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<document id="A3AC363FA3342EE831F7D6099301FF22" ID-CLB-Dataset="54546" ID-DOI="10.5852/ejt.2022.821.1799" ID-GBIF-Dataset="62057142-b60a-43c5-b7cd-947fc7615635" ID-ISSN="2118-9773" ID-Zenodo-Dep="6595468" ID-ZooBank="C85A43AA-6753-4851-8CEE-88AAE6DF069B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1653931612537" checkinUser="felipe" docAuthor="Ranasinghe, Sasanka, Eberle, Jonas, Athukorala, Namal, Benjamin, Suresh P. &amp; Ahrens, Dirk" docDate="2022" docId="03A99E29FF873077FD4CE8624D2C112F" docLanguage="en" docName="ejt-821_57-101.pdf" docOrigin="European Journal of Taxonomy 821 (1)" docSource="http://zoobank.org/c85a43aa-6753-4851-8cee-88aae6df069b" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Maladera haniel Ranasinghe &amp; Eberle &amp; Athukorala &amp; Benjamin &amp; Ahrens 2022, sp. nov." docType="treatment" docUuid="B68147DB-D51A-4D08-8E56-10D1C194889A" docUuidSource="ZooBank" docVersion="9" lastPageNumber="72" masterDocId="FF90E651FF8B3078FFC8EC494C7C155C" masterDocTitle="New species of Sericini from Sri Lanka (Coleoptera, Scarabaeidae). Part II" masterLastPageNumber="101" masterPageNumber="57" pageNumber="69" updateTime="1699338309283" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="499FA78BF3EACD5FD441C9ED566EC68D">New species of Sericini from Sri Lanka (Coleoptera, Scarabaeidae). Part II</mods:title>
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<mods:namePart id="EF0781741A6B682E46772688B7B4A818">Ranasinghe, Sasanka</mods:namePart>
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<mods:affiliation id="04A17EA35B461641BFE728A61DDC23D9">Zoologisches Forschungsmuseum A. Koenig, Adenauerallee 127, 53113 Bonn, Germany.</mods:affiliation>
<mods:nameIdentifier id="BDFAC04293A3F47CA3A79A77061322A3" type="email">lakmalisanky@gmail.com</mods:nameIdentifier>
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<mods:namePart id="8B4405590047B0DA8A4AE8B4D4309095">Eberle, Jonas</mods:namePart>
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<mods:affiliation id="C209BBCDF8451C9C8C15FA3BB595B00A">Zoologisches Forschungsmuseum A. Koenig, Adenauerallee 127, 53113 Bonn, Germany.</mods:affiliation>
<mods:nameIdentifier id="A57793A874443E7B4878D4AB0742C431" type="email">eberle.jonas@gmail.com</mods:nameIdentifier>
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<mods:namePart id="32F9173AF3E457025C6DFE51D15AE8B8">Athukorala, Namal</mods:namePart>
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<mods:affiliation id="5BBD9D29506A1750F32C14B73762328B">National Institute of Fundamental Studies, Hantana Road, 20000 Kandy, Sri Lanka.</mods:affiliation>
<mods:nameIdentifier id="763FF8D87166D5D86D93AB3FF11A7FD6" type="email">namal_athu@yahoo.com</mods:nameIdentifier>
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<mods:namePart id="93D5AFACDAB888BA82C2E7870821889F">Benjamin, Suresh P.</mods:namePart>
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<mods:affiliation id="85B4D1DA58CDA0EE9FAF7F07C09C1C25">Zoologisches Forschungsmuseum A. Koenig, Adenauerallee 127, 53113 Bonn, Germany. &amp; National Institute of Fundamental Studies, Hantana Road, 20000 Kandy, Sri Lanka.</mods:affiliation>
<mods:nameIdentifier id="FDEE76FDEACEC131A3C81D8BF0E89174" type="email">suresh.benjamin@gmail.com</mods:nameIdentifier>
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<mods:namePart id="D17854295050A69917E9E620E5145F8F">Ahrens, Dirk</mods:namePart>
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<mods:affiliation id="34F789A3EC893F1AE563676AEF363E77">Zoologisches Forschungsmuseum A. Koenig, Adenauerallee 127, 53113 Bonn, Germany.</mods:affiliation>
<mods:nameIdentifier id="757E6316205B93995D7051902A7B3653" type="email">ahrens.dirk_col@gmx.de,d.ahrens@leibniz-zfmk.de</mods:nameIdentifier>
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<mods:date id="0869B9BBEC2FBADA393DCD1D965012AA">2022</mods:date>
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<treatment id="03A99E29FF873077FD4CE8624D2C112F" ID-DOI="http://doi.org/10.5281/zenodo.7473426" ID-GBIF-Taxon="207927103" ID-Zenodo-Dep="7473426" LSID="urn:lsid:zoobank.org:act:B68147DB-D51A-4D08-8E56-10D1C194889A" httpUri="http://treatment.plazi.org/id/03A99E29FF873077FD4CE8624D2C112F" lastPageId="15" lastPageNumber="72" pageId="12" pageNumber="69">
<subSubSection id="C31A7CB4FF873074FD4CE8624FCC111A" box="[644,944,1067,1094]" pageId="12" pageNumber="69" type="nomenclature">
<paragraph id="8BBF2F3FFF873074FD4CE8624FCC111A" blockId="12.[365,1224,1067,1164]" box="[644,944,1067,1094]" pageId="12" pageNumber="69">
<heading id="D0F79853FF873074FD4CE8624FCC111A" box="[644,944,1067,1094]" centered="true" fontSize="11" level="2" pageId="12" pageNumber="69" reason="2">
<taxonomicName id="4C0054BCFF873074FD4CE8624F32111A" authority="Ranasinghe &amp; Eberle &amp; Athukorala &amp; Benjamin &amp; Ahrens, 2022" authorityName="Ranasinghe &amp; Eberle &amp; Athukorala &amp; Benjamin &amp; Ahrens" authorityYear="2022" box="[644,846,1067,1094]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="69" phylum="Arthropoda" rank="species" species="haniel" status="sp. nov.">
<emphasis id="B974F32DFF873074FD4CE8624F32111A" bold="true" box="[644,846,1067,1094]" italics="true" pageId="12" pageNumber="69">Maladera haniel</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2474E56FF873074FC9DE8654FCC111A" box="[853,944,1068,1094]" pageId="12" pageNumber="69" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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<subSubSection id="C31A7CB4FF873074FEA5E8074FF111D0" pageId="12" pageNumber="69" type="description">
<paragraph id="8BBF2F3FFF873074FEA5E80748B41135" blockId="12.[365,1224,1067,1164]" box="[365,1224,1102,1129]" pageId="12" pageNumber="69">
<uri id="FF91233DFF873074FEA5E80748B41135" box="[365,1224,1102,1129]" pageId="12" pageNumber="69">
urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
<paragraph id="8BBF2F3FFF873074FD6EE83B4FF111D0" blockId="12.[365,1224,1067,1164]" box="[678,909,1138,1164]" pageId="12" pageNumber="69">
<figureCitation id="133B33BAFF873074FD6EE83B4F5711D0" box="[678,811,1138,1164]" captionStart="Fig" captionStartId="14.[189,232,1797,1823]" captionTargetBox="[204,1387,285,1731]" captionTargetId="figure-15@14.[189,1398,265,1758]" captionTargetPageId="14" captionText="Fig. 4. A–D. Maladera haniel sp. nov., holotype, ♂ (ZFMK). E–H. M. kishi sp. nov., holotype, ♂ (ZFMK). I–L. M. windy sp. nov., holotype, ♂ (ZFMK). A, E, I. Aedeagus, left side lateral view. C, G, K. Aedeagus, right side lateral view. B, F, J. Parameres, dorsal view. D, H, L. Habitus (not to scale). Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6595477" httpUri="https://zenodo.org/record/6595477/files/figure.png" pageId="12" pageNumber="69">Figs 4A–D</figureCitation>
,
<figureCitation id="133B33BAFF873074FCFFE83B4F2111D0" box="[823,861,1138,1164]" captionStart="Fig" captionStartId="29.[189,232,1917,1943]" captionTargetBox="[189,1398,1076,1874]" captionTargetId="figure-244@29.[189,1398,1076,1874]" captionTargetPageId="29" captionText="Fig. 7. Distribution of ten new species. A. Selaserica fabriziae sp. nov. B. Sel. sororinitida sp. nov. C. Neoserica pophami sp. nov. D. Maladera haniel sp. nov. E. M. kishi sp. nov. F. M. windy sp. nov. G. M. karunaratnae sp. nov. H. M. hiyarensis sp. nov. I. M. dambullana sp. nov. J. M. deenstana sp. nov." figureDoi="http://doi.org/10.5281/zenodo.6595483" httpUri="https://zenodo.org/record/6595483/files/figure.png" pageId="12" pageNumber="69">7D</figureCitation>
,
<figureCitation id="133B33BAFF873074FCA1E83B4FF111D0" box="[873,909,1138,1164]" captionStart="Fig" captionStartId="30.[189,232,1885,1911]" captionTargetBox="[235,1353,265,1847]" captionTargetId="figure-16@30.[235,1353,265,1847]" captionTargetPageId="30" captionText="Fig. 8. Photographs of the habitats of the new species.A. Selaserica fabriziae sp. nov. B. Sel. sororinitida sp. nov. C. Neoserica pophami sp. nov. D. Maladera haniel sp. nov. and M. kishi sp. nov. E. M. windy sp. nov. F. M. karunaratnae sp. nov. G. M. hiyarensis sp. nov.H. M. dambullana sp. nov. I. M. deenstana sp. nov." figureDoi="http://doi.org/10.5281/zenodo.6595485" httpUri="https://zenodo.org/record/6595485/files/figure.png" pageId="12" pageNumber="69">8D</figureCitation>
</paragraph>
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<subSubSection id="C31A7CB4FF873074FF75E8FA4F9B1039" pageId="12" pageNumber="69" type="diagnosis">
<paragraph id="8BBF2F3FFF873074FF75E8FA4D4A1191" blockId="12.[189,310,1203,1229]" box="[189,310,1203,1229]" pageId="12" pageNumber="69">
<heading id="D0F79853FF873074FF75E8FA4D4A1191" bold="true" box="[189,310,1203,1229]" fontSize="11" level="3" pageId="12" pageNumber="69" reason="3">
<emphasis id="B974F32DFF873074FF75E8FA4D4A1191" bold="true" box="[189,310,1203,1229]" pageId="12" pageNumber="69">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BBF2F3FFF873074FF75E8AB4F9B1039" blockId="12.[188,1401,1250,1382]" pageId="12" pageNumber="69">
<taxonomicName id="4C0054BCFF873074FF75E8AB4DFD11A0" authority="Ranasinghe &amp; Eberle &amp; Athukorala &amp; Benjamin &amp; Ahrens, 2022" authorityName="Ranasinghe &amp; Eberle &amp; Athukorala &amp; Benjamin &amp; Ahrens" authorityYear="2022" box="[189,385,1250,1276]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="69" phylum="Arthropoda" rank="species" species="haniel" status="sp. nov.">
<emphasis id="B974F32DFF873074FF75E8AB4DFD11A0" box="[189,385,1250,1276]" italics="true" pageId="12" pageNumber="69">Maladera haniel</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2474E56FF873074FE42E8AB4D9911A0" box="[394,485,1250,1276]" pageId="12" pageNumber="69" rank="species">sp. nov.</taxonomicNameLabel>
is in external morphology very similar to
<taxonomicName id="4C0054BCFF873074FC10E8AA4D801043" authority="Ranasinghe, Eberle, Benjamin &amp; Ahrens, 2020" authorityName="Ranasinghe, Eberle, Benjamin &amp; Ahrens" authorityYear="2020" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="69" phylum="Arthropoda" rank="species" species="cervicornis">
<emphasis id="B974F32DFF873074FC10E8AA48FF11A0" box="[984,1155,1251,1276]" italics="true" pageId="12" pageNumber="69">M. cervicornis</emphasis>
<bibRefCitation id="EF9152CEFF873074FB43E8AB4D801043" author="Ranasinghe S. &amp; Eberle J. &amp; Benjamin S. P. &amp; Ahrens D." pageId="12" pageNumber="69" pagination="1 - 20" refId="ref31865" refString="Ranasinghe S., Eberle J., Benjamin S. P. &amp; Ahrens D. 2020. New species of Sericini from Sri Lanka (Coleoptera, Scarabaeidae). European Journal of Taxonomy 621: 1 - 20. https: // doi. org / 10.5852 / ejt. 2020.621" type="journal article" year="2020">Ranasinghe, Eberle, Benjamin &amp; Ahrens, 2020</bibRefCitation>
</taxonomicName>
, both having in common the tubercle on abdominal sternite III.
<taxonomicName id="4C0054BCFF873074FACDE94C4D7A101E" authority="Ranasinghe &amp; Eberle &amp; Athukorala &amp; Benjamin &amp; Ahrens, 2022" authorityName="Ranasinghe &amp; Eberle &amp; Athukorala &amp; Benjamin &amp; Ahrens" authorityYear="2022" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="69" phylum="Arthropoda" rank="species" species="haniel" status="sp. nov.">
<emphasis id="B974F32DFF873074FACDE94C4D7A101E" italics="true" pageId="12" pageNumber="69">Maladera haniel</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2474E56FF873074FEC5E9604D1A101F" box="[269,358,1321,1347]" pageId="12" pageNumber="69" rank="species">sp. nov.</taxonomicNameLabel>
differs in the setose pronotum, as well as the shape of parameres: the fused parameres are extremely long, as long as the rest of the basal part of the phallobase.
</paragraph>
</subSubSection>
<subSubSection id="C31A7CB4FF873074FF75E9C44960108A" box="[189,1308,1421,1494]" pageId="12" pageNumber="69" type="etymology">
<paragraph id="8BBF2F3FFF873074FF75E9C44D3810FB" blockId="12.[189,324,1421,1447]" box="[189,324,1421,1447]" pageId="12" pageNumber="69">
<heading id="D0F79853FF873074FF75E9C44D3810FB" bold="true" box="[189,324,1421,1447]" fontSize="11" level="3" pageId="12" pageNumber="69" reason="3">
<emphasis id="B974F32DFF873074FF75E9C44D3810FB" bold="true" box="[189,324,1421,1447]" pageId="12" pageNumber="69">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BBF2F3FFF873074FF75E9F24960108A" blockId="12.[189,1308,1467,1494]" box="[189,1308,1467,1494]" pageId="12" pageNumber="69">The new species is named for Suresh Benjamin’s son, Haniel P. Benjamin (noun in apposition).</paragraph>
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<paragraph id="8BBF2F3FFF873074FF75E9B44D10134B" blockId="12.[189,364,1533,1559]" box="[189,364,1533,1559]" pageId="12" pageNumber="69">
<heading id="D0F79853FF873074FF75E9B44D10134B" bold="true" box="[189,364,1533,1559]" fontSize="11" level="3" pageId="12" pageNumber="69" reason="3">
<emphasis id="B974F32DFF873074FF75E9B44D10134B" bold="true" box="[189,364,1533,1559]" pageId="12" pageNumber="69">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="8BBF2F3FFF873074FF22EA6248D713D0" blockId="12.[189,1398,1579,1676]" pageId="12" pageNumber="69">
<materialsCitation id="3B682562FF873074FF22EA6248D713D0" ID-GBIF-Occurrence="4003031429" collectingDate="2019-10-18" collectingMethod="Black light" collectionCode="ZFMK" collectorName="Eberle &amp; Bohacz &amp; Ranasinghe" country="Sri Lanka" county="Knuckles South" elevation="1108" latitude="7.33082" location="Sri Lanka" longLatPrecision="1" longitude="80.86203" municipality="Deenston" pageId="12" pageNumber="69" specimenCode="X-SR0251" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B974F32DFF873074FF22EA624D201319" bold="true" box="[234,348,1579,1605]" pageId="12" pageNumber="69">
<typeStatus id="54BB919DFF873074FF22EA624D201319" box="[234,348,1579,1605]" pageId="12" pageNumber="69" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="F3176FAFFF873074FF75EA064D201335" box="[189,348,1615,1641]" name="Sri Lanka" pageId="12" pageNumber="69">SRI LANKA</collectingCountry>
•
<specimenCount id="9D06E4B6FF873074FE48EA064DE51335" box="[384,409,1615,1641]" pageId="12" pageNumber="69" type="male">♂</specimenCount>
; “
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,
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•
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•
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•
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<paragraph id="8BBF2F3FFF863075FF75EE60485B171F" blockId="13.[189,1063,553,580]" box="[189,1063,553,580]" pageId="13" pageNumber="70">
MEASUREMENTS. Length: 8.0 mm, length of elytra:
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, width:
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HABITUS (
<figureCitation id="133B33BAFF863075FEE6EE234DF517D8" box="[302,393,618,645]" captionStart="Fig" captionStartId="14.[189,232,1797,1823]" captionTargetBox="[204,1387,285,1731]" captionTargetId="figure-15@14.[189,1398,265,1758]" captionTargetPageId="14" captionText="Fig. 4. A–D. Maladera haniel sp. nov., holotype, ♂ (ZFMK). E–H. M. kishi sp. nov., holotype, ♂ (ZFMK). I–L. M. windy sp. nov., holotype, ♂ (ZFMK). A, E, I. Aedeagus, left side lateral view. C, G, K. Aedeagus, right side lateral view. B, F, J. Parameres, dorsal view. D, H, L. Habitus (not to scale). Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6595477" httpUri="https://zenodo.org/record/6595477/files/figure.png" pageId="13" pageNumber="70">Fig. 4D</figureCitation>
) AND COLOURATION. Body short oval, dark brown, antenna yellow, dorsal surface shiny, finely densely setose.
</paragraph>
<paragraph id="8BBF2F3FFF863075FF75EE99490B117A" blockId="13.[189,1401,719,1062]" pageId="13" pageNumber="70">HEAD. Labroclypeus short and trapezoidal, wider than long, widest at base, lateral margins strongly convex and convergent to widely rounded anterior angles, lateral border and ocular canthus producing an indistinct blunt angle, margins weakly reflexed, anterior margin almost weakly sinuate medially; surface slightly convex, finely and densely punctate, distance between punctures smaller than their diameter, with numerous erect setae in larger punctures; frontoclypeal suture indistinctly incised and bluntly bent medially; smooth area in front of eye approximately three times as wide as long; ocular canthus long and narrow, minutely and superficially punctate, with a single short terminal seta. Frons with fine, dense punctures, with a few long erect setae in larger punctures, setae on disc less dense. Eyes large, ratio of diameter/interocular width: 0.79. Antenna yellow, with ten antennomeres; club with three antennomeres, as long as remaining antennomeres combined. Mentum elevated and anteriorly flattened.</paragraph>
<paragraph id="8BBF2F3FFF863075FF75E8074F791067" blockId="13.[188,1401,1101,1339]" pageId="13" pageNumber="70">PRONOTUM. Wide, widest at base, lateral margins in basal half straight and weakly convergent, in anterior half weakly convex and narrowed to anterior angles, anterior angles moderately produced and sharp, anterior marginal line fine and complete, anterior margin convexly produced medially; surface finely densely punctate, with moderately dense, short and fine setae and dense long erect setae being directed anteriorly; anterior and lateral borders sparsely setose, basal margin without marginal line; hypomeron ventrobasally carinate and slightly produced ventrally. Scutellum short and wide, triangular, with fine and dense punctures, with short, dense, fine setae.</paragraph>
<paragraph id="8BBF2F3FFF863075FF75E92B4DC21355" blockId="13.[188,1399,1378,1545]" pageId="13" pageNumber="70">ELYTRA. Short oval, widest shortly behind middle, striae distinctly impressed, finely and densely punctate, intervals weakly convex, with moderately fine, dense punctures and with dense, fine, short setae as well as with sparse long erect setae; epipleural edge fine, ending at the weakly convex external apical angle of elytra, epipleura densely setose; apical border narrowly membranous, apex covered with short microtrichomes.</paragraph>
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VENTRAL SURFACE. Shiny, thorax and metacoxa with large and dense punctures, sparsely setose, metacoxa with minute adjacent setae in the punctures except for numerous long setae laterally, apical margin weakly convex, without a wide rim of long white microtrichomes; each abdominal sternite, in addition to evenly distributed fine and dense punctures bearing each a fine seta, with a distinct transverse row of coarse punctures each bearing a long and more robust seta, 3
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sternite with a sharp median tubercle being half as high as sternite length, penultimate sternite apically with a shiny smooth chitinous border, which is a quarter as long as the sternite. Mesosternum between mesocoxae as wide as mesofemur, with a semi-circular ridge bearing robust setae. Ratio of length of metepisternum/metacoxa: 1/1.93. Pygidium moderately convex, moderately finely and densely punctate, without smooth midline, punctures with short and dense, adjacent setae as well as with moderately dense, long, erect setae.
</paragraph>
<caption id="DF7F7FB7FF853076FF75EB4C4DCB12D5" ID-DOI="http://doi.org/10.5281/zenodo.6595477" ID-Zenodo-Dep="6595477" httpUri="https://zenodo.org/record/6595477/files/figure.png" pageId="14" pageNumber="71" startId="14.[189,232,1797,1823]" targetBox="[204,1387,285,1731]" targetPageId="14">
<paragraph id="8BBF2F3FFF853076FF75EB4C4DCB12D5" blockId="14.[189,1399,1797,1930]" pageId="14" pageNumber="71">
<emphasis id="B974F32DFF853076FF75EB4C4D281243" bold="true" box="[189,340,1797,1823]" pageId="14" pageNumber="71">Fig. 4. A–D</emphasis>
.
<taxonomicName id="4C0054BCFF853076FEAEEB4C4E521243" authority="Ranasinghe &amp; Eberle &amp; Athukorala &amp; Benjamin &amp; Ahrens, 2022" authorityName="Ranasinghe &amp; Eberle &amp; Athukorala &amp; Benjamin &amp; Ahrens" authorityYear="2022" box="[358,558,1797,1823]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="71" phylum="Arthropoda" rank="species" species="haniel" status="sp. nov.">
<emphasis id="B974F32DFF853076FEAEEB4C4E521243" box="[358,558,1797,1823]" italics="true" pageId="14" pageNumber="71">Maladera haniel</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2474E56FF853076FDF2EB4F4EE7127C" box="[570,667,1798,1824]" pageId="14" pageNumber="71" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (ZFMK).
<emphasis id="B974F32DFF853076FC75EB4C4F8A1243" bold="true" box="[957,1014,1797,1823]" pageId="14" pageNumber="71">E–H</emphasis>
.
<taxonomicName id="4C0054BCFF853076FBC1EB4F48171243" authority="Ranasinghe &amp; Eberle &amp; Athukorala &amp; Benjamin &amp; Ahrens, 2022" authorityName="Ranasinghe &amp; Eberle &amp; Athukorala &amp; Benjamin &amp; Ahrens" authorityYear="2022" box="[1033,1131,1797,1823]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="71" phylum="Arthropoda" rank="species" species="kishi" status="sp. nov.">
<emphasis id="B974F32DFF853076FBC1EB4F48171243" box="[1033,1131,1797,1823]" italics="true" pageId="14" pageNumber="71">M. kishi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2474E56FF853076FBBEEB4F48AB127C" box="[1142,1239,1798,1824]" pageId="14" pageNumber="71" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (ZFMK).
<emphasis id="B974F32DFF853076FEF9EB604D22121F" bold="true" box="[305,350,1833,1859]" pageId="14" pageNumber="71">I–L</emphasis>
.
<taxonomicName id="4C0054BCFF853076FEA6EB604DA7121E" authority="Ranasinghe &amp; Eberle &amp; Athukorala &amp; Benjamin &amp; Ahrens, 2022" authorityName="Ranasinghe &amp; Eberle &amp; Athukorala &amp; Benjamin &amp; Ahrens" authorityYear="2022" box="[366,475,1832,1858]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="71" phylum="Arthropoda" rank="species" species="windy" status="sp. nov.">
<emphasis id="B974F32DFF853076FEA6EB604DA7121E" box="[366,475,1832,1858]" italics="true" pageId="14" pageNumber="71">M. windy</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2474E56FF853076FE2BEB604E3C121F" box="[483,576,1833,1859]" pageId="14" pageNumber="71" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (ZFMK).
<emphasis id="B974F32DFF853076FC9CEB614FD9121F" bold="true" box="[852,933,1832,1859]" pageId="14" pageNumber="71">A, E, I</emphasis>
. Aedeagus, left side lateral view.
<emphasis id="B974F32DFF853076FAFCEB614CAF123A" bold="true" pageId="14" pageNumber="71">C, G, K</emphasis>
. Aedeagus, right side lateral view.
<emphasis id="B974F32DFF853076FDB2EB054EB7123A" bold="true" box="[634,715,1868,1894]" pageId="14" pageNumber="71">B, F, J</emphasis>
. Parameres, dorsal view.
<emphasis id="B974F32DFF853076FBC8EB054822123A" bold="true" box="[1024,1118,1868,1894]" pageId="14" pageNumber="71">D, H, L</emphasis>
. Habitus (not to scale). Scale bars = 0.5 mm.
</paragraph>
</caption>
<paragraph id="8BBF2F3FFF843077FF75ED404F4117B0" blockId="15.[188,1399,265,748]" pageId="15" pageNumber="72">LEGS. Short and wide; femur with two longitudinal rows of setae, finely and densely punctate; metafemur shiny, anterior edge acute, lacking an adjacent serrated line, ventral surface densely punctate and setose, posterior ventral margin straight, only little widened in apical half, finely serrate apically, dorsally not serrated, glabrous. Metatibia short, widest at middle, posteriorly only very little narrowed, ratio width/ length: 1/2.5, dorsally sharply carinate, with two groups of spines, basal shortly behind middle, apical one at three quarters of metatibial length, basally beside dorsal margin with two single punctures with serrated margins, each bearing a single robust spine and beside them a longitudinal serrated line; lateral face almost flat, with dense, large punctures and with minute setae in the punctures; ventral margin with five strong spines equidistant from each other, medial face smooth, apex interiorly near tarsal articulation shallowly concave. Meso- and metatarsomeres finely and sparsely punctate but glabrous dorsally, ventrally with sparse, short setae; metatarsomeres ventrally with a strongly serrated ridge, beside which is a strong longitudinal carina; first metatarsomere as long as following two tarsomeres combined and as long as dorsal tibial spur. Protibia short, bidentate. All claws symmetrical, feebly curved and long, with normally developed basal tooth.</paragraph>
<paragraph id="8BBF2F3FFF843077FF75EF5D4DBA1672" blockId="15.[189,454,788,815]" box="[189,454,788,815]" pageId="15" pageNumber="72">
AEDEAGUS.
<figureCitation id="133B33BAFF843077FE8AEF5D4DBE1672" box="[322,450,788,814]" captionStart="Fig" captionStartId="14.[189,232,1797,1823]" captionTargetBox="[204,1387,285,1731]" captionTargetId="figure-15@14.[189,1398,265,1758]" captionTargetPageId="14" captionText="Fig. 4. A–D. Maladera haniel sp. nov., holotype, ♂ (ZFMK). E–H. M. kishi sp. nov., holotype, ♂ (ZFMK). I–L. M. windy sp. nov., holotype, ♂ (ZFMK). A, E, I. Aedeagus, left side lateral view. C, G, K. Aedeagus, right side lateral view. B, F, J. Parameres, dorsal view. D, H, L. Habitus (not to scale). Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6595477" httpUri="https://zenodo.org/record/6595477/files/figure.png" pageId="15" pageNumber="72">Fig. 4A–C</figureCitation>
.
</paragraph>
<paragraph id="8BBF2F3FFF843077FF75EF1C4D4F1633" blockId="15.[189,307,853,879]" box="[189,307,853,879]" pageId="15" pageNumber="72">
<heading id="D0F79853FF843077FF75EF1C4D4F1633" bold="true" box="[189,307,853,879]" fontSize="11" level="3" pageId="15" pageNumber="72" reason="3">
<emphasis id="B974F32DFF843077FF75EF1C4D4F1633" bold="true" box="[189,307,853,879]" pageId="15" pageNumber="72">Variation</emphasis>
</heading>
</paragraph>
<paragraph id="8BBF2F3FFF843077FF75EFCD487D16C3" blockId="15.[189,1025,900,927]" box="[189,1025,900,927]" pageId="15" pageNumber="72">
Length: 8.0–
<quantity id="4CF882DAFF843077FE9BEFCD4DCC16C3" box="[339,432,900,927]" metricMagnitude="-3" metricUnit="m" metricValue="8.2" pageId="15" pageNumber="72" unit="mm" value="8.2">8.2 mm</quantity>
, length of elytra:
<quantity id="4CF882DAFF843077FDB6EFCD4F7216C3" box="[638,782,900,927]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" metricValueMax="6.1" metricValueMin="5.5" pageId="15" pageNumber="72" unit="mm" value="5.8" valueMax="6.1" valueMin="5.5">5.5–6.1 mm</quantity>
, width:
<quantity id="4CF882DAFF843077FCA4EFCD4F8016C3" box="[876,1020,900,927]" metricMagnitude="-3" metricUnit="m" metricValue="4.55" metricValueMax="4.8" metricValueMin="4.3" pageId="15" pageNumber="72" unit="mm" value="4.55" valueMax="4.8" valueMin="4.3">4.3–4.8 mm</quantity>
.
</paragraph>
<paragraph id="8BBF2F3FFF843077FF22EF8F4D3916BC" blockId="15.[189,325,966,1027]" box="[234,325,966,992]" pageId="15" pageNumber="72">
<emphasis id="B974F32DFF843077FF22EF8F4D3916BC" bold="true" box="[234,325,966,992]" pageId="15" pageNumber="72">Female</emphasis>
</paragraph>
<paragraph id="8BBF2F3FFF843077FF75EFA04D45115F" blockId="15.[189,325,966,1027]" box="[189,313,1001,1027]" pageId="15" pageNumber="72">Unknown.</paragraph>
</subSubSection>
<subSubSection id="C31A7CB4FF843077FF75E8634D2C112F" box="[189,342,1066,1139]" pageId="15" pageNumber="72" type="distribution">
<paragraph id="8BBF2F3FFF843077FF75E8634D2A1118" blockId="15.[189,342,1066,1092]" box="[189,342,1066,1092]" pageId="15" pageNumber="72">
<heading id="D0F79853FF843077FF75E8634D2A1118" bold="true" box="[189,342,1066,1092]" fontSize="11" level="3" pageId="15" pageNumber="72" reason="3">
<emphasis id="B974F32DFF843077FF75E8634D2A1118" bold="true" box="[189,342,1066,1092]" pageId="15" pageNumber="72">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BBF2F3FFF843077FF75E8104D2C112F" blockId="15.[189,336,1112,1139]" box="[189,336,1112,1139]" pageId="15" pageNumber="72">
See
<figureCitation id="133B33BAFF843077FF27E8114D30112F" box="[239,332,1112,1139]" captionStart="Fig" captionStartId="29.[189,232,1917,1943]" captionTargetBox="[189,1398,1076,1874]" captionTargetId="figure-244@29.[189,1398,1076,1874]" captionTargetPageId="29" captionText="Fig. 7. Distribution of ten new species. A. Selaserica fabriziae sp. nov. B. Sel. sororinitida sp. nov. C. Neoserica pophami sp. nov. D. Maladera haniel sp. nov. E. M. kishi sp. nov. F. M. windy sp. nov. G. M. karunaratnae sp. nov. H. M. hiyarensis sp. nov. I. M. dambullana sp. nov. J. M. deenstana sp. nov." figureDoi="http://doi.org/10.5281/zenodo.6595483" httpUri="https://zenodo.org/record/6595483/files/figure.png" pageId="15" pageNumber="72">Fig. 7D</figureCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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