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248 lines
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<document id="924D7014C44E27F61C0BCDC1FF27E931" ID-DOI="10.11646/zootaxa.5523.1.9" ID-ISSN="1175-5326" ID-Zenodo-Dep="13934022" ID-ZooBank="6BB3EC47-EE6C-4286-BC85-0649F93513AC" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1728985959978" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2024" docId="E21C250D4B082F22FF4C2CDEFD01FAB5" docLanguage="en" docName="zootaxa.5523.1.9.pdf" docOrigin="Zootaxa 5523 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5523.1.9" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cephennomicrus tenebrosus Jałoszyński, 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="142" masterDocId="1E255D754B092F21FFDB295CFF86FFCC" masterDocTitle="A new species and new records of Cephennomicrus Reitter in Japan (Coleoptera, Staphylinidae, Scydmaeninae)" masterLastPageNumber="145" masterPageNumber="139" pageNumber="140" updateTime="1728986532874" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="F9DD58F97CF08768C3FF1582C6082731">A new species and new records of Cephennomicrus Reitter in Japan (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
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<mods:namePart id="37552903CB0F0AFC0F124FDE369DD029">Jałoszyński, Paweł</mods:namePart>
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<mods:title id="4D45617B498846DBFD7FE7DC224F6A0C">Zootaxa</mods:title>
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<mods:part id="73D31C279F68555F9D455CB34C2000FA">
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<mods:date id="72F4870AA24363A8A6C59813C1ADE7C6">2024</mods:date>
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<mods:number id="CA425ED16F999BD5248A0BE787CF59F1">2024-10-14</mods:number>
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<mods:number id="79CC7A029EA1A9F6794317AC4656AEEA">5523</mods:number>
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<mods:url id="F0F601EDFA661BB187A1D91D0BF26712">http://dx.doi.org/10.11646/zootaxa.5523.1.9</mods:url>
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<mods:identifier id="325F07933A145C7EE33F56BA69269EAF" type="DOI">10.11646/zootaxa.5523.1.9</mods:identifier>
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<mods:identifier id="BD532B51783BFAE65678C5807A027B61" type="ISSN">1175-5326</mods:identifier>
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<treatment id="E21C250D4B082F22FF4C2CDEFD01FAB5" ID-DOI="http://doi.org/10.5281/zenodo.13934051" ID-Zenodo-Dep="13934051" LSID="urn:lsid:plazi:treatment:E21C250D4B082F22FF4C2CDEFD01FAB5" httpUri="http://treatment.plazi.org/id/E21C250D4B082F22FF4C2CDEFD01FAB5" lastPageId="3" lastPageNumber="142" pageId="1" pageNumber="140">
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<subSubSection id="22AFC7904B082F20FF4C2CDEFDD3FA51" box="[151,597,1410,1437]" pageId="1" pageNumber="140" type="nomenclature">
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<paragraph id="6A0A941B4B082F20FF4C2CDEFDD3FA51" blockId="1.[151,597,1410,1473]" box="[151,597,1410,1437]" pageId="1" pageNumber="140">
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<heading id="314223774B082F20FF4C2CDEFDD3FA51" bold="true" box="[151,597,1410,1437]" fontSize="11" level="1" pageId="1" pageNumber="140" reason="1">
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<emphasis id="58C148094B082F20FF4C2CDEFDD3FA51" bold="true" box="[151,597,1410,1437]" pageId="1" pageNumber="140">
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<taxonomicName id="ADB5EF984B082F20FF4C2CDEFE76FA51" authorityName="Jałoszyński" authorityYear="2024" box="[151,496,1410,1437]" class="Insecta" family="Staphylinidae" genus="Cephennomicrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="140" phylum="Arthropoda" rank="species" species="tenebrosus" status="sp. nov.">
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<emphasis id="58C148094B082F20FF4C2CDEFE76FA51" bold="true" box="[151,496,1410,1437]" italics="true" pageId="1" pageNumber="140">Cephennomicrus tenebrosus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="43F2F5724B082F20FE2C2CDFFDD3FA51" box="[503,597,1411,1437]" pageId="1" pageNumber="140" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="22AFC7904B082F20FF4C2CFBFE8DFA0D" box="[151,267,1447,1473]" pageId="1" pageNumber="140" type="description">
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<paragraph id="6A0A941B4B082F20FF4C2CFBFE8DFA0D" blockId="1.[151,597,1410,1473]" box="[151,267,1447,1473]" pageId="1" pageNumber="140">
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(
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<figureCitation id="F28E889E4B082F20FF442CFBFE84FA0D" box="[159,258,1447,1473]" captionStart-0="FIGURES 1–7" captionStart-1="FIGURE 8" captionStartId-0="2.[151,264,1676,1700]" captionStartId-1="4.[151,250,1259,1283]" captionTargetBox-0="[155,1429,185,1644]" captionTargetBox-1="[151,1428,179,1229]" captionTargetId-0="figure-24@2.[151,1436,179,1650]" captionTargetId-1="figure-24@4.[151,1435,179,1233]" captionTargetPageId-0="2" captionTargetPageId-1="4" captionText-0="FIGURES 1–7. Cephennomicrus tenebrosus sp. nov., holotype (1–5), C. japonigenus Jałoszyński & Hoshina (6), and C. nagoanus Jałoszyński (7). Dorsal habitus (1), aedeagus in ventral (2, 4, 6, 7) and lateral (3, 5) views." captionText-1="FIGURE 8. Distribution of nominal Cephennomicrus species in Japan, with aedeagus illustrated for each species: a, C. tanakai Hoshina; b, C. japonigenus (Jałoszyński & Hoshina); c, C. cactiformis (Jałoszyński & Hoshina); d, C. ushimanus Jałoszyński; e, C. okinawanus (Jałoszyński, S. Arai & K.Arai); f, C. disjunctus (Jałoszyński, S. Arai & K. Arai); g, C. nagoanus Jałoszyński; h, C. aji Jałoszyński; i, C. tenebrosus Jałoszyński; j, C. nomurai (Jałoszyński & Hoshina); k, C. hobbiti (Jałoszyński & Hoshina); l, C. iriomotensis Jałoszyński; m, C. pseudojaponigenus Jałoszyński." figureDoi-0="http://doi.org/10.5281/zenodo.13934024" figureDoi-1="http://doi.org/10.5281/zenodo.13934026" httpUri-0="https://zenodo.org/record/13934024/files/figure.png" httpUri-1="https://zenodo.org/record/13934026/files/figure.png" pageId="1" pageNumber="140">Figs 1–8</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="22AFC7904B082F20FF4C2CB3FF79F9B9" pageId="1" pageNumber="140" type="materials_examined">
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<paragraph id="6A0A941B4B082F20FF4C2CB3FF79F9B9" blockId="1.[151,1437,1518,1977]" pageId="1" pageNumber="140">
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<emphasis id="58C148094B082F20FF4C2CB3FECEF9C4" bold="true" box="[151,328,1518,1545]" pageId="1" pageNumber="140">Type material.</emphasis>
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<materialsCitation id="DADD9E464B082F20FE882CB3FF79F9B9" collectingDate="2018-03-24" collectionCode="NSMT" collectorName="N. Tokushige & Japanese" country="Japan" location="Nishihara Undokoen" municipality="Nishihara-cho" pageId="1" pageNumber="140" specimenCount="1" specimenCount-male="1" stateProvince="Okinawa" typeStatus="holotype">
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<emphasis id="58C148094B082F20FE882CB3FD61F9C4" bold="true" box="[339,743,1518,1545]" pageId="1" pageNumber="140">
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<typeStatus id="B50E2AB94B082F20FE882CB3FE42F9C5" box="[339,452,1519,1545]" pageId="1" pageNumber="140" type="holotype">Holotype</typeStatus>
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:
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<collectingCountry id="12A2D48B4B082F20FE082CB2FDABF9C4" box="[467,557,1518,1544]" name="Japan" pageId="1" pageNumber="140">JAPAN</collectingCountry>
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(
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<collectingRegion id="A8715AF94B082F20FDE42CB2FD28F9C4" box="[575,686,1518,1544]" country="Japan" name="Okinawa" pageId="1" pageNumber="140">Okinawa</collectingRegion>
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Is.):
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</emphasis>
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<specimenCount id="7CB35F924B082F20FD292CB3FC8FF9C5" box="[754,777,1519,1545]" count="1" pageId="1" pageNumber="140" type="male">♂</specimenCount>
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, three labels: “JAPAN,
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<collectingRegion id="A8715AF94B082F20FBFB2CB3FB24F9C5" box="[1056,1186,1519,1545]" country="Japan" name="Okinawa" pageId="1" pageNumber="140">OKINAWA</collectingRegion>
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Is. /
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<collectingMunicipality id="8A6E0E614B082F20FB3F2CB3FA00F9C5" box="[1252,1414,1519,1545]" pageId="1" pageNumber="140">Nishihara-chō</collectingMunicipality>
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, /
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<location id="6F6AC2C04B082F20FF4C2F4FFE05F9E1" LSID="urn:lsid:plazi:treatment:E21C250D4B082F22FF4C2CDEFD01FAB5:6F6AC2C04B082F20FF4C2F4FFE05F9E1" box="[151,387,1555,1581]" country="Japan" municipality="Nishihara-cho" name="Nishihara Undokoen" pageId="1" pageNumber="140" stateProvince="Okinawa">Nishihara Undōkōen</location>
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/
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<date id="1E0BB2DB4B082F20FE462F4FFDA6F9E1" box="[413,544,1555,1581]" pageId="1" pageNumber="140" value="2018-03-24">
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<collectingDate id="0E4F4B334B082F20FE462F4FFDA6F9E1" box="[413,544,1555,1581]" pageId="1" pageNumber="140" value="2018-03-24">24 III 2018</collectingDate>
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</date>
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/ leg.
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<collectorName id="C740F1CD4B082F20FDB52F4FFC8FF9E1" box="[622,777,1555,1581]" pageId="1" pageNumber="140">N. Tokushige</collectorName>
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” [white, printed], the same data are repeated in
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<collectorName id="C740F1CD4B082F20FAE22F4FFA1BF9E1" box="[1337,1437,1555,1581]" pageId="1" pageNumber="140">Japanese</collectorName>
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on second label, “
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<taxonomicName id="ADB5EF984B082F20FEB02F6BFDE3F99D" authorityName="Reitter" authorityYear="1907" box="[363,613,1591,1617]" class="Insecta" family="Staphylinidae" genus="Cephennomicrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="140" phylum="Arthropoda" rank="genus">
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<emphasis id="58C148094B082F20FEB02F6BFDE3F99D" box="[363,613,1591,1617]" italics="true" pageId="1" pageNumber="140">CEPHENNOMICRUS</emphasis>
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</taxonomicName>
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/
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<taxonomicName id="ADB5EF984B082F20FD582F6BFD7DF99D" authorityName="Jałoszyński" authorityYear="2024" box="[643,763,1591,1617]" class="Insecta" family="Staphylinidae" genus="Cephennomicrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="140" phylum="Arthropoda" rank="species" species="tenebrosus">
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<emphasis id="58C148094B082F20FD582F6BFD7DF99D" box="[643,763,1591,1617]" italics="true" pageId="1" pageNumber="140">tenebrosus</emphasis>
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</taxonomicName>
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m. / P.
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<bibRefCitation id="0E24E9EA4B082F20FC842F6BFBACF99D" author="Jaloszynski, P." box="[863,1066,1591,1617]" pageId="1" pageNumber="140" pagination="465 - 472" refId="ref4189" refString="Jaloszynski, P. (2024) Cephenniini of the Philippines. Part 6. First record of Cephennomicrus Reitter from Mindanao, with three new species (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa, 5471 (4), 465 - 472. https: // doi. org / 10.11646 / zootaxa. 5471.4.5" type="journal article" year="2024">Jałoszyński, 2024</bibRefCitation>
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/
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<typeStatus id="B50E2AB94B082F20FB9C2F6BFB76F99D" box="[1095,1264,1591,1617]" pageId="1" pageNumber="140" type="holotype">HOLOTYPUS</typeStatus>
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” [red, printed] (
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<collectionCode id="0CA40CDE4B082F20FF442F07FF75F9B9" box="[159,243,1627,1653]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="1" pageNumber="140">NSMT</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="22AFC7904B082F20FF1C2F22FCBBF8BD" pageId="1" pageNumber="140" type="diagnosis">
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<paragraph id="6A0A941B4B082F20FF1C2F22FCBBF8BD" blockId="1.[151,1437,1518,1977]" pageId="1" pageNumber="140">
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<emphasis id="58C148094B082F20FF1C2F22FEC4F954" bold="true" box="[199,322,1662,1688]" pageId="1" pageNumber="140">Diagnosis.</emphasis>
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Member of
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<taxonomicName id="ADB5EF984B082F20FE3A2F23FC8AF954" authorityName="Ishigaki Is., Iriomote Is., Yonaguni Is." baseAuthorityName="Jaloszynski & Hoshina" baseAuthorityYear="2003" box="[481,780,1663,1689]" class="Insecta" family="Staphylinidae" genus="Cephennomicrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="140" phylum="Arthropoda" rank="species" species="nomurai">
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<emphasis id="58C148094B082F20FE3A2F23FC8AF954" box="[481,780,1663,1689]" italics="true" pageId="1" pageNumber="140">Cephennomicrus nomurai</emphasis>
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</taxonomicName>
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species group (based on median longitudinal pronotal groove) with dark brown body, in male frons weakly elevated and delimited laterally by narrow longitudinal impressions, trimerous antennal club so slender that not delimited from proximal antennomeres, with antennomere 11 conspicuously elongate, 2.5 times as long as broad; aedeagus in ventral view drop-shaped, apex with pair of elongate lateral lobes curved mesally, endophallus with three pairs of elongate sclerites: median pair with each sclerite approximately L-shaped and touching at middle, sublateral pair with each sclerite bifurcate, and lateral outer pair with each sclerite strongly elongate and curved mesally.
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</paragraph>
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</subSubSection>
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<subSubSection id="22AFC7904B082F22FF1C2E26FE18FC4D" lastPageId="3" lastPageNumber="142" pageId="1" pageNumber="140" type="description">
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<paragraph id="6A0A941B4B082F20FF1C2E26FD3DF875" blockId="1.[151,1437,1518,1977]" pageId="1" pageNumber="140">
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<emphasis id="58C148094B082F20FF1C2E26FEDFF858" bold="true" box="[199,345,1914,1940]" pageId="1" pageNumber="140">Description.</emphasis>
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Body of male (
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<figureCitation id="F28E889E4B082F20FDD02E27FDCAF859" box="[523,588,1915,1941]" captionStart="FIGURES 1–7" captionStartId="2.[151,264,1676,1700]" captionTargetBox="[155,1429,185,1644]" captionTargetId="figure-24@2.[151,1436,179,1650]" captionTargetPageId="2" captionText="FIGURES 1–7. Cephennomicrus tenebrosus sp. nov., holotype (1–5), C. japonigenus Jałoszyński & Hoshina (6), and C. nagoanus Jałoszyński (7). Dorsal habitus (1), aedeagus in ventral (2, 4, 6, 7) and lateral (3, 5) views." figureDoi="http://doi.org/10.5281/zenodo.13934024" httpUri="https://zenodo.org/record/13934024/files/figure.png" pageId="1" pageNumber="140">Fig. 1</figureCitation>
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) stout, strongly convex, dark brown with lighter tibiae, tarsi and antennae; setae distinctly lighter than cuticle; BL
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<quantity id="AD4D39FE4B082F20FD8B2EC3FD31F875" box="[592,695,1951,1977]" metricMagnitude="-3" metricUnit="m" metricValue="1.08" pageId="1" pageNumber="140" unit="mm" value="1.08">1.08 mm</quantity>
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.
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</paragraph>
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<caption id="3ECAC4934B0B2F23FF4C2FD0FB15F904" ID-DOI="http://doi.org/10.5281/zenodo.13934024" ID-Zenodo-Dep="13934024" httpUri="https://zenodo.org/record/13934024/files/figure.png" pageId="2" pageNumber="141" startId="2.[151,264,1676,1700]" targetBox="[155,1429,185,1644]" targetPageId="2" targetType="figure">
|
|||
|
|
<paragraph id="6A0A941B4B0B2F23FF4C2FD0FB15F904" blockId="2.[151,1437,1676,1736]" pageId="2" pageNumber="141">
|
|||
|
|
<emphasis id="58C148094B0B2F23FF4C2FD0FEB8F968" bold="true" box="[151,318,1676,1700]" pageId="2" pageNumber="141">FIGURES 1–7.</emphasis>
|
|||
|
|
<taxonomicName id="ADB5EF984B0B2F23FE922FD0FDEDF968" authorityName="Jałoszyński" authorityYear="2024" box="[329,619,1676,1700]" class="Insecta" family="Staphylinidae" genus="Cephennomicrus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="141" phylum="Arthropoda" rank="species" species="tenebrosus" status="sp. nov.">
|
|||
|
|
<emphasis id="58C148094B0B2F23FE922FD0FDEDF968" box="[329,619,1676,1700]" italics="true" pageId="2" pageNumber="141">Cephennomicrus tenebrosus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="58C148094B0B2F23FDAE2FD0FD4CF968" bold="true" box="[629,714,1676,1700]" pageId="2" pageNumber="141">
|
|||
|
|
<taxonomicNameLabel id="43F2F5724B0B2F23FDAE2FD0FD4CF968" box="[629,714,1676,1700]" pageId="2" pageNumber="141" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
, holotype (
|
|||
|
|
<emphasis id="58C148094B0B2F23FC9D2FD0FCEAF968" bold="true" box="[838,876,1676,1700]" pageId="2" pageNumber="141">1–5</emphasis>
|
|||
|
|
),
|
|||
|
|
<taxonomicName id="ADB5EF984B0B2F23FC5E2FD0FA98F968" authority="Jaloszynski & Hoshina" authorityName="Jaloszynski & Hoshina" box="[901,1310,1676,1700]" class="Insecta" family="Staphylinidae" genus="Cephennomicrus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="141" phylum="Arthropoda" rank="species" species="japonigenus">
|
|||
|
|
<emphasis id="58C148094B0B2F23FC5E2FD0FBA5F968" box="[901,1059,1676,1700]" italics="true" pageId="2" pageNumber="141">C. japonigenus</emphasis>
|
|||
|
|
Jałoszyński & Hoshina
|
|||
|
|
</taxonomicName>
|
|||
|
|
(
|
|||
|
|
<emphasis id="58C148094B0B2F23FAEA2FD0FAB8F968" bold="true" box="[1329,1342,1676,1700]" pageId="2" pageNumber="141">6</emphasis>
|
|||
|
|
), and
|
|||
|
|
<taxonomicName id="ADB5EF984B0B2F23FA5E2FD0FEF3F904" authority="Jaloszynski" authorityName="Jaloszynski" authorityYear="2009" class="Insecta" family="Staphylinidae" genus="Cephennomicrus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="141" phylum="Arthropoda" rank="species" species="nagoanus">
|
|||
|
|
<emphasis id="58C148094B0B2F23FA5E2FD0FF7CF904" italics="true" pageId="2" pageNumber="141">C. nagoanus</emphasis>
|
|||
|
|
Jałoszyński
|
|||
|
|
</taxonomicName>
|
|||
|
|
(
|
|||
|
|
<emphasis id="58C148094B0B2F23FE5F2FECFE17F904" bold="true" box="[388,401,1712,1736]" pageId="2" pageNumber="141">7</emphasis>
|
|||
|
|
). Dorsal habitus (
|
|||
|
|
<emphasis id="58C148094B0B2F23FD9C2FECFDD2F904" bold="true" box="[583,596,1712,1736]" pageId="2" pageNumber="141">1</emphasis>
|
|||
|
|
), aedeagus in ventral (
|
|||
|
|
<emphasis id="58C148094B0B2F23FCE02FECFC12F904" bold="true" box="[827,916,1712,1736]" pageId="2" pageNumber="141">2, 4, 6, 7</emphasis>
|
|||
|
|
) and lateral (
|
|||
|
|
<emphasis id="58C148094B0B2F23FBC72FECFBC4F904" bold="true" box="[1052,1090,1712,1736]" pageId="2" pageNumber="141">3, 5</emphasis>
|
|||
|
|
) views.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="6A0A941B4B0B2F23FF1C2E46FED2F824" blockId="2.[151,1437,1818,2025]" pageId="2" pageNumber="141">
|
|||
|
|
Head broadest at large, strongly convex and coarsely faceted eyes, HL
|
|||
|
|
<quantity id="AD4D39FE4B0B2F23FC3D2E46FBCAF8F9" box="[998,1100,1818,1845]" metricMagnitude="-4" metricUnit="m" metricValue="1.3" pageId="2" pageNumber="141" unit="mm" value="0.13">0.13 mm</quantity>
|
|||
|
|
, HW
|
|||
|
|
<quantity id="AD4D39FE4B0B2F23FB552E46FB73F8F9" box="[1166,1269,1818,1845]" metricMagnitude="-4" metricUnit="m" metricValue="2.6" pageId="2" pageNumber="141" unit="mm" value="0.26">0.26 mm</quantity>
|
|||
|
|
; frontal glands indiscernible; frons weakly elevated and laterally delimited by narrow impressions; supraantennal tubercles distinct but weakly elevated. Punctures on frons and vertex inconspicuous, fine; setae sparse, short and suberect. Antennae conspicuously slender, with trimerous clubs so slender that apparently not delimited,AnL
|
|||
|
|
<quantity id="AD4D39FE4B0B2F23FBAA2EDAFB50F86D" box="[1137,1238,1926,1953]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="141" unit="mm" value="0.6">0.60 mm</quantity>
|
|||
|
|
; all antennomeres elongate, 1–2, 7, 10 and 11 each strongly so, remaining antennomeres slightly elongate, antennomere 11 2.5 times as long as broad.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="6A0A941B4B0A2F22FF1C29CBFF77FE45" blockId="3.[151,1437,151,1401]" pageId="3" pageNumber="142">
|
|||
|
|
Pronotum bell-shaped, broadest between middle and anterior third and distinctly narrowing posteriorly; PL
|
|||
|
|
<quantity id="AD4D39FE4B0A2F22FAB029CBFF43FF19" metricMagnitude="-4" metricUnit="m" metricValue="3.3" pageId="3" pageNumber="142" unit="mm" value="0.33">0.33 mm</quantity>
|
|||
|
|
, PW
|
|||
|
|
<quantity id="AD4D39FE4B0A2F22FF2629E7FEE4FF19" box="[253,354,187,213]" metricMagnitude="-4" metricUnit="m" metricValue="3.9" pageId="3" pageNumber="142" unit="mm" value="0.39">0.39 mm</quantity>
|
|||
|
|
; anterior margin weakly convex; lateral margins strongly rounded in anterior half, slightly concave behind middle and nearly straight in front of hind corners; hind corners nearly right-angled and blunt; posterior margin weakly arcuate. Pronotal disc with slightly microserrate lateral carinae, sharply marked median longitudinal groove, narrow transverse antebasal groove connecting small inner pits, and small outer pits slightly anterad inner pits. Punctures on pronotal disc fine, inconspicuous; setae moderately dense, short and weakly suberect, macrosetae lacking.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="6A0A941B4B0A2F22FF1C28CFFF77FDF1" blockId="3.[151,1437,151,1401]" pageId="3" pageNumber="142">
|
|||
|
|
Elytra together oval, broadest slightly in front of middle; EL
|
|||
|
|
<quantity id="AD4D39FE4B0A2F22FC5F28CFFC6AFE61" box="[900,1004,403,429]" metricMagnitude="-4" metricUnit="m" metricValue="6.3" pageId="3" pageNumber="142" unit="mm" value="0.63">0.63 mm</quantity>
|
|||
|
|
, EW
|
|||
|
|
<quantity id="AD4D39FE4B0A2F22FBF628CEFB13FE61" box="[1069,1173,402,429]" metricMagnitude="-4" metricUnit="m" metricValue="5.3" pageId="3" pageNumber="142" unit="mm" value="0.53">0.53 mm</quantity>
|
|||
|
|
, EI 1.19; humeral calli distinct; subhumeral lines lacking; sides of elytra strongly rounded; elytral apices separately rounded. Punctures more distinct than those on pronotum and dense, separated by spaces subequal to their diameters, but punctures are superficial and with diffuse margins so they appear indistinct. Setae similar to those on pronotum, macrosetae lacking.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="6A0A941B4B0A2F22FF1C2B1BFDB4FDAD" blockId="3.[151,1437,151,1401]" box="[199,562,583,609]" pageId="3" pageNumber="142">Hind wings long and functional.</paragraph>
|
|||
|
|
<paragraph id="6A0A941B4B0A2F22FF1C2B37FD5EFD49" blockId="3.[151,1437,151,1401]" box="[199,728,619,645]" pageId="3" pageNumber="142">Legs moderately long and slender, unmodified.</paragraph>
|
|||
|
|
<paragraph id="6A0A941B4B0A2F22FF1C2BD3FE74FC91" blockId="3.[151,1437,151,1401]" pageId="3" pageNumber="142">
|
|||
|
|
Aedeagus (
|
|||
|
|
<figureCitation id="F28E889E4B0A2F22FE902BD3FE30FD65" box="[331,438,654,681]" captionStart="FIGURES 1–7" captionStartId="2.[151,264,1676,1700]" captionTargetBox="[155,1429,185,1644]" captionTargetId="figure-24@2.[151,1436,179,1650]" captionTargetPageId="2" captionText="FIGURES 1–7. Cephennomicrus tenebrosus sp. nov., holotype (1–5), C. japonigenus Jałoszyński & Hoshina (6), and C. nagoanus Jałoszyński (7). Dorsal habitus (1), aedeagus in ventral (2, 4, 6, 7) and lateral (3, 5) views." figureDoi="http://doi.org/10.5281/zenodo.13934024" httpUri="https://zenodo.org/record/13934024/files/figure.png" pageId="3" pageNumber="142">Figs 2–5</figureCitation>
|
|||
|
|
) stout, drop-shaped; AeL
|
|||
|
|
<quantity id="AD4D39FE4B0A2F22FD282BD3FCE6FD65" box="[755,864,655,681]" metricMagnitude="-4" metricUnit="m" metricValue="2.8" pageId="3" pageNumber="142" unit="mm" value="0.28">0.28 mm</quantity>
|
|||
|
|
; median lobe in ventral view broadest in basal region, with subtriangular blunt apex flanked by elongate lateral lobes curved mesally; endophallus complex and symmetrical, with six elongate sclerites: pair of median L-shaped sclerites touching at middle, submedian lateral pair each composed of two fused elongate sclerites, so that each pair appears as bifurcate, and long outer lateral sclerites curved mesally. Parameres slender, their apices not reaching apex of median lobe, each paramere with one apical and two subapical setae.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="6A0A941B4B0A2F22FF1C2A3BFE18FC4D" blockId="3.[151,1437,151,1401]" box="[199,414,871,897]" pageId="3" pageNumber="142">Female. Unknown.</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="22AFC7904B0A2F22FF1C2AD6FDFFFC69" box="[199,633,906,933]" pageId="3" pageNumber="142" type="distribution">
|
|||
|
|
<paragraph id="6A0A941B4B0A2F22FF1C2AD6FDFFFC69" blockId="3.[151,1437,151,1401]" box="[199,633,906,933]" pageId="3" pageNumber="142">
|
|||
|
|
<emphasis id="58C148094B0A2F22FF1C2AD6FEDFFC68" bold="true" box="[199,345,906,932]" pageId="3" pageNumber="142">Distribution</emphasis>
|
|||
|
|
.
|
|||
|
|
<collectingRegion id="A8715AF94B0A2F22FEBC2AD7FE4AFC69" box="[359,460,907,933]" country="Japan" name="Okinawa" pageId="3" pageNumber="142">Okinawa</collectingRegion>
|
|||
|
|
Island (
|
|||
|
|
<figureCitation id="F28E889E4B0A2F22FDFC2AD7FDEAFC69" box="[551,620,907,933]" captionStart="FIGURE 8" captionStartId="4.[151,250,1259,1283]" captionTargetBox="[151,1428,179,1229]" captionTargetId="figure-24@4.[151,1435,179,1233]" captionTargetPageId="4" captionText="FIGURE 8. Distribution of nominal Cephennomicrus species in Japan, with aedeagus illustrated for each species: a, C. tanakai Hoshina; b, C. japonigenus (Jałoszyński & Hoshina); c, C. cactiformis (Jałoszyński & Hoshina); d, C. ushimanus Jałoszyński; e, C. okinawanus (Jałoszyński, S. Arai & K.Arai); f, C. disjunctus (Jałoszyński, S. Arai & K. Arai); g, C. nagoanus Jałoszyński; h, C. aji Jałoszyński; i, C. tenebrosus Jałoszyński; j, C. nomurai (Jałoszyński & Hoshina); k, C. hobbiti (Jałoszyński & Hoshina); l, C. iriomotensis Jałoszyński; m, C. pseudojaponigenus Jałoszyński." figureDoi="http://doi.org/10.5281/zenodo.13934026" httpUri="https://zenodo.org/record/13934026/files/figure.png" pageId="3" pageNumber="142">Fig. 8</figureCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="22AFC7904B0A2F22FF1C2AF3FA08FC05" box="[199,1422,943,969]" pageId="3" pageNumber="142" type="etymology">
|
|||
|
|
<paragraph id="6A0A941B4B0A2F22FF1C2AF3FA08FC05" blockId="3.[151,1437,151,1401]" box="[199,1422,943,969]" pageId="3" pageNumber="142">
|
|||
|
|
<emphasis id="58C148094B0A2F22FF1C2AF3FEC8FC05" bold="true" box="[199,334,943,969]" pageId="3" pageNumber="142">Etymology.</emphasis>
|
|||
|
|
The adjective
|
|||
|
|
<taxonomicName id="ADB5EF984B0A2F22FE282AF3FDEDFC05" authorityName="Jałoszyński" authorityYear="2024" box="[499,619,943,969]" class="Insecta" family="Staphylinidae" genus="Cephennomicrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="142" phylum="Arthropoda" rank="species" species="tenebrosus">
|
|||
|
|
<emphasis id="58C148094B0A2F22FE282AF3FDEDFC05" box="[499,619,943,969]" italics="true" pageId="3" pageNumber="142">tenebrosus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(Latin: dark, gloomy) alludes to the unusually dark body pigmentation.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="22AFC7904B0A2F22FF1C2A8FFD01FAB5" pageId="3" pageNumber="142" type="discussion">
|
|||
|
|
<paragraph id="6A0A941B4B0A2F22FF1C2A8FFF55FBB1" blockId="3.[151,1437,151,1401]" pageId="3" pageNumber="142">
|
|||
|
|
<emphasis id="58C148094B0A2F22FF1C2A8FFEBDFC21" bold="true" box="[199,315,979,1005]" pageId="3" pageNumber="142">Remarks.</emphasis>
|
|||
|
|
Adults of this species are conspicuously darkly pigmented, but only the aedeagus offers unambiguous features for identification. Among Japanese species with similarly drop-shaped aedeagi, the one of
|
|||
|
|
<taxonomicName id="ADB5EF984B0A2F22FB312AABFA04FBDD" authorityName="Jałoszyński" authorityYear="2024" box="[1258,1410,1015,1041]" class="Insecta" family="Staphylinidae" genus="Cephennomicrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="142" phylum="Arthropoda" rank="species" species="tenebrosus">
|
|||
|
|
<emphasis id="58C148094B0A2F22FB312AABFA04FBDD" box="[1258,1410,1015,1041]" italics="true" pageId="3" pageNumber="142">C. tenebrosus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
is most conspicuous because of the complex symmetrical set of endophallic sclerites containing a sublateral bifurcate pair. Moreover, this is the only known
|
|||
|
|
<taxonomicName id="ADB5EF984B0A2F22FD972D63FC8CFB95" authorityName="Reitter" authorityYear="1907" box="[588,778,1087,1113]" class="Insecta" family="Staphylinidae" genus="Cephennomicrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="142" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="58C148094B0A2F22FD972D63FC8CFB95" box="[588,778,1087,1113]" italics="true" pageId="3" pageNumber="142">Cephennomicrus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
with lateral lobes flanking the apex of aedeagus in ventral view.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="6A0A941B4B0A2F22FF1C2DDBFC90FAFD" blockId="3.[151,1437,151,1401]" pageId="3" pageNumber="142">
|
|||
|
|
I have seen a female specimen of
|
|||
|
|
<taxonomicName id="ADB5EF984B0A2F22FDE92DDBFD76FB6D" authorityName="Reitter" authorityYear="1907" box="[562,752,1159,1185]" class="Insecta" family="Staphylinidae" genus="Cephennomicrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="142" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="58C148094B0A2F22FDE92DDBFD76FB6D" box="[562,752,1159,1185]" italics="true" pageId="3" pageNumber="142">Cephennomicrus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
collected on Amami Ôshima Island, which strongly resembles
|
|||
|
|
<taxonomicName id="ADB5EF984B0A2F22FF4C2DF7FEA9FB09" authorityName="Jałoszyński" authorityYear="2024" box="[151,303,1195,1221]" class="Insecta" family="Staphylinidae" genus="Cephennomicrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="142" phylum="Arthropoda" rank="species" species="tenebrosus">
|
|||
|
|
<emphasis id="58C148094B0A2F22FF4C2DF7FEA9FB09" box="[151,303,1195,1221]" italics="true" pageId="3" pageNumber="142">C. tenebrosus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
in a dark body pigmentation and body length and proportions. However, its antennae are less slender, especially the antennomere 11 is less than twice as long as wide. It cannot be excluded that the unusual elongation of antennae is a male dimorphic character, but without either females from
|
|||
|
|
<collectingRegion id="A8715AF94B0A2F22FC342DAFFBD2FAC1" box="[1007,1108,1267,1293]" country="Japan" name="Okinawa" pageId="3" pageNumber="142">Okinawa</collectingRegion>
|
|||
|
|
Is. or males from the distant Amami Ôshima it is not possible to verify this suspicion.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="6A0A941B4B0A2F22FF1C2C67FD01FAB5" blockId="3.[151,1437,151,1401]" pageId="3" pageNumber="142">
|
|||
|
|
Norihide Tokushige kindly provided further collecting details for the
|
|||
|
|
<typeStatus id="B50E2AB94B0A2F22FC142C67FBB3FA99" box="[975,1077,1339,1365]" pageId="3" pageNumber="142" type="holotype">holotype</typeStatus>
|
|||
|
|
: the specimen was found within the Nishihara Undôkôen Park under a stone.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|