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<document id="9D95D4DBDB4FFD4B94A17329A92791C6" ID-DOI="10.11606/1807-0205/2018.58.58" ID-ISSN="1807-0205" ID-Zenodo-Dep="5384147" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630298591680" checkinUser="valdenar" docAuthor="Biffi, Gabriel &amp; Constantin, Robert" docDate="2018" docId="03821672FFD21D4953B9FF31F981FD0C" docLanguage="en" docName="148343-Article Text-322524-1-10-20181129.pdf" docOrigin="Papéis Avulsos de Zoologia (Pap. Avulsos Zool., S. Paulo) 58" docSource="http://dx.doi.org/10.11606/1807-0205/2018.58.58" docStyle="DocumentStyle:6844D5473582683DD13A9A75B001759F.4:PapAvZool.2018-.journal_article" docStyleId="6844D5473582683DD13A9A75B001759F" docStyleName="PapAvZool.2018-.journal_article" docStyleVersion="4" docTitle="Macromalthinus quadratithorax Biffi &amp; Constantin 2018, sp. nov." docType="treatment" docVersion="4" lastPageNumber="14" masterDocId="FFBB6E0AFFDE1D44500DFFEBFD7DFF9F" masterDocTitle="A revision of Macromalthinus Pic, 1919 with description of six new species (Coleoptera: Cantharidae: Chauliognathinae)" masterLastPageNumber="22" masterPageNumber="1" pageNumber="13" updateTime="1699219721757" updateUser="plazi" zenodo-license-document="CC-BY-NC-4.0" zenodo-license-figures="CC-BY-NC-4.0">
<mods:mods id="D98088026E35A5206220130242C2C806" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="9730B82DCEFA075112FCD726605A8F4D">
<mods:title id="705178151FA1945CC0ADC3415201FA7F">A revision of Macromalthinus Pic, 1919 with description of six new species (Coleoptera: Cantharidae: Chauliognathinae)</mods:title>
</mods:titleInfo>
<mods:name id="CF3A4FDD4C1A2B8782D4AB3B4B74F0B4" type="personal">
<mods:role id="52165764C399BE56BA12F76EF94398FE">
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<mods:namePart id="3D5EC8860A7D9327ECC36EA0E1BB4C44">Biffi, Gabriel</mods:namePart>
</mods:name>
<mods:name id="ADCCEA02F65E093D515A00665A19897D" type="personal">
<mods:role id="FF8BD93DED27A796345E2D59EAC87AFC">
<mods:roleTerm id="1B360396E8A6BB5DF16978FEBFA8D292">Author</mods:roleTerm>
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<mods:namePart id="D01F8B7429F6F6615A08EB100B5AA135">Constantin, Robert</mods:namePart>
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<mods:typeOfResource id="F6B3BC08E5FE4FB0C5AA3A073125AA14">text</mods:typeOfResource>
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<mods:title id="678B588ACCC3F4593D1ED0BA0EEE10BB">Papéis Avulsos de Zoologia</mods:title>
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<mods:part id="35E8ED9503538C7E5BFD4ADD31D8D49A">
<mods:date id="46DEF45AB34DE820BE36E2FE5A423D9C">2018</mods:date>
<mods:detail id="6753C88A85DBDBDDBC3E8AEBECFB7086" type="series">
<mods:title id="7ABA4F2B2E962AFF40B52B519AAAF4CC">Pap. Avulsos Zool., S. Paulo</mods:title>
</mods:detail>
<mods:detail id="18AF0A71457D0721CBA787608F873AFC" type="pubDate">
<mods:number id="3D8E4CE1574C0B5F4D2735E2B24A82C3">2018-11-29</mods:number>
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<mods:detail id="671C0A2A6EAD03C785E86363937C7B77" type="volume">
<mods:number id="DB6360666025682C15A33126E993B44F">58</mods:number>
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<mods:url id="D7CC2FE88764B53648F87AC0C227A6F5">http://dx.doi.org/10.11606/1807-0205/2018.58.58</mods:url>
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<mods:classification id="758579FD5B226B185C3EF544F6004D0C">journal article</mods:classification>
<mods:identifier id="F89DE5C9FC8FD1698A02DD7EBD36C530" type="DOI">10.11606/1807-0205/2018.58.58</mods:identifier>
<mods:identifier id="42F111E9299A4D8B89394E350001CF4F" type="ISSN">1807-0205</mods:identifier>
<mods:identifier id="23A7516579CFE88279FB99FAA274762A" type="Zenodo-Dep">5384147</mods:identifier>
<mods:identifier id="9DEC84469273633B790E9BBD7E588D66" type="ZooBank">97C262C2-6F02-4916-9533-43653FE5561F</mods:identifier>
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<treatment id="03821672FFD21D4953B9FF31F981FD0C" ID-DOI="http://doi.org/10.5281/zenodo.5466709" ID-Zenodo-Dep="5466709" LSID="urn:lsid:plazi:treatment:03821672FFD21D4953B9FF31F981FD0C" httpUri="http://treatment.plazi.org/id/03821672FFD21D4953B9FF31F981FD0C" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
<subSubSection id="C331F4EFFFD21D4853B9FF31F8FBFF6C" box="[948,1414,218,243]" pageId="12" pageNumber="13" type="nomenclature">
<paragraph id="8B94A764FFD21D4853B9FF31F8FBFF6C" blockId="12.[869,1494,218,563]" box="[948,1414,218,243]" pageId="12" pageNumber="13">
<heading id="D0DC1008FFD21D4853B9FF31F8FBFF6C" box="[948,1414,218,243]" centered="true" fontSize="10" level="2" pageId="12" pageNumber="13" reason="2">
<emphasis id="B95F7B76FFD21D4853B9FF31F8FBFF6C" bold="true" box="[948,1414,218,243]" pageId="12" pageNumber="13">
<taxonomicName id="4C2BDCE7FFD21D4853B9FF31F85AFF6C" authority="Biffi &amp; Constantin, 2018" authorityName="Biffi &amp; Constantin" authorityYear="2018" box="[948,1319,218,243]" class="Insecta" family="Cantharidae" genus="Macromalthinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="quadratithorax" status="sp. nov.">
<emphasis id="B95F7B76FFD21D4853B9FF31F85AFF6C" bold="true" box="[948,1319,218,243]" italics="true" pageId="12" pageNumber="13">Macromalthinus quadratithorax</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26CC60DFFD21D485520FF31F8FBFF6C" box="[1325,1414,218,243]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C331F4EFFFD21D4853D0FF11F823FE8C" box="[989,1374,250,275]" pageId="12" pageNumber="13" type="description">
<paragraph id="8B94A764FFD21D4853D0FF11F823FE8C" blockId="12.[869,1494,218,563]" box="[989,1374,250,275]" pageId="12" pageNumber="13">
<emphasis id="B95F7B76FFD21D4853D0FF11F823FE8C" bold="true" box="[989,1374,250,275]" pageId="12" pageNumber="13">
(
<figureCitation id="1310BBE1FFD21D4853E9FF11F94FFE8C" box="[996,1074,250,275]" captionStart="Figures 1-6" captionStartId="3.[94,162,2074,2099]" captionTargetBox="[152,1356,206,2041]" captionTargetId="figure-1@3.[151,1360,204,2044]" captionTargetPageId="3" captionText="Figures 1-6:Dorsal habitus ofMacromalthinus species.(1) M.guyanensis sp.nov.(2)M.luteoapicalissp.nov.(3)M.orapuensis sp.nov.(4) M.piceiventrissp.nov. (5) M.quadratithorax sp.nov.(6)M.xerophilus sp.nov.Scale bars:2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5384149" httpUri="https://zenodo.org/record/5384149/files/figure.png" pageId="12" pageNumber="13">Figs. 5</figureCitation>
,
<figureCitation id="1310BBE1FFD21D485431FF11F926FE8C" box="[1084,1115,250,275]" captionStart="Figures 7-12" captionStartId="4.[869,937,1217,1242]" captionTargetBox="[869,1493,204,1187]" captionTargetId="figure-697@4.[869,1493,204,1187]" captionTargetPageId="4" captionText="Figures 7-12: Pronotum of Macromalthinus species. (7) M. guyanensis sp. nov. (8) M. luteoapicalis sp. nov. (9) M. orapuensis sp. nov. (10) M. piceiventris sp. nov. (11) M. quadratithorax sp. nov. (12) M. xerophilus sp. nov. Scale bars:1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5384151" httpUri="https://zenodo.org/record/5384151/files/figure.png" pageId="12" pageNumber="13">11</figureCitation>
,
<figureCitation id="1310BBE1FFD21D48546BFF11F9FBFE8C" box="[1126,1158,250,275]" captionStart="Figures 13-18" captionStartId="5.[94,162,2074,2099]" captionTargetBox="[97,1416,991,2040]" captionTargetId="figure-354@5.[94,1417,986,2044]" captionTargetPageId="5" captionText="Figures 13-18: Fore femur of Macromalthinus species. (13) M. guyanensis sp. nov. (14) M. luteoapicalis sp. nov.(15) M. orapuensis sp.nov.(16) M. piceiventris sp.nov.(17) M. quadratithoraxsp.nov.(18) M. xerophilus sp.nov.Scale bars:0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5384153" httpUri="https://zenodo.org/record/5384153/files/figure.png" pageId="12" pageNumber="13">17</figureCitation>
,
<figureCitation id="1310BBE1FFD21D48549DFF11F9CDFE8C" box="[1168,1200,250,275]" captionStart="Figures 19-24" captionStartId="6.[170,238,2074,2099]" captionTargetBox="[172,1491,1272,2042]" captionTargetId="figure-506@6.[170,1493,1269,2044]" captionTargetPageId="6" captionText="Figures 19-24:Ventrite VII of males of Macromalthinus species.(19) M.guyanensis sp.nov.(20) M.luteoapicalis sp.nov.(21)M. orapuensis sp.nov.(22) M.piceiventris sp.nov.(23) M.quadratithorax sp.nov.(24) M.xerophilus sp.nov.Scale bars:0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5384156" httpUri="https://zenodo.org/record/5384156/files/figure.png" pageId="12" pageNumber="13">23</figureCitation>
,
<figureCitation id="1310BBE1FFD21D4854B7FF11F9A7FE8C" box="[1210,1242,250,275]" captionStart="Figures 25-30" captionStartId="7.[794,862,2010,2035]" captionTargetBox="[798,1414,205,1979]" captionTargetId="figure-491@7.[794,1418,204,1980]" captionTargetPageId="7" captionText="Figures 25-30: Ventrite VIII of males of Macromalthinus species. (25) M. guyanensis sp. nov. (26) M. luteoapicalis sp. nov. (27) M. orapuensis sp.nov.(28)M.piceiventrissp.nov.(29) M.quadratithoraxsp.nov.(30) M.xerophilus sp.nov.Scale bars:0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5384158" httpUri="https://zenodo.org/record/5384158/files/figure.png" pageId="12" pageNumber="13">29</figureCitation>
,
<figureCitation id="1310BBE1FFD21D4854E9FF11F879FE8C" box="[1252,1284,250,275]" captionStart="Figures 31-36" captionStartId="8.[170,238,2042,2067]" captionTargetBox="[171,1491,674,2008]" captionTargetId="figure-183@8.[170,1493,670,2012]" captionTargetPageId="8" captionText="Figures 31-36: Aedeagus of Macromalthinus species. (31) M. guyanensis sp. nov. (32) M. luteoapicalis sp. nov. (33) M. orapuensis sp. nov. (34) M. piceiventris sp.nov.(35) M. quadratithorax sp.nov.(36) M. xerophilus sp.nov. Abbreviations:pt = prolongation of tegmen, sp = setiferous prolongation,rp = right paramere, lp = left paramere,ml = median lobe.Scale bars:0.5mm." figureDoi="http://doi.org/10.5281/zenodo.5384160" httpUri="https://zenodo.org/record/5384160/files/figure.png" pageId="12" pageNumber="13">35</figureCitation>
,
<figureCitation id="1310BBE1FFD21D485503FF11F850FE8C" box="[1294,1325,250,275]" captionStart="Figures 37-42" captionStartId="9.[94,162,2074,2099]" captionTargetBox="[94,1414,1424,2040]" captionTargetId="figure-592@9.[94,1417,1420,2044]" captionTargetPageId="9" captionText="Figures 37-42:VentriteVII offemales ofMacromalthinusspecies.(37) M.guyanensis sp.nov.(38)M.luteoapicalissp.nov.(39) M.orapuensissp.nov.(40)M.piceiventris sp.nov.(41) M. quadratithoraxsp.nov.(42) M. xerophilus sp.nov.Scale bars:0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5384162" httpUri="https://zenodo.org/record/5384162/files/figure.png" pageId="12" pageNumber="13">41</figureCitation>
,
<figureCitation id="1310BBE1FFD21D485535FF11F82BFE8C" box="[1336,1366,250,275]" captionStart="Figures 43-48" captionStartId="10.[869,937,2010,2035]" captionTargetBox="[873,1490,1055,1976]" captionTargetId="figure-666@10.[869,1493,1054,1980]" captionTargetPageId="10" captionText="Figures 43-48: Coxites of Macromalthinus species. (43) M. guyanensis sp. nov. (44) M. luteoapicalis sp. nov. (45) M. orapuensis sp. nov. (46) M. piceiventris sp. nov. (47) M. quadratithorax sp. nov. (48) M. xerophilus sp. nov. Scale bars:0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5384164" httpUri="https://zenodo.org/record/5384164/files/figure.png" pageId="12" pageNumber="13">47</figureCitation>
)
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C331F4EFFFD21D485368FED1F820FDAC" pageId="12" pageNumber="13" type="materials_examined">
<paragraph id="8B94A764FFD21D485368FED1F820FDAC" blockId="12.[869,1494,218,563]" pageId="12" pageNumber="13">
<emphasis id="B95F7B76FFD21D485368FED1F96CFECC" bold="true" box="[869,1041,314,339]" pageId="12" pageNumber="13">Type material:</emphasis>
<materialsCitation id="3B43AD39FFD21D485414FED1F873FE0C" collectingDate="1975-07" collectionCode="NHMB" collectorName="J. Price" country="Trinidad and Tobago" location="Blanchisseuse Rd." municipality="Simla" pageId="12" pageNumber="13" specimenCount="1" specimenCount-male="1" stateProvince="Arima" typeStatus="holotype">
<typeStatus id="549019C6FFD21D485414FED1F9E8FECC" box="[1049,1173,314,339]" pageId="12" pageNumber="13" type="holotype">HOLOTYPE</typeStatus>
:
<specimenCount id="9D2D6CEDFFD21D4854ADFEDCF9CEFECD" box="[1184,1203,311,338]" pageId="12" pageNumber="13" type="male">♂</specimenCount>
:
<collectingCountry id="F33CE7F4FFD21D4854B2FED1F8AFFECC" box="[1215,1490,314,339]" name="Trinidad and Tobago" pageId="12" pageNumber="13">TRINIDAD AND TOBAGO</collectingCountry>
.
<emphasis id="B95F7B76FFD21D485368FEB1FEBEFEEC" box="[869,963,346,371]" italics="true" pageId="12" pageNumber="13">Trinidad,</emphasis>
<collectingMunicipality id="6BF03D1EFFD21D4853D5FEB1F965FEEC" box="[984,1048,346,371]" pageId="12" pageNumber="13">Simla</collectingMunicipality>
,
<collectingRegion id="49EF6986FFD21D485422FEB1F909FEEC" box="[1071,1140,346,371]" country="Trinidad and Tobago" name="Arima" pageId="12" pageNumber="13">Arima</collectingRegion>
,
<location id="8EF4F1BFFFD21D485481FEB1F81CFEEC" LSID="urn:lsid:plazi:treatment:03821672FFD21D4953B9FF31F981FD0C:8EF4F1BFFFD21D485481FEB1F81CFEEC" box="[1164,1377,346,371]" country="Trinidad and Tobago" municipality="Simla" name="Blanchisseuse Rd." pageId="12" pageNumber="13" stateProvince="Arima">Blanchisseuse Rd.</location>
,
<date id="FF9581A4FFD21D485575FEB1F8AFFEEC" box="[1400,1490,346,371]" pageId="12" pageNumber="13" value="1975-07">
<collectingDate id="EFD1784CFFD21D485575FEB1F8AFFEEC" box="[1400,1490,346,371]" pageId="12" pageNumber="13" value="1975-07">VII.1975</collectingDate>
</date>
,
<collectingMethod id="526ADF73FFD21D485368FE91F974FE0C" box="[869,1033,378,403]" pageId="12" pageNumber="13">Blacklight trap</collectingMethod>
,
<collectorName id="26DEC2B2FFD21D48541EFE91F922FE0C" box="[1043,1119,378,403]" pageId="12" pageNumber="13">J. Price</collectorName>
(
<collectionCode id="ED3A3FA1FFD21D485466FE91F9CBFE0C" box="[1131,1206,378,403]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="12" pageNumber="13">NHMB</collectionCode>
) (
<figureCitation id="1310BBE1FFD21D4854C4FE91F87AFE0C" box="[1225,1287,378,403]" captionStart="Figures 1-6" captionStartId="3.[94,162,2074,2099]" captionTargetBox="[152,1356,206,2041]" captionTargetId="figure-1@3.[151,1360,204,2044]" captionTargetPageId="3" captionText="Figures 1-6:Dorsal habitus ofMacromalthinus species.(1) M.guyanensis sp.nov.(2)M.luteoapicalissp.nov.(3)M.orapuensis sp.nov.(4) M.piceiventrissp.nov. (5) M.quadratithorax sp.nov.(6)M.xerophilus sp.nov.Scale bars:2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5384149" httpUri="https://zenodo.org/record/5384149/files/figure.png" pageId="12" pageNumber="13">Fig. 5</figureCitation>
)
</materialsCitation>
.
<materialsCitation id="3B43AD39FFD21D485514FE91F8AFFE4C" collectingDate="1969-04" collectionCode="NHMB" collectorName="W. I., S. Grande &amp; H &amp; A Howden Collection" country="Trinidad and Tobago" location="Turure Rd." pageId="12" pageNumber="13" specimenCount="6" specimenCount-female="6" typeStatus="paratype">
<typeStatus id="549019C6FFD21D485514FE91F8EAFE0C" box="[1305,1431,378,403]" pageId="12" pageNumber="13" type="paratype">PARATYPES</typeStatus>
(
<specimenCount id="9D2D6CEDFFD21D4855A9FE91F8B5FE0D" box="[1444,1480,375,403]" pageId="12" pageNumber="13" type="female">4 ♀</specimenCount>
).
<collectingCountry id="F33CE7F4FFD21D485368FE71F90BFE2C" box="[869,1142,410,435]" name="Trinidad and Tobago" pageId="12" pageNumber="13">TRINIDAD AND TOBAGO</collectingCountry>
.
<emphasis id="B95F7B76FFD21D48548DFE71F9A0FE2C" box="[1152,1245,410,435]" italics="true" pageId="12" pageNumber="13">Trinidad,</emphasis>
<collectorName id="26DEC2B2FFD21D4854EEFE71F8F9FE2C" box="[1251,1412,410,435]" pageId="12" pageNumber="13">W.I., S. Grande</collectorName>
,
<location id="8EF4F1BFFFD21D485580FE71FEF6FE4C" LSID="urn:lsid:plazi:treatment:03821672FFD21D4953B9FF31F981FD0C:8EF4F1BFFFD21D485580FE71FEF6FE4C" country="Trinidad and Tobago" name="Turure Rd." pageId="12" pageNumber="13">Turure Rd.</location>
,
<date id="FF9581A4FFD21D485396FE51FE90FE4C" box="[923,1005,442,467]" pageId="12" pageNumber="13" value="1969-04">
<collectingDate id="EFD1784CFFD21D485396FE51FE90FE4C" box="[923,1005,442,467]" pageId="12" pageNumber="13" value="1969-04">IV.1969</collectingDate>
</date>
,
<collectorName id="26DEC2B2FFD21D4853F3FE51F848FE4C" box="[1022,1333,442,467]" pageId="12" pageNumber="13">H &amp; A Howden Collection</collectorName>
(
<specimenCount id="9D2D6CEDFFD21D485544FE51F809FE4D" box="[1353,1396,439,467]" pageId="12" pageNumber="13" type="female">2 ♀</specimenCount>
<collectionCode id="ED3A3FA1FFD21D48558CFE51F8B1FE4C" box="[1409,1484,442,467]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="12" pageNumber="13">NHMB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B43AD39FFD21D485368FE31F925FD8C" collectingDate="1969-08-15" collectionCode="NHMB" collectorName="H &amp; A Howden" country="Trinidad and Tobago" location="Morne Bleu" pageId="12" pageNumber="13" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<emphasis id="B95F7B76FFD21D485368FE31FEBEFE6C" box="[869,963,474,499]" italics="true" pageId="12" pageNumber="13">
<collectingCountry id="F33CE7F4FFD21D485368FE31FEC3FE6C" box="[869,958,474,499]" name="Trinidad and Tobago" pageId="12" pageNumber="13">Trinidad</collectingCountry>
,
</emphasis>
W.I.,
<location id="8EF4F1BFFFD21D485406FE31F9E8FE6C" LSID="urn:lsid:plazi:treatment:03821672FFD21D4953B9FF31F981FD0C:8EF4F1BFFFD21D485406FE31F9E8FE6C" box="[1035,1173,474,499]" country="Trinidad and Tobago" name="Morne Bleu" pageId="12" pageNumber="13">Morne Bleu</location>
, 2700’,
<date id="FF9581A4FFD21D4854FEFE31F808FE6C" box="[1267,1397,474,499]" pageId="12" pageNumber="13" value="1969-08-15">
<collectingDate id="EFD1784CFFD21D4854FEFE31F808FE6C" box="[1267,1397,474,499]" pageId="12" pageNumber="13" value="1969-08-15">15.VIII.1969</collectingDate>
</date>
,
<collectorName id="26DEC2B2FFD21D48558BFE31FEBEFD8C" pageId="12" pageNumber="13">H &amp; A Howden</collectorName>
(
<specimenCount id="9D2D6CEDFFD21D4853D5FE11FE80FD8D" box="[984,1021,503,531]" pageId="12" pageNumber="13" type="female">1 ♀</specimenCount>
<collectionCode id="ED3A3FA1FFD21D48540AFE11F92FFD8C" box="[1031,1106,506,531]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="12" pageNumber="13">NHMB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B43AD39FFD21D48546BFE11F824FDAC" collectingDate="1969-08-08" collectionCode="MZSP" collectorName="H &amp; A Howden" country="Trinidad and Tobago" location="Morne Bleu" pageId="12" pageNumber="13" specimenCode="MZSP 35641" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<emphasis id="B95F7B76FFD21D48546BFE11F9BEFD8C" box="[1126,1219,506,531]" italics="true" pageId="12" pageNumber="13">
<collectingCountry id="F33CE7F4FFD21D48546BFE11F9C2FD8C" box="[1126,1215,506,531]" name="Trinidad and Tobago" pageId="12" pageNumber="13">Trinidad</collectingCountry>
,
</emphasis>
W.I.,
<location id="8EF4F1BFFFD21D48550EFE11F8F7FD8C" LSID="urn:lsid:plazi:treatment:03821672FFD21D4953B9FF31F981FD0C:8EF4F1BFFFD21D48550EFE11F8F7FD8C" box="[1283,1418,506,531]" country="Trinidad and Tobago" name="Morne Bleu" pageId="12" pageNumber="13">Morne Bleu</location>
, 2700’,
<date id="FF9581A4FFD21D485368FDF1FE9AFDAC" box="[869,999,538,563]" pageId="12" pageNumber="13" value="1969-08-08">
<collectingDate id="EFD1784CFFD21D485368FDF1FE9AFDAC" box="[869,999,538,563]" pageId="12" pageNumber="13" value="1969-08-08">08.VIII.1969</collectingDate>
</date>
,
<collectorName id="26DEC2B2FFD21D4853FDFDF1F9EDFDAC" box="[1008,1168,538,563]" pageId="12" pageNumber="13">H &amp; A Howden</collectorName>
(
<specimenCount id="9D2D6CEDFFD21D4854ADFDF1F9BDFDAD" box="[1184,1216,535,563]" pageId="12" pageNumber="13" type="female">1 ♀</specimenCount>
<specimenCode id="DB8D0F1FFFD21D4854CBFDF1F82EFDAC" box="[1222,1363,538,563]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="12" pageNumber="13" type="Museum">MZSP 35641</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C331F4EFFFD21D495368FDB1FFD9FEAC" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13" type="diagnosis">
<paragraph id="8B94A764FFD21D485368FDB1F8C6FCEC" blockId="12.[869,1494,602,947]" pageId="12" pageNumber="13">
<emphasis id="B95F7B76FFD21D485368FDB1F916FDEC" bold="true" box="[869,1131,602,627]" pageId="12" pageNumber="13">Differential diagnosis:</emphasis>
Pronotum subquadrate, entirely pale yellow; elytra greyish brown with yellowish to lightbrown apical patch; males: fore femur slightly swollen, with distinct, rounded ventral tooth; ventrite VIII without ridges; posterior margin notched at left side and acute tip; aedeagus with right prolongation of tegmen short;left setiferous prolongation short and wide, bearing thick, short apical setae; females: ventrite VII short, wide, lateral margins nearly parallel, posterior margin slightly projected.
</paragraph>
<paragraph id="8B94A764FFD21D495386FC91FFD9FEAC" blockId="12.[869,1494,602,947]" lastBlockId="13.[94,718,218,307]" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
<taxonomicName id="4C2BDCE7FFD21D485386FC91F99AFC0C" authority="Biffi &amp; Constantin, 2018" authorityName="Biffi &amp; Constantin" authorityYear="2018" box="[907,1255,890,915]" class="Insecta" family="Cantharidae" genus="Macromalthinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="quadratithorax" status="sp. nov.">
<emphasis id="B95F7B76FFD21D485386FC91F99AFC0C" box="[907,1255,890,915]" italics="true" pageId="12" pageNumber="13">Macromalthinus quadratithorax</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26CC60DFFD21D4854FDFC91F835FC0C" box="[1264,1352,890,915]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
differs from the other northern South American species by having the pronotum subquadrate and greyish-brown elytra (the other species have pronotum strongly constricted near anterior angles and dark-brown to black elytra).
</paragraph>
</subSubSection>
<caption id="DF54F7ECFFD21D485368F811F9CCF7CC" ID-DOI="http://doi.org/10.5281/zenodo.5384168" ID-Zenodo-Dep="5384168" httpUri="https://zenodo.org/record/5384168/files/figure.png" pageId="12" pageNumber="13" startId="12.[869,937,2042,2067]" targetBox="[869,1493,978,2012]" targetPageId="12">
<paragraph id="8B94A764FFD21D485368F811F9CCF7CC" blockId="12.[869,1494,2042,2131]" pageId="12" pageNumber="13">
Figures 55-58:
<emphasis id="B95F7B76FFD21D4853F5F811F99AF78C" bold="true" box="[1016,1255,2042,2067]" italics="true" pageId="12" pageNumber="13">
<taxonomicName id="4C2BDCE7FFD21D4853F5F811F99EF78C" authorityName="Pic" authorityYear="1919" box="[1016,1251,2042,2067]" class="Insecta" family="Cantharidae" genus="Macromalthinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="maximiceps">Macromalthinus maximiceps</taxonomicName>
,
</emphasis>
female. (55) Ventrite VII. (56) Apex of abdomen, ventral view. (57) Coxite. (58) Left wing. Scale bars: 55-56:0.5 mm;57:0.2 mm; 58: 2.0mm.
</paragraph>
</caption>
<subSubSection id="C331F4EFFFD31D495053FEB1FE00FE4C" pageId="13" pageNumber="14" type="description">
<paragraph id="8B94A764FFD31D495053FEB1FC80FD2C" blockId="13.[94,719,346,691]" pageId="13" pageNumber="14">
<emphasis id="B95F7B76FFD31D495053FEB1FC28FEEC" bold="true" box="[94,341,346,371]" pageId="13" pageNumber="14">Description: Length:</emphasis>
<quantity id="4CD30A81FFD31D49516CFEB1FC98FEEC" box="[353,485,346,371]" metricMagnitude="-3" metricUnit="m" metricValue="6.05" metricValueMax="6.4" metricValueMin="5.7" pageId="13" pageNumber="14" unit="mm" value="6.05" valueMax="6.4" valueMin="5.7">5.7-6.4 mm</quantity>
.
<emphasis id="B95F7B76FFD31D4951F9FEB1FFF5FEEC" bold="true" box="[500,648,346,371]" pageId="13" pageNumber="14">Colouration:</emphasis>
Head black, frons and clypeus dark brown; antennae dark brown, antennomeres I-III testaceous ventrally, antennomeres IX-XI light brown; labium and maxillae light brown, labial and maxillary palpi testaceous to light brown; pronotum entirely pale yellow; elytra greyish brown, with a yellowish to light-brown apical patch; wings dark brown; legs pale yellow with dark-brown spots on the apex of femora dorsally, dorsal surface of tibiae and distal tarsomeres; thorax and abdominal tergites dark brown, abdominal ventrites light brown (
<figureCitation id="1310BBE1FFD31D4951B9FD71FC8FFD2C" box="[436,498,666,691]" captionStart="Figures 1-6" captionStartId="3.[94,162,2074,2099]" captionTargetBox="[152,1356,206,2041]" captionTargetId="figure-1@3.[151,1360,204,2044]" captionTargetPageId="3" captionText="Figures 1-6:Dorsal habitus ofMacromalthinus species.(1) M.guyanensis sp.nov.(2)M.luteoapicalissp.nov.(3)M.orapuensis sp.nov.(4) M.piceiventrissp.nov. (5) M.quadratithorax sp.nov.(6)M.xerophilus sp.nov.Scale bars:2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5384149" httpUri="https://zenodo.org/record/5384149/files/figure.png" pageId="13" pageNumber="14">Fig. 5</figureCitation>
).
</paragraph>
<paragraph id="8B94A764FFD31D495053FD31FF16F8CC" blockId="13.[94,719,730,1875]" pageId="13" pageNumber="14">
<emphasis id="B95F7B76FFD31D495053FD31FDD6FD6C" bold="true" box="[94,171,730,755]" pageId="13" pageNumber="14">Males:</emphasis>
Head large, 1.3 times wider than the pronotum, temples elongate, strongly tapering posteriorly, vertex concave. Eyes large, oblong, very prominent. Antennae filiform, slender, covered with thin short setae; antennomere I elongate, II very short, 0.4 the length of I, III twice longer than II, IV-XI nearly same length as I. Pronotum (
<figureCitation id="1310BBE1FFD31D495068FC71FDD0FC2C" box="[101,173,922,947]" captionStart="Figures 7-12" captionStartId="4.[869,937,1217,1242]" captionTargetBox="[869,1493,204,1187]" captionTargetId="figure-697@4.[869,1493,204,1187]" captionTargetPageId="4" captionText="Figures 7-12: Pronotum of Macromalthinus species. (7) M. guyanensis sp. nov. (8) M. luteoapicalis sp. nov. (9) M. orapuensis sp. nov. (10) M. piceiventris sp. nov. (11) M. quadratithorax sp. nov. (12) M. xerophilus sp. nov. Scale bars:1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5384151" httpUri="https://zenodo.org/record/5384151/files/figure.png" pageId="13" pageNumber="14">Fig. 11</figureCitation>
) subquadrate, 1.1 times wider than long; anterior margin slightly arched, lateral margins almost parallel, slightly constricted near anterior angles; pronotal surface elevated antero-medially and near posterior angles, forming a V-shaped shallow groove posteriorly. Elytra short, rugulose, densely covered with short and thin setae; sutures broadly dehiscent from posterior half; apex swollen dorsally. Legs slender; fore femur (
<figureCitation id="1310BBE1FFD31D49523FFB91FF00FB0C" box="[562,637,1146,1171]" captionStart="Figures 13-18" captionStartId="5.[94,162,2074,2099]" captionTargetBox="[97,1416,991,2040]" captionTargetId="figure-354@5.[94,1417,986,2044]" captionTargetPageId="5" captionText="Figures 13-18: Fore femur of Macromalthinus species. (13) M. guyanensis sp. nov. (14) M. luteoapicalis sp. nov.(15) M. orapuensis sp.nov.(16) M. piceiventris sp.nov.(17) M. quadratithoraxsp.nov.(18) M. xerophilus sp.nov.Scale bars:0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5384153" httpUri="https://zenodo.org/record/5384153/files/figure.png" pageId="13" pageNumber="14">Fig. 17</figureCitation>
) slightly swollen,with weak,rounded ventral tooth; femora,tibiae and tarsomeres increasing in length from fore to hind legs. Tarsal claws simple, not toothed. Abdominal tergites and sternites sclerotized; ventrites densely covered with long thin setae; glandular pores not prominent; distal margin of ventrite VI arched and slightly sclerotized; ventrite VII (
<figureCitation id="1310BBE1FFD31D4950E4FAB1FC4AFAEC" box="[233,311,1370,1395]" captionStart="Figures 19-24" captionStartId="6.[170,238,2074,2099]" captionTargetBox="[172,1491,1272,2042]" captionTargetId="figure-506@6.[170,1493,1269,2044]" captionTargetPageId="6" captionText="Figures 19-24:Ventrite VII of males of Macromalthinus species.(19) M.guyanensis sp.nov.(20) M.luteoapicalis sp.nov.(21)M. orapuensis sp.nov.(22) M.piceiventris sp.nov.(23) M.quadratithorax sp.nov.(24) M.xerophilus sp.nov.Scale bars:0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5384156" httpUri="https://zenodo.org/record/5384156/files/figure.png" pageId="13" pageNumber="14">Fig. 23</figureCitation>
) H-shaped, lateral margins sinuous, broader anteriorly, with broad membranous area anteriorly and deep posterior notch; ventrite VIII (
<figureCitation id="1310BBE1FFD31D49523AFA71FFFCFA2C" box="[567,641,1434,1459]" captionStart="Figures 25-30" captionStartId="7.[794,862,2010,2035]" captionTargetBox="[798,1414,205,1979]" captionTargetId="figure-491@7.[794,1418,204,1980]" captionTargetPageId="7" captionText="Figures 25-30: Ventrite VIII of males of Macromalthinus species. (25) M. guyanensis sp. nov. (26) M. luteoapicalis sp. nov. (27) M. orapuensis sp.nov.(28)M.piceiventrissp.nov.(29) M.quadratithoraxsp.nov.(30) M.xerophilus sp.nov.Scale bars:0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5384158" httpUri="https://zenodo.org/record/5384158/files/figure.png" pageId="13" pageNumber="14">Fig. 29</figureCitation>
) asymmetrical, oblong, strongly convex, without ridges; distal margin notched at left side, tip acute. Aedeagus (
<figureCitation id="1310BBE1FFD31D495273FA31FFB5FA6C" box="[638,712,1498,1523]" captionStart="Figures 31-36" captionStartId="8.[170,238,2042,2067]" captionTargetBox="[171,1491,674,2008]" captionTargetId="figure-183@8.[170,1493,670,2012]" captionTargetPageId="8" captionText="Figures 31-36: Aedeagus of Macromalthinus species. (31) M. guyanensis sp. nov. (32) M. luteoapicalis sp. nov. (33) M. orapuensis sp. nov. (34) M. piceiventris sp.nov.(35) M. quadratithorax sp.nov.(36) M. xerophilus sp.nov. Abbreviations:pt = prolongation of tegmen, sp = setiferous prolongation,rp = right paramere, lp = left paramere,ml = median lobe.Scale bars:0.5mm." figureDoi="http://doi.org/10.5281/zenodo.5384160" httpUri="https://zenodo.org/record/5384160/files/figure.png" pageId="13" pageNumber="14">Fig. 35</figureCitation>
) with right prolongation of tegmen short, projected dorsally, without apophysis on its dorsal surface; left setiferous prolongation short, wide, bearing thick, short apical setae; right paramere elongate, slightly curved, with slight constriction near apex; left paramere elongate, narrow, flat, apex rounded; median lobe straight, broad, apex membranous.
<emphasis id="B95F7B76FFD31D495134F951FF02F94C" bold="true" box="[313,639,1722,1747]" pageId="13" pageNumber="14">
Measurements of
<typeStatus id="549019C6FFD31D495201F951FF06F94C" box="[524,635,1722,1747]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
:
</emphasis>
TL: 6.0; AL: 4.8; OL: 0.51; HW: 1.12; IOW: 0.57; IAW: 0.2; PL: 0.86; PW: 0.93; EL: 1.8; EW: 1.24.
<emphasis id="B95F7B76FFD31D495182F911FDF6F8AC" bold="true" pageId="13" pageNumber="14">Length of each antennomere:</emphasis>
aI: 0.37; aII: 0.18; aIII: 0.30; aIV: 0.43; aV: 0.45; aVI: 0.47; aVII: 0.47; aVIII: 0.48; aIX: 0.48; aX: 0.43; aXI: 0.46.
</paragraph>
<paragraph id="8B94A764FFD31D495053F891FE00FE4C" blockId="13.[94,719,1914,2131]" lastBlockId="13.[794,1418,218,467]" pageId="13" pageNumber="14">
<emphasis id="B95F7B76FFD31D495053F891FDC6F80C" bold="true" box="[94,187,1914,1939]" pageId="13" pageNumber="14">Female:</emphasis>
similar to male; eyes smaller, less prominent; fore femora slender, without intumescence or tooth; ventrite VI entire, not notched posteriorly; ventrite VII (
<figureCitation id="1310BBE1FFD31D495068F831FDD2F86C" box="[101,175,2010,2035]" captionStart="Figures 37-42" captionStartId="9.[94,162,2074,2099]" captionTargetBox="[94,1414,1424,2040]" captionTargetId="figure-592@9.[94,1417,1420,2044]" captionTargetPageId="9" captionText="Figures 37-42:VentriteVII offemales ofMacromalthinusspecies.(37) M.guyanensis sp.nov.(38)M.luteoapicalissp.nov.(39) M.orapuensissp.nov.(40)M.piceiventris sp.nov.(41) M. quadratithoraxsp.nov.(42) M. xerophilus sp.nov.Scale bars:0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5384162" httpUri="https://zenodo.org/record/5384162/files/figure.png" pageId="13" pageNumber="14">Fig. 41</figureCitation>
) short, wide, lateral margins arched, slightly narrowed near angles, posterior margin slightly projected, strongly sinuous, forming two rounded lobes; base of coxites (
<figureCitation id="1310BBE1FFD31D4950B7F7D1FC7BF7CC" box="[186,262,2106,2131]" captionStart="Figures 43-48" captionStartId="10.[869,937,2010,2035]" captionTargetBox="[873,1490,1055,1976]" captionTargetId="figure-666@10.[869,1493,1054,1980]" captionTargetPageId="10" captionText="Figures 43-48: Coxites of Macromalthinus species. (43) M. guyanensis sp. nov. (44) M. luteoapicalis sp. nov. (45) M. orapuensis sp. nov. (46) M. piceiventris sp. nov. (47) M. quadratithorax sp. nov. (48) M. xerophilus sp. nov. Scale bars:0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5384164" httpUri="https://zenodo.org/record/5384164/files/figure.png" pageId="13" pageNumber="14">Fig. 47</figureCitation>
) elongate, slender, apex very short, with short and broad projection behind styles bearing few thin setae; styles short, slightly wider apically, bearing thick setae.
<emphasis id="B95F7B76FFD31D4953A1FEF1FEF1FECC" bold="true" pageId="13" pageNumber="14">
Measurements of medium size female
<typeStatus id="549019C6FFD31D495317FED1FEF4FECC" box="[794,905,314,339]" pageId="13" pageNumber="14" type="paratype">paratype</typeStatus>
:
</emphasis>
TL: 5.6; AL: 5.12; OL: 0.55; HW: 1.15; IOW: 0.58; IAW: 0.21; PL: 0.87; PW: 0.97; EL: 1.70; EW: 1.22.
<emphasis id="B95F7B76FFD31D495515FEB1F97DFE0C" bold="true" pageId="13" pageNumber="14">Length of each antennomere:</emphasis>
aI: 0.46; aII: 0.12; aIII: 0.38; aIV: 0.51; aV: 0.53; aVI: 0.53; aVII: 0.53; aVIII: 0.55; aIX: 0.52; aX: 0.47; aXI: 0.47.
</paragraph>
</subSubSection>
<subSubSection id="C331F4EFFFD31D495317FE11F875FDCC" pageId="13" pageNumber="14" type="etymology">
<paragraph id="8B94A764FFD31D495317FE11F875FDCC" blockId="13.[794,1417,506,595]" pageId="13" pageNumber="14">
<emphasis id="B95F7B76FFD31D495317FE11FEDCFD8C" bold="true" box="[794,929,506,531]" pageId="13" pageNumber="14">Etymology:</emphasis>
The specific epithet is derivative of the Latin words
<emphasis id="B95F7B76FFD31D495369FDF1FEAFFDAC" box="[868,978,538,563]" italics="true" pageId="13" pageNumber="14">quadratus</emphasis>
(square) +
<emphasis id="B95F7B76FFD31D49545DFDF1F9EBFDAC" box="[1104,1174,538,563]" italics="true" pageId="13" pageNumber="14">thorax</emphasis>
(pronotum), referring to the shape of the pronotum in this species.
</paragraph>
</subSubSection>
<subSubSection id="C331F4EFFFD31D495317FD91F981FD0C" box="[794,1276,634,659]" pageId="13" pageNumber="14" type="distribution">
<paragraph id="8B94A764FFD31D495317FD91F981FD0C" blockId="13.[794,1276,634,659]" box="[794,1276,634,659]" pageId="13" pageNumber="14">
<emphasis id="B95F7B76FFD31D495317FD91FECCFD0C" bold="true" box="[794,945,634,659]" pageId="13" pageNumber="14">Distribution:</emphasis>
<collectingCountry id="F33CE7F4FFD31D4953BBFD91F9E7FD0C" box="[950,1178,634,659]" name="Trinidad and Tobago" pageId="13" pageNumber="14">Trinidad and Tobago</collectingCountry>
(
<figureCitation id="1310BBE1FFD31D4954A8FD91F98FFD0C" box="[1189,1266,634,659]" captionStart="Figures 66-69" captionStartId="19.[94,162,2042,2067]" captionTargetBox="[151,1360,431,2012]" captionTargetId="figure-79@19.[151,1360,431,2012]" captionTargetPageId="19" captionText="Figures 66-69: Distribution of Macromalthinus species. (66) Distribution of Macromalthinus species in South America. (67) Distribution of M. brasiliensis in Brazil.(68)Distribution ofM.maximiceps inFrench Guiana.(69)Distribution of M.guyanensis sp.nov.,andM.orapuensis sp.nov.inFrench Guiana.Background maps of Figs.66-67 by NaturalEarth (available at:www.naturalearthdata.com), and Figs.68-69 byEric Gaba." figureDoi="http://doi.org/10.5281/zenodo.5384172" httpUri="https://zenodo.org/record/5384172/files/figure.png" pageId="13" pageNumber="14">Fig. 66</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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