219 lines
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219 lines
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<document ID-DOI="10.3897/zookeys.19.221" ID-GBIF-Dataset="e6ea8eea-7156-4785-8313-f4bd88dadad1" ID-ISSN="1313–2970" ID-Zenodo-Dep="576503" ID-ZooBank="C473F9F6-1AE7-4B3F-B17F-CA1C2709010C" checkinTime="1587956031210" checkinUser="plazi" docAuthor="Wesener, Thomas, Enghoff, Henrik & Sierwald, Petra" docDate="2009" docId="1772122E7840FF8EFF013A27AB0AE96A" docLanguage="en" docName="ZK_article_2037.pdf" docOrigin="ZooKeys 19 (19)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@11b73717" docStyleName="zookeys.2008.journal_article" docTitle="Caprobolus Wesener & Enghoff & Sierwald 2009, gen. n." docType="treatment" docUuid="23C9C435-1012-44AE-8FF1-82C471F2A0E4" docUuidSource="ZooBank" docVersion="6" lastPageNumber="73" masterDocId="EB4B6A567807FFC6FF8D3F74AF59EA51" masterDocTitle="Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda)" masterLastPageNumber="128" masterPageNumber="1" pageNumber="72" updateTime="1636630017160" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Wesener, Thomas</mods:namePart>
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<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
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<mods:affiliation>Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, IL 60605, U. S. A. & Zoologisches Forschungsmuseum Alexander Koenig, Museumsmeile Bonn, Adenauerallee 160, D- 53113 Bonn, Germany. & B 54 D 3623309</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Enghoff, Henrik</mods:namePart>
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<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
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<mods:affiliation>Natural History Museum of Denmark, Universitetsparken 15, DK- 2100 Copenhagen, Denmark</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sierwald, Petra</mods:namePart>
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<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
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<mods:affiliation>Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, IL 60605, U. S. A.</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2009</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2009-09-04</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>19</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>19</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>128</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.3897/zookeys.19.221</mods:identifier>
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<mods:identifier type="GBIF-Dataset">e6ea8eea-7156-4785-8313-f4bd88dadad1</mods:identifier>
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<mods:identifier type="ISSN">1313–2970</mods:identifier>
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<mods:identifier type="Zenodo-Dep">576503</mods:identifier>
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<mods:identifier type="ZooBank">C473F9F6-1AE7-4B3F-B17F-CA1C2709010C</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3791427" ID-GBIF-Taxon="164184223" ID-Zenodo-Dep="3791427" LSID="urn:lsid:zoobank.org:act:23C9C435-1012-44AE-8FF1-82C471F2A0E4" httpUri="http://treatment.plazi.org/id/1772122E7840FF8EFF013A27AB0AE96A" lastPageId="72" lastPageNumber="73" pageId="71" pageNumber="72">
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<subSubSection box="[140,477,1363,1390]" pageId="71" pageNumber="72" type="nomenclature">
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<paragraph blockId="71.[140,961,1363,1425]" box="[140,477,1363,1390]" pageId="71" pageNumber="72">
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<heading box="[140,477,1363,1390]" fontSize="11" level="2" pageId="71" pageNumber="72" reason="3">
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<emphasis bold="true" box="[140,477,1363,1390]" pageId="71" pageNumber="72">
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<taxonomicName authority="Wesener" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[140,384,1363,1390]" class="Diplopoda" family="Pachybolidae" genus="Caprobolus" kingdom="Animalia" order="Spirobolida" pageId="71" pageNumber="72" phylum="Arthropoda" rank="genus" status="gen. n.">
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<emphasis bold="true" box="[140,276,1363,1390]" italics="true" pageId="71" pageNumber="72">Caprobolus</emphasis>
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Wesener
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</taxonomicName>
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,
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<taxonomicNameLabel box="[396,477,1363,1389]" pageId="71" pageNumber="72" rank="genus">gen. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[140,961,1398,1425]" pageId="71" pageNumber="72" type="description">
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<paragraph blockId="71.[140,961,1363,1425]" box="[140,961,1398,1425]" pageId="71" pageNumber="72">
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<uri box="[140,961,1398,1425]" pageId="71" pageNumber="72">
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urn:lsid:zoobank.org:act:
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<uuid box="[423,961,1398,1425]" pageId="71" pageNumber="72">
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23C9C435-
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<date box="[557,654,1398,1425]" pageId="71" pageNumber="72" value="1012" valueMax="1044">1012-44</date>
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AE-8FF1-82C471F2A0E4
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</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSection lastPageId="72" lastPageNumber="73" pageId="71" pageNumber="72" type="multiple">
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<subSubSection box="[140,761,1469,1496]" pageId="71" pageNumber="72" type="type_taxon">
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<paragraph blockId="71.[140,761,1469,1496]" box="[140,761,1469,1496]" pageId="71" pageNumber="72">
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<emphasis bold="true" box="[140,287,1469,1495]" pageId="71" pageNumber="72">
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<typeStatus box="[140,198,1469,1495]" pageId="71" pageNumber="72">Type</typeStatus>
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species
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</emphasis>
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:
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<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[302,555,1469,1495]" class="Diplopoda" family="Pachybolidae" genus="Caprobolus" kingdom="Animalia" order="Spirobolida" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="andringitra" status="sp. n.">
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<emphasis box="[302,555,1469,1495]" italics="true" pageId="71" pageNumber="72">Caprobolus andringitra</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[562,626,1470,1496]" pageId="71" pageNumber="72" rank="species">sp. n.</taxonomicNameLabel>
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, monotypic
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="72" lastPageNumber="73" pageId="71" pageNumber="72" type="diagnosis">
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<paragraph blockId="71.[140,1108,1539,1672]" lastBlockId="72.[140,1108,167,827]" lastPageId="72" lastPageNumber="73" pageId="71" pageNumber="72">
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<emphasis bold="true" box="[140,259,1539,1565]" inLineHeading="true" pageId="71" pageNumber="72" reason="1">Diagnosis</emphasis>
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: mandible with a very short molar plate (
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<figureCitation box="[743,845,1539,1566]" captionStart="Figure 40" captionStartId="73.[140,212,1526,1548]" captionTargetBox="[154,1094,165,1507]" captionTargetId="figure@73.[154,1095,165,1507]" captionTargetPageId="73" captionText="Figure 40. Caprobolus andringitra sp. n. male paratype, SEM. A antenna, overview B antenna, apical view on disc C antennomere 5 with sensilla basiconica D gnathochilarium, ventral view E endochilarium detail F left mandible, mesal view G mandible, detail of pectinate lamella. 3iT = 3-combed internal tooth; cp = central pads of endochilarium; Endo = endochilarium; eT = external tooth; L = ledge; LT = lateral tooth; Me = mentum; mp = molar plate; pL = pectinate lamella; sb = sensilla basiconica; ST = stipites." figureDoi="http://doi.org/10.5281/zenodo.3767328" httpUri="https://zenodo.org/record/3767328/files/figure.png" pageId="71" pageNumber="72">Fig. 40F</figureCitation>
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), so far a unique character (but the shape of the mandible is only known for a few genera of
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<taxonomicName authorityName="Cook" authorityYear="1895" box="[960,1096,1575,1602]" class="Diplopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirobolida" pageId="71" pageNumber="72" phylum="Arthropoda" rank="order">Spirobolida</taxonomicName>
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). Anterior and posterior gonopods of
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<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[560,682,1610,1636]" class="Diplopoda" family="Pachybolidae" genus="Caprobolus" kingdom="Animalia" order="Spirobolida" pageId="71" pageNumber="72" phylum="Arthropoda" rank="genus">
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<emphasis box="[560,682,1610,1636]" italics="true" pageId="71" pageNumber="72">Caprobolus</emphasis>
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</taxonomicName>
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are unique for
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<taxonomicName authorityName="Cook" authorityYear="1895" box="[865,999,1610,1636]" class="Diplopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirobolida" pageId="71" pageNumber="72" phylum="Arthropoda" rank="order">Spirobolida</taxonomicName>
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. Anterior gonopods with a conspicuously enlarged, slender sternite (
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<figureCitation box="[804,908,1645,1671]" captionStart="Figure 41" captionStartId="74.[140,212,1526,1548]" captionTargetBox="[232,1016,176,1498]" captionTargetId="figure@74.[223,1024,165,1506]" captionTargetPageId="74" captionText="Figure 41. Caprobolus andringitra sp. n., male holotype. A anterior gonopod, anterior view B anterior gonopod, posterior view C right posterior gonopod, posterior view D right posterior gonopod, telopodite, posterior view E right posterior gonopod, telopodite, anterior view. cp = internal coxite process; Cx = coxite; rp = retrorse process; SC = free sperm canal; St = sternite; T = telopodite; T-ibr = inner branch of telopodite; T-mbr =main branch of telopodite; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820823" httpUri="https://zenodo.org/record/3820823/files/figure.png" pageId="71" pageNumber="72">Fig. 41A</figureCitation>
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), but sternite still much wider than in
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<taxonomicName authority="(Wesener and Enghof, 2009)" baseAuthorityName="Wesener and Enghof" baseAuthorityYear="2009" box="[387,870,167,193]" class="Diplopoda" family="Pachybolidae" genus="Spiromimus" kingdom="Animalia" order="Spirobolida" pageId="72" pageNumber="73" phylum="Arthropoda" rank="genus">
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<emphasis box="[387,517,167,193]" italics="true" pageId="72" pageNumber="73">Spiromimus</emphasis>
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(Wesener and Enghof, 2009)
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</taxonomicName>
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. Telopodite of anterior gonopods prominently enlarged into unique, long, slender, sabre-like processes (
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<figureCitation box="[148,251,237,263]" captionStart="Figure 41" captionStartId="74.[140,212,1526,1548]" captionTargetBox="[232,1016,176,1498]" captionTargetId="figure@74.[223,1024,165,1506]" captionTargetPageId="74" captionText="Figure 41. Caprobolus andringitra sp. n., male holotype. A anterior gonopod, anterior view B anterior gonopod, posterior view C right posterior gonopod, posterior view D right posterior gonopod, telopodite, posterior view E right posterior gonopod, telopodite, anterior view. cp = internal coxite process; Cx = coxite; rp = retrorse process; SC = free sperm canal; St = sternite; T = telopodite; T-ibr = inner branch of telopodite; T-mbr =main branch of telopodite; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820823" httpUri="https://zenodo.org/record/3820823/files/figure.png" pageId="72" pageNumber="73">Fig. 41B</figureCitation>
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). Basis of telopodite process projecting into a slender, well-rounded retrorse process (
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<figureCitation box="[236,338,272,298]" captionStart="Figure 41" captionStartId="74.[140,212,1526,1548]" captionTargetBox="[232,1016,176,1498]" captionTargetId="figure@74.[223,1024,165,1506]" captionTargetPageId="74" captionText="Figure 41. Caprobolus andringitra sp. n., male holotype. A anterior gonopod, anterior view B anterior gonopod, posterior view C right posterior gonopod, posterior view D right posterior gonopod, telopodite, posterior view E right posterior gonopod, telopodite, anterior view. cp = internal coxite process; Cx = coxite; rp = retrorse process; SC = free sperm canal; St = sternite; T = telopodite; T-ibr = inner branch of telopodite; T-mbr =main branch of telopodite; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820823" httpUri="https://zenodo.org/record/3820823/files/figure.png" pageId="72" pageNumber="73">Fig. 41B</figureCitation>
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), which is present in numerous genera of
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<taxonomicName authorityName="Cook" authorityYear="1895" box="[812,946,272,298]" class="Diplopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirobolida" pageId="72" pageNumber="73" phylum="Arthropoda" rank="order">Spirobolida</taxonomicName>
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. Telopodite of posterior gonopod longer than, but as wide as, coxite (
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<figureCitation box="[773,881,307,334]" captionStart="Figure 41" captionStartId="74.[140,212,1526,1548]" captionTargetBox="[232,1016,176,1498]" captionTargetId="figure@74.[223,1024,165,1506]" captionTargetPageId="74" captionText="Figure 41. Caprobolus andringitra sp. n., male holotype. A anterior gonopod, anterior view B anterior gonopod, posterior view C right posterior gonopod, posterior view D right posterior gonopod, telopodite, posterior view E right posterior gonopod, telopodite, anterior view. cp = internal coxite process; Cx = coxite; rp = retrorse process; SC = free sperm canal; St = sternite; T = telopodite; T-ibr = inner branch of telopodite; T-mbr =main branch of telopodite; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820823" httpUri="https://zenodo.org/record/3820823/files/figure.png" pageId="72" pageNumber="73">Fig. 41C</figureCitation>
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). Telopodite basally deeply divided into two branches of almost similar width and length (
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<figureCitation box="[953,1038,343,369]" captionStart="Figure 41" captionStartId="74.[140,212,1526,1548]" captionTargetBox="[232,1016,176,1498]" captionTargetId="figure@74.[223,1024,165,1506]" captionTargetPageId="74" captionText="Figure 41. Caprobolus andringitra sp. n., male holotype. A anterior gonopod, anterior view B anterior gonopod, posterior view C right posterior gonopod, posterior view D right posterior gonopod, telopodite, posterior view E right posterior gonopod, telopodite, anterior view. cp = internal coxite process; Cx = coxite; rp = retrorse process; SC = free sperm canal; St = sternite; T = telopodite; T-ibr = inner branch of telopodite; T-mbr =main branch of telopodite; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820823" httpUri="https://zenodo.org/record/3820823/files/figure.png" pageId="72" pageNumber="73">Figs 41</figureCitation>
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C–E). Sperm canal is free, starting at basis of telopodite, only slightly attached to shorter, lateral main branch (
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<figureCitation box="[374,476,413,439]" captionStart="Figure 41" captionStartId="74.[140,212,1526,1548]" captionTargetBox="[232,1016,176,1498]" captionTargetId="figure@74.[223,1024,165,1506]" captionTargetPageId="74" captionText="Figure 41. Caprobolus andringitra sp. n., male holotype. A anterior gonopod, anterior view B anterior gonopod, posterior view C right posterior gonopod, posterior view D right posterior gonopod, telopodite, posterior view E right posterior gonopod, telopodite, anterior view. cp = internal coxite process; Cx = coxite; rp = retrorse process; SC = free sperm canal; St = sternite; T = telopodite; T-ibr = inner branch of telopodite; T-mbr =main branch of telopodite; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820823" httpUri="https://zenodo.org/record/3820823/files/figure.png" pageId="72" pageNumber="73">Fig. 41E</figureCitation>
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). Sperm canal projecting as a flagellum above telopodite branches (
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<figureCitation box="[254,361,448,474]" captionStart="Figure 41" captionStartId="74.[140,212,1526,1548]" captionTargetBox="[232,1016,176,1498]" captionTargetId="figure@74.[223,1024,165,1506]" captionTargetPageId="74" captionText="Figure 41. Caprobolus andringitra sp. n., male holotype. A anterior gonopod, anterior view B anterior gonopod, posterior view C right posterior gonopod, posterior view D right posterior gonopod, telopodite, posterior view E right posterior gonopod, telopodite, anterior view. cp = internal coxite process; Cx = coxite; rp = retrorse process; SC = free sperm canal; St = sternite; T = telopodite; T-ibr = inner branch of telopodite; T-mbr =main branch of telopodite; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820823" httpUri="https://zenodo.org/record/3820823/files/figure.png" pageId="72" pageNumber="73">Fig. 41D</figureCitation>
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). Shares a reddish colour pattern and a yellowish dorsal stripe with
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<taxonomicName authority="de Saussure & Zehntner, 1901" authorityName="de Saussure & Zehntner" authorityYear="1901" box="[140,631,483,510]" class="Diplopoda" family="Pachybolidae" genus="Spiromimus" kingdom="Animalia" order="Spirobolida" pageId="72" pageNumber="73" phylum="Arthropoda" rank="genus">
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<emphasis box="[140,270,484,510]" italics="true" pageId="72" pageNumber="73">Spiromimus</emphasis>
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<bibRefCitation author="de Saussure & Zehntner" box="[278,631,483,510]" firstAuthor="de Saussure" pageId="72" pageNumber="73" pagination="429 - 460" refId="ref56960" refString="de Saussure H, Zehntner L (1901) Myriopoden aus Madagaskar und Zansibar, gesammelt von Dr. A. Voeltzkow. Abhandlungen der Senckenbergischen naturforschenden Gesellschaft 26: 429 - 460." type="journal article" year="1901">de Saussure & Zehntner, 1901</bibRefCitation>
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</taxonomicName>
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,
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<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[645,779,483,509]" class="Diplopoda" family="Pachybolidae" genus="Alluviobolus" kingdom="Animalia" order="Spirobolida" pageId="72" pageNumber="73" phylum="Arthropoda" rank="genus" status="gen. n.">
|
|||
|
|
<emphasis box="[645,779,483,509]" italics="true" pageId="72" pageNumber="73">Alluviobolus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel box="[787,865,484,510]" pageId="72" pageNumber="73" rank="genus">gen. n.</taxonomicNameLabel>
|
|||
|
|
and
|
|||
|
|
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[924,1051,483,509]" class="Diplopoda" family="Pachybolidae" genus="Ostinobolus" kingdom="Animalia" order="Spirobolida" pageId="72" pageNumber="73" phylum="Arthropoda" rank="genus" status="gen. n.">
|
|||
|
|
<emphasis box="[924,1051,483,509]" italics="true" pageId="72" pageNumber="73">Ostinobolus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel pageId="72" pageNumber="73" rank="genus">gen. n.</taxonomicNameLabel>
|
|||
|
|
Gnathochilarium with a subdivided mentum and a single sclerotized ledge on each stipites (
|
|||
|
|
<figureCitation box="[235,336,554,580]" captionStart="Figure 39" captionStartId="72.[140,212,1526,1548]" captionTargetBox="[140,1108,927,1507]" captionTargetId="figure@72.[140,1108,927,1506]" captionTargetPageId="72" captionText="Figure 39. Caprobolus andringitra sp. n., A–C male holotype, D–G male paratype A head, lateral view B legs 1–7, ventral view C telson D head, detail of eye and mandible E gnathochilarium, ventral view F right midbody leg, overview G tarsus of midbody leg, SEM. av = anal valves; Co = collum; Gn = gnathochilarium; L = ledge; LL = lamella lingualis; Md = mandible; Me = mentum; Pre = preanal ring; re = rectangular extension; ST = stipites. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767326" httpUri="https://zenodo.org/record/3767326/files/figure.png" pageId="72" pageNumber="73">Fig. 39E</figureCitation>
|
|||
|
|
), like in
|
|||
|
|
<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[438,576,554,580]" class="Diplopoda" family="Pachybolidae" genus="Flagellobolus" kingdom="Animalia" order="Spirobolida" pageId="72" pageNumber="73" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis box="[438,576,554,580]" italics="true" pageId="72" pageNumber="73">Flagellobolus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[592,735,554,580]" class="Diplopoda" family="Pachybolidae" genus="Riotintobolus" kingdom="Animalia" order="Spirobolida" pageId="72" pageNumber="73" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis box="[592,735,554,580]" italics="true" pageId="72" pageNumber="73">Riotintobolus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[751,944,554,580]" class="Diplopoda" family="Pachybolidae" genus="Pseudocentrobolus" kingdom="Animalia" order="Spirobolida" pageId="72" pageNumber="73" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis box="[751,944,554,580]" italics="true" pageId="72" pageNumber="73">Pseudocentrobolus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[959,1099,554,580]" class="Diplopoda" family="Pachybolidae" genus="Granitobolus" kingdom="Animalia" order="Spirobolida" pageId="72" pageNumber="73" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis box="[959,1099,554,580]" italics="true" pageId="72" pageNumber="73">Granitobolus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[140,274,589,615]" class="Diplopoda" family="Pachybolidae" genus="Alluviobolus" kingdom="Animalia" order="Spirobolida" pageId="72" pageNumber="73" phylum="Arthropoda" rank="genus" status="gen. n.">
|
|||
|
|
<emphasis box="[140,274,589,615]" italics="true" pageId="72" pageNumber="73">Alluviobolus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel box="[281,358,590,616]" pageId="72" pageNumber="73" rank="genus">gen. n.</taxonomicNameLabel>
|
|||
|
|
,
|
|||
|
|
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[373,500,589,615]" class="Diplopoda" family="Pachybolidae" genus="Ostinobolus" kingdom="Animalia" order="Spirobolida" pageId="72" pageNumber="73" phylum="Arthropoda" rank="genus" status="gen. n.">
|
|||
|
|
<emphasis box="[373,500,589,615]" italics="true" pageId="72" pageNumber="73">Ostinobolus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel box="[507,584,590,616]" pageId="72" pageNumber="73" rank="genus">gen. n.</taxonomicNameLabel>
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection pageId="72" pageNumber="73" type="distribution">
|
|||
|
|
<paragraph blockId="72.[140,1108,167,827]" pageId="72" pageNumber="73">
|
|||
|
|
<emphasis bold="true" box="[185,487,624,650]" pageId="72" pageNumber="73">
|
|||
|
|
<emphasis box="[185,336,624,650]" inLineHeading="true" pageId="72" pageNumber="73" reason="1">Distribution</emphasis>
|
|||
|
|
and ecology
|
|||
|
|
</emphasis>
|
|||
|
|
: only known from the rainforests of Andringitra, close by Ivohibe and Vevembe, located at the edge of the high plateau towards the east coast in the southern half of
|
|||
|
|
<collectingCountry box="[410,543,695,721]" name="Madagascar" pageId="72" pageNumber="73">Madagascar</collectingCountry>
|
|||
|
|
(
|
|||
|
|
<figureCitation box="[560,647,695,721]" captionStart="Figure 30" captionStartId="56.[140,210,1654,1676]" captionTargetBox="[129,1120,1092,1644]" captionTargetId="figure@56.[140,1108,1098,1642]" captionTargetPageId="56" captionText="Figure 30. Distribution of the genera Pseudocentrobolus gen. n., Granitobolus gen. n. and Caprobolus gen.n." figureDoi="http://doi.org/10.5281/zenodo.3820818" httpUri="https://zenodo.org/record/3820818/files/figure.png" pageId="72" pageNumber="73">Fig. 30</figureCitation>
|
|||
|
|
). The only known species of
|
|||
|
|
<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[986,1108,695,721]" class="Diplopoda" family="Pachybolidae" genus="Caprobolus" kingdom="Animalia" order="Spirobolida" pageId="72" pageNumber="73" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis box="[986,1108,695,721]" italics="true" pageId="72" pageNumber="73">Caprobolus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
was collected in Andringitra together with a
|
|||
|
|
<taxonomicName authorityName="de Saussure & Zehntner" authorityYear="1901" box="[643,773,730,756]" class="Diplopoda" family="Pachybolidae" genus="Spiromimus" kingdom="Animalia" order="Spirobolida" pageId="72" pageNumber="73" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis box="[643,773,730,756]" italics="true" pageId="72" pageNumber="73">Spiromimus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
species.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection pageId="72" pageNumber="73" type="etymology">
|
|||
|
|
<paragraph blockId="72.[140,1108,167,827]" pageId="72" pageNumber="73">
|
|||
|
|
<emphasis bold="true" box="[185,315,765,791]" inLineHeading="true" pageId="72" pageNumber="73" reason="1">Etymology</emphasis>
|
|||
|
|
:
|
|||
|
|
<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[329,451,765,791]" class="Diplopoda" family="Pachybolidae" genus="Caprobolus" kingdom="Animalia" order="Spirobolida" pageId="72" pageNumber="73" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis box="[329,451,765,791]" italics="true" pageId="72" pageNumber="73">Caprobolus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, masculine, composed of
|
|||
|
|
<emphasis box="[741,808,766,792]" italics="true" pageId="72" pageNumber="73">Capro</emphasis>
|
|||
|
|
-, formed after
|
|||
|
|
<emphasis box="[977,1100,766,791]" italics="true" pageId="72" pageNumber="73">capricornus</emphasis>
|
|||
|
|
, which refers to the tropic of Capricorn where this genus was first collected, and
|
|||
|
|
<emphasis box="[1036,1100,800,826]" italics="true" pageId="72" pageNumber="73">-bolus</emphasis>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</subSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|