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148 lines
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<document ID-DOI="10.1206/0003-0090.457.1.1" ID-ISSN="0003-0090" ID-Zenodo-Dep="6971356" approvalRequired="59" approvalRequired_for_taxonomicNames="59" checkinTime="1659882917898" checkinUser="felipe" docAuthor="Beck, Robin M. D., Voss, Robert S. & Jansa, Sharon A." docDate="2022" docId="03EFDD5DF7706965D985FC061B67FB7F" docLanguage="en" docName="BulAmeMusNatHist.2022.457.1-350.pdf" docOrigin="Bulletin of the American Museum of Natural History 2022 (457)" docSource="https://bioone.org/journals/bulletin-of-the-american-museum-of-natural-history/volume-457/issue-1/0003-0090.457.1.1/Craniodental-Morphology-and-Phylogeny-of-Marsupials/10.1206/0003-0090.457.1.1.full" docStyle="DocumentStyle:915933466F796C9C739DF4DB6B8DCFA6.8:BulAmeMusNatHis.2011-.journal_article.1cover.type1" docStyleId="915933466F796C9C739DF4DB6B8DCFA6" docStyleName="BulAmeMusNatHis.2011-.journal_article.1cover.type1" docStyleVersion="8" docTitle="Ilaria undetermined" docType="treatment" docVersion="5" lastPageNumber="331" masterDocId="FFD6A525F63B6829DA0BFFAE1A62FFC7" masterDocTitle="Craniodental Morphology And Phylogeny Of Marsupials" masterLastPageNumber="353" masterPageNumber="1" pageNumber="331" updateTime="1659987007554" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Craniodental Morphology And Phylogeny Of Marsupials</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Beck, Robin M. D.</mods:namePart>
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<mods:affiliation>School of Science, Engineering and Environment University of Salford, U. K. & School of Biological, Earth & Environmental Sciences University of New South Wales, Australia & Division of Vertebrate Zoology (Mammalogy) American Museum of Natural History</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Voss, Robert S.</mods:namePart>
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<mods:affiliation>Division of Vertebrate Zoology (Mammalogy) American Museum of Natural History</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Jansa, Sharon A.</mods:namePart>
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<mods:affiliation>Bell Museum and Department of Ecology, Evolution, and Behavior University of Minnesota</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
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<mods:title>Bulletin of the American Museum of Natural History</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-06-28</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>2022</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>457</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>353</mods:end>
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<mods:url>https://bioone.org/journals/bulletin-of-the-american-museum-of-natural-history/volume-457/issue-1/0003-0090.457.1.1/Craniodental-Morphology-and-Phylogeny-of-Marsupials/10.1206/0003-0090.457.1.1.full</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.1206/0003-0090.457.1.1</mods:identifier>
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<mods:identifier type="ISSN">0003-0090</mods:identifier>
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<mods:identifier type="Zenodo-Dep">6971356</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6974501" ID-Zenodo-Dep="6974501" LSID="urn:lsid:plazi:treatment:03EFDD5DF7706965D985FC061B67FB7F" httpUri="http://treatment.plazi.org/id/03EFDD5DF7706965D985FC061B67FB7F" lastPageId="332" lastPageNumber="331" pageId="331" pageNumber="331">
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<subSubSection box="[910,988,936,960]" pageId="331" pageNumber="331" type="nomenclature">
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<paragraph blockId="331.[910,988,936,960]" box="[910,988,936,960]" pageId="331" pageNumber="331">
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<heading box="[910,988,936,960]" centered="true" fontSize="10" level="2" pageId="331" pageNumber="331" reason="4">
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<emphasis bold="true" box="[910,988,936,960]" pageId="331" pageNumber="331">
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†
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<taxonomicName box="[923,988,936,960]" class="Mammalia" family="Ilariidae" genus="Ilaria" kingdom="Animalia" order="Diprotodontia" pageId="331" pageNumber="331" phylum="Chordata" rank="species">
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<emphasis bold="true" box="[923,988,936,960]" italics="true" pageId="331" pageNumber="331">Ilaria</emphasis>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="331" pageNumber="331" type="synonymic_list">
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<paragraph blockId="331.[684,1215,986,1738]" pageId="331" pageNumber="331">
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SPECIES SCORED: †
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<taxonomicName authorityName="Tedford & Woodburne" authorityYear="1987" box="[921,1097,986,1010]" class="Mammalia" family="Ilariidae" genus="Ilaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="331" pageNumber="331" phylum="Chordata" rank="species">
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<emphasis box="[921,1097,986,1010]" italics="true" pageId="331" pageNumber="331">Ilaria illumidens</emphasis>
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</taxonomicName>
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(
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<typeStatus box="[1111,1160,986,1010]" pageId="331" pageNumber="331">type</typeStatus>
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species), †
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<taxonomicName authorityName="Woodburne" authorityYear="1984" box="[761,865,1019,1043]" class="Mammalia" family="Macropodidae" genus="Wakiewakie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="331" pageNumber="331" phylum="Chordata" rank="species">
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<emphasis box="[761,865,1019,1043]" italics="true" pageId="331" pageNumber="331">I. lawsoni</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="331" pageNumber="331" type="materials_examined">
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<paragraph blockId="331.[684,1215,986,1738]" pageId="331" pageNumber="331">
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GEOLOGICAL PROVENANCE OF SCORED
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<collectionCode box="[684,745,1085,1109]" pageId="331" pageNumber="331">SPECIMENS</collectionCode>
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: AMNH site B, Zone B (Pinpa Local Fauna) Namba Formation, Lake Pinpa, South Australia (†
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<taxonomicName authorityName="Tedford & Woodburne" authorityYear="1987" box="[812,945,1151,1175]" class="Mammalia" family="Ilariidae" genus="Ilaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="331" pageNumber="331" phylum="Chordata" rank="species">
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<emphasis box="[812,945,1151,1175]" italics="true" pageId="331" pageNumber="331">I. illumidens</emphasis>
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</taxonomicName>
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); AMNH
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<collectingCountry box="[1058,1122,1152,1176]" name="Thailand" pageId="331" pageNumber="331">SIAM</collectingCountry>
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locality, unit 6, Zone B (Ditjimanka Local Fauna), Lake Palankarinna, Etadunna Formation, South Australia (†
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<taxonomicName authorityName="Woodburne" authorityYear="1984" box="[771,875,1251,1275]" class="Mammalia" family="Macropodidae" genus="Wakiewakie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="331" pageNumber="331" phylum="Chordata" rank="species">
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<emphasis box="[771,875,1251,1275]" italics="true" pageId="331" pageNumber="331">I. lawsoni</emphasis>
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</taxonomicName>
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);
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</paragraph>
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</subSubSection>
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<subSubSection pageId="331" pageNumber="331" type="description">
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<paragraph blockId="331.[684,1215,986,1738]" pageId="331" pageNumber="331">AGE OF SCORED SPECIMENS: Based on palaeomagnetic data, Metzger and Retallack (2010) estimated the Etadunna Formation to span 26.1– 23.6 Mya. Zone A of the Etadunna Formation and the Pinpa Local Fauna of the Namba Formation appear to be stratigraphic equivalents (Woodburne et al., 1994). However, in the absence of radiometric dates, we have assumed the entire span of the late Oligocene (Chattian; Cohen et al., 2013 [updated]) for this terminal.</paragraph>
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<paragraph blockId="331.[684,1215,986,1738]" box="[717,1197,1615,1639]" pageId="331" pageNumber="331">
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ASSIGNED AGE
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<collectionCode box="[888,964,1615,1639]" pageId="331" pageNumber="331">RANGE</collectionCode>
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:
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<geoCoordinate box="[978,1050,1615,1639]" degrees="27.820" direction="north" orientation="latitude" pageId="331" pageNumber="331" precision="55" value="27.82">27.820</geoCoordinate>
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<geoCoordinate box="[1049,1135,1615,1639]" degrees="23.030" direction="west" orientation="longitude" pageId="331" pageNumber="331" precision="55" value="-23.03">–23.030</geoCoordinate>
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Mya.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="332" lastPageNumber="332" pageId="331" pageNumber="331" type="discussion">
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<paragraph blockId="331.[684,1215,986,1738]" lastBlockId="332.[108,637,226,1208]" lastPageId="332" lastPageNumber="332" pageId="331" pageNumber="331">
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REMARKS: †
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<taxonomicName authorityName="Tedford & Woodburne" authorityYear="1987" box="[847,1022,1647,1671]" class="Mammalia" family="Ilariidae" genus="Ilaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="331" pageNumber="331" phylum="Chordata" rank="species">
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<emphasis box="[847,1022,1647,1671]" italics="true" pageId="331" pageNumber="331">Ilaria illumidens</emphasis>
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</taxonomicName>
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is represented by craniodental and associated postcranial elements of several individuals, while the slightly larger †
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<taxonomicName box="[1197,1213,1714,1737]" pageId="331" pageNumber="331">
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<emphasis box="[1197,1213,1714,1737]" italics="true" pageId="331" pageNumber="331">I.</emphasis>
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</taxonomicName>
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<emphasis box="[108,187,226,250]" italics="true" pageId="332" pageNumber="332">lawsoni</emphasis>
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is known from a single partial right dentary only (Tedford and Woodburne, 1987). Both species are characterized by large size (estimated body mass of †
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<taxonomicName authorityName="Tedford & Woodburne" authorityYear="1987" box="[274,449,325,349]" class="Mammalia" family="Ilariidae" genus="Ilaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="332" pageNumber="332" phylum="Chordata" rank="species">
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<emphasis box="[274,449,325,349]" italics="true" pageId="332" pageNumber="332">Ilaria illumidens</emphasis>
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</taxonomicName>
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is ~
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<quantity box="[495,572,325,349]" metricMagnitude="2" metricUnit="kg" metricValue="2.15" pageId="332" pageNumber="332" unit="kg" value="215.0">215 kg</quantity>
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; Beck et al., 2020) and several unusual craniodental features. The upper molars are strongly selenodont, while the lower molars are incipiently lophodont with prominent neomorphic cuspids present; Tedford and Woodburne (1987) identified the “central cuspid” between the metaconid and protoconid, and a second between the hypconid and entoconid, as neomorphic, but could not rule out the possibility that it is in fact the lingual cuspids (the putative “metaconid” and “entoconid”) are the neomorphs (Tedford and Woodburne, 1987: 415), and we have treated the homologies of these structures as unknown in this study. †
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<taxonomicName authorityName="Tedford & Woodburne" authorityYear="1987" box="[508,566,755,779]" class="Mammalia" family="Ilariidae" genus="Ilaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="332" pageNumber="332" phylum="Chordata" rank="genus">
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<emphasis box="[508,566,755,779]" italics="true" pageId="332" pageNumber="332">Ilaria</emphasis>
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</taxonomicName>
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is also unusual in that the mandibular symphysis is fused, a feature otherwise seen only in a few other vombatiforms (see char. 97). Another ilariid, the much smaller (estimated body mass ~
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<quantity box="[512,574,887,911]" metricMagnitude="1" metricUnit="kg" metricValue="1.6" pageId="332" pageNumber="332" unit="kg" value="16.0">16 kg</quantity>
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; Beck et al., 2020) †
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<taxonomicName authorityName="Pledge" authorityYear="1987" box="[247,430,921,944]" class="Mammalia" family="Ilariidae" genus="Kuterintja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="332" pageNumber="332" phylum="Chordata" rank="species">
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<emphasis box="[247,430,921,944]" italics="true" pageId="332" pageNumber="332">Kuterintja ngama</emphasis>
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</taxonomicName>
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, is known from the Ngama Local Fauna (Zone D of the Etadunna Formation; Pledge, 1987b; Woodburne et al., 1994) and the Riversleigh Faunal Zone A White Hunter Site (Myers and Archer, 1997), both of which are late Oligocene (see above); however, †
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<taxonomicName authorityName="Pledge" authorityYear="1987" class="Mammalia" family="Ilariidae" genus="Kuterintja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="332" pageNumber="332" phylum="Chordata" rank="species">
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<emphasis italics="true" pageId="332" pageNumber="332">K. ngama</emphasis>
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</taxonomicName>
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specimens is currently represented by fragmentary dental material only, and so has not been included here.
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</paragraph>
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</subSubSection>
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</treatment>
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</document>
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