142 lines
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142 lines
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.347.6002" ID-GBIF-Dataset="4523286c-586e-4e1e-b33f-6836d3165a18" ID-PMC="PMC3822444" ID-Pensoft-Pub="1313-2970-347-1" ID-PubMed="24222723" ID-ZBK="52232D18DD784A84882CACA428B4A9D2" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-347-1" ModsDocOrigin="ZooKeys 347" ModsDocTitle="The Doryctinae (Braconidae) of Costa Rica: genera and species of the tribe Heterospilini" checkinTime="1451246744844" checkinUser="pensoft" docAuthor="Marsh, Paul M., Wild, Alexander L. & Whitfield, James B." docDate="2013" docId="003F79B0CAE8FA742C50E31753B1F8B0" docLanguage="en" docName="ZooKeys 347: 1-474" docOrigin="ZooKeys 347" docSource="http://dx.doi.org/10.3897/zookeys.347.6002" docTitle="Pioscelus Muesebeck & Walkley" docType="treatment" docVersion="4" lastPageNumber="19" masterDocId="FF80331FA362FFFA435DFFBEFFDF333B" masterDocTitle="The Doryctinae (Braconidae) of Costa Rica: genera and species of the tribe Heterospilini" masterLastPageNumber="474" masterPageNumber="1" pageNumber="18" updateTime="1668156762379" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The Doryctinae (Braconidae) of Costa Rica: genera and species of the tribe Heterospilini</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Marsh, Paul M.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Wild, Alexander L.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Whitfield, James B.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2013</mods:date>
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<mods:detail type="volume">
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<mods:number>347</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>474</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.347.6002</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.347.6002</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-347-1</mods:identifier>
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<mods:identifier type="ZBK">52232D18DD784A84882CACA428B4A9D2</mods:identifier>
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<mods:identifier type="ZooBank">52232D18DD784A84882CACA428B4A9D2</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152049387" LSID="urn:lsid:plazi:treatment:003F79B0CAE8FA742C50E31753B1F8B0" httpUri="http://treatment.plazi.org/id/003F79B0CAE8FA742C50E31753B1F8B0" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
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<subSubSection pageId="17" pageNumber="18" type="nomenclature">
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<paragraph pageId="17" pageNumber="18">
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Genus
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<taxonomicName LSID="http://species-id.net/wiki/Pioscelus" authority="Muesebeck & Walkley" class="Insecta" family="Braconidae" genus="Pioscelus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pioscelus" order="Hymenoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">Pioscelus Muesebeck & Walkley</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="17" pageNumber="18" type="reference_group">
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<paragraph pageId="17" pageNumber="18">
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<taxonomicName class="Insecta" family="Braconidae" genus="Pioscelus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pioscelus" order="Hymenoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">Pioscelus</taxonomicName>
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Muesebeck & Walkley 1951: 180.
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</paragraph>
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<paragraph pageId="17" pageNumber="18">
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<taxonomicName class="Insecta" family="Braconidae" genus="Amazondoryctes" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Amazondoryctes" order="Hymenoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">Amazondoryctes</taxonomicName>
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Barbalho & Penteado-Dias, in
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<bibRefCitation author="Barbalho, SM" journalOrPublisher="Journal of Hymenoptera Research" pageId="435" pageNumber="436" pagination="139 - 153" title="Descriptions of new genera from Brazil in the tribes Heterospilini and Spathiini with similar wing venation (Hymenoptera: Braconidae, Doryctinae)." volume="8" year="1999">Barbalho et al. 1999</bibRefCitation>
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: 142. Synonymized by
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<bibRefCitation author="Belokobylskij, SA" journalOrPublisher="Insect Systematics and Evolution" pageId="435" pageNumber="436" pagination="149 - 178" title="Neoheterospilus gen. n., a new genus of the tribe Heterospilini (Hymenoptera: Braconidae, Doryctinae) with highly modified ovipositor and a worldwide distribution." url="10.1163/187631206788831119" volume="37" year="2006">Belokobylskij 2006</bibRefCitation>
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: 150.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="17" pageNumber="18" type="type species">
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<paragraph pageId="17" pageNumber="18">Type species.</paragraph>
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<paragraph pageId="17" pageNumber="18">
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<taxonomicName class="Insecta" genus="Hedysomus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Hedysomus wichitus" order="Hymenoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="wichitus">Hedysomus wichitus</taxonomicName>
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Viereck.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18" type="diagnosis">
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<paragraph pageId="17" pageNumber="18">Diagnosis</paragraph>
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<paragraph lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
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(taken from
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<bibRefCitation author="Marsh, PM" journalOrPublisher="Memoirs of the American Entomological Institute" pageId="436" pageNumber="437" title="The Doryctinae of Costa Rica (excluding the genus Heterospilus)." volume="70" year="2002">Marsh 2002</bibRefCitation>
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). Body size: 2.5-4.5 mm; head cubicle; occipital carina meeting hypostomal carina; frons slightly excavated; mesoscutum declivous anteriorly; propodeum often horizontal and barely declivous posteriorly; precoxal sulcus usually as long as mesopleuron; fore tibia with single row of short spines along anterior edge; hind coxa without distinct tooth or tubercle at base (Fig. 10D) although smoothly angled; fore wing vein r-m present, vein 2RS absent or represented partially by weakly infuscate line, first subdiscal cell open at apex; hind wing vein M+CU equal to or shorter than vein 1M; first metasomal tergum longer then apical width; second metasomal tergum with either 2 converging median grooves which set off triangular
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<pageBreakToken pageId="18" pageNumber="19" start="start">area</pageBreakToken>
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(Fig. 10A) or 2 median parallel carinae (Fig. 10B); basal sternal plate of first metasomal segment 0.25-0.33 length of tergum, dorsope distinct or weak.
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</paragraph>
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<caption pageId="18" pageNumber="19">
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<paragraph pageId="18" pageNumber="19">
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Figure 10. A
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<taxonomicName class="Insecta" family="Braconidae" genus="Pioscelus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pioscelus" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">Pioscelus</taxonomicName>
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sp. showing typical sculpturing on the second metasomal tergum B
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<taxonomicName class="Insecta" family="Braconidae" genus="Pioscelus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pioscelus" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">Pioscelus</taxonomicName>
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sp. showing the
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<taxonomicName class="Insecta" family="Braconidae" genus="Amazondoryctes" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Amazondoryctes" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">Amazondoryctes</taxonomicName>
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style of sculpturing on the second metasomal tergum C hind coxa of
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<taxonomicName class="Insecta" family="Braconidae" genus="Heterospilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heterospilus" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">Heterospilus</taxonomicName>
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sp. D hind coxa of
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<taxonomicName class="Insecta" family="Braconidae" genus="Pioscelus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pioscelus" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">Pioscelus</taxonomicName>
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sp.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="18" pageNumber="19" type="distribution">
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<paragraph pageId="18" pageNumber="19">Distribution.</paragraph>
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<paragraph pageId="18" pageNumber="19">Nearctic and Neotropical Regions.</paragraph>
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</subSubSection>
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<subSubSection pageId="18" pageNumber="19" type="biology">
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<paragraph pageId="18" pageNumber="19">Biology.</paragraph>
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<paragraph pageId="18" pageNumber="19">Unknown.</paragraph>
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</subSubSection>
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<subSubSection pageId="18" pageNumber="19" type="comments">
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<paragraph pageId="18" pageNumber="19">Comments.</paragraph>
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<paragraph pageId="18" pageNumber="19">
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The genus
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<taxonomicName class="Insecta" family="Braconidae" genus="Amazondoryctes" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Amazondoryctes" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">Amazondoryctes</taxonomicName>
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was synonymized with
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<taxonomicName class="Insecta" family="Braconidae" genus="Pioscelus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pioscelus" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">Pioscelus</taxonomicName>
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by
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<bibRefCitation author="Belokobylskij, SA" journalOrPublisher="Insect Systematics and Evolution" pageId="435" pageNumber="436" pagination="149 - 178" title="Neoheterospilus gen. n., a new genus of the tribe Heterospilini (Hymenoptera: Braconidae, Doryctinae) with highly modified ovipositor and a worldwide distribution." url="10.1163/187631206788831119" volume="37" year="2006">Belokobylskij (2006)</bibRefCitation>
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. In the molecular analysis (
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<bibRefCitation author="Wild, AL" journalOrPublisher="PLoS ONE" pageId="437" pageNumber="438" title="Fast-evolving homoplastic traits are best for species identification in a group of Neotropical wasps." url="10.1371/journal.pone.0074837" volume="8" year="2013">Wild et al. 2013</bibRefCitation>
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) specimens identified as
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<taxonomicName class="Insecta" family="Braconidae" genus="Amazondoryctes" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Amazondoryctes" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">Amazondoryctes</taxonomicName>
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emerged basally within
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<taxonomicName class="Insecta" family="Braconidae" genus="Heterospilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heterospilus" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">Heterospilus</taxonomicName>
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but no specimens identified as
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<taxonomicName class="Insecta" family="Braconidae" genus="Pioscelus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pioscelus" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">Pioscelus</taxonomicName>
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were available for analysis. Thus, the exact placement of these two genera within the
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<taxonomicName lsidName="" pageId="18" pageNumber="19" rank="tribe" tribe="Heterospilini">Heterospilini</taxonomicName>
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is still in question.
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</paragraph>
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<paragraph pageId="18" pageNumber="19">
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Two species occur in Costa Rica,
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<taxonomicName class="Insecta" family="Braconidae" genus="Pioscelus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pioscelus mesomphalus" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="mesomphalus">Pioscelus mesomphalus</taxonomicName>
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Marsh and
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<taxonomicName class="Insecta" family="Braconidae" genus="Pioscelus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pioscelus costaricensis" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="costaricensis">Pioscelus costaricensis</taxonomicName>
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(Marsh), comb. n. Descriptions for each species can be found in
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<bibRefCitation author="Marsh, PM" journalOrPublisher="Memoirs of the American Entomological Institute" pageId="436" pageNumber="437" title="The Doryctinae of Costa Rica (excluding the genus Heterospilus)." volume="70" year="2002">Marsh 2002</bibRefCitation>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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</document>
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