treatments-xml/data/03/EF/87/03EF87E2FF8DA637FF33B94DFEC8FDAF.xml

<document id="83243101553AB4244DC7AE7C9F22F45D" ID-DOI="10.11646/zootaxa.5296.2.6" ID-ISSN="1175-5326" ID-Zenodo-Dep="7973330" ID-ZooBank="ADFA6458-9358-4E88-8715-BF181D5E0B54" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1685086619639" checkinUser="plazi" docAuthor="Paiva, Nathália De Oliveira, Rodrigues, Juliana Mourão Dos Santos, Franco, Cleilton Lima &amp; Moreira, Felipe Ferraz Figueiredo" docDate="2023" docId="03EF87E2FF8DA637FF33B94DFEC8FDAF" docLanguage="en" docName="zootaxa.5296.2.6.pdf" docOrigin="Zootaxa 5296 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5296.2.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rhagovelia leilae Paiva, Rodrigues &amp; Moreira 2023, sp. nov." docType="treatment" docVersion="4" lastPageNumber="256" masterDocId="FFD6FF9AFF8CA630FFA4BF5FFFD2FFCE" masterDocTitle="Description of a new species and new records of Rhagovelia Mayr, 1865 (Hemiptera Heteroptera: Veliidae) from northern and northeastern Brazil" masterLastPageNumber="264" masterPageNumber="250" pageNumber="251" updateTime="1685367487908" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="7E1086558FDE9A7927BE8B07F58058BC">Description of a new species and new records of Rhagovelia Mayr, 1865 (Hemiptera Heteroptera: Veliidae) from northern and northeastern Brazil</mods:title>
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<mods:namePart id="527FA6747C56CD65B3DE7641A2520888">Paiva, Nathália De Oliveira</mods:namePart>
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<mods:affiliation id="D0AF371053D4A5061D4F4DA4E1BB89F7">Fundação Oswaldo Cruz, Instituto Oswaldo Cruz, Laboratório de Entomologia, Avenida Brasil, 4365, Pavilhão Mourisco, sala 214, Manguinhos, Rio de Janeiro, RJ, CEP 21041 - 250, Brazil. &amp; nathaliapaiva. 93 @ gmail. com; https: // orcid. org / 0000 - 0002 - 1693 - 2617</mods:affiliation>
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<mods:namePart id="87C337860710C89CE83025F992AACF62">Rodrigues, Juliana Mourão Dos Santos</mods:namePart>
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<mods:affiliation id="3368058A205914705D89EB4382A9330E">Fundação Oswaldo Cruz, Instituto Oswaldo Cruz, Laboratório de Entomologia, Avenida Brasil, 4365, Pavilhão Mourisco, sala 214, Manguinhos, Rio de Janeiro, RJ, CEP 21041 - 250, Brazil. &amp; julianamourao @ yahoo. com. br; https: // orcid. org / 0000 - 0003 - 2872 - 138 X</mods:affiliation>
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<mods:namePart id="FA1DCFF6505D66FEAFE8F5BD187CC6C7">Franco, Cleilton Lima</mods:namePart>
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<mods:namePart id="B2E8C96A077D9EEB7D73E20B814AA521">Moreira, Felipe Ferraz Figueiredo</mods:namePart>
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<mods:affiliation id="7191E2C1CDDFA25BA89F49F567A09C89">Fundação Oswaldo Cruz, Instituto Oswaldo Cruz, Laboratório de Entomologia, Avenida Brasil, 4365, Pavilhão Mourisco, sala 214, Manguinhos, Rio de Janeiro, RJ, CEP 21041 - 250, Brazil. &amp; ppmeiameiameia @ gmail. com; https: // orcid. org / 0000 - 0002 - 6692 - 0323</mods:affiliation>
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<taxonomicName id="4C464D77FF8DA631FF33B94DFD07F9E2" authority="Paiva, Rodrigues &amp; Moreira" authorityName="Paiva, Rodrigues &amp; Moreira" authorityYear="2023" box="[151,725,1554,1581]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="251" phylum="Arthropoda" rank="species" species="leilae" status="sp. nov.">
<emphasis id="B932EAE6FF8DA631FF33B94DFEBAF9E3" bold="true" box="[151,360,1554,1581]" italics="true" pageId="1" pageNumber="251">Rhagovelia leilae</emphasis>
Paiva, Rodrigues &amp; Moreira
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, 
<taxonomicNameLabel id="A201579DFF8DA631FD47B94CFC92F9E3" box="[739,832,1555,1581]" pageId="1" pageNumber="251" rank="species">sp. nov.</taxonomicNameLabel>
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<figureCitation id="137D2A71FF8DA631FF33B968FF0EF99F" box="[151,220,1591,1617]" captionStart="FIGURE 1" captionStartId="2.[152,255,1934,1958]" captionTargetBox="[152,1436,345,1911]" captionTargetId="figure-106@2.[151,1436,345,1911]" captionTargetPageId="2" captionText="FIGURE 1. Rhagovelia leilae Paiva, Rodrigues &amp; Moreira, sp. nov., male holotype. A. Habitus, dorsal view. B. Habitus, ventral view. C. Hind femur; ventrolateral view. D−F. Abdominal segment VIII. D. Dorsal view. E. Lateral view. F. Ventral view. G. Proctiger, dorsal view. H. Genital capsule, lateral view. Scale bars: 2.0 mm (A, B), 0.2 mm (C−H)." figureDoi="http://doi.org/10.5281/zenodo.7973332" httpUri="https://zenodo.org/record/7973332/files/figure.png" pageId="1" pageNumber="251">Figs 1</figureCitation>
, 
<figureCitation id="137D2A71FF8DA631FF4EB968FEDDF99F" box="[234,271,1591,1617]" captionStart="FIGURE 2" captionStartId="3.[152,255,651,675]" captionTargetBox="[199,1388,185,624]" captionTargetId="figure-20@3.[196,1391,181,627]" captionTargetPageId="3" captionText="FIGURE 2. Parameres. A. Rhagovelia leilae Paiva, Rodrigues &amp; Moreira, sp. nov., male holotype. B. Rhagovelia fontanalis Bacon, 1948 [modified from Bacon (1956)]." figureDoi="http://doi.org/10.5281/zenodo.7973334" httpUri="https://zenodo.org/record/7973334/files/figure.png" pageId="1" pageNumber="251">2A</figureCitation>
, 
<figureCitation id="137D2A71FF8DA631FEBFB969FE95F99F" box="[283,327,1590,1617]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStart-2="FIGURE 5" captionStartId-0="4.[152,255,1414,1438]" captionStartId-1="5.[152,255,1564,1588]" captionStartId-2="6.[152,255,1194,1218]" captionTargetBox-0="[152,1436,185,1389]" captionTargetBox-1="[151,1435,182,1540]" captionTargetBox-2="[154,1433,420,1171]" captionTargetId-0="figure-17@4.[151,1436,181,1389]" captionTargetId-1="figure-20@5.[151,1436,181,1540]" captionTargetId-2="figure-142@6.[151,1436,417,1171]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="FIGURE 3. Rhagovelia leilae Paiva, Rodrigues &amp; Moreira, sp. nov., macropterous male paratype. A. Habitus, dorsal view. B. Habitus, ventral view. C. Hind femur; ventrolateral view. D. Proctiger, dorsal view. E. Genital capsule, lateral view. Scale bars: 2.0 mm (A, B), 0.5 mm (C), 0.1 mm (D, E)." captionText-1="FIGURE 4. Rhagovelia leilae Paiva, Rodrigues &amp; Moreira, sp. nov., apterous female paratypes, habitus. A, B. Specimen from the state of Maranhão. A. Dorsal view. B. Ventral view. C, D. Specimen from the state of Alagoas. C. Dorsal view. D. Ventral view. Scale bars: 2.0 mm." captionText-2="FIGURE 5. Geographic distribution of Rhagovelia leilae Paiva, Rodrigues &amp; Moreira, sp. nov. Red star shows the type locality in Carolina municipality, Maranhão state (highlighted in pink). Green stars show the records from Estação Ecológica de Murici (highlighted in dark green), Murici municipality, Alagoas state (highlighted in light green)." figureDoi-0="http://doi.org/10.5281/zenodo.7973336" figureDoi-1="http://doi.org/10.5281/zenodo.7973338" figureDoi-2="http://doi.org/10.5281/zenodo.7973340" httpUri-0="https://zenodo.org/record/7973336/files/figure.png" httpUri-1="https://zenodo.org/record/7973338/files/figure.png" httpUri-2="https://zenodo.org/record/7973340/files/figure.png" pageId="1" pageNumber="251">3−5</figureCitation>
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Type material. 
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. 
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— 
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<collectingRegion id="4982F816FF8DA631FDFDB920FD05F957" box="[601,727,1663,1689]" country="Brazil" name="Maranhao" pageId="1" pageNumber="251">Maranhão</collectingRegion>
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• 
<taxonomicName id="4C464D77FF8DA631FD5EB920FC76F957" authority=", Sitio" authorityName="Sitio" box="[762,932,1663,1689]" class="Insecta" family="Cetoniidae" genus="Carolina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="251" phylum="Arthropoda" rank="genus">
<collectingMunicipality id="6B9DAC8EFF8DA631FD5EB920FC8FF957" box="[762,861,1663,1689]" pageId="1" pageNumber="251">Carolina</collectingMunicipality>
, 
<location id="8E99602FFF8DA631FCCBB920FBE5F957" LSID="urn:lsid:plazi:treatment:03EF87E2FF8DA637FF33B94DFEC8FDAF:8E99602FFF8DA631FCCBB920FBE5F957" box="[879,1079,1663,1689]" country="Brazil" municipality="Carolina" name="Sitio do Antonio" pageId="1" pageNumber="251" stateProvince="Maranhao">Sítio do Antônio</location>
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, riacho; -07.1270°, -47.3783°; 
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<collectingDate id="EFBCE9DCFF8DA631FF33B9FCFED7F973" box="[151,261,1699,1725]" pageId="1" pageNumber="251" value="2022-01-29">29.I.2022</collectingDate>
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; 
<collectorName id="26B35322FF8DA631FEB1B9FCFE4DF973" box="[277,415,1699,1725]" pageId="1" pageNumber="251">C.L. Franco</collectorName>
leg.; 1 apterous 
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, 
<specimenCode id="DBE09E8FFF8DA631FD09B9FCFC87F973" box="[685,853,1699,1725]" pageId="1" pageNumber="251">CEIOC 82816</specimenCode>
.
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<materialsCitation id="3B2E3CA9FF8DA631FCC6B9FCFC89F92F" collectionCode="CEIOC" country="Brazil" location="Maranhao" pageId="1" pageNumber="251" specimenCode="CEIOC 82817" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Maranhao" typeStatus="paratype">
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<typeStatus id="54FD8856FF8DA631FCC6B9FCFC0EF973" box="[866,988,1699,1725]" pageId="1" pageNumber="251">Paratypes</typeStatus>
.
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<collectingCountry id="F3517664FF8DA631FC4FB9FCFB82F973" box="[1003,1104,1699,1725]" name="Brazil" pageId="1" pageNumber="251">BRAZIL</collectingCountry>
— 
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<collectingRegion id="4982F816FF8DA631FBC8B9FCFB38F973" box="[1132,1258,1699,1725]" country="Brazil" name="Maranhao" pageId="1" pageNumber="251">Maranhão</collectingRegion>
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• same data as holotype; 1 apterous 
<specimenCount id="9D40FD7DFF8DA631FE2CB998FE13F92F" box="[392,449,1735,1761]" pageId="1" pageNumber="251" type="male">male</specimenCount>
, 2 apterous 
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, 
<specimenCode id="DBE09E8FFF8DA631FD16B998FC85F92F" box="[690,855,1735,1761]" pageId="1" pageNumber="251">CEIOC 82817</specimenCode>
.
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<materialsCitation id="3B2E3CA9FF8DA631FCFEB998FA4EF8E7" collectingDate="2019-05-22" collectionCode="CEIOC" collectorName="J. M. S. Rodrigues &amp; W. Sousa &amp; F. F. F. Moreira" elevation="361" location="Estac &quot; o" municipality="Murici" pageId="1" pageNumber="251" specimenCode="CEIOC 82269" specimenCount="1" specimenCount-male="1" stateProvince="Alagoas">
— 
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<collectingRegion id="4982F816FF8DA631FCDCB999FC05F92E" box="[888,983,1734,1760]" country="Brazil" name="Alagoas" pageId="1" pageNumber="251">Alagoas</collectingRegion>
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• 
<collectingMunicipality id="6B9DAC8EFF8DA631FC56B998FB93F92F" box="[1010,1089,1735,1761]" pageId="1" pageNumber="251">Murici</collectingMunicipality>
, 
<location id="8E99602FFF8DA631FBEAB998FB76F92F" LSID="urn:lsid:plazi:treatment:03EF87E2FF8DA637FF33B94DFEC8FDAF:8E99602FFF8DA631FBEAB998FB76F92F" box="[1102,1188,1735,1761]" municipality="Murici" name="Estac &quot; o" pageId="1" pageNumber="251" stateProvince="Alagoas">Estaç ã o</location>
<location id="8E99602FFF8DA631FB09B998FA4AF92F" LSID="urn:lsid:plazi:treatment:03EF87E2FF8DA637FF33B94DFEC8FDAF:8E99602FFF8DA631FB09B998FA4AF92F" box="[1197,1432,1735,1761]" municipality="Murici" name="Ecologica de Murici" pageId="1" pageNumber="251" stateProvince="Alagoas">Ecológica de Murici</location>
, 
<location id="8E99602FFF8DA631FF33B9B4FEF8F8CB" LSID="urn:lsid:plazi:treatment:03EF87E2FF8DA637FF33B94DFEC8FDAF:8E99602FFF8DA631FF33B9B4FEF8F8CB" box="[151,298,1771,1797]" municipality="Murici" name="Vale Socorro" pageId="1" pageNumber="251" stateProvince="Alagoas">Vale Socorró</location>
, 
<location id="8E99602FFF8DA631FE97B9B4FDF6F8CB" LSID="urn:lsid:plazi:treatment:03EF87E2FF8DA637FF33B94DFEC8FDAF:8E99602FFF8DA631FE97B9B4FDF6F8CB" box="[307,548,1771,1797]" municipality="Murici" name="Cachoeira do Socorro" pageId="1" pageNumber="251" stateProvince="Alagoas">Cachoeira do Socorró</location>
[photograph provided by 
<bibRefCitation id="EFD74B05FF8DA631FCE4B9B4FB90F8CB" author="Rodrigues, J. M. S. &amp; Nery, L. &amp; Rodrigues, H. D. D. &amp; Moreira, F. F. F." box="[832,1090,1771,1797]" pageId="1" pageNumber="251" pagination="103 - 159" refId="ref9537" refString="Rodrigues, J. M. S., Nery, L., Rodrigues, H. D. D. &amp; Moreira, F. F. F. (2021) Survey of the semiaquatic bugs (Hemiptera: Heteroptera: Gerromorpha) from Alagoas and Sergipe, Northeast Brazil. Zootaxa, 4958 (1), 103 - 159. https: // doi. org / 10.11646 / zootaxa. 4958.1.9" type="journal article" year="2021">
Rodrigues 
<emphasis id="B932EAE6FF8DA631FC1DB9B4FC3DF8CB" box="[953,1007,1771,1797]" italics="true" pageId="1" pageNumber="251">et al.</emphasis>
(2021)
</bibRefCitation>
: fig. 20]; -09.2360°, -35.8609°; 
<quantity id="4CBE9B11FF8DA631FF33B850FF32F8E7" box="[151,224,1807,1833]" metricMagnitude="2" metricUnit="m" metricValue="3.61" pageId="1" pageNumber="251" unit="m" value="361.0">
<elevation id="006BD1C7FF8DA631FF33B850FF32F8E7" box="[151,224,1807,1833]" metricMagnitude="2" metricUnit="m" metricValue="3.61" pageId="1" pageNumber="251" unit="m" value="361.0">361 m</elevation>
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a.s.l.; 
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<collectingDate id="EFBCE9DCFF8DA631FE8AB850FE71F8E7" box="[302,419,1807,1833]" pageId="1" pageNumber="251" value="2019-05-22">22.V.2019</collectingDate>
</date>
; 
<collectorName id="26B35322FF8DA631FE15B850FDAAF8E7" box="[433,632,1807,1833]" pageId="1" pageNumber="251">J.M.S. Rodrigues</collectorName>
, 
<collectorName id="26B35322FF8DA631FD21B850FC65F8E7" box="[645,951,1807,1833]" pageId="1" pageNumber="251">W. Sousa &amp; F.F.F. Moreira</collectorName>
leg.; 1 macropterous 
<specimenCount id="9D40FD7DFF8DA631FB17B850FB3EF8E7" box="[1203,1260,1807,1833]" pageId="1" pageNumber="251" type="male">male</specimenCount>
, 
<specimenCode id="DBE09E8FFF8DA631FB5DB850FA4EF8E7" box="[1273,1436,1807,1833]" pageId="1" pageNumber="251">CEIOC 82269</specimenCode>
</materialsCitation>
<materialsCitation id="3B2E3CA9FF8DA631FF33B86CFE92F85B" collectingDate="2018-04-28" collectionCode="CEIOC" collectorName="C. F. B. Floriano &amp; J. M. S. Rodrigues &amp; O. M. Magalh &quot; es" elevation="320" location="Estac &quot; o" municipality="Murici" pageId="1" pageNumber="251" specimenCode="CEIOC 82268" specimenCount="1" specimenCount-female="1">
• 
<collectingMunicipality id="6B9DAC8EFF8DA631FF08B86CFF29F883" box="[172,251,1843,1869]" pageId="1" pageNumber="251">Murici</collectingMunicipality>
, 
<location id="8E99602FFF8DA631FEADB86CFEB2F883" LSID="urn:lsid:plazi:treatment:03EF87E2FF8DA637FF33B94DFEC8FDAF:8E99602FFF8DA631FEADB86CFEB2F883" box="[265,352,1843,1869]" municipality="Murici" name="Estac &quot; o" pageId="1" pageNumber="251">Estaç ã o</location>
<location id="8E99602FFF8DA631FECFB86CFD84F883" LSID="urn:lsid:plazi:treatment:03EF87E2FF8DA637FF33B94DFEC8FDAF:8E99602FFF8DA631FECFB86CFD84F883" box="[363,598,1843,1869]" municipality="Murici" name="Ecologica de Murici" pageId="1" pageNumber="251">Ecológica de Murici</location>
[photograph provided by 
<bibRefCitation id="EFD74B05FF8DA631FC2DB86CFB49F883" author="Rodrigues, J. M. S. &amp; Nery, L. &amp; Rodrigues, H. D. D. &amp; Moreira, F. F. F." box="[905,1179,1843,1869]" pageId="1" pageNumber="251" pagination="103 - 159" refId="ref9537" refString="Rodrigues, J. M. S., Nery, L., Rodrigues, H. D. D. &amp; Moreira, F. F. F. (2021) Survey of the semiaquatic bugs (Hemiptera: Heteroptera: Gerromorpha) from Alagoas and Sergipe, Northeast Brazil. Zootaxa, 4958 (1), 103 - 159. https: // doi. org / 10.11646 / zootaxa. 4958.1.9" type="journal article" year="2021">
Rodrigues 
<emphasis id="B932EAE6FF8DA631FBA3B86CFB90F883" box="[1031,1090,1843,1869]" italics="true" pageId="1" pageNumber="251">et al.</emphasis>
(2021)
</bibRefCitation>
: fig. 19]; -09.2379°, - 35.8641°; 
<quantity id="4CBE9B11FF8DA631FEA9B808FE87F8BF" box="[269,341,1879,1905]" metricMagnitude="2" metricUnit="m" metricValue="3.2" pageId="1" pageNumber="251" unit="m" value="320.0">
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a.s.l.; 
<date id="FFF81034FF8DA631FE04B808FDCDF8BF" box="[416,543,1879,1905]" pageId="1" pageNumber="251" value="2018-04-28">
<collectingDate id="EFBCE9DCFF8DA631FE04B808FDCDF8BF" box="[416,543,1879,1905]" pageId="1" pageNumber="251" value="2018-04-28">28.IV.2018</collectingDate>
</date>
, 
<collectorName id="26B35322FF8DA631FD8EB808FD0EF8BF" box="[554,732,1879,1905]" pageId="1" pageNumber="251">C.F.B. Floriano</collectorName>
, 
<collectorName id="26B35322FF8DA631FD4CB808FC79F8BF" box="[744,939,1879,1905]" pageId="1" pageNumber="251">J.M.S. Rodrigues</collectorName>
&amp; 
<collectorName id="26B35322FF8DA631FC74B808FB5FF8BF" box="[976,1165,1879,1905]" pageId="1" pageNumber="251">O.M. Magalh ã es</collectorName>
leg.; 1 apterous 
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, 
<specimenCode id="DBE09E8FFF8DA631FF33B824FEEEF85B" box="[151,316,1915,1941]" pageId="1" pageNumber="251">CEIOC 82268</specimenCode>
.
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</paragraph>
</subSubSection>
<subSubSection id="C35C657FFF8DA635FF63B8C1FCB7F852" lastPageId="5" lastPageNumber="255" pageId="1" pageNumber="251" type="description">
<paragraph id="8BF936F4FF8DA632FF63B8C1FD48FE8F" blockId="1.[151,1436,1662,2013]" lastBlockId="2.[151,1437,151,321]" lastPageId="2" lastPageNumber="252" pageId="1" pageNumber="251">
<emphasis id="B932EAE6FF8DA631FF63B8C1FDC1F877" bold="true" box="[199,531,1950,1977]" pageId="1" pageNumber="251">Description. Apterous male</emphasis>
(
<figureCitation id="137D2A71FF8DA631FD81B8C0FDBDF877" box="[549,623,1951,1977]" captionStart="FIGURE 1" captionStartId="2.[152,255,1934,1958]" captionTargetBox="[152,1436,345,1911]" captionTargetId="figure-106@2.[151,1436,345,1911]" captionTargetPageId="2" captionText="FIGURE 1. Rhagovelia leilae Paiva, Rodrigues &amp; Moreira, sp. nov., male holotype. A. Habitus, dorsal view. B. Habitus, ventral view. C. Hind femur; ventrolateral view. D−F. Abdominal segment VIII. D. Dorsal view. E. Lateral view. F. Ventral view. G. Proctiger, dorsal view. H. Genital capsule, lateral view. Scale bars: 2.0 mm (A, B), 0.2 mm (C−H)." figureDoi="http://doi.org/10.5281/zenodo.7973332" httpUri="https://zenodo.org/record/7973332/files/figure.png" pageId="1" pageNumber="251">Figs 1</figureCitation>
, 
<figureCitation id="137D2A71FF8DA631FD24B8C0FD74F877" box="[640,678,1951,1977]" captionStart="FIGURE 2" captionStartId="3.[152,255,651,675]" captionTargetBox="[199,1388,185,624]" captionTargetId="figure-20@3.[196,1391,181,627]" captionTargetPageId="3" captionText="FIGURE 2. Parameres. A. Rhagovelia leilae Paiva, Rodrigues &amp; Moreira, sp. nov., male holotype. B. Rhagovelia fontanalis Bacon, 1948 [modified from Bacon (1956)]." figureDoi="http://doi.org/10.5281/zenodo.7973334" httpUri="https://zenodo.org/record/7973334/files/figure.png" pageId="1" pageNumber="251">2A</figureCitation>
). 
<typeStatus id="54FD8856FF8DA631FD19B8C0FCF7F877" box="[701,805,1951,1977]" pageId="1" pageNumber="251">Holotype</typeStatus>
(
<typeStatus id="54FD8856FF8DA631FC93B8C0FC72F877" box="[823,928,1951,1977]" pageId="1" pageNumber="251">Paratype</typeStatus>
). BL 3.08 (3.08); HL 0.32 (0.32); HW 0.80 (0.72); ANT I 0.65 (0.72); ANT II 0.35 (0.35); ANT III 0.37 (0.37); ANT IV 0.42 (0.42); EYE 0.24 (0.24); PL 0.20 (0.20); PW 0.80 (0.80); FORELEG, FEM 0.76 (0.84); TIB 0.80 (0.84); TAR I 0.02 (0.02); TAR II 0.02 (0.02); TAR III 0.20 (0.20); MIDLEG, FEM 1.36 (1.40); TIB 0.96 (0.96); TAR I 0.04 (0.04); TAR II 0.44 (0.44); TAR III 0.66 (0.66); HINDLEG, FEM 1.12 (1.12); TIB 1.28 (1.32); TAR I 0.04 (0.04); TAR II 0.12 (0.10); TAR III 0.28 (0.28); ABDT I 0.12 (0.12); ABDT II 0.16 (0.16); ABDT III 0.16 (0.16); ABDT IV 0.16 (0.16); ABDT V 0.20 (0.20); ABDT VI 0.20 (0.20); ABDT VII 0.36 (0.36).
</paragraph>
<caption id="DF39667CFF8EA632FF3CB8D1FB0FF821" ID-DOI="http://doi.org/10.5281/zenodo.7973332" ID-Zenodo-Dep="7973332" httpUri="https://zenodo.org/record/7973332/files/figure.png" pageId="2" pageNumber="252" startId="2.[152,255,1934,1958]" targetBox="[152,1436,345,1911]" targetPageId="2" targetType="figure">
<paragraph id="8BF936F4FF8EA632FF3CB8D1FB0FF821" blockId="2.[151,1436,1934,2031]" pageId="2" pageNumber="252">
<emphasis id="B932EAE6FF8EA632FF3CB8D1FEC9F868" bold="true" box="[152,283,1934,1958]" pageId="2" pageNumber="252">FIGURE 1.</emphasis>
<taxonomicName id="4C464D77FF8EA632FE81B8D1FCDFF868" authority="Paiva, Rodrigues &amp; Moreira" authorityName="Paiva, Rodrigues &amp; Moreira" authorityYear="2023" box="[293,781,1934,1959]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="252" phylum="Arthropoda" rank="species" species="leilae" status="sp. nov.">
<emphasis id="B932EAE6FF8EA632FE81B8D1FE0BF868" box="[293,473,1934,1958]" italics="true" pageId="2" pageNumber="252">Rhagovelia leilae</emphasis>
Paiva, Rodrigues &amp; Moreira
</taxonomicName>
, 
<emphasis id="B932EAE6FF8EA632FCBEB8D0FCBFF869" bold="true" box="[794,877,1935,1959]" pageId="2" pageNumber="252">
<taxonomicNameLabel id="A201579DFF8EA632FCBEB8D0FCBFF869" box="[794,877,1935,1959]" pageId="2" pageNumber="252" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male holotype. 
<emphasis id="B932EAE6FF8EA632FB84B8D1FBEBF868" bold="true" box="[1056,1081,1934,1958]" pageId="2" pageNumber="252">A.</emphasis>
Habitus, dorsal view. 
<emphasis id="B932EAE6FF8EA632FA83B8D0FAECF869" bold="true" box="[1319,1342,1935,1959]" pageId="2" pageNumber="252">B.</emphasis>
Habitus, ventral view. 
<emphasis id="B932EAE6FF8EA632FE80B8EDFEEFF804" bold="true" box="[292,317,1970,1994]" pageId="2" pageNumber="252">C.</emphasis>
Hind femur; ventrolateral view. 
<emphasis id="B932EAE6FF8EA632FD36B8ECFD15F805" bold="true" box="[658,711,1971,1995]" pageId="2" pageNumber="252">D−F.</emphasis>
Abdominal segment VIII. 
<emphasis id="B932EAE6FF8EA632FC7AB8ECFC25F805" bold="true" box="[990,1015,1971,1995]" pageId="2" pageNumber="252">D.</emphasis>
Dorsal view. 
<emphasis id="B932EAE6FF8EA632FB2DB8ECFB72F805" bold="true" box="[1161,1184,1971,1995]" pageId="2" pageNumber="252">E.</emphasis>
Lateral view. 
<emphasis id="B932EAE6FF8EA632FA93B8ECFA98F805" bold="true" box="[1335,1354,1971,1995]" pageId="2" pageNumber="252">F.</emphasis>
Ventral view. 
<emphasis id="B932EAE6FF8EA632FF77B889FF3FF821" bold="true" box="[211,237,2006,2031]" pageId="2" pageNumber="252">G.</emphasis>
Proctiger, dorsal view. 
<emphasis id="B932EAE6FF8EA632FE79B888FE25F821" bold="true" box="[477,503,2007,2031]" pageId="2" pageNumber="252">H.</emphasis>
Genital capsule, lateral view. Scale bars: 2.0 mm (
<emphasis id="B932EAE6FF8EA632FC5EB889FBDEF820" bold="true" box="[1018,1036,2006,2030]" pageId="2" pageNumber="252">A</emphasis>
, 
<emphasis id="B932EAE6FF8EA632FBBDB888FBF8F821" bold="true" box="[1049,1066,2007,2031]" pageId="2" pageNumber="252">B</emphasis>
), 0.2 mm (
<emphasis id="B932EAE6FF8EA632FB3FB889FB1DF820" bold="true" box="[1179,1231,2006,2030]" pageId="2" pageNumber="252">C−H</emphasis>
).
</paragraph>
</caption>
<caption id="DF39667CFF8FA633FF3CBDD4FD8BFD09" ID-DOI="http://doi.org/10.5281/zenodo.7973334" ID-Zenodo-Dep="7973334" httpUri="https://zenodo.org/record/7973334/files/figure.png" pageId="3" pageNumber="253" startId="3.[152,255,651,675]" targetBox="[199,1388,185,624]" targetPageId="3" targetType="figure">
<paragraph id="8BF936F4FF8FA633FF3CBDD4FD8BFD09" blockId="3.[151,1436,651,711]" pageId="3" pageNumber="253">
<emphasis id="B932EAE6FF8FA633FF3CBDD4FECBFD6D" bold="true" box="[152,281,651,675]" pageId="3" pageNumber="253">FIGURE 2.</emphasis>
Parameres. 
<emphasis id="B932EAE6FF8FA633FE33BDD4FE62FD6D" bold="true" box="[407,432,651,675]" pageId="3" pageNumber="253">A.</emphasis>
<taxonomicName id="4C464D77FF8FA633FE13BDD4FC47FD6D" authority="Paiva, Rodrigues &amp; Moreira" authorityName="Paiva, Rodrigues &amp; Moreira" authorityYear="2023" box="[439,917,651,675]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="253" phylum="Arthropoda" rank="species" species="leilae" status="sp. nov.">
<emphasis id="B932EAE6FF8FA633FE13BDD4FDB8FD6D" box="[439,618,651,675]" italics="true" pageId="3" pageNumber="253">Rhagovelia leilae</emphasis>
Paiva, Rodrigues &amp; Moreira
</taxonomicName>
, 
<emphasis id="B932EAE6FF8FA633FC04BDD4FC23FD6D" bold="true" box="[928,1009,651,675]" pageId="3" pageNumber="253">
<taxonomicNameLabel id="A201579DFF8FA633FC04BDD4FC23FD6D" box="[928,1009,651,675]" pageId="3" pageNumber="253" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male holotype. 
<emphasis id="B932EAE6FF8FA633FB3ABDD4FB67FD6D" bold="true" box="[1182,1205,651,675]" pageId="3" pageNumber="253">B.</emphasis>
<taxonomicName id="4C464D77FF8FA633FB18BDD4FECAFD09" authority="Bacon, 1948" authorityName="Bacon" authorityYear="1948" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="253" phylum="Arthropoda" rank="species" species="fontanalis">
<emphasis id="B932EAE6FF8FA633FB18BDD4FA4EFD6D" box="[1212,1436,651,675]" italics="true" pageId="3" pageNumber="253">Rhagovelia fontanalis</emphasis>
<bibRefCitation id="EFD74B05FF8FA633FF33BDF0FECAFD09" author="Bacon, J. A." box="[151,280,687,711]" pageId="3" pageNumber="253" pagination="71 - 87" refId="ref8408" refString="Bacon, J. A. (1948) Some new species of Rhagovelia (Hemiptera, Veliidae). Journal of the Kansas Entomological Society, 21 (3), 71 - 87. [https: // www. jstor. org / stable / 25081858]" type="journal article" year="1948">Bacon, 1948</bibRefCitation>
</taxonomicName>
[modified from 
<bibRefCitation id="EFD74B05FF8FA633FE1BBDF0FD9DFD09" author="Bacon, J. A." box="[447,591,687,711]" pageId="3" pageNumber="253" pagination="695 - 913" refId="ref8458" refString="Bacon, J. A. (1956) A taxonomic study of the genus Rhagovelia (Hemiptera, Veliidae) of the Western Hemisphere. University of Kansas Science Bulletin, 38, 695 - 913. [https: // www. biodiversitylibrary. org / page / 28892618]" type="journal article" year="1956">Bacon (1956)</bibRefCitation>
].
</paragraph>
</caption>
<paragraph id="8BF936F4FF8FA633FF63BC46FA55FA59" blockId="3.[151,1437,793,2044]" pageId="3" pageNumber="253">General color black, covered by golden pubescence. Head dorsally black, covered by short setae, longitudinal midline and a pair of oblique indentations at base impressed and shiny. Eyes grayish-red. Antenniferous tubercle shiny dark-brown. Antennomeres covered by short and medium-sized setae; antennomere I shiny yellow at the base, turning brown then black towards apex, with few thick long black setae; antennomeres II–IV black; II with few thick long black setae. Buccula black; labium dark-brown. Venter of head black. Pronotum mostly black, whitish behind head, with small orange mark behind vertex, covered by medium-sized dark setae. Meso- and metanota, and pro-, meso- and metapleura grayish-black, covered by short golden pubescence and medium-sized black setae. Proacetabulum yellow, with a light-brown spot medially; mesoacetabulum grayish at base, yellowish laterally and mesally, with dark-brown mark medially; metacetabulum yellow. Pro-, meso- and metasterna grayish-blue, covered by short golden setae. Fore and hind coxae yellow; middle coxa light-brown close to acetabulum, then yellow. Fore and hind trochanters yellow, with medium-sized golden setae; middle trochanter dark-brown to black. Fore femur mostly black, dorsum yellow at basal tip, venter yellow at basal 1/3; middle and hind femora, and all tibiae black, covered by short golden setae and several medium-sized black setae. Fore and hind tarsus dark brown, covered by medium-sized brown setae; middle tarsus dark-brown to black, densely covered by medium and long brown setae, with some longer brown setae on anterior surface. Abdominal medio- and laterotergites black, covered by golden pubescence; mediotergites II−VII with shiny black marks increasing in size towards posterior segments, covering almost entire mediotergite VII. Abdominal sterna II−VII grayish-blue, covered by short golden setae; VII with wide orange-brown subquadrate area medially. Abdominal segment VIII shiny black dorsally, orange-brown ventrally.</paragraph>
<paragraph id="8BF936F4FF8FA633FF63BAFEFEB1F979" blockId="3.[151,1437,793,2044]" pageId="3" pageNumber="253">
Head short, compact. Antennomere I thickest, curved laterally; II–III cylindrical; IV fusiform. Labium robust, reaching mesosternum, slightly surpassing fore coxae. Pronotum shorter medially than dorsal eye length, with posterior margin slightly concave. Mesonotum slightly elevated centrally, posterior margin widely rounded. Metanotum short at midline, posterior margin slightly concave centrally. Fore trochanter unarmed. Fore femur thicker at middle than fore tibia; fore tibia thicker at apex, with grasping comb evident. Middle femur without flattening or constriction, thicker basally. Hind femur not surpassing apex of terminalia, thinner at base than middle femur, armed with two medium-sized spines followed by 4−5 smaller spines towards apex (
<figureCitation id="137D2A71FF8FA633FB1CB926FAC0F95D" box="[1208,1298,1657,1683]" captionStart="FIGURE 1" captionStartId="2.[152,255,1934,1958]" captionTargetBox="[152,1436,345,1911]" captionTargetId="figure-106@2.[151,1436,345,1911]" captionTargetPageId="2" captionText="FIGURE 1. Rhagovelia leilae Paiva, Rodrigues &amp; Moreira, sp. nov., male holotype. A. Habitus, dorsal view. B. Habitus, ventral view. C. Hind femur; ventrolateral view. D−F. Abdominal segment VIII. D. Dorsal view. E. Lateral view. F. Ventral view. G. Proctiger, dorsal view. H. Genital capsule, lateral view. Scale bars: 2.0 mm (A, B), 0.2 mm (C−H)." figureDoi="http://doi.org/10.5281/zenodo.7973332" httpUri="https://zenodo.org/record/7973332/files/figure.png" pageId="3" pageNumber="253">Fig. 1C</figureCitation>
). Hind tibia straight, unarmed.
</paragraph>
<paragraph id="8BF936F4FF8FA633FF63B99EFE41F8A5" blockId="3.[151,1437,793,2044]" pageId="3" pageNumber="253">
Abdominal laterotergites slightly elevated; lateral margin slightly sinuous; abdomen wider between segments IV and V. Lengths of abdominal sterna on midline decreasing from II–IV; IV–V subequal, shorter than VI; VII longest, II−III weakly compressed laterally. Abdominal segment VIII cylindrical; posterodorsal margin almost straight (
<figureCitation id="137D2A71FF8FA633FACEB856FF72F889" captionStart="FIGURE 1" captionStartId="2.[152,255,1934,1958]" captionTargetBox="[152,1436,345,1911]" captionTargetId="figure-106@2.[151,1436,345,1911]" captionTargetPageId="2" captionText="FIGURE 1. Rhagovelia leilae Paiva, Rodrigues &amp; Moreira, sp. nov., male holotype. A. Habitus, dorsal view. B. Habitus, ventral view. C. Hind femur; ventrolateral view. D−F. Abdominal segment VIII. D. Dorsal view. E. Lateral view. F. Ventral view. G. Proctiger, dorsal view. H. Genital capsule, lateral view. Scale bars: 2.0 mm (A, B), 0.2 mm (C−H)." figureDoi="http://doi.org/10.5281/zenodo.7973332" httpUri="https://zenodo.org/record/7973332/files/figure.png" pageId="3" pageNumber="253">Figs 1</figureCitation>
D−E). Proctiger with lateral angles and apex rounded (
<figureCitation id="137D2A71FF8FA633FCACB872FCB3F889" box="[776,865,1837,1863]" captionStart="FIGURE 1" captionStartId="2.[152,255,1934,1958]" captionTargetBox="[152,1436,345,1911]" captionTargetId="figure-106@2.[151,1436,345,1911]" captionTargetPageId="2" captionText="FIGURE 1. Rhagovelia leilae Paiva, Rodrigues &amp; Moreira, sp. nov., male holotype. A. Habitus, dorsal view. B. Habitus, ventral view. C. Hind femur; ventrolateral view. D−F. Abdominal segment VIII. D. Dorsal view. E. Lateral view. F. Ventral view. G. Proctiger, dorsal view. H. Genital capsule, lateral view. Scale bars: 2.0 mm (A, B), 0.2 mm (C−H)." figureDoi="http://doi.org/10.5281/zenodo.7973332" httpUri="https://zenodo.org/record/7973332/files/figure.png" pageId="3" pageNumber="253">Fig. 1G</figureCitation>
). Paramere short, suboval, depressed at the median portion (
<figureCitation id="137D2A71FF8FA633FF53B80EFE81F8A5" box="[247,339,1873,1899]" captionStart="FIGURE 1" captionStartId="2.[152,255,1934,1958]" captionTargetBox="[152,1436,345,1911]" captionTargetId="figure-106@2.[151,1436,345,1911]" captionTargetPageId="2" captionText="FIGURE 1. Rhagovelia leilae Paiva, Rodrigues &amp; Moreira, sp. nov., male holotype. A. Habitus, dorsal view. B. Habitus, ventral view. C. Hind femur; ventrolateral view. D−F. Abdominal segment VIII. D. Dorsal view. E. Lateral view. F. Ventral view. G. Proctiger, dorsal view. H. Genital capsule, lateral view. Scale bars: 2.0 mm (A, B), 0.2 mm (C−H)." figureDoi="http://doi.org/10.5281/zenodo.7973332" httpUri="https://zenodo.org/record/7973332/files/figure.png" pageId="3" pageNumber="253">Figs 1H</figureCitation>
, 
<figureCitation id="137D2A71FF8FA633FEC4B80EFE54F8A5" box="[352,390,1873,1899]" captionStart="FIGURE 2" captionStartId="3.[152,255,651,675]" captionTargetBox="[199,1388,185,624]" captionTargetId="figure-20@3.[196,1391,181,627]" captionTargetPageId="3" captionText="FIGURE 2. Parameres. A. Rhagovelia leilae Paiva, Rodrigues &amp; Moreira, sp. nov., male holotype. B. Rhagovelia fontanalis Bacon, 1948 [modified from Bacon (1956)]." figureDoi="http://doi.org/10.5281/zenodo.7973334" httpUri="https://zenodo.org/record/7973334/files/figure.png" pageId="3" pageNumber="253">2A</figureCitation>
).
</paragraph>
<paragraph id="8BF936F4FF8FA633FF63B82AFC97F835" blockId="3.[151,1437,793,2044]" pageId="3" pageNumber="253">
<emphasis id="B932EAE6FF8FA633FF63B82AFE63F841" bold="true" box="[199,433,1909,1935]" pageId="3" pageNumber="253">Macropterous male</emphasis>
(
<figureCitation id="137D2A71FF8FA633FE64B82AFDD7F841" box="[448,517,1909,1935]" captionStart="FIGURE 3" captionStartId="4.[152,255,1414,1438]" captionTargetBox="[152,1436,185,1389]" captionTargetId="figure-17@4.[151,1436,181,1389]" captionTargetPageId="4" captionText="FIGURE 3. Rhagovelia leilae Paiva, Rodrigues &amp; Moreira, sp. nov., macropterous male paratype. A. Habitus, dorsal view. B. Habitus, ventral view. C. Hind femur; ventrolateral view. D. Proctiger, dorsal view. E. Genital capsule, lateral view. Scale bars: 2.0 mm (A, B), 0.5 mm (C), 0.1 mm (D, E)." figureDoi="http://doi.org/10.5281/zenodo.7973336" httpUri="https://zenodo.org/record/7973336/files/figure.png" pageId="3" pageNumber="253">Fig. 3</figureCitation>
). 
<materialsCitation id="3B2E3CA9FF8FA633FDBFB829FA8EF85E" box="[539,1372,1909,1936]" collectionCode="III" pageId="3" pageNumber="253" specimenCount="1" typeStatus="paratype">
<typeStatus id="54FD8856FF8FA633FDBFB829FD53F85E" box="[539,641,1910,1936]" pageId="3" pageNumber="253">Paratype</typeStatus>
. BL 3.20; HL 0.32; HW 0.80; ANT I 0.72; ANT II 0.32; ANT 
<collectionCode id="ED57AE31FF8FA633FAE4B829FA8EF85E" box="[1344,1372,1910,1936]" pageId="3" pageNumber="253">III</collectionCode>
</materialsCitation>
0.32; ANT IV 
<materialsCitation id="3B2E3CA9FF8FA633FF58B8C6FC8BF87A" box="[252,857,1945,1972]" collectionCode="FORELEG, FEM" pageId="3" pageNumber="253" specimenCode="PL 1.28, PW 1.32" specimenCount="1" typeStatus="paratype">
0.45; EYE 0.28; 
<specimenCode id="DBE09E8FFF8FA633FE1EB8C5FDC4F87D" box="[442,534,1945,1972]" collectionCode="PL" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="3" pageNumber="253" type="Museum">PL 1.28</specimenCode>
; 
<specimenCode id="DBE09E8FFF8FA633FD86B8C5FD55F87D" box="[546,647,1945,1972]" collectionCode="PW" country="Thailand" name="Paleontological Collections" pageId="3" pageNumber="253">PW 1.32</specimenCode>
; 
<collectionCode id="ED57AE31FF8FA633FD37B8C6FCC6F87D" box="[659,788,1945,1971]" pageId="3" pageNumber="253">FORELEG</collectionCode>
, 
<collectionCode id="ED57AE31FF8FA633FCBBB8C5FC8BF87A" box="[799,857,1946,1972]" pageId="3" pageNumber="253">FEM</collectionCode>
</materialsCitation>
0.68; TIB 0.72; TAR I -; TAR II -; TAR III - 
<materialsCitation id="3B2E3CA9FF8FA633FACEB8C6FD39F819" collectionCode="MIDLEG" pageId="3" pageNumber="253" specimenCode="FEM 1.36" specimenCount="1" typeStatus="paratype">
[lost before they could be measured]; 
<collectionCode id="ED57AE31FF8FA633FE5BB8E2FDBDF819" box="[511,623,1981,2007]" pageId="3" pageNumber="253">MIDLEG</collectionCode>
, 
<specimenCode id="DBE09E8FFF8FA633FDDCB8E1FD39F819" box="[632,747,1981,2008]" collectionCode="FEM" pageId="3" pageNumber="253">FEM 1.36</specimenCode>
</materialsCitation>
; TIB 0.96; TAR I 0.04; TAR II 0.50; TAR III 
<materialsCitation id="3B2E3CA9FF8FA633FB7FB8E2FEDEF835" collectionCode="HINDLEG" pageId="3" pageNumber="253" specimenCode="FEM 1.08" specimenCount="1" typeStatus="paratype">
0.70; 
<collectionCode id="ED57AE31FF8FA633FABDB8E2FA4AF819" box="[1305,1432,1981,2007]" pageId="3" pageNumber="253">HINDLEG</collectionCode>
, 
<specimenCode id="DBE09E8FFF8FA633FF33B8BDFEDEF835" box="[151,268,2017,2044]" collectionCode="FEM" pageId="3" pageNumber="253">FEM 1.08</specimenCode>
</materialsCitation>
; TIB 1.28; TAR I 0.04; TAR II 0.16; TAR III 0.30.
</paragraph>
<caption id="DF39667CFF88A634FF3CBAD9FD84FA28" ID-DOI="http://doi.org/10.5281/zenodo.7973336" ID-Zenodo-Dep="7973336" httpUri="https://zenodo.org/record/7973336/files/figure.png" pageId="4" pageNumber="254" startId="4.[152,255,1414,1438]" targetBox="[152,1436,185,1389]" targetPageId="4" targetType="figure">
<paragraph id="8BF936F4FF88A634FF3CBAD9FD84FA28" blockId="4.[151,1437,1414,1510]" pageId="4" pageNumber="254">
<emphasis id="B932EAE6FF88A634FF3CBAD9FECAFA50" bold="true" box="[152,280,1414,1438]" pageId="4" pageNumber="254">FIGURE 3.</emphasis>
<taxonomicName id="4C464D77FF88A634FEBABAD9FD2AFA50" authority="Paiva, Rodrigues &amp; Moreira" authorityName="Paiva, Rodrigues &amp; Moreira" authorityYear="2023" box="[286,760,1414,1438]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="254" phylum="Arthropoda" rank="species" species="leilae" status="sp. nov.">
<emphasis id="B932EAE6FF88A634FEBABAD9FE02FA50" box="[286,464,1414,1438]" italics="true" pageId="4" pageNumber="254">Rhagovelia leilae</emphasis>
Paiva, Rodrigues &amp; Moreira
</taxonomicName>
, 
<emphasis id="B932EAE6FF88A634FCA6BAD9FC80FA50" bold="true" box="[770,850,1414,1438]" pageId="4" pageNumber="254">
<taxonomicNameLabel id="A201579DFF88A634FCA6BAD9FC80FA50" box="[770,850,1414,1438]" pageId="4" pageNumber="254" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, macropterous male paratype. 
<emphasis id="B932EAE6FF88A634FB2FBAD9FB71FA50" bold="true" box="[1163,1187,1414,1438]" pageId="4" pageNumber="254">A.</emphasis>
Habitus, dorsal view. 
<emphasis id="B932EAE6FF88A634FA21BAD9FA4EFA50" bold="true" box="[1413,1436,1414,1438]" pageId="4" pageNumber="254">B.</emphasis>
Habitus, ventral view. 
<emphasis id="B932EAE6FF88A634FEDFBAF5FE46FA0C" bold="true" box="[379,404,1450,1474]" pageId="4" pageNumber="254">C.</emphasis>
Hind femur; ventrolateral view. 
<emphasis id="B932EAE6FF88A634FD7BBAF5FD25FA0C" bold="true" box="[735,759,1450,1474]" pageId="4" pageNumber="254">D.</emphasis>
Proctiger, dorsal view. 
<emphasis id="B932EAE6FF88A634FC42BAF5FC2FFA0C" bold="true" box="[998,1021,1450,1474]" pageId="4" pageNumber="254">E.</emphasis>
Genital capsule, lateral view. Scale bars: 2.0 mm (
<emphasis id="B932EAE6FF88A634FF57BA91FED7FA28" bold="true" box="[243,261,1486,1510]" pageId="4" pageNumber="254">A</emphasis>
, 
<emphasis id="B932EAE6FF88A634FEB6BA91FEF1FA28" bold="true" box="[274,291,1486,1510]" pageId="4" pageNumber="254">B</emphasis>
), 0.5 mm (
<emphasis id="B932EAE6FF88A634FE31BA91FE75FA28" bold="true" box="[405,423,1486,1510]" pageId="4" pageNumber="254">C</emphasis>
), 0.1 mm (
<emphasis id="B932EAE6FF88A634FDBCBA91FD9AFA28" bold="true" box="[536,584,1486,1510]" pageId="4" pageNumber="254">D, E</emphasis>
).
</paragraph>
</caption>
<paragraph id="8BF936F4FF88A634FF63B969FE93F8EB" blockId="4.[151,1437,1590,2042]" pageId="4" pageNumber="254">
Similar to apterous male in general color, pilosity and structure, with some exceptions. Middle coxa dark-brown to black. Dorsum of fore femur entirely black; venter black, with yellow mark at basal tip. Abdominal segment VIII shiny black dorsally, orange-brown to dark-brown ventrally. Pronotum long, subpentagonal; posterior lobe with several punctures; posterior margin slightly pointed. Wings present, but autotomized above abdominal segment VI. Forewing covered by short black setae along anterior margin, with two long basal cells; base of one apical cell visible despite autotomy. Hind femur armed with two medium-sized spines followed by 3 smaller spines towards apex (
<figureCitation id="137D2A71FF88A634FF7FB854FEE6F8EB" box="[219,308,1803,1829]" captionStart="FIGURE 3" captionStartId="4.[152,255,1414,1438]" captionTargetBox="[152,1436,185,1389]" captionTargetId="figure-17@4.[151,1436,181,1389]" captionTargetPageId="4" captionText="FIGURE 3. Rhagovelia leilae Paiva, Rodrigues &amp; Moreira, sp. nov., macropterous male paratype. A. Habitus, dorsal view. B. Habitus, ventral view. C. Hind femur; ventrolateral view. D. Proctiger, dorsal view. E. Genital capsule, lateral view. Scale bars: 2.0 mm (A, B), 0.5 mm (C), 0.1 mm (D, E)." figureDoi="http://doi.org/10.5281/zenodo.7973336" httpUri="https://zenodo.org/record/7973336/files/figure.png" pageId="4" pageNumber="254">Fig. 3C</figureCitation>
).
</paragraph>
<paragraph id="8BF936F4FF88A634FF63B871FF02F834" blockId="4.[151,1437,1590,2042]" pageId="4" pageNumber="254">
<emphasis id="B932EAE6FF88A634FF63B871FE5DF886" bold="true" box="[199,399,1838,1864]" pageId="4" pageNumber="254">Apterous female</emphasis>
(
<figureCitation id="137D2A71FF88A634FE07B870FE3FF887" box="[419,493,1839,1865]" captionStart="FIGURE 4" captionStartId="5.[152,255,1564,1588]" captionTargetBox="[151,1435,182,1540]" captionTargetId="figure-20@5.[151,1436,181,1540]" captionTargetPageId="5" captionText="FIGURE 4. Rhagovelia leilae Paiva, Rodrigues &amp; Moreira, sp. nov., apterous female paratypes, habitus. A, B. Specimen from the state of Maranhão. A. Dorsal view. B. Ventral view. C, D. Specimen from the state of Alagoas. C. Dorsal view. D. Ventral view. Scale bars: 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7973338" httpUri="https://zenodo.org/record/7973338/files/figure.png" pageId="4" pageNumber="254">Fig. 4</figureCitation>
). 
<typeStatus id="54FD8856FF88A634FDA2B870FDA5F887" box="[518,631,1839,1865]" pageId="4" pageNumber="254">Paratypes</typeStatus>
. BL 3.20−3.36; HL 0.32−0.36; HW 0.80; ANT I 0.62−0.67; ANT II 0.32−0.35; ANT III 0.35; ANT IV 0.42; EYE 0.24−0.28; PL 0.20−0.24; PW 0.80; FORE LEG, FEM 0.84; TIB 0.84; TAR I 0.02; TAR II 0.02−0.04; TAR III 0.22−0.24; MID 
<materialsCitation id="3B2E3CA9FF88A634FCBEB829FE73F87D" collectionCode="LEG" pageId="4" pageNumber="254" specimenCode="FEM 1.40, 1.44" specimenCount="1" typeStatus="paratype">
<collectionCode id="ED57AE31FF88A634FCBEB829FC81F85E" box="[794,851,1910,1936]" pageId="4" pageNumber="254">LEG</collectionCode>
, 
<specimenCode id="DBE09E8FFF88A634FCFBB829FC01F85E" box="[863,979,1910,1936]" collectionCode="FEM" pageId="4" pageNumber="254">FEM 1.40</specimenCode>
− 
<specimenCode id="DBE09E8FFF88A634FC47B829FBC4F85E" box="[995,1046,1910,1936]" collectionCode="LEG" pageId="4" pageNumber="254">1.44</specimenCode>
; TIB 0.96−1.00; TAR I 0.04−0.06; TAR II 0.44−0.48; TAR
</materialsCitation>
III 0.66−0.68; 
<materialsCitation id="3B2E3CA9FF88A634FDEDB8C5FA4EF87D" box="[585,1436,1945,1972]" collectionCode="HIND, LEG" pageId="4" pageNumber="254" specimenCode="FEM 1.08" specimenCount="1" typeStatus="paratype">
<collectionCode id="ED57AE31FF88A634FDEDB8C5FD1CF87D" box="[585,718,1945,1972]" pageId="4" pageNumber="254">HIND LEG</collectionCode>
, 
<specimenCode id="DBE09E8FFF88A634FD7CB8C5FC9EF87D" box="[728,844,1945,1972]" collectionCode="FEM" pageId="4" pageNumber="254">FEM 1.08</specimenCode>
; TIB 1.24−1.28; TAR I 0.04; TAR II 0.10−0.12; TAR
</materialsCitation>
III 0.28; ABDT I 0.12; ABDT II 0.16; ABDT III 0.20; ABDT IV 0.20; ABDT V 0.24; ABDT VI 0.24; ABDT VII 0.32.
</paragraph>
<caption id="DF39667CFF89A635FF3CB943FE4FF9B2" ID-DOI="http://doi.org/10.5281/zenodo.7973338" ID-Zenodo-Dep="7973338" httpUri="https://zenodo.org/record/7973338/files/figure.png" pageId="5" pageNumber="255" startId="5.[152,255,1564,1588]" targetBox="[151,1435,182,1540]" targetPageId="5" targetType="figure">
<paragraph id="8BF936F4FF89A635FF3CB943FE4FF9B2" blockId="5.[151,1436,1564,1660]" pageId="5" pageNumber="255">
<emphasis id="B932EAE6FF89A635FF3CB943FECAF9FA" bold="true" box="[152,280,1564,1588]" pageId="5" pageNumber="255">FIGURE 4.</emphasis>
<taxonomicName id="4C464D77FF89A635FEBAB943FD24F9FA" authority="Paiva, Rodrigues &amp; Moreira" authorityName="Paiva, Rodrigues &amp; Moreira" authorityYear="2023" box="[286,758,1564,1588]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="255" phylum="Arthropoda" rank="species" species="leilae" status="sp. nov.">
<emphasis id="B932EAE6FF89A635FEBAB943FE1DF9FA" box="[286,463,1564,1588]" italics="true" pageId="5" pageNumber="255">Rhagovelia leilae</emphasis>
Paiva, Rodrigues &amp; Moreira
</taxonomicName>
, 
<emphasis id="B932EAE6FF89A635FD5BB943FC9DF9FA" bold="true" box="[767,847,1564,1588]" pageId="5" pageNumber="255">
<taxonomicNameLabel id="A201579DFF89A635FD5BB943FC9DF9FA" box="[767,847,1564,1588]" pageId="5" pageNumber="255" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, apterous female paratypes, habitus. 
<emphasis id="B932EAE6FF89A635FB61B943FB29F9FA" bold="true" box="[1221,1275,1564,1588]" pageId="5" pageNumber="255">A, B.</emphasis>
Specimen from the state of Maranh ã o. 
<emphasis id="B932EAE6FF89A635FE26B91FFE49F996" bold="true" box="[386,411,1600,1624]" pageId="5" pageNumber="255">A.</emphasis>
Dorsal view. 
<emphasis id="B932EAE6FF89A635FD8DB91FFD93F996" bold="true" box="[553,577,1600,1624]" pageId="5" pageNumber="255">B.</emphasis>
Ventral view. 
<emphasis id="B932EAE6FF89A635FD71B91FFCDFF996" bold="true" box="[725,781,1600,1624]" pageId="5" pageNumber="255">C, D.</emphasis>
Specimen from the state of Alagoas. 
<emphasis id="B932EAE6FF89A635FB29B91FFB77F996" bold="true" box="[1165,1189,1600,1624]" pageId="5" pageNumber="255">C.</emphasis>
Dorsal view. 
<emphasis id="B932EAE6FF89A635FA90B91FFA9FF996" bold="true" box="[1332,1357,1600,1624]" pageId="5" pageNumber="255">D.</emphasis>
Ventral view. Scale bars: 2.0 mm.
</paragraph>
</caption>
<paragraph id="8BF936F4FF89A635FF63B991FCB7F852" blockId="5.[151,1437,1742,2021]" pageId="5" pageNumber="255">Similar to apterous male in general color, pilosity and structure, with following exceptions. Middle coxa light-brown close to acetabulum, then dark-brown, or more uniformly dark-brown to black. Dorsum of fore femur black, with yellow mark at basal tip; venter black, with yellow mark on basal 1/3 to almost 1/2. Hind femur armed with 4−5 spines, the first one longer than the others. Posterolateral angle of abdominal laterotergite VII with tuft of black setae. Abdominal laterotergites horizontal to slightly elevated. Contrasting subquadrate area of abdominal sternum VII mostly dark-brown to black, orange-brown only posteriorly.</paragraph>
</subSubSection>
<subSubSection id="C35C657FFF89A635FF63B8F8FAE6F80E" box="[199,1332,1958,1985]" pageId="5" pageNumber="255" type="etymology">
<paragraph id="8BF936F4FF89A635FF63B8F8FAE6F80E" blockId="5.[151,1437,1742,2021]" box="[199,1332,1958,1985]" pageId="5" pageNumber="255">
<emphasis id="B932EAE6FF89A635FF63B8F8FE9CF80F" bold="true" box="[199,334,1959,1985]" pageId="5" pageNumber="255">Etymology.</emphasis>
The new species is named in honor of Leila de Oliveira Paiva, mother of the first author.
</paragraph>
</subSubSection>
<subSubSection id="C35C657FFF89A637FF63B895FEC8FDAF" lastPageId="7" lastPageNumber="257" pageId="5" pageNumber="255" type="discussion">
<paragraph id="8BF936F4FF89A636FF63B895FB57FE47" blockId="5.[151,1437,1742,2021]" lastBlockId="6.[151,1437,151,393]" lastPageId="6" lastPageNumber="256" pageId="5" pageNumber="255">
<emphasis id="B932EAE6FF89A635FF63B895FE9DF82A" bold="true" box="[199,335,1994,2020]" pageId="5" pageNumber="255">Comments.</emphasis>
<taxonomicName id="4C464D77FF89A635FEF2B895FCB3F82A" authority="Paiva, Rodrigues &amp; Moreira" authorityName="Paiva, Rodrigues &amp; Moreira" authorityYear="2023" box="[342,865,1994,2021]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="255" phylum="Arthropoda" rank="species" species="leilae" status="sp. nov.">
<emphasis id="B932EAE6FF89A635FEF2B895FDC8F82A" box="[342,538,1994,2020]" italics="true" pageId="5" pageNumber="255">Rhagovelia leilae</emphasis>
Paiva, Rodrigues &amp; Moreira
</taxonomicName>
, 
<emphasis id="B932EAE6FF89A635FCC8B894FC17F82B" bold="true" box="[876,965,1995,2021]" pageId="5" pageNumber="255">
<taxonomicNameLabel id="A201579DFF89A635FCC8B894FC17F82B" box="[876,965,1995,2021]" pageId="5" pageNumber="255" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
belongs to the 
<taxonomicName id="4C464D77FF89A635FBD4B894FB3AF82A" authorityName="Uhler" authorityYear="1894" box="[1136,1256,1995,2020]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="255" phylum="Arthropoda" rank="species" species="angustipes">
<emphasis id="B932EAE6FF89A635FBD4B894FB3AF82A" box="[1136,1256,1995,2020]" italics="true" pageId="5" pageNumber="255">angustipes</emphasis>
</taxonomicName>
complex, based on the pronotum of the apterous form shorter than the dorsal length of the eye, with the posterior margin slightly concave. Because of the tarsal formula 3-3-3 and the occurrence in freshwater, it is certainly not part of the 
<taxonomicName id="4C464D77FF8AA636FAFCBFE4FA4EFF1B" baseAuthorityName="Champion" baseAuthorityYear="1898" box="[1368,1436,187,213]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="256" phylum="Arthropoda" rank="species" species="salina">
<emphasis id="B932EAE6FF8AA636FAFCBFE4FA4EFF1B" box="[1368,1436,187,213]" italics="true" pageId="6" pageNumber="256">salina</emphasis>
</taxonomicName>
group. However, it cannot be assigned to either the 
<taxonomicName id="4C464D77FF8AA636FD45BF80FC9BFF37" authorityName="Bacon" authorityYear="1948" box="[737,841,223,249]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="256" phylum="Arthropoda" rank="species" species="bisignata">
<emphasis id="B932EAE6FF8AA636FD45BF80FC9BFF37" box="[737,841,223,249]" italics="true" pageId="6" pageNumber="256">bisignata</emphasis>
</taxonomicName>
or 
<taxonomicName id="4C464D77FF8AA636FCD5BF80FC3DFF37" authorityName="Drake &amp; Harris" authorityYear="1933" box="[881,1007,223,249]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="256" phylum="Arthropoda" rank="species" species="hambletoni">
<emphasis id="B932EAE6FF8AA636FCD5BF80FC3DFF37" box="[881,1007,223,249]" italics="true" pageId="6" pageNumber="256">hambletoni</emphasis>
</taxonomicName>
groups because the wings of the only macropterous specimen available are autotomized, making it impossible to know for certain if there are three or four closed cells on the forewing. Although species with unknown condition of the forewings were previously treated tentatively as part of the 
<taxonomicName id="4C464D77FF8AA636FE02BE14FDDCFEAB" authorityName="Bacon" authorityYear="1948" box="[422,526,331,357]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="256" phylum="Arthropoda" rank="species" species="bisignata">
<emphasis id="B932EAE6FF8AA636FE02BE14FDDCFEAB" box="[422,526,331,357]" italics="true" pageId="6" pageNumber="256">bisignata</emphasis>
</taxonomicName>
group (
<bibRefCitation id="EFD74B05FF8AA636FDC6BE14FCC1FEAB" author="Polhemus, D. A." box="[610,787,331,357]" pageId="6" pageNumber="256" refId="ref9495" refString="Polhemus, D. A. (1997) Systematics of the genus Rhagovelia Mayr (Heteroptera: Veliidae) in the Western Hemisphere (exclusive of the angustipes complex). Entomological Society of America, Lanham, 386 pp." type="book" year="1997">Polhemus 1997</bibRefCitation>
; 
<bibRefCitation id="EFD74B05FF8AA636FCBABE14FB84FEAB" author="Padilla-Gil, D. N. &amp; Moreira, F. F. F." box="[798,1110,331,357]" pageId="6" pageNumber="256" pagination="409 - 424" refId="ref9432" refString="Padilla-Gil, D. N. &amp; Moreira, F. F. F. (2013) Checklist, taxonomy and distribution of the Rhagovelia Mayr, 1865 (Hemiptera: Heteroptera: Veliidae) of the Americas. Zootaxa, 3640 (3), 409 - 424. https: // doi. org / 10.11646 / zootaxa. 3640.3.5" type="journal article" year="2013">Padilla-Gil &amp; Moreira 2013</bibRefCitation>
), we consider it more prudent to treat them (including the new species) as 
<emphasis id="B932EAE6FF8AA636FD20BE30FCCDFE47" box="[644,799,367,393]" italics="true" pageId="6" pageNumber="256">incertae sedis</emphasis>
within the 
<taxonomicName id="4C464D77FF8AA636FC3ABE30FBC4FE46" authorityName="Uhler" authorityYear="1894" box="[926,1046,367,392]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="256" phylum="Arthropoda" rank="species" species="angustipes">
<emphasis id="B932EAE6FF8AA636FC3ABE30FBC4FE46" box="[926,1046,367,392]" italics="true" pageId="6" pageNumber="256">angustipes</emphasis>
</taxonomicName>
complex.
</paragraph>
<caption id="DF39667CFF8AA636FF3CBBF5FBE2FAC4" ID-DOI="http://doi.org/10.5281/zenodo.7973340" ID-Zenodo-Dep="7973340" httpUri="https://zenodo.org/record/7973340/files/figure.png" pageId="6" pageNumber="256" startId="6.[152,255,1194,1218]" targetBox="[154,1433,420,1171]" targetPageId="6" targetType="figure">
<paragraph id="8BF936F4FF8AA636FF3CBBF5FBE2FAC4" blockId="6.[151,1436,1194,1290]" pageId="6" pageNumber="256">
<emphasis id="B932EAE6FF8AA636FF3CBBF5FEC5FB0C" bold="true" box="[152,279,1194,1218]" pageId="6" pageNumber="256">FIGURE 5.</emphasis>
Geographic distribution of 
<taxonomicName id="4C464D77FF8AA636FD88BBF5FBD3FB0C" authority="Paiva, Rodrigues &amp; Moreira" authorityName="Paiva, Rodrigues &amp; Moreira" authorityYear="2023" box="[556,1025,1194,1218]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="256" phylum="Arthropoda" rank="species" species="leilae" status="sp. nov.">
<emphasis id="B932EAE6FF8AA636FD88BBF5FD0FFB0C" box="[556,733,1194,1218]" italics="true" pageId="6" pageNumber="256">Rhagovelia leilae</emphasis>
Paiva, Rodrigues &amp; Moreira
</taxonomicName>
, 
<emphasis id="B932EAE6FF8AA636FBAEBBF5FB8BFB0C" bold="true" box="[1034,1113,1194,1218]" pageId="6" pageNumber="256">
<taxonomicNameLabel id="A201579DFF8AA636FBAEBBF5FB8BFB0C" box="[1034,1113,1194,1218]" pageId="6" pageNumber="256" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Red star shows the type locality in Carolina municipality, Maranh ã o state (highlighted in pink). Green stars show the records from Estaç ã o Ecológica de Murici (highlighted in dark green), Murici municipality, Alagoas state (highlighted in light green).
</paragraph>
</caption>
<paragraph id="8BF936F4FF8AA636FF63BA02FB6FF958" blockId="6.[151,1437,1372,2011]" pageId="6" pageNumber="256">
Among South American species of the 
<taxonomicName id="4C464D77FF8AA636FDDFBA03FD31FAB8" authorityName="Bacon" authorityYear="1948" box="[635,739,1372,1398]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="256" phylum="Arthropoda" rank="species" species="bisignata">
<emphasis id="B932EAE6FF8AA636FDDFBA03FD31FAB8" box="[635,739,1372,1398]" italics="true" pageId="6" pageNumber="256">bisignata</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C464D77FF8AA636FCBCBA03FC44FAB8" authorityName="Drake &amp; Harris" authorityYear="1933" box="[792,918,1372,1398]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="256" phylum="Arthropoda" rank="species" species="hambletoni">
<emphasis id="B932EAE6FF8AA636FCBCBA03FC44FAB8" box="[792,918,1372,1398]" italics="true" pageId="6" pageNumber="256">hambletoni</emphasis>
</taxonomicName>
groups, 
<taxonomicName id="4C464D77FF8AA636FC52BA03FF26FA54" authority="Paiva, Rodrigues &amp; Moreira" authorityName="Paiva, Rodrigues &amp; Moreira" authorityYear="2023" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="256" phylum="Arthropoda" rank="species" species="leilae" status="sp. nov.">
<emphasis id="B932EAE6FF8AA636FC52BA03FB68FAB8" box="[1014,1210,1372,1398]" italics="true" pageId="6" pageNumber="256">Rhagovelia leilae</emphasis>
Paiva, Rodrigues &amp; Moreira
</taxonomicName>
, 
<emphasis id="B932EAE6FF8AA636FEA4BADEFE88FA55" bold="true" box="[256,346,1409,1435]" pageId="6" pageNumber="256">
<taxonomicNameLabel id="A201579DFF8AA636FEA4BADEFE88FA55" box="[256,346,1409,1435]" pageId="6" pageNumber="256" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be diagnosed by the following combination of characteristics: 1) body length 
<quantity id="4CBE9B11FF8AA636FB53BADEFA4FFA55" box="[1271,1437,1409,1435]" metricMagnitude="-3" metricUnit="m" metricValue="3.14" metricValueMax="3.2" metricValueMin="3.08" pageId="6" pageNumber="256" unit="mm" value="3.14" valueMax="3.2" valueMin="3.08">3.08−3.20 mm</quantity>
in the males and 
<quantity id="4CBE9B11FF8AA636FEF7BAFBFE25FA71" box="[339,503,1444,1471]" metricMagnitude="-3" metricUnit="m" metricValue="3.28" metricValueMax="3.36" metricValueMin="3.2" pageId="6" pageNumber="256" unit="mm" value="3.28" valueMax="3.36" valueMin="3.2">3.20−3.36 mm</quantity>
in the females; 2) fore and hind coxae and trochanters yellow; 3) fore femur mostly black, with variable basal yellow marks; 4) middle trochanter dark-brown to black; 5) abdominal mediotergites II−VII with shiny black areas increasing towards the posterior segments; 6) abdominal sterna grayish-blue, with a large contrasting area medially on segment VII; 7) all trochanters and tibiae unarmed in both sexes; 8) hind femur not reaching apex of the terminalia, with a row of 6−7 spines in the males and 4−5 spines in the females; 9) abdominal laterotergites horizontal to slightly elevated, with a tuft of black setae at the posterolateral angle of the last segment in females; and 10) paramere short, suboval, and depressed at the median portion.
</paragraph>
<paragraph id="8BF936F4FF8AA636FF63B9FFFCBDF8A1" blockId="6.[151,1437,1372,2011]" pageId="6" pageNumber="256">
Although the fore femur and middle coxa vary between our material from Maranh ã o (lighter pattern) and that from 
<collectingRegion id="4982F816FF8AA636FF76B99BFEFCF910" box="[210,302,1732,1758]" country="Brazil" name="Alagoas" pageId="6" pageNumber="256">Alagoas</collectingRegion>
(darker pattern), we consider this to be of intraspecific nature only, because there are many similarities in other aspects of color and structure shared by them. Variations on the color of leg segments or on the pattern of shiny abdominal areas are already known for other species of the 
<taxonomicName id="4C464D77FF8AA636FCD0B852FC3EF8E8" authorityName="Uhler" authorityYear="1894" box="[884,1004,1805,1830]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="256" phylum="Arthropoda" rank="species" species="angustipes">
<emphasis id="B932EAE6FF8AA636FCD0B852FC3EF8E8" box="[884,1004,1805,1830]" italics="true" pageId="6" pageNumber="256">angustipes</emphasis>
</taxonomicName>
complex, such as 
<taxonomicName id="4C464D77FF8AA636FB1DB852FF1DF885" authority="Uhler, 1894" authorityName="Uhler" authorityYear="1894" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="256" phylum="Arthropoda" rank="species" species="angustipes">
<emphasis id="B932EAE6FF8AA636FB1DB852FA82F8E8" box="[1209,1360,1805,1830]" italics="true" pageId="6" pageNumber="256">R. angustipes</emphasis>
Uhler, 1894
</taxonomicName>
(
<bibRefCitation id="EFD74B05FF8AA636FF7BB86EFDBAF884" author="De Kort-Gommers, M. &amp; Nieser, N." box="[223,616,1840,1867]" pageId="6" pageNumber="256" pagination="72 - 87" refId="ref8654" refString="De Kort-Gommers, M. &amp; Nieser, N. (1969) Records of Antillean water-striders. Studies on the Fauna of Curacao and other Caribbean Islands, 30 (112), 72 - 87. [https: // repository. naturalis. nl / pub / 506095]" type="journal article" year="1969">De Kort-Gommers &amp; Nieser 1969</bibRefCitation>
; 
<bibRefCitation id="EFD74B05FF8AA636FDD1B86FFC6DF885" author="Galindo-Malagon, X. A. &amp; Morales, I. &amp; Moreira, F. F. F." box="[629,959,1840,1867]" pageId="6" pageNumber="256" pagination="167 - 225" refId="ref8804" refString="Galindo-Malagon, X. A., Morales, I. &amp; Moreira, F. F. F. (2021) Revision of the Rhagovelia angustipes complex (Insecta: Hemiptera: Veliidae) from Colombia. Zootaxa, 4958 (1), 167 - 225. https: // doi. org / 10.11646 / zootaxa. 4958.1.11" type="journal article" year="2021">
Galindo-Malagón 
<emphasis id="B932EAE6FF8AA636FCE1B86EFCACF884" box="[837,894,1840,1866]" italics="true" pageId="6" pageNumber="256">et al.</emphasis>
2021
</bibRefCitation>
) and 
<taxonomicName id="4C464D77FF8AA636FBA5B86EFA91F885" authority="Champion, 1898" authorityName="Champion" authorityYear="1898" box="[1025,1347,1840,1867]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="256" phylum="Arthropoda" rank="species" species="tenuipes">
<emphasis id="B932EAE6FF8AA636FBA5B86EFBACF884" box="[1025,1150,1841,1866]" italics="true" pageId="6" pageNumber="256">R. tenuipes</emphasis>
<bibRefCitation id="EFD74B05FF8AA636FB23B86FFA91F885" author="Champion, G. C." box="[1159,1347,1840,1867]" pageId="6" pageNumber="256" refId="ref8535" refString="Champion, G. C. (1898) Insecta: Rhynchota (Hemiptera - Heteroptera). Vol. II. In: Godwin, F. &amp; Salvin, O. (Eds.), Biologia Centrali-Americana. pp. i - xvi + 1 - 416, 22 pls." type="journal volume" year="1898">Champion, 1898</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="EFD74B05FF8AA636FAF7B86EFF00F8A1" author="Bacon, J. A." pageId="6" pageNumber="256" pagination="695 - 913" refId="ref8458" refString="Bacon, J. A. (1956) A taxonomic study of the genus Rhagovelia (Hemiptera, Veliidae) of the Western Hemisphere. University of Kansas Science Bulletin, 38, 695 - 913. [https: // www. biodiversitylibrary. org / page / 28892618]" type="journal article" year="1956">Bacon 1956</bibRefCitation>
; 
<bibRefCitation id="EFD74B05FF8AA636FF7AB80BFDDCF8A1" author="Nieser, N. &amp; Polhemus, D. A." box="[222,526,1876,1903]" pageId="6" pageNumber="256" pagination="53 - 76" refId="ref9365" refString="Nieser, N. &amp; Polhemus, D. A. (1999) Four new species of Rhagovelia (Heteroptera: Veliidae) from Minas Gerais (Brazil), with a key to the regional species of the angustipes complex. Aquatic Insects, 21 (1), 53 - 76. https: // doi. org / 10.1076 / aqin. 21.1.53.4540" type="journal article" year="1999">Nieser and Polhemus 1999</bibRefCitation>
; 
<bibRefCitation id="EFD74B05FF8AA636FDBEB80BFCB1F8A1" author="Galindo-Malagon, X. A. &amp; Morales, I. &amp; Moreira, F. F. F." box="[538,867,1876,1903]" pageId="6" pageNumber="256" pagination="167 - 225" refId="ref8804" refString="Galindo-Malagon, X. A., Morales, I. &amp; Moreira, F. F. F. (2021) Revision of the Rhagovelia angustipes complex (Insecta: Hemiptera: Veliidae) from Colombia. Zootaxa, 4958 (1), 167 - 225. https: // doi. org / 10.11646 / zootaxa. 4958.1.11" type="journal article" year="2021">
Galindo-Malagón 
<emphasis id="B932EAE6FF8AA636FD4CB80AFCF2F8A0" box="[744,800,1876,1902]" italics="true" pageId="6" pageNumber="256">et al.</emphasis>
2021
</bibRefCitation>
).
</paragraph>
<paragraph id="8BF936F4FF8AA637FF63B827FEC8FDAF" blockId="6.[151,1437,1372,2011]" lastBlockId="7.[151,1437,151,609]" lastPageId="7" lastPageNumber="257" pageId="6" pageNumber="256">
Running our specimens through the keys to the 
<taxonomicName id="4C464D77FF8AA636FD53B826FCBDF85C" authorityName="Uhler" authorityYear="1894" box="[759,879,1913,1938]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="256" phylum="Arthropoda" rank="species" species="angustipes">
<emphasis id="B932EAE6FF8AA636FD53B826FCBDF85C" box="[759,879,1913,1938]" italics="true" pageId="6" pageNumber="256">angustipes</emphasis>
</taxonomicName>
complex from southeastern and southern 
<collectingCountry id="F3517664FF8AA636FAFCB827FA4EF85C" box="[1368,1436,1912,1938]" name="Brazil" pageId="6" pageNumber="256">Brazil</collectingCountry>
(
<bibRefCitation id="EFD74B05FF8AA636FF3BB8C3FE6EF879" author="Nieser, N. &amp; Polhemus, D. A." box="[159,444,1948,1975]" pageId="6" pageNumber="256" pagination="53 - 76" refId="ref9365" refString="Nieser, N. &amp; Polhemus, D. A. (1999) Four new species of Rhagovelia (Heteroptera: Veliidae) from Minas Gerais (Brazil), with a key to the regional species of the angustipes complex. Aquatic Insects, 21 (1), 53 - 76. https: // doi. org / 10.1076 / aqin. 21.1.53.4540" type="journal article" year="1999">Nieser &amp; Polhemus 1999</bibRefCitation>
) and from 
<collectingCountry id="F3517664FF8AA636FD9CB8C3FD74F878" box="[568,678,1948,1974]" name="Colombia" pageId="6" pageNumber="256">Colombia</collectingCountry>
(
<bibRefCitation id="EFD74B05FF8AA636FD11B8C3FC2EF879" author="Galindo-Malagon, X. A. &amp; Morales, I. &amp; Moreira, F. F. F." box="[693,1020,1948,1975]" pageId="6" pageNumber="256" pagination="167 - 225" refId="ref8804" refString="Galindo-Malagon, X. A., Morales, I. &amp; Moreira, F. F. F. (2021) Revision of the Rhagovelia angustipes complex (Insecta: Hemiptera: Veliidae) from Colombia. Zootaxa, 4958 (1), 167 - 225. https: // doi. org / 10.11646 / zootaxa. 4958.1.11" type="journal article" year="2021">
Galindo-Malagón 
<emphasis id="B932EAE6FF8AA636FC20B8C2FC6FF878" box="[900,957,1948,1974]" italics="true" pageId="6" pageNumber="256">et al.</emphasis>
2021
</bibRefCitation>
) ends in no logical results. However, they key to 
<taxonomicName id="4C464D77FF8AA636FEB3B89EFDE4F815" authority="Bacon, 1948" authorityName="Bacon" authorityYear="1948" box="[279,566,1984,2011]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="256" phylum="Arthropoda" rank="species" species="fontanalis">
<emphasis id="B932EAE6FF8AA636FEB3B89EFE77F814" box="[279,421,1984,2010]" italics="true" pageId="6" pageNumber="256">R. fontanalis</emphasis>
<bibRefCitation id="EFD74B05FF8AA636FE0EB89EFDE4F815" author="Bacon, J. A." box="[426,566,1985,2011]" pageId="6" pageNumber="256" pagination="71 - 87" refId="ref8408" refString="Bacon, J. A. (1948) Some new species of Rhagovelia (Hemiptera, Veliidae). Journal of the Kansas Entomological Society, 21 (3), 71 - 87. [https: // www. jstor. org / stable / 25081858]" type="journal article" year="1948">Bacon, 1948</bibRefCitation>
</taxonomicName>
when using the key provided by 
<bibRefCitation id="EFD74B05FF8AA636FC06B89EFBE8F815" author="Bacon, J. A." box="[930,1082,1984,2011]" pageId="6" pageNumber="256" pagination="695 - 913" refId="ref8458" refString="Bacon, J. A. (1956) A taxonomic study of the genus Rhagovelia (Hemiptera, Veliidae) of the Western Hemisphere. University of Kansas Science Bulletin, 38, 695 - 913. [https: // www. biodiversitylibrary. org / page / 28892618]" type="journal article" year="1956">Bacon (1956)</bibRefCitation>
for his 
<taxonomicName id="4C464D77FF8AA636FB28B89EFAD6F814" authorityName="Uhler" authorityYear="1894" box="[1164,1284,1985,2010]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="256" phylum="Arthropoda" rank="species" species="angustipes">
<emphasis id="B932EAE6FF8AA636FB28B89EFAD6F814" box="[1164,1284,1985,2010]" italics="true" pageId="6" pageNumber="256">angustipes</emphasis>
</taxonomicName>
group, which is indeed a species similar to the material at hand. The new species and 
<taxonomicName id="4C464D77FF8BA637FC67BFC8FB81FF7F" authorityName="Bacon" authorityYear="1948" box="[963,1107,151,177]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="257" phylum="Arthropoda" rank="species" species="fontanalis">
<emphasis id="B932EAE6FF8BA637FC67BFC8FB81FF7F" box="[963,1107,151,177]" italics="true" pageId="7" pageNumber="257">R. fontanalis</emphasis>
</taxonomicName>
share the following features: 1) base of antennomere I, and fore and hind trochanters yellow; 2) presence of broad, shiny black spots on the dorsum of abdominal mediotergites II−VII; 3) abdominal sternum VII partially orange-brown in males; and 4) all trochanters and tibiae unarmed in both sexes. Nonetheless, there are several differences between them, including: 1) body length considerably different (
<quantity id="4CBE9B11FF8BA637FDF0BE78FD6FFE8F" box="[596,701,295,321]" metricMagnitude="-3" metricUnit="m" metricValue="3.08" pageId="7" pageNumber="257" unit="mm" value="3.08">3.08 mm</quantity>
[apterous male], 
<quantity id="4CBE9B11FF8BA637FC2DBE78FC22FE8F" box="[905,1008,295,321]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="7" pageNumber="257" unit="mm" value="3.2">3.20 mm</quantity>
[macropterous male], 
<quantity id="4CBE9B11FF8BA637FB51BE78FA4FFE8F" box="[1269,1437,295,321]" metricMagnitude="-3" metricUnit="m" metricValue="3.28" metricValueMax="3.36" metricValueMin="3.2" pageId="7" pageNumber="257" unit="mm" value="3.28" valueMax="3.36" valueMin="3.2">3.20−3.36 mm</quantity>
[apterous female] in the new species, 
<emphasis id="B932EAE6FF8BA637FDE0BE14FD89FEAA" box="[580,603,331,356]" italics="true" pageId="7" pageNumber="257">vs</emphasis>
. 
<quantity id="4CBE9B11FF8BA637FDCEBE15FD1DFEAB" box="[618,719,330,357]" metricMagnitude="-3" metricUnit="m" metricValue="3.45" pageId="7" pageNumber="257" unit="mm" value="3.45">3.45 mm</quantity>
[apterous male], 
<quantity id="4CBE9B11FF8BA637FC32BE14FC29FEAB" box="[918,1019,331,357]" metricMagnitude="-3" metricUnit="m" metricValue="4.33" pageId="7" pageNumber="257" unit="mm" value="4.33">4.33 mm</quantity>
[macropterous male], 3.86 [apterous female] in 
<taxonomicName id="4C464D77FF8BA637FEB6BE30FE73FE47" authorityName="Bacon" authorityYear="1948" box="[274,417,367,393]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="257" phylum="Arthropoda" rank="species" species="fontanalis">
<emphasis id="B932EAE6FF8BA637FEB6BE30FE73FE47" box="[274,417,367,393]" italics="true" pageId="7" pageNumber="257">R. fontanalis</emphasis>
</taxonomicName>
); 2) antennomeres II−III subequal in length in the new species, 
<emphasis id="B932EAE6FF8BA637FBD2BE30FB5FFE46" box="[1142,1165,367,392]" italics="true" pageId="7" pageNumber="257">vs</emphasis>
. II shorter than III in 
<taxonomicName id="4C464D77FF8BA637FA20BE30FED5FE63" authorityName="Bacon" authorityYear="1948" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="257" phylum="Arthropoda" rank="species" species="fontanalis">
<emphasis id="B932EAE6FF8BA637FA20BE30FED5FE63" italics="true" pageId="7" pageNumber="257">R. fontanalis</emphasis>
</taxonomicName>
; and 3) paramere longer and more acute at apex in 
<taxonomicName id="4C464D77FF8BA637FC98BECCFC18FE63" authorityName="Bacon" authorityYear="1948" box="[828,970,403,429]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="257" phylum="Arthropoda" rank="species" species="fontanalis">
<emphasis id="B932EAE6FF8BA637FC98BECCFC18FE63" box="[828,970,403,429]" italics="true" pageId="7" pageNumber="257">R. fontanalis</emphasis>
</taxonomicName>
(compare 
<figureCitation id="137D2A71FF8BA637FBE4BECCFB24FE63" box="[1088,1270,403,429]" captionStart="FIGURE 2" captionStartId="3.[152,255,651,675]" captionTargetBox="[199,1388,185,624]" captionTargetId="figure-20@3.[196,1391,181,627]" captionTargetPageId="3" captionText="FIGURE 2. Parameres. A. Rhagovelia leilae Paiva, Rodrigues &amp; Moreira, sp. nov., male holotype. B. Rhagovelia fontanalis Bacon, 1948 [modified from Bacon (1956)]." figureDoi="http://doi.org/10.5281/zenodo.7973334" httpUri="https://zenodo.org/record/7973334/files/figure.png" pageId="7" pageNumber="257">Figs. 2A and 2B</figureCitation>
). Furthermore, 
<bibRefCitation id="EFD74B05FF8BA637FF33BEE8FEE2FE1F" author="Bacon, J. A." box="[151,304,439,465]" pageId="7" pageNumber="257" pagination="71 - 87" refId="ref8408" refString="Bacon, J. A. (1948) Some new species of Rhagovelia (Hemiptera, Veliidae). Journal of the Kansas Entomological Society, 21 (3), 71 - 87. [https: // www. jstor. org / stable / 25081858]" type="journal article" year="1948">Bacon (1948)</bibRefCitation>
did not mention the presence of a tuft of black setae on the posterolateral angle of female abdominal laterotergite VII, which is a distinctive feature in our specimens. Finally, 
<taxonomicName id="4C464D77FF8BA637FC68BE84FB89FE3B" authorityName="Bacon" authorityYear="1948" box="[972,1115,475,501]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="257" phylum="Arthropoda" rank="species" species="fontanalis">
<emphasis id="B932EAE6FF8BA637FC68BE84FB89FE3B" box="[972,1115,475,501]" italics="true" pageId="7" pageNumber="257">R. fontanalis</emphasis>
</taxonomicName>
is endemic to 
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, where it occurs between about 450 and 
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a.s.l. (
<bibRefCitation id="EFD74B05FF8BA637FDDDBEA0FCD1FDD7" author="Bacon, J. A." box="[633,771,511,537]" pageId="7" pageNumber="257" pagination="71 - 87" refId="ref8408" refString="Bacon, J. A. (1948) Some new species of Rhagovelia (Hemiptera, Veliidae). Journal of the Kansas Entomological Society, 21 (3), 71 - 87. [https: // www. jstor. org / stable / 25081858]" type="journal article" year="1948">Bacon 1948</bibRefCitation>
, 
<bibRefCitation id="EFD74B05FF8BA637FCA8BEA1FC95FDD7" author="Bacon, J. A." box="[780,839,510,537]" pageId="7" pageNumber="257" pagination="695 - 913" refId="ref8458" refString="Bacon, J. A. (1956) A taxonomic study of the genus Rhagovelia (Hemiptera, Veliidae) of the Western Hemisphere. University of Kansas Science Bulletin, 38, 695 - 913. [https: // www. biodiversitylibrary. org / page / 28892618]" type="journal article" year="1956">1956</bibRefCitation>
; 
<bibRefCitation id="EFD74B05FF8BA637FCF6BEA0FBA6FDD7" author="Cordeiro, I. R. S. &amp; Moreira, F. F. F." box="[850,1140,510,537]" pageId="7" pageNumber="257" pagination="4913" refId="ref8589" refString="Cordeiro, I. R. S. &amp; Moreira, F. F. F. (2015) New distributional data on aquatic and semiaquatic bugs (Hemiptera: Heteroptera: Gerromorpha &amp; Nepomorpha) from South America. Biodiversity Data Journal, 3, e 4913. https: // doi. org / 10.3897 / BDJ. 3. e 4913" type="journal article" year="2015">Cordeiro &amp; Moreira 2015</bibRefCitation>
); whereas, the new species has been collected exclusively in Cerrado and Atlantic Forest areas of northeastern 
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, between about 300 and 
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a.s.l.
</paragraph>
</subSubSection>
</treatment>
</document>