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<document id="E45BF7AB3AAE1B7543657E6F76E5B8FD" ID-DOI="10.1206/0003-0090.457.1.1" ID-ISSN="0003-0090" ID-Zenodo-Dep="6971356" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1659882917898" checkinUser="felipe" docAuthor="Beck, Robin M. D., Voss, Robert S. & Jansa, Sharon A." docDate="2022" docId="03EFDD5DF6E968FBDABEFF4C19C5FD0E" docLanguage="en" docName="BulAmeMusNatHist.2022.457.1-350.pdf" docOrigin="Bulletin of the American Museum of Natural History 2022 (457)" docSource="https://bioone.org/journals/bulletin-of-the-american-museum-of-natural-history/volume-457/issue-1/0003-0090.457.1.1/Craniodental-Morphology-and-Phylogeny-of-Marsupials/10.1206/0003-0090.457.1.1.full" docStyle="DocumentStyle:915933466F796C9C739DF4DB6B8DCFA6.9:BulAmeMusNatHis.2011-.journal_article.1cover.type1" docStyleId="915933466F796C9C739DF4DB6B8DCFA6" docStyleName="BulAmeMusNatHis.2011-.journal_article.1cover.type1" docStyleVersion="9" docTitle="Peramelemorphia Ameghino 1889" docType="treatment" docVersion="11" lastPageNumber="210" masterDocId="FFD6A525F63B6829DA0BFFAE1A62FFC7" masterDocTitle="Craniodental Morphology And Phylogeny Of Marsupials" masterLastPageNumber="353" masterPageNumber="1" pageNumber="210" updateTime="1699350301078" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="2DE9F125664181902946B5A2FC8ADF98">Craniodental Morphology And Phylogeny Of Marsupials</mods:title>
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<mods:namePart id="AAEB5B72E46D0A05AD3749F7640D24B2">Beck, Robin M. D.</mods:namePart>
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<mods:affiliation id="B1C1FB6EF8936044658542BCE14D2281">School of Science, Engineering and Environment University of Salford, U. K. & School of Biological, Earth & Environmental Sciences University of New South Wales, Australia & Division of Vertebrate Zoology (Mammalogy) American Museum of Natural History</mods:affiliation>
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<mods:namePart id="AADCE5EB7E079371A4304A1940008B03">Voss, Robert S.</mods:namePart>
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<mods:affiliation id="2CF0152A3CB728B324C48AA9FDBDD923">Division of Vertebrate Zoology (Mammalogy) American Museum of Natural History</mods:affiliation>
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<mods:namePart id="2A8FB06D0B5BB3C570181F6B6DDF6861">Jansa, Sharon A.</mods:namePart>
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<mods:affiliation id="EF202904B299ABF2769B3894966F27FC">Bell Museum and Department of Ecology, Evolution, and Behavior University of Minnesota</mods:affiliation>
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<mods:date id="06299CD4DFC47772BBA7351D30A0DFD7">2022</mods:date>
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<mods:number id="1D078D7331B9834B20CB0D79632A1530">2022-06-28</mods:number>
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<mods:number id="1F5A7B9CFDE05CF0B1BC4A02334839B9">457</mods:number>
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<mods:url id="D6A1F17A729291B42EA9BF7781D19D66">https://bioone.org/journals/bulletin-of-the-american-museum-of-natural-history/volume-457/issue-1/0003-0090.457.1.1/Craniodental-Morphology-and-Phylogeny-of-Marsupials/10.1206/0003-0090.457.1.1.full</mods:url>
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<mods:identifier id="B0B90E98EB8CDF79CEB3EFF9AB164AE3" type="DOI">10.1206/0003-0090.457.1.1</mods:identifier>
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<mods:identifier id="DDB7A73302993E7AD0E601CBCF04CC3D" type="ISSN">0003-0090</mods:identifier>
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<mods:identifier id="D08E0B45D1ADFBD11A933C88AA5DB39F" type="Zenodo-Dep">6971356</mods:identifier>
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<treatment id="03EFDD5DF6E968FBDABEFF4C19C5FD0E" ID-DOI="http://doi.org/10.5281/zenodo.7036151" ID-Zenodo-Dep="7036151" LSID="urn:lsid:plazi:treatment:03EFDD5DF6E968FBDABEFF4C19C5FD0E" httpUri="http://treatment.plazi.org/id/03EFDD5DF6E968FBDABEFF4C19C5FD0E" lastPageNumber="210" pageId="210" pageNumber="210">
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<subSubSection id="C35C3FC0F6E968FBDABEFF4C1851FF3D" box="[181,563,226,250]" pageId="210" pageNumber="210" type="nomenclature">
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<paragraph id="8BF96C4BF6E968FBDABEFF4C1851FF3D" blockId="210.[181,563,226,250]" box="[181,563,226,250]" pageId="210" pageNumber="210">
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<heading id="D0B1DB27F6E968FBDABEFF4C1851FF3D" box="[181,563,226,250]" centered="true" fontSize="9" level="2" pageId="210" pageNumber="210" reason="2">
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<taxonomicName id="4C4617C8F6E968FBDABEFF4C1851FF3D" ID-CoL="3S9" authority="Ameghino, 1889" authorityName="Ameghino" authorityYear="1889" box="[181,563,226,250]" class="Mammalia" kingdom="Animalia" order="Peramelemorphia" pageId="210" pageNumber="210" phylum="Chordata" rank="order">Peramelemorphia Ameghino, 1889</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C35C3FC0F6E968FBDA87FE8D1A94FE9B" pageId="210" pageNumber="210" type="synonymic_list">
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<paragraph id="8BF96C4BF6E968FBDA87FE8D1A94FE9B" blockId="210.[107,638,291,1737]" pageId="210" pageNumber="210">
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CONTENTS: †
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<taxonomicName id="4C4617C8F6E968FBDB10FE8D1B17FEFC" box="[283,373,291,315]" class="Mammalia" genus="Bulungu" higherTaxonomySource="GBIF" kingdom="Animalia" order="Peramelemorphia" pageId="210" pageNumber="210" phylum="Chordata" rank="genus">
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<emphasis id="B932B059F6E968FBDB10FE8D1B17FEFC" box="[283,373,291,315]" italics="true" pageId="210" pageNumber="210">Bulungu</emphasis>
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</taxonomicName>
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, †
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<taxonomicName id="4C4617C8F6E968FBDB84FE8D181AFEFC" authority=", Perameloidea" authorityName="Perameloidea" box="[399,632,291,315]" class="Mammalia" genus="Galadi" higherTaxonomySource="GBIF" kingdom="Animalia" order="Peramelemorphia" pageId="210" pageNumber="210" phylum="Chordata" rank="genus">
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<emphasis id="B932B059F6E968FBDB84FE8D1BB5FEFC" box="[399,471,291,315]" italics="true" pageId="210" pageNumber="210">Galadi</emphasis>
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, Perameloidea
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</taxonomicName>
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, and †
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<emphasis id="B932B059F6E968FBDAA1FEEA1A94FE9B" box="[170,246,324,348]" italics="true" pageId="210" pageNumber="210">
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<taxonomicName id="4C4617C8F6E968FBDAA1FEEA1A90FE9B" baseAuthorityName="Muirhead and Filan" baseAuthorityYear="1995" box="[170,242,324,348]" class="Mammalia" family="Yaralidae" genus="Yarala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Peramelemorphia" pageId="210" pageNumber="210" phylum="Chordata" rank="genus">Yarala</taxonomicName>
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.
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="C35C3FC0F6E968FBDA87FECB1A8AFCEC" pageId="210" pageNumber="210" type="description">
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<paragraph id="8BF96C4BF6E968FBDA87FECB1ACEFE59" blockId="210.[107,638,291,1737]" pageId="210" pageNumber="210">STEM AGE: 43.7 Mya (95% HPD: 39.6–47.4 Mya).</paragraph>
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<paragraph id="8BF96C4BF6E968FBDA87FE091ACFFE27" blockId="210.[107,638,291,1737]" pageId="210" pageNumber="210">CROWN AGE: 31.4 Mya (95% HPD: 25.3–37.9 Mya).</paragraph>
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<paragraph id="8BF96C4BF6E968FBDA87FE471A8AFCEC" blockId="210.[107,638,291,1737]" pageId="210" pageNumber="210">UNAMBIGUOUS CRANIODENTAL SYNAPOMORPHIES: Facial exposure of jugal deeply bifid, with distinct anterodorsal and anteroventral processes (char. 20: 0→1; ci = 1.000); lambdoid sesamoids present (char. 89: 0→1; ci = 1.000); M1 preparacrista absent or indistinct (char. 138: 0→1; ci = 0.333); M1 pseudopreparacrista present (char. 139: 0→1; ci = 0.333); and hypoconulid notch in anterior cingulid of m2–4 absent (char. 158: 0→1; ci = 0.500).</paragraph>
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</subSubSection>
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<subSubSection id="C35C3FC0F6E968FBDA87FC9A19C5FD0E" pageId="210" pageNumber="210" type="discussion">
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<paragraph id="8BF96C4BF6E968FBDA87FC9A1BDCF941" blockId="210.[107,638,291,1737]" pageId="210" pageNumber="210">
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COMMENTS: Our estimate for the divergence between
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<taxonomicName id="4C4617C8F6E968FBDAC5FCFB1BE9FCAA" authorityName="Ameghino" authorityYear="1889" box="[206,395,853,877]" class="Mammalia" kingdom="Animalia" order="Peramelemorphia" pageId="210" pageNumber="210" phylum="Chordata" rank="order">Peramelemorphia</taxonomicName>
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and
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<taxonomicName id="4C4617C8F6E968FBDBCDFCFB181EFCAA" baseAuthorityName="Kealy and Beck" baseAuthorityYear="2017" box="[454,636,853,877]" class="Mammalia" kingdom="Animalia" order="Dasyuromorphia" pageId="210" pageNumber="210" phylum="Chordata" rank="order">Dasyuromorphia</taxonomicName>
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dates to the middle Eocene. As mentioned above (see Agreodontia), isolated dental remains from the early Eocene Tingamarra Local Fauna have been identified as putatively peramelemorphian, which would predate our estimate, but the Tingamarran specimens cannot be unequivocally referred to the
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<taxonomicName id="4C4617C8F6E968FBDA66FB921B4DFB93" authorityName="Ameghino" authorityYear="1889" box="[109,303,1084,1108]" class="Mammalia" kingdom="Animalia" order="Peramelemorphia" pageId="210" pageNumber="210" phylum="Chordata" rank="order">Peramelemorphia</taxonomicName>
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. Instead, the oldest definitive peramelemorphians are late Oligocene (
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<bibRefCitation id="EFD711BAF6E968FBD808FBF31A91FB50" author="Travouillon" etAl="et al." firstAuthor="Travouillon" pageId="210" pageNumber="210" pagination="13" refId="ref230544" refString="Travouillon, K. J., R. M. D. Beck, S. J. Hand, and M. Archer. 2013 a. The oldest fossil record of bandicoots (Marsupialia; Peramelemorphia) from the late Oligocene of Australia. Palaeontologia Electronica 16 (2): 13 A." type="journal article" year="2013" yearSuffix="a">
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Travouillon et al., 2013
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<bibRefCitation id="EFD711BAF6E968FBDAE3FBD11A91FB50" author="a" box="[232,243,1151,1175]" etAl="et al." firstAuthor="a" pageId="210" pageNumber="210" pagination="13" refId="ref230544" refString="Travouillon, K. J., R. M. D. Beck, S. J. Hand, and M. Archer. 2013 a. The oldest fossil record of bandicoots (Marsupialia; Peramelemorphia) from the late Oligocene of Australia. Palaeontologia Electronica 16 (2): 13 A." type="journal article" year="2021" yearSuffix="a">a</bibRefCitation>
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</bibRefCitation>
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, 2021;
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<bibRefCitation id="EFD711BAF6E968FBDB4DFBD01AC1FB70" author="Warburton and Travouillon" firstAuthor="Warburton" pageId="210" pageNumber="210" pagination="151 - 181" refId="ref232688" refString="Warburton, N. M., and K. J. Travouillon. 2016. The biology and palaeontology of the Peramelemorphia: a review of current knowledge and future research directions. Australian Journal of Zoology 64 (3): 151 - 181." type="journal article" year="2016">Warburton and Travouillon, 2016</bibRefCitation>
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). The three fossils included here (†
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<taxonomicName id="4C4617C8F6E968FBD815FB311814FB70" box="[542,630,1183,1207]" class="Mammalia" genus="Bulungu" higherTaxonomySource="GBIF" kingdom="Animalia" order="Peramelemorphia" pageId="210" pageNumber="210" phylum="Chordata" rank="genus">
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<emphasis id="B932B059F6E968FBD815FB311814FB70" box="[542,630,1183,1207]" italics="true" pageId="210" pageNumber="210">Bulungu</emphasis>
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</taxonomicName>
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, †
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<taxonomicName id="4C4617C8F6E968FBDA72FB6E1ADDFB1F" box="[121,191,1216,1240]" class="Mammalia" genus="Galadi" higherTaxonomySource="GBIF" kingdom="Animalia" order="Peramelemorphia" pageId="210" pageNumber="210" phylum="Chordata" rank="genus">
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<emphasis id="B932B059F6E968FBDA72FB6E1ADDFB1F" box="[121,191,1216,1240]" italics="true" pageId="210" pageNumber="210">Galadi</emphasis>
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</taxonomicName>
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, †
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<taxonomicName id="4C4617C8F6E968FBDAD7FB6E1B7DFB1F" baseAuthorityName="Muirhead and Filan" baseAuthorityYear="1995" box="[220,287,1216,1240]" class="Mammalia" family="Yaralidae" genus="Yarala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Peramelemorphia" pageId="210" pageNumber="210" phylum="Chordata" rank="genus">
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<emphasis id="B932B059F6E968FBDAD7FB6E1B7DFB1F" box="[220,287,1216,1240]" italics="true" pageId="210" pageNumber="210">Yarala</emphasis>
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</taxonomicName>
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) are late Oligocene to Miocene taxa (
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<bibRefCitation id="EFD711BAF6E968FBDAACFB4C1BA3FB3E" author="Muirhead, J. & S. Filan" box="[167,449,1249,1274]" pageId="210" pageNumber="210" pagination="127 - 134" refId="ref217527" refString="Muirhead, J., and S. Filan. 1995. Yarala burchfieldi (Peramelemorphia) from Oligo-Miocene deposits of Riversleigh, northwestern Queensland. Journal of Paleontology 69 (1): 127 - 134." type="journal article" year="1995">Muirhead and Filan, 1995</bibRefCitation>
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;
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<bibRefCitation id="EFD711BAF6E968FBDBC5FB4C181BFB3D" author="Muirhead, J." box="[462,633,1250,1274]" pageId="210" pageNumber="210" pagination="512 - 523" refId="ref217438" refString="Muirhead, J. 2000. Yaraloidea (Marsupialia, Peramelemorphia), a new superfamily of marsupial and a description and analysis of the cranium of the Miocene of Yarala burchfieldi. Journal of Paleontology 74 (3): 512 - 523." type="journal article" year="2000">Muirhead, 2000</bibRefCitation>
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;
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<bibRefCitation id="EFD711BAF6E968FBDA67FAAD1B12FADC" author="Travouillon" box="[108,368,1283,1307]" etAl="et al." firstAuthor="Travouillon" pageId="210" pageNumber="210" pagination="1528 - 1546" refId="ref230485" refString="Travouillon, K. J., Y. Gurovich, R. M. D. Beck, and J. Muirhead. 2010. An exceptionally well-preserved short-snouted bandicoot (Marsupialia; Peramelemorphia) from Riversleigh's Oligo-Miocene deposits, northwestern Queensland, Australia. Journal of Vertebrate Paleontology 30 (5): 1528 - 1546." type="journal article" year="2010">Travouillon et al., 2010</bibRefCitation>
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,
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<bibRefCitation id="EFD711BAF6E968FBDB75FAAD1BA3FADC" author="Travouillon" box="[382,449,1283,1307]" firstAuthor="Travouillon" pageId="210" pageNumber="210" pagination="153 - 168" refId="ref230601" refString="Travouillon, K. J., Y. Gurovich, M. Archer, S. J. Hand, and J. Muirhead. 2013 b. The genus Galadi: three new bandicoots (Marsupialia, Peramelemorphia) from Riversleigh's Miocene deposits, Northwestern Queensland, Australia. Journal of Vertebrate Paleontology 33 (1): 153 - 168." type="journal article" year="2013" yearSuffix="b">2013b</bibRefCitation>
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;
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<bibRefCitation id="EFD711BAF6E968FBDBDBFAAD1AC2FAFB" author="Gurovich, Y. & K. J. Travouillon & R. M. D. Beck & J. Muirhead & M. Archer" pageId="210" pageNumber="210" pagination="265 - 290" refId="ref208146" refString="Gurovich, Y., K. J. Travouillon, R. M. D. Beck, J. Muirhead, and M. Archer. 2014. Biogeographical implications of a new mouse-sized fossil bandicoot (Marsupialia: Peramelemorphia) occupying a dasyurid-like ecological niche across Australia. Journal of Systematic Palaeontology 12 (3): 265 - 290." type="journal article" year="2014">Gurovich et al., 2014</bibRefCitation>
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;
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<bibRefCitation id="EFD711BAF6E968FBDAA7FA8A1873FAFB" author="Warburton and Travouillon" box="[172,529,1316,1340]" firstAuthor="Warburton" pageId="210" pageNumber="210" pagination="151 - 181" refId="ref232688" refString="Warburton, N. M., and K. J. Travouillon. 2016. The biology and palaeontology of the Peramelemorphia: a review of current knowledge and future research directions. Australian Journal of Zoology 64 (3): 151 - 181." type="journal article" year="2016">Warburton and Travouillon, 2016</bibRefCitation>
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) and they are recovered outside the peramelemorphian crown clade (Perameloidea; see below) in our analyses, as in previous phylogenetic studies (
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<bibRefCitation id="EFD711BAF6E968FBDBCDFA291AC0FA07" author="Travouillon" etAl="et al." firstAuthor="Travouillon" pageId="210" pageNumber="210" pagination="1528 - 1546" refId="ref230485" refString="Travouillon, K. J., Y. Gurovich, R. M. D. Beck, and J. Muirhead. 2010. An exceptionally well-preserved short-snouted bandicoot (Marsupialia; Peramelemorphia) from Riversleigh's Oligo-Miocene deposits, northwestern Queensland, Australia. Journal of Vertebrate Paleontology 30 (5): 1528 - 1546." type="journal article" year="2010">Travouillon et al., 2010</bibRefCitation>
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,
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<bibRefCitation id="EFD711BAF6E968FBDAB9FA061A91FA07" author="Travouillon" box="[178,243,1448,1472]" firstAuthor="Travouillon" pageId="210" pageNumber="210" pagination="13" refId="ref230544" refString="Travouillon, K. J., R. M. D. Beck, S. J. Hand, and M. Archer. 2013 a. The oldest fossil record of bandicoots (Marsupialia; Peramelemorphia) from the late Oligocene of Australia. Palaeontologia Electronica 16 (2): 13 A." type="journal article" year="2013" yearSuffix="a">2013a</bibRefCitation>
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,
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<bibRefCitation id="EFD711BAF6E968FBDB0FFA061B24FA07" author="Travouillon" box="[260,326,1448,1472]" firstAuthor="Travouillon" pageId="210" pageNumber="210" pagination="153 - 168" refId="ref230601" refString="Travouillon, K. J., Y. Gurovich, M. Archer, S. J. Hand, and J. Muirhead. 2013 b. The genus Galadi: three new bandicoots (Marsupialia, Peramelemorphia) from Riversleigh's Miocene deposits, Northwestern Queensland, Australia. Journal of Vertebrate Paleontology 33 (1): 153 - 168." type="journal article" year="2013" yearSuffix="b">2013b</bibRefCitation>
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,
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<bibRefCitation id="EFD711BAF6E968FBDB5CFA061BFAFA07" author="Travouillon" box="[343,408,1448,1472]" firstAuthor="Travouillon" pageId="210" pageNumber="210" pagination="375 - 382" refId="ref230665" refString="Travouillon, K. J., S. J. Hand, M. Archer, and K. H. Black. 2014 a. Earliest modern bandicoot and bilby (Marsupialia, Peramelidae and Thylacomyidae) from the Miocene of the Riversleigh World Heritage Area, northwestern Queensland, Australia. Journal of Vertebrate Paleontology 34 (2): 375 - 382." type="journal article" year="2014" yearSuffix="a">
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2014
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<bibRefCitation id="EFD711BAF6E968FBDB86FA061BFAFA07" author="a" box="[397,408,1448,1472]" firstAuthor="a" pageId="210" pageNumber="210" pagination="1408" refId="ref237143" refString="Yates, A. M. 2015 b. New craniodental remains of Wakaleo alcootaensis (Diprotodontia: Thylacoleonidae) a carnivorous marsupial from the late Miocene Alcoota Local Fauna of the Northern Territory, Australia. PeerJ 3: e 1408." type="journal article" year="2015" yearSuffix="b">a</bibRefCitation>
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</bibRefCitation>
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, 2015
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<bibRefCitation id="EFD711BAF6E968FBDBD5FA061850FA07" author="b" box="[478,562,1448,1472]" firstAuthor="b" pageId="210" pageNumber="210" pagination="657 - 671" refId="ref224780" refString="Rosa, B. B., G. A. R. Melo, and M. S. Barbeitos. 2019. Homoplasy-based partitioning outperforms alternatives in Bayesian analysis of discrete morphological data. Systematic Biology 68 (4): 657 - 671." type="journal article" year="2019">b, 2019</bibRefCitation>
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, 2021;
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<bibRefCitation id="EFD711BAF6E968FBDA67FA671B20FA26" author="Gurovich, Y. & K. J. Travouillon & R. M. D. Beck & J. Muirhead & M. Archer" box="[108,322,1481,1505]" pageId="210" pageNumber="210" pagination="265 - 290" refId="ref208146" refString="Gurovich, Y., K. J. Travouillon, R. M. D. Beck, J. Muirhead, and M. Archer. 2014. Biogeographical implications of a new mouse-sized fossil bandicoot (Marsupialia: Peramelemorphia) occupying a dasyurid-like ecological niche across Australia. Journal of Systematic Palaeontology 12 (3): 265 - 290." type="journal article" year="2014">Gurovich et al., 2014</bibRefCitation>
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;
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<bibRefCitation id="EFD711BAF6E968FBDB40FA671825FA26" author="Chamberlain, P. M. & K. J. Travouillon & M. Archer & S. J. Hand" box="[331,583,1481,1505]" pageId="210" pageNumber="210" pagination="197 - 206" refId="ref197024" refString="Chamberlain, P. M., K. J. Travouillon, M. Archer, and S. J. Hand. 2015. Kutjamarcoot brevirostrum gen. et sp. nov., a new short-snouted, early Miocene bandicoot (Marsupialia: Peramelemorphia) from the Kutjamarpu Local Fauna (Wipajiri Formation) in South Australia. Alcheringa: an Australasian Journal of Palaeontology 40 (2): 197 - 206." type="journal article" year="2015">Chamberlain et al., 2015</bibRefCitation>
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;
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<bibRefCitation id="EFD711BAF6E968FBD85BFA641BF6F9C5" author="Travouillon and Phillips" firstAuthor="Travouillon" pageId="210" pageNumber="210" pagination="224 - 256" refId="ref230320" refString="Travouillon, K. J., and M. J. Phillips. 2018. Total evidence analysis of the phylogenetic relationships of bandicoots and bilbies (Marsupialia: Peramelemorphia): reassessment of two species and description of a new species. Zootaxa 4378 (2): 224 - 256." type="journal article" year="2018">Travouillon and Phillips, 2018</bibRefCitation>
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). Putative early perameloids are known from the middle Miocene (
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<bibRefCitation id="EFD711BAF6E968FBDA78F9821B0CF983" author="Travouillon" box="[115,366,1580,1604]" etAl="et al." firstAuthor="Travouillon" pageId="210" pageNumber="210" pagination="375 - 382" refId="ref230665" refString="Travouillon, K. J., S. J. Hand, M. Archer, and K. H. Black. 2014 a. Earliest modern bandicoot and bilby (Marsupialia, Peramelidae and Thylacomyidae) from the Miocene of the Riversleigh World Heritage Area, northwestern Queensland, Australia. Journal of Vertebrate Paleontology 34 (2): 375 - 382." type="journal article" year="2014" yearSuffix="a">Travouillon et al., 2014a</bibRefCitation>
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; see Perameloidea below), but these were not included here because they are only known from isolated teeth.
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</paragraph>
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<paragraph id="8BF96C4BF6E968FBDA86F93E19C5FD0E" blockId="210.[107,638,291,1737]" lastBlockId="210.[684,1212,226,713]" pageId="210" pageNumber="210">
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A recent study by
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<bibRefCitation id="EFD711BAF6E968FBDB8FF9211AD4F90E" author="Abello, M. A. & A. M. Candela" pageId="210" pageNumber="210" pagination="419 - 444" refId="ref188032" refString="Abello, M. A., and A. M. Candela. 2019. Paleobiology of Argyrolagus (Marsupialia, Argyrolagidae): an astonishing case of bipedalism among South American mammals. Journal of Mammalian Evolution 27: 419 - 444." type="journal article" year="2019">Abello and Candela (2019)</bibRefCitation>
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included morphological and total evidence phylogenetic analyses that placed members of the South American family †
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<taxonomicName id="4C4617C8F6E968FBD8BCFE8A193CFEFB" authorityName="Amenghino" authorityYear="1904" box="[695,862,292,316]" class="Mammalia" family="Argyrolagidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Paucituberculata" pageId="210" pageNumber="210" phylum="Chordata" rank="family">Argyrolagidae</taxonomicName>
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in a clade that was sister to
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<taxonomicName id="4C4617C8F6E968FBD8A7FEEB1917FE9A" authorityName="Ameghino" authorityYear="1889" box="[684,885,325,349]" class="Mammalia" kingdom="Animalia" order="Peramelemorphia" pageId="210" pageNumber="210" phylum="Chordata" rank="order">Peramelemorphia</taxonomicName>
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(represented by
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<taxonomicName id="4C4617C8F6E968FBDE32FEEB1EDEFE9A" authorityName="Lesson" authorityYear="1842" box="[1081,1212,325,349]" class="Mammalia" family="Peramelidae" genus="Echymipera" kingdom="Animalia" order="Peramelemorphia" pageId="210" pageNumber="210" phylum="Chordata" rank="genus">
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<emphasis id="B932B059F6E968FBDE32FEEB1EDEFE9A" box="[1081,1212,325,349]" italics="true" pageId="210" pageNumber="210">Echymipera</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C4617C8F6E968FBD8D5FEC8192EFEB9" authorityName="E. Geoffroy Saint-Hilaire" authorityYear="1804" box="[734,844,358,382]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="210" pageNumber="210" phylum="Chordata" rank="genus">
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<emphasis id="B932B059F6E968FBD8D5FEC8192EFEB9" box="[734,844,358,382]" italics="true" pageId="210" pageNumber="210">Perameles</emphasis>
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</taxonomicName>
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), but the authors acknowledged that this geographically implausible relationship might reflect convergent evolution of similar derived tarsal features; based on available evidence, argyrolagids are most likely to be paucituberculatans (
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<bibRefCitation id="EFD711BAF6E968FBD9BAFDA51EDAFDE4" author="Sanchez-Villagra, M. R." box="[945,1208,523,547]" pageId="210" pageNumber="210" pagination="481 - 496" refId="ref225799" refString="Sanchez-Villagra, M. R. 2001. The phylogenetic relationships of argyrolagid marsupials. Zoological Journal of the Linnean Society 131: 481 - 496." type="journal article" year="2001">Sánchez-Villagra, 2001</bibRefCitation>
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;
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<bibRefCitation id="EFD711BAF6E968FBD8A7FD82195AFD83" author="Beck, R. M. D." box="[684,824,556,580]" pageId="210" pageNumber="210" pagination="373 - 414" refId="ref192706" refString="Beck, R. M. D. 2017 a. The skull of Epidolops ameghinoi from the early Eocene Itaborai fauna, southeastern Brazil, and the affinities of the extinct marsupialiform order Polydolopimorphia. Journal of Mammalian Evolution 24 (4): 373 - 414." type="journal article" year="2017">Beck, 2017a</bibRefCitation>
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;
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<bibRefCitation id="EFD711BAF6E968FBD94FFD821E32FD83" author="Zimicz and Goin" box="[836,1104,556,580]" firstAuthor="Zimicz" pageId="210" pageNumber="210" pagination="975 - 992" refId="ref237724" refString="Zimicz, A. N., and F. J. Goin. 2020. A reassessment of the genus Groeberia Patterson, 1952 (Mammalia, Metatheria): functional and phylogenetic implications. Journal of Systematic Palaeontology 18 (12): 975 - 992." type="journal article" year="2020">Zimicz and Goin, 2020</bibRefCitation>
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) or polydolopimorphians (
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<bibRefCitation id="EFD711BAF6E968FBD983FDE31E04FDA2" author="Goin, F. J. & A. M. Candela & M. A. Abello & E. V. Oliveira" box="[904,1126,589,613]" pageId="210" pageNumber="210" pagination="867 - 884" refId="ref206706" refString="Goin, F. J., A. M. Candela, M. A. Abello, and E. V. Oliveira. 2009 a. Earliest South American paucituberculatans and their significance in the understanding of ' pseudodiprotodont' marsupial radiations. Zoological Journal of the Linnean Society 155: 867 - 884." type="journal article" year="2009">Goin et al., 2009a</bibRefCitation>
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;
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<bibRefCitation id="EFD711BAF6E968FBDE7CFDE3199CFD41" author="Chornogubsky, L. & F. J. Goin" pageId="210" pageNumber="210" pagination="983238" refId="ref197262" refString="Chornogubsky, L., and F. J. Goin. 2015. A review of the molar morphology and phylogenetic affinities of Sillustania quechuense (Metatheria, Polydolopimorphia, Sillustaniidae), from the early Paleogene of Laguna Umayo, southeastern Peru. Journal of Vertebrate Paleontology 35 (6): e 983238." type="journal article" year="2015">Chornogubsky and Goin, 2015</bibRefCitation>
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;
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<bibRefCitation id="EFD711BAF6E968FBDE1FFDC0193CFD6F" author="Carneiro, L. M. & E. V. Oliveira" pageId="210" pageNumber="210" pagination="355 - 372" refId="ref196481" refString="Carneiro, L. M., and E. V. Oliveira. 2017 a. Systematic affinities of the extinct metatherian Eobrasilia coutoi Simpson, 1947, a South American early Eocene Stagodontidae: implications for " Eobrasiliinae. " Revista Brasileira de Paleontologia 20 (3): 355 - 372." type="journal article" year="2017">Carneiro and Oliveira, 2017a</bibRefCitation>
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,
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<bibRefCitation id="EFD711BAF6E968FBD967FD2119D1FD60" author="Carneiro, L. M. & E. V. Oliveira" box="[876,947,655,679]" pageId="210" pageNumber="210" pagination="497 - 507" refId="ref196536" refString="Carneiro, L. M., and E. V. Oliveira. 2017 b. The Eocene South American metatherian Zeusdelphys complicatus is not a protodidelphid but a hatcheriform: Paleobiogeographic implications. Acta Palaeontologica Polonica 62 (3): 497 - 507." type="journal article" year="2017">2017b</bibRefCitation>
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;
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<bibRefCitation id="EFD711BAF6E968FBD9C9FD211E13FD6F" author="Carneiro, L. M." box="[962,1137,655,680]" pageId="210" pageNumber="210" pagination="88 - 96" refId="ref196389" refString="Carneiro, L. M. 2018. A new species of Varalphadon (Mammalia, Metatheria, Sparassodonta) from the upper Cenomanian of southern Utah, North America: Phylogenetic and biogeographic insights. Cretaceous Research 84: 88 - 96." type="journal article" year="2018">Carneiro, 2018</bibRefCitation>
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,
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<bibRefCitation id="EFD711BAF6E968FBDE74FD3E1ED5FD6F" author="Carneiro, L. M." box="[1151,1207,656,680]" pageId="210" pageNumber="210" pagination="20180440" refId="ref196433" refString="Carneiro, L. M. 2019. A new protodidelphid (Mammalia, Marsupialia, Didelphimorphia) from the Itaborai Basin and its implications for the evolution of the Protodidelphidae. Anais da Academia Brasileira de Ciencias 91 (Suppl. 2): e 20180440." type="journal article" year="2019">2019</bibRefCitation>
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;
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<bibRefCitation id="EFD711BAF6E968FBD8A7FD1E19F9FD0E" author="Carneiro, L. M. & E. V. Oliveira & F. J. Goin" box="[684,923,688,713]" pageId="210" pageNumber="210" pagination="120 - 131" refId="ref196583" refString="Carneiro, L. M., E. V. Oliveira, and F. J. Goin. 2018. Austropediomys marshalli gen. et sp. nov., a new Pediomyoidea (Mammalia, Metatheria) from the Paleogene of Brazil: paleobiogeographic implications. Revista Brasileira de Paleontologia 21 (2): 120 - 131." type="journal article" year="2018">Carneiro et al., 2018</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document>
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