr>
<tr pageId="0" pageNumber="139">
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">TMP</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">2.1</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">1.6</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">2.2</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">1.9</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">1.9</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">1.7</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">1.9</td>
</tr>
<tr pageId="0" pageNumber="139">
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">LAHL</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">11.7</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">10.4</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">10.8</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">10.9</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">11.5</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">11.6</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">11.9</td>
</tr>
<tr pageId="0" pageNumber="139">
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">HLL</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">37.4</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">37.6</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">40.7</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">40.3</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">38.8</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">39.7</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">41.3</td>
</tr>
<tr pageId="0" pageNumber="139">
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">TIB</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">12.1</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">11.4</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">12.6</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">12.3</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">12.1</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">13.2</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">12.9</td>
</tr>
<tr pageId="0" pageNumber="139">
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">FOT</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">11.0</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">10.9</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">11.2</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">11.3</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">10.6</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">11.6</td>
<td colspan="1" pageId="0" pageNumber="139" rowspan="1">11.8</td>
</tr>
</table>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="139" type="natural history">
<paragraph pageId="0" pageNumber="139">Natural history.</paragraph>
<paragraph pageId="0" pageNumber="139">
All specimens were collected at night in small streams in Basha Nature Reserve approximately 900 m elevation (Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Habitat at the type locality of Leptobrachella bashaensis sp. nov., Basha Nature Reserve, Congjiang, Guizhou province, China." figureDoi="10.3897/zookeys.1008.56412.figure8" httpUri="https://binary.pensoft.net/fig/492945" pageId="0" pageNumber="139">8</figureCitation>
). Calling males were found along the streams, perching on large rocks, in rocky crevices, or under dead wood. Insect-like calls could be heard in June. The breeding season of this species is likely to occur from June to July, as females collected during these months were gravid, and males were heard calling only from June to the beginning of July. During both surveys, the number of males observed was much greater than females (males:females = 12:1).
</paragraph>
<caption doi="10.3897/zookeys.1008.56412.figure8" httpUri="https://binary.pensoft.net/fig/492945" pageId="0" pageNumber="139" start="Figure 8" startId="F8">
<paragraph pageId="0" pageNumber="139">
<emphasis bold="true" pageId="0" pageNumber="139">Figure 8.</emphasis>
Habitat at the type locality of
<taxonomicName authorityName="Lyu &amp; Dai &amp; Wei &amp; He &amp; Yuan &amp; Shi &amp; Zhou &amp; Ran &amp; Kuang &amp; Guo &amp; Wei &amp; Yuan" authorityYear="2020" class="Amphibia" family="Megophryidae" genus="Leptobrachella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptobrachella bashaensis" order="Anura" pageId="0" pageNumber="139" phylum="Chordata" rank="species" species="bashaensis">
<emphasis italics="true" pageId="0" pageNumber="139">Leptobrachella bashaensis</emphasis>
</taxonomicName>
sp. nov., Basha Nature Reserve, Congjiang, Guizhou province, China.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="139" type="comparisons">
<paragraph pageId="0" pageNumber="139">Comparisons.</paragraph>
<paragraph pageId="0" pageNumber="139">
<taxonomicName authorityName="Lyu &amp; Dai &amp; Wei &amp; He &amp; Yuan &amp; Shi &amp; Zhou &amp; Ran &amp; Kuang &amp; Guo &amp; Wei &amp; Yuan" authorityYear="2020" class="Amphibia" family="Megophryidae" genus="Leptobrachella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptobrachella bashaensis" order="Anura" pageId="0" pageNumber="139" phylum="Chordata" rank="species" species="bashaensis">
<emphasis italics="true" pageId="0" pageNumber="139">Leptobrachella bashaensis</emphasis>
</taxonomicName>
sp. nov. differs from all known
<taxonomicName authorityName="Smith" authorityYear="1925" class="Amphibia" family="Megophryidae" genus="Leptobrachella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptobrachella" order="Anura" pageId="0" pageNumber="139" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="139">Leptobrachella</emphasis>
</taxonomicName>
species distributed north of the Isthmus of Kra by a combination of male body size, externally distinct tympanum, and black spots on the flank, plus ventral coloration, degree of webbing and fringing on the toes, iris coloration, and dorsal skin texture.
</paragraph>
<paragraph pageId="0" pageNumber="139">
Care was taken to differentiate
<taxonomicName authorityName="Lyu &amp; Dai &amp; Wei &amp; He &amp; Yuan &amp; Shi &amp; Zhou &amp; Ran &amp; Kuang &amp; Guo &amp; Wei &amp; Yuan" authorityYear="2020" class="Amphibia" family="Megophryidae" genus="Leptobrachella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptobrachella bashaensis" order="Anura" pageId="0" pageNumber="139" phylum="Chordata" rank="species" species="bashaensis">
<emphasis italics="true" pageId="0" pageNumber="139">Leptobrachella bashaensis</emphasis>
</taxonomicName>
sp. nov. from the five most morphologically and molecularly similar species,
<taxonomicName lsidName="L. maoershanensis" pageId="0" pageNumber="139" rank="species" species="maoershanensis">
<emphasis italics="true" pageId="0" pageNumber="139">L. maoershanensis</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="L. laui" pageId="0" pageNumber="139" rank="species" species="laui">
<emphasis italics="true" pageId="0" pageNumber="139">L. laui</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="L. mangshanensis" pageId="0" pageNumber="139" rank="species" species="mangshanensis">
<emphasis italics="true" pageId="0" pageNumber="139">L. mangshanensis</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="L. yunkaiensis" pageId="0" pageNumber="139" rank="species" species="yunkaiensis">
<emphasis italics="true" pageId="0" pageNumber="139">L. yunkaiensis</emphasis>
</taxonomicName>
, and
<taxonomicName lsidName="L. liui" pageId="0" pageNumber="139" rank="species" species="liui">
<emphasis italics="true" pageId="0" pageNumber="139">L. liui</emphasis>
</taxonomicName>
from China.
</paragraph>
<paragraph pageId="0" pageNumber="139">
<taxonomicName authorityName="Lyu &amp; Dai &amp; Wei &amp; He &amp; Yuan &amp; Shi &amp; Zhou &amp; Ran &amp; Kuang &amp; Guo &amp; Wei &amp; Yuan" authorityYear="2020" class="Amphibia" family="Megophryidae" genus="Leptobrachella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptobrachella bashaensis" order="Anura" pageId="0" pageNumber="139" phylum="Chordata" rank="species" species="bashaensis">
<emphasis italics="true" pageId="0" pageNumber="139">Leptobrachella bashaensis</emphasis>
</taxonomicName>
sp. nov. differs from its sister taxon
<taxonomicName lsidName="L. maoershanensis" pageId="0" pageNumber="139" rank="species" species="maoershanensis">
<emphasis italics="true" pageId="0" pageNumber="139">L. maoershanensis</emphasis>
</taxonomicName>
by the following: smaller body size (SVL 22.9-25.6 mm in males, 27.1 mm in female vs 25.2-30.4 mm in males, 28.6 mm in female); bicolored iris with bright orange in upper half, fading to silver in lower half (vs typically bright orange-red in upper half, fading to silver in lower half); vertical and distinct narrow pupil (vs vertical and wide pupil); narrow lateral fringes present only on II and III toes (vs narrow lateral fringes present on all toes).
</paragraph>
<paragraph pageId="0" pageNumber="139">
<taxonomicName authorityName="Lyu &amp; Dai &amp; Wei &amp; He &amp; Yuan &amp; Shi &amp; Zhou &amp; Ran &amp; Kuang &amp; Guo &amp; Wei &amp; Yuan" authorityYear="2020" class="Amphibia" family="Megophryidae" genus="Leptobrachella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptobrachella bashaensis" order="Anura" pageId="0" pageNumber="139" phylum="Chordata" rank="species" species="bashaensis">
<emphasis italics="true" pageId="0" pageNumber="139">Leptobrachella bashaensis</emphasis>
</taxonomicName>
sp. nov. differs from
<taxonomicName lsidName="L. laui" pageId="0" pageNumber="139" rank="species" species="laui">
<emphasis italics="true" pageId="0" pageNumber="139">L. laui</emphasis>
</taxonomicName>
in having slightly a smaller body size (SVL 22.9-25.6 mm in males, 27.1 mm in female vs 24.8-26.7 mm in males, 28.1 mm in female); bicolored iris with bright orange in upper half, fading to silver in lower half (vs uniformly coppery orange with fine black reticulations throughout); vertical and distinct narrow pupil (vs vertical and wide pupil); narrow lateral fringes present only on II and III toes (vs wide lateral fringes present on all toes); distinct black spots on the flank (vs indistinct black spots on the flank).
</paragraph>
<paragraph pageId="0" pageNumber="139">
<taxonomicName authorityName="Lyu &amp; Dai &amp; Wei &amp; He &amp; Yuan &amp; Shi &amp; Zhou &amp; Ran &amp; Kuang &amp; Guo &amp; Wei &amp; Yuan" authorityYear="2020" class="Amphibia" family="Megophryidae" genus="Leptobrachella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptobrachella bashaensis" order="Anura" pageId="0" pageNumber="139" phylum="Chordata" rank="species" species="bashaensis">
<emphasis italics="true" pageId="0" pageNumber="139">Leptobrachella bashaensis</emphasis>
</taxonomicName>
sp. nov. differs from
<taxonomicName lsidName="L. mangshanensis" pageId="0" pageNumber="139" rank="species" species="mangshanensis">
<emphasis italics="true" pageId="0" pageNumber="139">L. mangshanensis</emphasis>
</taxonomicName>
by having slightly smaller body size (SVL 22.9-25.6 mm in males, 27.1 mm in female vs 22.2-27.8 mm in males, 30.2 mm in female); bicolored iris with bright orange in upper half, fading to silver in lower half (vs bright orange in upper half, greyish cream in lower half); narrow lateral fringes present only on II and III toes (vs weak lateral fringes on toes).
</paragraph>
<paragraph pageId="0" pageNumber="139">
<taxonomicName authorityName="Lyu &amp; Dai &amp; Wei &amp; He &amp; Yuan &amp; Shi &amp; Zhou &amp; Ran &amp; Kuang &amp; Guo &amp; Wei &amp; Yuan" authorityYear="2020" class="Amphibia" family="Megophryidae" genus="Leptobrachella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptobrachella bashaensis" order="Anura" pageId="0" pageNumber="139" phylum="Chordata" rank="species" species="bashaensis">
<emphasis italics="true" pageId="0" pageNumber="139">Leptobrachella bashaensis</emphasis>
</taxonomicName>
sp. nov. differs from
<taxonomicName lsidName="L. yunkaiensis" pageId="0" pageNumber="139" rank="species" species="yunkaiensis">
<emphasis italics="true" pageId="0" pageNumber="139">L. yunkaiensis</emphasis>
</taxonomicName>
by having smaller body size (SVL 22.9-25.6 mm in males, 27.1 mm in female vs 25.9-29.3 mm in males, 34.0-35.3 mm in females); bicolored iris with bright orange in upper half, fading to silver in lower half (vs coppery orange in upper half and silver in lower half); narrow lateral fringes present only on II and III toes (vs wide lateral fringes present on all toes); distinct black spots on the flank (vs indistinct black spots on the flank).
</paragraph>
<paragraph pageId="0" pageNumber="139">
<taxonomicName authorityName="Lyu &amp; Dai &amp; Wei &amp; He &amp; Yuan &amp; Shi &amp; Zhou &amp; Ran &amp; Kuang &amp; Guo &amp; Wei &amp; Yuan" authorityYear="2020" class="Amphibia" family="Megophryidae" genus="Leptobrachella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptobrachella bashaensis" order="Anura" pageId="0" pageNumber="139" phylum="Chordata" rank="species" species="bashaensis">
<emphasis italics="true" pageId="0" pageNumber="139">Leptobrachella bashaensis</emphasis>
</taxonomicName>
sp. nov. differs from
<taxonomicName lsidName="L. liui" pageId="0" pageNumber="139" rank="species" species="liui">
<emphasis italics="true" pageId="0" pageNumber="139">L. liui</emphasis>
</taxonomicName>
in having slightly smaller body size (SVL 22.9-25.6 mm in males, 27.1 mm in female vs 23.0-28.7 mm in males, 23.1-28.1 mm in females); vertical and distinctly narrow pupil (vs vertical and wide pupil); narrow lateral fringes present only on II and III toes (vs wide lateral fringes present on all toes).
</paragraph>
</subSubSection>
</treatment>
</document>
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