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148 lines
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<document ID-DOI="http://dx.doi.org/10.3897/fr.25.83781" ID-Pensoft-Pub="2193-0074-1-99" ID-Pensoft-UUID="2F5D6AE2EEB55A17ACF1623B06B4EA8D" ID-ZooBank="7A16698D4F1848D29D9651A6E0CC15AC" ModsDocID="2193-0074-25-1-99" checkinTime="1667366289781" checkinUser="pensoft" docAuthor="Vasilyan, Davit, Cernansky, Andrej, Szyndlar, Zbigniew & Moers, Thomas" docDate="2022" docId="FC89A98367C45B4798C21B729E4C7F14" docLanguage="en" docName="FossilRecord 25(1): 99-145" docOrigin="Fossil Record 25 (1)" docPubDate="2022-05-10" docSource="http://dx.doi.org/10.3897/fr.25.83781" docTitle="Chelotriton indet." docType="treatment" docVersion="1" id="2F5D6AE2EEB55A17ACF1623B06B4EA8D" lastPageNumber="99" masterDocId="2F5D6AE2EEB55A17ACF1623B06B4EA8D" masterDocTitle="Amphibian and reptilian fauna from the early Miocene of Echzell, Germany" masterLastPageNumber="145" masterPageNumber="99" pageNumber="99" updateTime="1667366289781" updateUser="pensoft">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Amphibian and reptilian fauna from the early Miocene of Echzell, Germany</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Vasilyan, Davit</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-8712-0678</mods:nameIdentifier>
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<mods:affiliation>JURASSICA Museum, Route de Fontenais 21. 2900 Porrentruy, Switzerland & Department of Geosciences, University of Fribourg, Chemin du musee 6, 1700, Fribourg, Switzerland</mods:affiliation>
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<mods:nameIdentifier type="email">davit.vasilyan@jurassica.ch</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Cernansky, Andrej</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-8920-2503</mods:nameIdentifier>
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<mods:affiliation>Department of Ecology, Laboratory of Evolutionary Biology, Faculty of Natural Sciences, Comenius University in Bratislava, Mlynska dolina, Ilkovicova 6, 842 15 Bratislava, Slovakia</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Szyndlar, Zbigniew</mods:namePart>
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<mods:affiliation>Institute of Systematics and Evolution of Animals, Polish Academy of Sciences, Slawkowska 17, 31 - 016 Krakow, Poland</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Moers, Thomas</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-2268-5824</mods:nameIdentifier>
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<mods:affiliation>Department of Palaeobiology, Swedish Museum of Natural History, P. O. Box 50007, 104 05 Stockholm, Sweden & Bolin Centre for Climate Research, Stockholm University, Stockholm, Sweden</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Fossil Record</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-05-10</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>25</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>1</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>99</mods:start>
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<mods:end>145</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/fr.25.83781</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/fr.25.83781</mods:identifier>
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<mods:identifier type="Pensoft-Pub">2193-0074-1-99</mods:identifier>
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<mods:identifier type="ZooBank">7A16698D4F1848D29D9651A6E0CC15AC</mods:identifier>
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<mods:identifier type="Pensoft-UUID">2F5D6AE2EEB55A17ACF1623B06B4EA8D</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:FC89A98367C45B4798C21B729E4C7F14" httpUri="http://treatment.plazi.org/id/FC89A98367C45B4798C21B729E4C7F14" lastPageNumber="99" pageId="0" pageNumber="99">
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<subSubSection pageId="0" pageNumber="99" type="nomenclature">
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<paragraph pageId="0" pageNumber="99">
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<taxonomicName LSID="FC89A983-67C4-5B47-98C2-1B729E4C7F14" authority="indet." authorityName="indet." family="Salamandridae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="" order="Caudata" pageId="0" pageNumber="99" rank="family">Salamandridae indet.</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="99" type="description">
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<paragraph pageId="0" pageNumber="99">
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Vertebrae (A-E. HLDM-Ez 2059; F-J. HLDM-Ez 2060) and ribs (K. HLDM-Ez 2054; L. HLDM-Ez 2053; M. HLDM-Ez 2055) of Chelotriton sp. Frontal (N, O. HLDM-Ez 2038) and ribs (P, Q. HLDM-Ez 2036; R, S. HLDM-Ez 2037) remains of Salamandridae indet. from Echzell. Bones in (A, H, N) dorsal, (B, I) lateral, (C, J, O) ventral, (D, F) anterior, (E, G) posterior, (K, L, M, P, S) posterior / anterior and (Q, R) medial views. Scale bars: 1 mm." figureDoi="10.3897/fr.25.83781.figure6" httpUri="https://binary.pensoft.net/fig/682226" pageId="0" pageNumber="99">Fig. 6N-S</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="99" type="materials_examined">
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<paragraph pageId="0" pageNumber="99">Material.</paragraph>
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<paragraph pageId="0" pageNumber="99">One frontal, HLMD-Ez 2038, seven trunk vertebrae, HLMD-Ez 2034, 20 caudal vertebrae, HLMD-Ez 2035, nine vertebrae, HLMD-Ez 2051, 2052, 2062, two ribs HLMD-Ez 2036, HLMD-Ez 2037, two extremity bones HLMD-Ez 2052.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="99" type="description">
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<paragraph pageId="0" pageNumber="99">Description and remarks.</paragraph>
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<paragraph pageId="0" pageNumber="99">
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A single frontal, with a length of 6 mm, displays a flat dorsal surface (Fig.
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Vertebrae (A-E. HLDM-Ez 2059; F-J. HLDM-Ez 2060) and ribs (K. HLDM-Ez 2054; L. HLDM-Ez 2053; M. HLDM-Ez 2055) of Chelotriton sp. Frontal (N, O. HLDM-Ez 2038) and ribs (P, Q. HLDM-Ez 2036; R, S. HLDM-Ez 2037) remains of Salamandridae indet. from Echzell. Bones in (A, H, N) dorsal, (B, I) lateral, (C, J, O) ventral, (D, F) anterior, (E, G) posterior, (K, L, M, P, S) posterior / anterior and (Q, R) medial views. Scale bars: 1 mm." figureDoi="10.3897/fr.25.83781.figure6" httpUri="https://binary.pensoft.net/fig/682226" pageId="0" pageNumber="99">6N</figureCitation>
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). In dorsal view, the most anterior portion of the bone possesses an articulation facet with the parietal that is covered by parallel to each other ridges. Comparison with recent similar-sized species shows most similarities with
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<taxonomicName authority="salamandra" class="Amphibia" family="Salamandridae" genus="Salamandra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Salamandra salamandra" order="Anura" pageId="0" pageNumber="99" phylum="Chordata" rank="species" species="salamandra">
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<emphasis italics="true" pageId="0" pageNumber="99">Salamandra salamandra</emphasis>
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</taxonomicName>
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(MJSN-OS 806). However, due to the lack of comprehensive studies of the skull bones among salamandrids, an allocation of the bone to the family
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<taxonomicName authorityName="Goldfuss" authorityYear="1820" class="Amphibia" family="Salamandridae" kingdom="Animalia" lsidName="" order="Caudata" pageId="0" pageNumber="99" phylum="Chordata" rank="family">Salamandridae</taxonomicName>
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is more appropriate.
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</paragraph>
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<paragraph pageId="0" pageNumber="99">
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The vertebrae are poorly preserved. They show ophistocoelous morphology partially with complex structures of haemal and neural processes, characteristic of the caudal region of the vertebral column (
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<bibRefCitation author="Duellman, WE" journalOrPublisher="The Johns Hopkins University Press, Baltimor-London" pageId="0" pageNumber="99" refId="B35" refString="Duellman, WE, Trueb, L, 1994. Biology of Amphibia. The Johns Hopkins University Press, Baltimor-London" title="Biology of Amphibia." year="1994">Duellman and Trueb 1994</bibRefCitation>
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). On the one hand, the poor preservation and, on the other hand, the poor knowledge on osteological differences of the caudal region in
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<taxonomicName class="Amphibia" family="Salamandridae" genus="Chelotriton" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Chelotriton" order="Caudata" pageId="0" pageNumber="99" phylum="Chordata" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="99">Chelotriton</emphasis>
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</taxonomicName>
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and
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<taxonomicName class="Amphibia" family="Salamandridae" genus="Salamandra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Salamandra" order="Anura" pageId="0" pageNumber="99" phylum="Chordata" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="99">Salamandra</emphasis>
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</taxonomicName>
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genera (both present in the fossil locality) make it at the moment impossible to identify the vertebrae correctly.
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</paragraph>
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<paragraph pageId="0" pageNumber="99">
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Nine small-sized opistocoelous vertebrae are available in the material. They have variable preservation; however, a large number of structures/characters are missing for further identification. Considering the vertebra sizes as well as available similar-sized salamander taxa present in the material, most probably, they represent remains of
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<taxonomicName class="Amphibia" family="Salamandridae" genus="Lissotriton" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lissotriton" order="Anura" pageId="0" pageNumber="99" phylum="Chordata" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="99">Lissotriton</emphasis>
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</taxonomicName>
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,
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<taxonomicName class="Amphibia" family="Salamandridae" genus="Mertensiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mertensiella" order="Caudata" pageId="0" pageNumber="99" phylum="Chordata" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="99">Mertensiella</emphasis>
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</taxonomicName>
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or
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<taxonomicName class="Amphibia" family="Salamandridae" genus="Chioglossa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chioglossa" order="Caudata" pageId="0" pageNumber="99" phylum="Chordata" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="99">Chioglossa</emphasis>
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</taxonomicName>
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. The juvenile and most distal caudal vertebra of
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<taxonomicName class="Amphibia" family="Salamandridae" genus="Salamandra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Salamandra" order="Anura" pageId="0" pageNumber="99" phylum="Chordata" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="99">Salamandra</emphasis>
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</taxonomicName>
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and
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<taxonomicName class="Amphibia" family="Salamandridae" genus="Chelotriton" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Chelotriton" order="Caudata" pageId="0" pageNumber="99" phylum="Chordata" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="99">Chelotriton</emphasis>
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</taxonomicName>
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can be excluded because in the former form the juvenile vertebrae are not fully ossified, whereas in the latter the dorsal tip of the neural crest possesses a ornamented surface, which is missing here.
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</paragraph>
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<paragraph pageId="0" pageNumber="99">
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Two bicapitate rips are present (Fig.
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Vertebrae (A-E. HLDM-Ez 2059; F-J. HLDM-Ez 2060) and ribs (K. HLDM-Ez 2054; L. HLDM-Ez 2053; M. HLDM-Ez 2055) of Chelotriton sp. Frontal (N, O. HLDM-Ez 2038) and ribs (P, Q. HLDM-Ez 2036; R, S. HLDM-Ez 2037) remains of Salamandridae indet. from Echzell. Bones in (A, H, N) dorsal, (B, I) lateral, (C, J, O) ventral, (D, F) anterior, (E, G) posterior, (K, L, M, P, S) posterior / anterior and (Q, R) medial views. Scale bars: 1 mm." figureDoi="10.3897/fr.25.83781.figure6" httpUri="https://binary.pensoft.net/fig/682226" pageId="0" pageNumber="99">6P-S</figureCitation>
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). Their lateral portion does not possess any process, which allows to exclude them from
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<taxonomicName class="Amphibia" family="Salamandridae" genus="Chelotriton" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Chelotriton" order="Caudata" pageId="0" pageNumber="99" phylum="Chordata" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="99">Chelotriton</emphasis>
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</taxonomicName>
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. Most probable, they should belong to the genus
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<taxonomicName class="Amphibia" family="Salamandridae" genus="Salamandra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Salamandra" order="Anura" pageId="0" pageNumber="99" phylum="Chordata" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="99">Salamandra</emphasis>
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</taxonomicName>
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, which has a similar rib morphology and is represented in the material by large-sized individuals as well.
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</paragraph>
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</subSubSection>
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</treatment>
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</document>
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