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123 lines
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<document id="C5CE2E501CF0C2AFFF0886308BE569DB" ID-DOI="10.11646/zootaxa.4048.4.1" ID-GBIF-Dataset="bbb51c92-e184-489d-aba8-18d1dc9a25cd" ID-ISSN="1175-5326" ID-Zenodo-Dep="234023" ID-ZooBank="71E2B8DA-825E-429F-BAA0-F582702B4A80" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461348015003" checkinUser="plazi" docAuthor="López-García, Margarita M., Gasca-Álvarez, Héctor J. & Amat-García, Germán" docDate="2015" docId="990DA529FF94FF8DFF464683159CF9CB" docLanguage="en" docName="zt04048p492.pdf" docOrigin="Zootaxa 4048 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Euetheola Bates 1888" docType="treatment" docVersion="11" lastPageNumber="461" masterDocId="6534DD51FF9EFF87FFD1425F1659FFE6" masterDocTitle="The scarab beetle tribe Pentodontini (Coleoptera: Scarabaeidae: Dynastinae) of Colombia: taxonomy, natural history, and distribution" masterLastPageNumber="492" masterPageNumber="451" pageNumber="461" updateTime="1698621022009" updateUser="plazi">
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<mods:titleInfo id="DF32E8C980B94AEFA67CC7A22C58AD85">
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<mods:title id="4F333D91C7C6702006BD56D7C7E217D7">The scarab beetle tribe Pentodontini (Coleoptera: Scarabaeidae: Dynastinae) of Colombia: taxonomy, natural history, and distribution</mods:title>
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<mods:namePart id="CD54B546127A21BA1ABAA9B9BEAD4746">López-García, Margarita M.</mods:namePart>
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<mods:namePart id="A7995D7B2A22EABD0ED81F0C9D98A359">Gasca-Álvarez, Héctor J.</mods:namePart>
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<mods:namePart id="A13B455CE6ECAD6FEE7A70185EBF0FD0">Amat-García, Germán</mods:namePart>
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<mods:typeOfResource id="0A819ECC17992A5F4C147040F71E9343">text</mods:typeOfResource>
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<mods:title id="19A9FB88E0E46D4CC07F6A6B98DBFBFD">Zootaxa</mods:title>
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<mods:date id="65284C3B5CFCB5583371A7DB1D4E4CB0">2015</mods:date>
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<mods:detail id="DB214EC41C08E93BA657788376C32C1F" type="volume">
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<mods:number id="2676D2AC34F1C97115D19C9A59D7D257">4048</mods:number>
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<mods:number id="B91F2B8E15671BEB2CF14B1739C31478">4</mods:number>
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<mods:identifier id="A1E674C5DD5B1A94B09E5288C2881F5E" type="DOI">10.11646/zootaxa.4048.4.1</mods:identifier>
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<treatment id="990DA529FF94FF8DFF464683159CF9CB" ID-DOI="http://doi.org/10.5281/zenodo.5699348" ID-GBIF-Taxon="119645639" ID-Zenodo-Dep="5699348" LSID="urn:lsid:plazi:treatment:990DA529FF94FF8DFF464683159CF9CB" httpUri="http://treatment.plazi.org/id/990DA529FF94FF8DFF464683159CF9CB" lastPageNumber="461" pageId="10" pageNumber="461">
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<subSubSection id="59BE47B4FF94FF8DFF46468317F1FB10" box="[151,424,1244,1270]" pageId="10" pageNumber="461" type="nomenclature">
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<paragraph id="111B143FFF94FF8DFF46468317F1FB10" blockId="10.[151,424,1244,1270]" box="[151,424,1244,1270]" pageId="10" pageNumber="461">
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<heading id="4A53A353FF94FF8DFF46468317F1FB10" bold="true" box="[151,424,1244,1270]" fontSize="11" level="1" pageId="10" pageNumber="461" reason="1">
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<taxonomicName id="D6A46FBCFF94FF8DFF46468317F1FB10" authority="Bates, 1888" authorityName="Bates" authorityYear="1888" box="[151,424,1244,1270]" class="Insecta" family="Dynastidae" genus="Euetheola" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="461" phylum="Arthropoda" rank="genus">
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<emphasis id="23D0C82DFF94FF8DFF46468317F1FB10" bold="true" box="[151,424,1244,1270]" pageId="10" pageNumber="461">
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<emphasis id="23D0C82DFF94FF8DFF464683174AFB10" bold="true" box="[151,275,1244,1270]" italics="true" pageId="10" pageNumber="461">Euetheola</emphasis>
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<bibRefCitation id="753569CEFF94FF8DFECB468317F1FB10" author="Bates" box="[282,424,1244,1270]" pageId="10" pageNumber="461" refString="Bates, H. W. (1888) Pectinicornia and Lamellicornia, Family Dynastinae. In: Godman, F. D. & Salvin, O. (Eds), Biologia Centrali-Americana, Insecta, Coleoptera, Vol. II, Part 2, 296 - 342." type="book chapter" year="1888">Bates, 1888</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="59BE47B4FF94FF8DFF46474215A7FAD5" box="[151,1022,1309,1331]" pageId="10" pageNumber="461" type="reference_group">
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<paragraph id="111B143FFF94FF8DFF46474215A7FAD5" blockId="10.[151,1022,1309,1331]" box="[151,1022,1309,1331]" pageId="10" pageNumber="461">
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<treatmentCitationGroup id="31B43311FF94FF8DFF46474215A7FAD5" box="[151,1022,1309,1331]" pageId="10" pageNumber="461">
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<taxonomicName id="D6A46FBCFF94FF8DFF46474217F2FAD5" authority="Bates 1888: 314" authorityName="Bates" authorityPageNumber="314" authorityYear="1888" box="[151,427,1309,1331]" class="Insecta" family="Dynastidae" genus="Euetheola" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="461" phylum="Arthropoda" rank="genus">
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<emphasis id="23D0C82DFF94FF8DFF46474216A7FAD5" box="[151,254,1309,1331]" italics="true" pageId="10" pageNumber="461">Euetheola</emphasis>
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<treatmentCitation id="9005322EFF94FF8DFED5474117F2FAD5" author="Bates" box="[260,427,1309,1331]" page="314" pageId="10" pageNumber="461" year="1888">
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<bibRefCitation id="753569CEFF94FF8DFED547411721FAD5" author="Bates" box="[260,376,1309,1331]" pageId="10" pageNumber="461" refString="Bates, H. W. (1888) Pectinicornia and Lamellicornia, Family Dynastinae. In: Godman, F. D. & Salvin, O. (Eds), Biologia Centrali-Americana, Insecta, Coleoptera, Vol. II, Part 2, 296 - 342." type="book chapter" year="1888">Bates 1888</bibRefCitation>
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: 314
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</treatmentCitation>
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</taxonomicName>
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.
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<typeStatus id="CE1FAA9DFF94FF8DFE64474117BEFAD5" box="[437,487,1310,1331]" pageId="10" pageNumber="461">Type</typeStatus>
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species:
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<taxonomicName id="D6A46FBCFF94FF8DFD95474215A0FAD5" authority="Burmeister 1847: 93" authorityName="Burmeister" authorityPageNumber="93" authorityYear="1847" box="[580,1017,1309,1331]" class="Insecta" family="Dynastidae" genus="Heteronychus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="461" phylum="Arthropoda" rank="species" species="humilis">
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<emphasis id="23D0C82DFF94FF8DFD9547421578FAD5" box="[580,801,1309,1331]" italics="true" pageId="10" pageNumber="461">Heteronychus humilis</emphasis>
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<treatmentCitation id="9005322EFF94FF8DFCF6474215A0FAD5" author="Burmeister" box="[807,1017,1309,1331]" page="93" pageId="10" pageNumber="461" year="1847">
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<bibRefCitation id="753569CEFF94FF8DFCF64742158DFAD5" author="Burmeister" box="[807,980,1309,1331]" pageId="10" pageNumber="461" refString="Burmeister, H. (1847) Handbuch der Entomologie. Coleoptera Lamellicornia, Xylophila et Pectinicornia, Band 5. Enslin, Berlin, Germany." type="book" year="1847">Burmeister 1847</bibRefCitation>
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: 93
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</treatmentCitation>
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</taxonomicName>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="59BE47B4FF94FF8DFF4647001553FA03" pageId="10" pageNumber="461" type="diagnosis">
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<paragraph id="111B143FFF94FF8DFF4647001553FA03" blockId="10.[151,1437,1375,1581]" pageId="10" pageNumber="461">
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<emphasis id="23D0C82DFF94FF8DFF4647001748FA9E" bold="true" box="[151,273,1375,1400]" pageId="10" pageNumber="461">Diagnosis.</emphasis>
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The species of this genus are differentiated from other pentodontines by having the mandibles hidden under the clypeus and the pronotum without an apical fovea or tubercle. The species of
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<taxonomicName id="D6A46FBCFF94FF8DFBBF47DB12B9FA7B" box="[1134,1248,1412,1437]" class="Insecta" family="Dynastidae" genus="Euetheola" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="461" phylum="Arthropoda" rank="genus">
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<emphasis id="23D0C82DFF94FF8DFBBF47DB12B9FA7B" box="[1134,1248,1412,1437]" italics="true" pageId="10" pageNumber="461">Euetheola</emphasis>
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</taxonomicName>
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can be mistaken for some
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<taxonomicName id="D6A46FBCFF94FF8DFEDE47F717C8FA59" box="[271,401,1448,1471]" class="Insecta" family="Dynastidae" genus="Stenocrates" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="461" phylum="Arthropoda" rank="genus">
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<emphasis id="23D0C82DFF94FF8DFEDE47F717C8FA59" box="[271,401,1448,1471]" italics="true" pageId="10" pageNumber="461">Stenocrates</emphasis>
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</taxonomicName>
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species (
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<taxonomicName id="D6A46FBCFF94FF8DFDD747F814EAFA26" box="[518,691,1447,1472]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="461" phylum="Arthropoda" rank="tribe" tribe="Cyclocephalini">Cyclocephalini</taxonomicName>
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) by their general appearance and by having a frontal carina. However,
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<taxonomicName id="D6A46FBCFF94FF8DFEDB479217D5FA02" box="[266,396,1485,1508]" class="Insecta" family="Dynastidae" genus="Stenocrates" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="461" phylum="Arthropoda" rank="genus">
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<emphasis id="23D0C82DFF94FF8DFEDB479217D5FA02" box="[266,396,1485,1508]" italics="true" pageId="10" pageNumber="461">Stenocrates</emphasis>
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</taxonomicName>
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species do not have clypeal teeth.
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</paragraph>
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</subSubSection>
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<subSubSection id="59BE47B4FF94FF8DFF1647B0159CF9CB" pageId="10" pageNumber="461" type="distribution">
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<paragraph id="111B143FFF94FF8DFF1647B0159CF9CB" blockId="10.[151,1437,1375,1581]" pageId="10" pageNumber="461">
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<emphasis id="23D0C82DFF94FF8DFF1647B01739F9EE" bold="true" box="[199,352,1519,1544]" pageId="10" pageNumber="461">Distribution.</emphasis>
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<taxonomicName id="D6A46FBCFF94FF8DFEB647B01780F9EE" box="[359,473,1519,1544]" class="Insecta" family="Dynastidae" genus="Euetheola" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="461" phylum="Arthropoda" rank="genus">
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<emphasis id="23D0C82DFF94FF8DFEB647B01780F9EE" box="[359,473,1519,1544]" italics="true" pageId="10" pageNumber="461">Euetheola</emphasis>
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</taxonomicName>
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contains four species widely distributed in the New World, ranging from the southern
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<collectingCountry id="69B354AFFF94FF8DFF46444B17EBF9CB" box="[151,434,1556,1581]" name="United States of America" pageId="10" pageNumber="461">United States of America</collectingCountry>
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to South
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<collectingCountry id="69B354AFFF94FF8DFDCF444B14D8F9CB" box="[542,641,1556,1581]" name="United States of America" pageId="10" pageNumber="461">America</collectingCountry>
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, and two occur in
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<collectingCountry id="69B354AFFF94FF8DFC9E444B1598F9CB" box="[847,961,1556,1581]" name="Colombia" pageId="10" pageNumber="461">Colombia</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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</document>
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