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<document id="2E7745D6BB0FA98FEA50F5F86B23F8DE" ID-DOI="10.11646/zootaxa.3919.2.1" ID-GBIF-Dataset="8a0f71e3-5416-41c5-8e7d-b4cc574b7340" ID-ISSN="1175-5326" ID-Zenodo-Dep="245246" ID-ZooBank="8CB77F9E-A35E-43E2-91F7-7822AE421B33" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460990631097" checkinUser="plazi" docAuthor="Bartsch, Ilse" docDate="2015" docId="03C887E5FFF6FF8FFF12A5ABFB4BFAD5" docLanguage="en" docName="zt03919p259.pdf" docOrigin="Zootaxa 3919 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Simognathus Trouessart 1889" docType="treatment" docVersion="9" lastPageNumber="230" masterDocId="FFF1FF9DFFEBFF92FF85A71FFFE8FF86" masterDocTitle="The genital area of Halacaridae (Acari), life stages and development of morphological characters and implication on the classification" masterLastPageNumber="259" masterPageNumber="201" pageNumber="230" updateTime="1698561660865" updateUser="plazi">
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<mods:title id="F8C9BBCB0626A0D931B600D12A5A1C67">The genital area of Halacaridae (Acari), life stages and development of morphological characters and implication on the classification</mods:title>
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<mods:namePart id="1D2EE2341ECDC4C949D607FF5F421673">Bartsch, Ilse</mods:namePart>
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<subSubSection id="C37B6578FFF6FF8FFF12A5ABFECFFD69" pageId="29" pageNumber="230" type="nomenclature">
<paragraph id="8BDE36F3FFF6FF8FFF12A5ABFDE5FD48" blockId="29.[151,525,692,751]" box="[151,525,692,718]" pageId="29" pageNumber="230">
<heading id="D096819FFFF6FF8FFF12A5ABFDE5FD48" bold="true" box="[151,525,692,718]" fontSize="11" level="1" pageId="29" pageNumber="230" reason="1">
<taxonomicName id="4C614D70FFF6FF8FFF12A5ABFDE5FD48" authority="Trouessart, 1889" authorityName="Trouessart" authorityYear="1889" box="[151,525,692,718]" class="Arachnida" family="Halacaridae" genus="Simognathus" kingdom="Animalia" order="Trombidiformes" pageId="29" pageNumber="230" phylum="Arthropoda" rank="genus">
<emphasis id="B915EAE1FFF6FF8FFF12A5ABFDE5FD48" bold="true" box="[151,525,692,718]" pageId="29" pageNumber="230">
<emphasis id="B915EAE1FFF6FF8FFF12A5ABFEDEFD48" bold="true" box="[151,310,692,718]" italics="true" pageId="29" pageNumber="230">Simognathus</emphasis>
Trouessart, 1889
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<paragraph id="8BDE36F3FFF6FF8FFF12A5C9FECFFD69" blockId="29.[151,525,692,751]" box="[151,295,726,751]" pageId="29" pageNumber="230">(Figs 7175)</paragraph>
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<subSubSection id="C37B6578FFF6FF8FFF12A401FBBDFB61" pageId="29" pageNumber="230" type="materials_examined">
<paragraph id="8BDE36F3FFF6FF8FFF12A401FD3EFCB1" blockId="29.[151,1436,798,1363]" box="[151,726,798,823]" pageId="29" pageNumber="230">
<emphasis id="B915EAE1FFF6FF8FFF12A401FEDAFCB1" bold="true" box="[151,306,798,823]" pageId="29" pageNumber="230">
<typeStatus id="54DA8851FFF6FF8FFF12A401FF3AFCB1" box="[151,210,798,823]" pageId="29" pageNumber="230">Type</typeStatus>
species.
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<taxonomicName id="4C614D70FFF6FF8FFEBCA401FD3AFCB1" authority="Hodge, 1863" authorityName="Hodge" authorityYear="1863" box="[313,722,798,823]" class="Arachnida" family="Halacaridae" genus="Pachygnathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="29" pageNumber="230" phylum="Arthropoda" rank="species" species="minutus">
<emphasis id="B915EAE1FFF6FF8FFEBCA401FDDFFCB0" box="[313,567,798,823]" italics="true" pageId="29" pageNumber="230">Pachygnathus minutus</emphasis>
<bibRefCitation id="EFF04B02FFF6FF8FFDBBA401FD3AFCB1" author="Hodge" box="[574,722,798,823]" pageId="29" pageNumber="230" refString="Hodge, G. (1863) Contributions to the marine zoology of Seaham Harbour. On some undescribed marine Acari. Transactions of the Tyneside Naturalists' Field Club, 5 (4), 298 - 303." type="journal article" year="1863">Hodge, 1863</bibRefCitation>
</taxonomicName>
.
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<paragraph id="8BDE36F3FFF6FF8FFF42A45DFAE7FBB5" blockId="29.[151,1436,798,1363]" pageId="29" pageNumber="230">
<emphasis id="B915EAE1FFF6FF8FFF42A45DFEF5FCDD" bold="true" box="[199,285,834,859]" pageId="29" pageNumber="230">Adults.</emphasis>
In both female and male GP and AP fused. Female GA with four to five pairs of pgs (rarely and unilaterally with three, exceptionally two pgs), subgenital setae lacking. Three small internal acetabula-like structures on either side of GO, situated in margin of large sacculus (Fig. 71). Ovipositor short, not extending beyond GO. Apical genital spines slender, spiniform, not sclerotized; number and arrangement not known. GO of male smaller than that of female; GA with about 1025 pairs of pgs; of these zero to three pairs outlying, the others arranged in a ring around GO. In some species pgs setiform and smooth in others delicately plumose (
<bibRefCitation id="EFF04B02FFF6FF8FFAC0A4E9FEDAFBB5" author="Bartsch" pageId="29" pageNumber="230" refString="Bartsch, I. (1994 b) The genus Simognathus (Acari: Halacaridae), description of six new species from southern Australia and a tabular key to all species. Acarologia, 35, 135 - 152." type="journal article" year="1994" yearSuffix="b">Bartsch 1994b: fig 58</bibRefCitation>
). Genital sclerites with three pairs of short sgs. Epimeral pores within epimeral vesicles.
</paragraph>
<paragraph id="8BDE36F3FFF6FF8FFF42A321FBBDFB61" blockId="29.[151,1436,798,1363]" pageId="29" pageNumber="230">
<emphasis id="B915EAE1FFF6FF8FFF42A321FED4FBD1" bold="true" box="[199,316,1086,1111]" pageId="29" pageNumber="230">Juveniles.</emphasis>
With deuto-, protonymph and larva, all stages with epimeral vesicles. In both deuto- and protonymph GP and AP separated. Deutonymphal GP with one (Figs 73, 74;
<bibRefCitation id="EFF04B02FFF6FF8FFB84A37DFB40FBFD" author="Bartsch" box="[1025,1192,1122,1147]" pageId="29" pageNumber="230" refString="Bartsch, I. (1977 b) Interstitielle Fauna von Galapagos. XX. Halacaridae (Acari). Mikrofauna des Meeresbodens, 65, 1 - 108." type="journal article" year="1977" yearSuffix="b">Bartsch 1977b</bibRefCitation>
: fig 380) or two pairs of pgs (
<bibRefCitation id="EFF04B02FFF6FF8FFF6FA399FE19FB19" author="Bartsch" box="[234,497,1158,1183]" pageId="29" pageNumber="230" refString="Bartsch, I. (1977 b) Interstitielle Fauna von Galapagos. XX. Halacaridae (Acari). Mikrofauna des Meeresbodens, 65, 1 - 108." type="journal article" year="1977" yearSuffix="b">Bartsch 1977b: fig 386</bibRefCitation>
); the latter close together in posterior part of GP. Deutonymphs without sgs. Genital acetabula small, two pairs present, these distinct or very small (Figs 73, 74). Protonymphal GP with zero to one pair of pgs (Figs 72, 75;
<bibRefCitation id="EFF04B02FFF6FF8FFE2EA3D1FDBAFB61" author="Bartsch" box="[427,594,1230,1255]" pageId="29" pageNumber="230" refString="Bartsch, I. (1977 b) Interstitielle Fauna von Galapagos. XX. Halacaridae (Acari). Mikrofauna des Meeresbodens, 65, 1 - 108." type="journal article" year="1977" yearSuffix="b">Bartsch 1977b</bibRefCitation>
: fig. 388) but no sgs. Single pair of gac small.
</paragraph>
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<subSubSection id="C37B6578FFF6FF8FFF42A3EDFB4BFAD5" pageId="29" pageNumber="230" type="discussion">
<paragraph id="8BDE36F3FFF6FF8FFF42A3EDFB4BFAD5" blockId="29.[151,1436,798,1363]" pageId="29" pageNumber="230">
<emphasis id="B915EAE1FFF6FF8FFF42A3EDFED3FA8D" bold="true" box="[199,315,1266,1291]" pageId="29" pageNumber="230">Remarks.</emphasis>
Forty-four species are described (
<bibRefCitation id="EFF04B02FFF6FF8FFD44A3EDFC81FA8D" author="Bartsch" box="[705,873,1266,1291]" pageId="29" pageNumber="230" refString="Bartsch, I. (2009 a) Checklist of marine and freshwater halacarid mite genera and species (Halacaridae: Acari) with notes on synonyms, habitats, distribution and descriptions of the taxa. Zootaxa, 1998, 1 - 170." type="journal article" year="2009" yearSuffix="a">Bartsch 2009a</bibRefCitation>
,
<bibRefCitation id="EFF04B02FFF6FF8FFCF3A3ECFC57FA8D" author="Bartsch" box="[886,959,1267,1291]" pageId="29" pageNumber="230" refString="Bartsch, I. (2009 c) Psammobiont halacarid mites (Acari: Halacaridae) from St John's Island, Singapore and remarks on the halacarid fauna of the Malay Peninsula. The Raffles Bulletin of Zoology, Supplement, 22, 173 - 201." type="journal article" year="2009" yearSuffix="c">2009c</bibRefCitation>
). Many of them live in sandy deposits, in beaches or sand-filled pockets amongst a macroflora and fauna. The genus is spread world-wide but more species are known from the tropical and warm-temperate south than from the north (
<bibRefCitation id="EFF04B02FFF6FF8FFC6AA225FB7FFAD5" author="Bartsch" box="[1007,1175,1338,1363]" pageId="29" pageNumber="230" refString="Bartsch, I. (2009 a) Checklist of marine and freshwater halacarid mite genera and species (Halacaridae: Acari) with notes on synonyms, habitats, distribution and descriptions of the taxa. Zootaxa, 1998, 1 - 170." type="journal article" year="2009" yearSuffix="a">Bartsch 2009a</bibRefCitation>
).
</paragraph>
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