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<document id="88513DDEB6937294C4B56CC199C3C340" ID-DOI="10.5281/zenodo.214842" ID-GBIF-Dataset="a6e682b5-3b8a-4b94-8439-1ce58e6204b8" ID-ISSN="1175-5326" ID-Zenodo-Dep="214842" ID-ZooBank="F6BA89F6-D9F7-4A42-BD3D-7E49C1453848" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460543484391" checkinUser="plazi" docAuthor="Cardona, Luisa &amp; Campos, Martha R." docDate="2012" docId="03A3EE5DA5452244FF63FE58A572FE4E" docLanguage="en" docName="zt03515p088.pdf" docOrigin="Zootaxa 3515" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Phallangothelphusa martensis Cardona &amp; Campos, 2012, new species" docType="treatment" docVersion="7" lastPageNumber="88" masterDocId="FF9A9625A5442241FFF4FFF5A660FF81" masterDocTitle="A new species of freshwater crab of the genus Phallangothelphusa Pretzmann, 1965, from Colombia (Crustacea: Decapoda: Pseudothelphusidae)" masterLastPageNumber="88" masterPageNumber="83" pageNumber="84" updateTime="1698317380925" updateUser="plazi">
<mods:mods id="AABDD5552F30B0B7FA93557C094DAB25" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="0F2D91D02082B42A01673F382AD38693">
<mods:title id="2731EF8CEB64AC1662A372D2C20EDC1F">A new species of freshwater crab of the genus Phallangothelphusa Pretzmann, 1965, from Colombia (Crustacea: Decapoda: Pseudothelphusidae)</mods:title>
</mods:titleInfo>
<mods:name id="FDB279B4C755A0C33585B51E14BDED04" type="personal">
<mods:role id="1D363BB57FB2ECC8F868D3AFFDADA885">
<mods:roleTerm id="83E9AEE54D8807B51287C428FC059B85">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C002D413BB2B592AE8582CDE949BDF3C">Cardona, Luisa</mods:namePart>
</mods:name>
<mods:name id="4853C8D639F2056DB9FF91AE8A66A870" type="personal">
<mods:role id="0631909DF5D2E6F538960C484A573E90">
<mods:roleTerm id="CEF7F9E27DB13277DC3E6B496618D092">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3CD6800C0AA595658A3CCFF92082D37A">Campos, Martha R.</mods:namePart>
</mods:name>
<mods:typeOfResource id="EFB7CACE8B88F8FC078E6676DDDA54E6">text</mods:typeOfResource>
<mods:relatedItem id="C05AF4221D8E764009C1A0A5C114F208" type="host">
<mods:titleInfo id="D9000E1C45B5719374F9703EDEAAF1EA">
<mods:title id="F83D7A40125846BD7B6858B334D657E6">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="5BD1841C47336886DFADD5E30830A1A2">
<mods:date id="F29C5939D98D595890D380C75B034A12">2012</mods:date>
<mods:detail id="7E0BE8C44D7B6C6C33F08F8EF4DD1938" type="volume">
<mods:number id="3C956D69F84DD182F499CAD8D724BEAF">3515</mods:number>
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<mods:classification id="52E656FF6B77808916A73B5CEB7D956D">journal article</mods:classification>
<mods:identifier id="7BD7BD00160FD895C73C46B2CE793237" type="DOI">10.5281/zenodo.214842</mods:identifier>
<mods:identifier id="A77C696354E07ADBA89BC72657D38FF6" type="GBIF-Dataset">a6e682b5-3b8a-4b94-8439-1ce58e6204b8</mods:identifier>
<mods:identifier id="E0C697FF79D7B25B29DBF70277D6C8DC" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="06B887083C23A6489205BB54A97E7662" type="Zenodo-Dep">214842</mods:identifier>
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</mods:mods>
<treatment id="03A3EE5DA5452244FF63FE58A572FE4E" ID-DOI="http://doi.org/10.5281/zenodo.6172409" ID-GBIF-Taxon="119528450" ID-Zenodo-Dep="6172409" LSID="urn:lsid:plazi:treatment:03A3EE5DA5452244FF63FE58A572FE4E" httpUri="http://treatment.plazi.org/id/03A3EE5DA5452244FF63FE58A572FE4E" lastPageId="5" lastPageNumber="88" pageId="1" pageNumber="84">
<subSubSection id="C3100CC0A5452240FF63FE58A771FE66" pageId="1" pageNumber="84" type="nomenclature">
<paragraph id="8BB55F4BA5452240FF63FE58A4C6FE46" blockId="1.[151,678,429,487]" box="[151,678,429,455]" pageId="1" pageNumber="84">
<heading id="D0FDE827A5452240FF63FE58A4C6FE46" bold="true" box="[151,678,429,455]" fontSize="11" level="1" pageId="1" pageNumber="84" reason="1">
<emphasis id="B97E8359A5452240FF63FE58A4C6FE46" bold="true" box="[151,678,429,455]" pageId="1" pageNumber="84">
<taxonomicName id="4C0A24C8A5452240FF63FE58A467FE46" ID-CoL="4FRSY" box="[151,519,429,455]" class="Malacostraca" family="Pseudothelphusidae" genus="Phallangothelphusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="84" phylum="Arthropoda" rank="species" species="martensis" status="sp. nov.">
<emphasis id="B97E8359A5452240FF63FE58A467FE46" bold="true" box="[151,519,429,455]" italics="true" pageId="1" pageNumber="84">Phallangothelphusa martensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A24D3E22A5452240FDE2FE58A4C6FE46" box="[534,678,429,455]" pageId="1" pageNumber="84" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB55F4BA5452240FF63FE3AA771FE66" blockId="1.[151,678,429,487]" box="[151,273,463,487]" pageId="1" pageNumber="84">
(
<figureCitation id="133143CEA5452240FF6BFE3AA768FE66" box="[159,264,463,487]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="2.[151,250,1943,1966]" captionStartId-1="3.[151,250,1917,1940]" captionStartId-2="4.[151,250,1027,1050]" captionTargetBox-0="[228,1335,193,1925]" captionTargetBox-1="[208,1363,199,1874]" captionTargetBox-2="[163,1430,196,987]" captionTargetId-0="figure@2.[223,1358,193,1927]" captionTargetId-1="figure@3.[208,1367,193,1877]" captionTargetId-2="figure@4.[151,1436,193,1006]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURE 1. Phallangothelphusa martensis new species, male holotype, 1 male, cl 17.3 mm, cb 30.1 mm, ICN-MHN-CR 2656. A, habitus; B, frontal view of carapace; C, opening of left efferent branchial channel, external view; D, chela of large cheliped, external view; E, left third maxilliped, external view." captionText-1="FIGURE 2. Phallangothelphusa martensis new species, male holotype, ICN-MHN-CR 2656. A, left first gonopod, caudal view; B, same, lateral view; C, same, mesial view; D, same, cephalic view; E, apex distal view; F, second gonopod, caudal view; G, distal part of second gonopod. 1, distal projection of mesial process; 2, curvature; 3, proximal projection of mesial process; 4, lateral process." captionText-2="FIGURE 3. Phallangothelphusa martensis new species, male holotype, ICN-MHN-CR 2656, left first gonopod: A, apex distal view; Phallangothelphusa dispar (Zimmer, 1912), ICN-MHN-CR 0 732, left first gonopod: B, apex, distal view. 1, distal projection of mesial process; 2, curvature; 3, proximal projection of mesial process; 4, lateral process." httpUri-0="https://zenodo.org/record/214843/files/figure.png" httpUri-1="https://zenodo.org/record/214844/files/figure.png" httpUri-2="https://zenodo.org/record/214845/files/figure.png" pageId="1" pageNumber="84">Figs. 1‒3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3100CC0A5452240FF63FDEDA48DFCFD" pageId="1" pageNumber="84" type="materials_examined">
<paragraph id="8BB55F4BA5452240FF63FDEDA72DFDFB" blockId="1.[151,1437,535,2011]" pageId="1" pageNumber="84">
<emphasis id="B97E8359A5452240FF63FDEDA76BFDB1" bold="true" box="[151,267,536,560]" pageId="1" pageNumber="84">
<typeStatus id="54B1E1E9A5452240FF63FDEDA766FDB1" box="[151,262,536,560]" pageId="1" pageNumber="84" type="holotype">Holotype</typeStatus>
.
</emphasis>
Municipio San Martín, Vereda La Unión, La Batea Stream, elevation
<quantity id="4CF2F2AEA5452240FB96FDE2A2D1FDAE" box="[1122,1201,535,559]" metricMagnitude="2" metricUnit="m" metricValue="2.9" pageId="1" pageNumber="84" unit="m" value="290.0">290 m</quantity>
, Cesar Department,
<collectingCountry id="F31D1FDBA5452240FF63FDCAA768FDD6" box="[151,264,575,599]" name="Colombia" pageId="1" pageNumber="84">Colombia</collectingCountry>
, 7° 59´42.4ʺ N, 73° 24´20.9ʺ W,
<date id="FFB4798BA5452240FD71FDCAA575FDD6" box="[645,789,575,599]" pageId="1" pageNumber="84" value="2011-11-03">3 Nov. 2011</date>
, leg. L. Cardona,
<specimenCount id="9D0C94C2A5452240FC17FDCAA253FDD6" box="[995,1075,575,599]" pageId="1" pageNumber="84" type="male">1 male</specimenCount>
, cl
<quantity id="4CF2F2AEA5452240FBA9FDCAA2A5FDD6" box="[1117,1221,575,599]" metricMagnitude="-2" metricUnit="m" metricValue="1.73" pageId="1" pageNumber="84" unit="mm" value="17.3">17.3 mm</quantity>
, cb
<quantity id="4CF2F2AEA5452240FB02FDCAA336FDD6" box="[1270,1366,575,599]" metricMagnitude="-2" metricUnit="m" metricValue="3.01" pageId="1" pageNumber="84" unit="mm" value="30.1">30.1mm</quantity>
, ICN- MHN-CR 2656.
</paragraph>
<paragraph id="8BB55F4BA5452240FF31FD72A49DFD43" blockId="1.[151,1437,535,2011]" pageId="1" pageNumber="84">
<emphasis id="B97E8359A5452240FF31FD72A724FD1E" bold="true" box="[197,324,647,671]" pageId="1" pageNumber="84">
<typeStatus id="54B1E1E9A5452240FF31FD72A75FFD1E" box="[197,319,647,671]" pageId="1" pageNumber="84" type="paratype">Paratypes</typeStatus>
.
</emphasis>
Same locality data as
<typeStatus id="54B1E1E9A5452240FDCAFD72A4C3FD1E" box="[574,675,647,671]" pageId="1" pageNumber="84" type="holotype">holotype</typeStatus>
,
<specimenCount id="9D0C94C2A5452240FD5AFD72A567FD1E" box="[686,775,647,671]" pageId="1" pageNumber="84" type="male">2 males</specimenCount>
, cl 16.5 and
<quantity id="4CF2F2AEA5452240FC60FD72A59BFD1E" box="[916,1019,647,671]" metricMagnitude="-2" metricUnit="m" metricValue="1.51" pageId="1" pageNumber="84" unit="mm" value="15.1">15.1 mm</quantity>
, cb 28.4 and
<quantity id="4CF2F2AEA5452240FB7AFD72A295FD1E" box="[1166,1269,647,671]" metricMagnitude="-2" metricUnit="m" metricValue="2.55" pageId="1" pageNumber="84" unit="mm" value="25.5">25.5 mm</quantity>
,
<specimenCount id="9D0C94C2A5452240FAF4FD72A31DFD1E" box="[1280,1405,647,671]" pageId="1" pageNumber="84" type="juvenile">5 juveniles</specimenCount>
, cl
<quantity id="4CF2F2AEA5452240FF6DFD5FA750FD43" box="[153,304,682,706]" metricMagnitude="-3" metricUnit="m" metricValue="8.95" metricValueMax="10.1" metricValueMin="7.8" pageId="1" pageNumber="84" unit="mm" value="8.95" valueMax="10.1" valueMin="7.8">7.8–10.1 mm</quantity>
, cb
<quantity id="4CF2F2AEA5452240FEA9FD5FA462FD43" box="[349,514,682,706]" metricMagnitude="-2" metricUnit="m" metricValue="1.48" metricValueMax="1.67" metricValueMin="1.29" pageId="1" pageNumber="84" unit="mm" value="14.8" valueMax="16.7" valueMin="12.9">12.9–16.7 mm</quantity>
, ICN-MHN-CR 2666.
</paragraph>
<paragraph id="8BB55F4BA5452240FF31FD3BA48DFCFD" blockId="1.[151,1437,535,2011]" pageId="1" pageNumber="84">
<emphasis id="B97E8359A5452240FF31FD3BA45AFD66" bold="true" box="[197,570,718,743]" pageId="1" pageNumber="84">Additional material examined:</emphasis>
Municipio San Martín, Vereda Alto de la Raya, stream by Rosmira Farm, elevation
<quantity id="4CF2F2AEA5452240FEFDFD00A735FC8C" box="[265,341,757,781]" metricMagnitude="2" metricUnit="m" metricValue="3.2" pageId="1" pageNumber="84" unit="m" value="320.0">320 m</quantity>
, César Department,
<collectingCountry id="F31D1FDBA5452240FDB6FD00A4D3FC8C" box="[578,691,757,781]" name="Colombia" pageId="1" pageNumber="84">Colombia</collectingCountry>
, 8° 4´7.2ʺ N, 73° 25´8.06ʺ W,
<date id="FFB4798BA5452240FBEFFD00A2DCFC8C" box="[1051,1212,757,782]" pageId="1" pageNumber="84" value="2011-08-26">26 Aug. 2011</date>
, leg. L. Cardona,
<specimenCount id="9D0C94C2A5452240FA7AFD00A6B2FCB0" pageId="1" pageNumber="84" type="male">1 male</specimenCount>
, cl
<quantity id="4CF2F2AEA5452240FF0FFCECA702FCB0" box="[251,354,793,817]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" pageId="1" pageNumber="84" unit="mm" value="15.5">15.5 mm</quantity>
, cb
<quantity id="4CF2F2AEA5452240FE66FCECA799FCB0" box="[402,505,793,817]" metricMagnitude="-2" metricUnit="m" metricValue="2.63" pageId="1" pageNumber="84" unit="mm" value="26.3">26.3 mm</quantity>
, ICN-MHN-CR 2654.— Municipio San Martín, Vereda Alto de la Raya, La Raya Stream, elevation
<quantity id="4CF2F2AEA5452240FE9DFCB5A7D5FCD9" box="[361,437,832,856]" metricMagnitude="2" metricUnit="m" metricValue="2.9" pageId="1" pageNumber="84" unit="m" value="290.0">290 m</quantity>
, César Department,
<collectingCountry id="F31D1FDBA5452240FD56FCB5A573FCD9" box="[674,787,832,856]" name="Colombia" pageId="1" pageNumber="84">Colombia</collectingCountry>
, 8° 3´8ʺ N, 73° 25´9ʺ W,
<date id="FFB4798BA5452240FBC5FCB5A2ABFCD9" box="[1073,1227,832,856]" pageId="1" pageNumber="84" value="2007-03-10">10 Mar 2007</date>
, leg.
<collectingCountry id="F31D1FDBA5452240FAF8FCB4A340FCD9" box="[1292,1312,833,856]" name="Italy" pageId="1" pageNumber="84">Y</collectingCountry>
. Muñoz,
<specimenCount id="9D0C94C2A5452240FA7AFCB5A6B2FCFD" pageId="1" pageNumber="84" type="male">1 male</specimenCount>
, cl 15.0 mm, cb
<quantity id="4CF2F2AEA5452240FE7FFC91A791FCFD" box="[395,497,868,892]" metricMagnitude="-2" metricUnit="m" metricValue="2.62" pageId="1" pageNumber="84" unit="mm" value="26.2">26.2 mm</quantity>
, ICN-MHN-CR 2443.
</paragraph>
</subSubSection>
<subSubSection id="C3100CC0A5452240FF31FC7DA40BFBD6" pageId="1" pageNumber="84" type="diagnosis">
<paragraph id="8BB55F4BA5452240FF31FC7DA40BFBD6" blockId="1.[151,1437,535,2011]" pageId="1" pageNumber="84">
<emphasis id="B97E8359A5452240FF31FC7DA75FFC20" bold="true" box="[197,319,904,929]" pageId="1" pageNumber="84">Diagnosis.</emphasis>
Third maxilliped with exognath 0.9 times length of ischium. Lateral side of first male gonopod with subdistal constriction, prominent ridge running parallel to caudal lobe; mesial process formed by 2 projections: distal, proximal, curvature between them greater than 90°, distal projection triangular in caudal view, with basal acute spine, directed downwards in distal view; proximal projection rectangular with notch on basal portion; lateral process nearly semicircular, inwardly turned, parallel to axis of gonopod (60%), remaining 40% projected mesially; apex outline oval in distal view.
</paragraph>
</subSubSection>
<subSubSection id="C3100CC0A5452245FF31FB97A5B5F9CA" lastPageId="4" lastPageNumber="87" pageId="1" pageNumber="84" type="description">
<paragraph id="8BB55F4BA5452240FF31FB97A5EBFA3E" blockId="1.[151,1437,535,2011]" pageId="1" pageNumber="84">
<emphasis id="B97E8359A5452240FF31FB97A443FBFA" bold="true" box="[197,547,1122,1147]" pageId="1" pageNumber="84">
Description of the
<typeStatus id="54B1E1E9A5452240FE41FB96A47EFBFA" box="[437,542,1123,1147]" pageId="1" pageNumber="84" type="holotype">holotype</typeStatus>
.
</emphasis>
Carapace (
<figureCitation id="133143CEA5452240FD5BFB96A494FBFA" box="[687,756,1123,1147]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1966]" captionTargetBox="[228,1335,193,1925]" captionTargetId="figure@2.[223,1358,193,1927]" captionTargetPageId="2" captionText="FIGURE 1. Phallangothelphusa martensis new species, male holotype, 1 male, cl 17.3 mm, cb 30.1 mm, ICN-MHN-CR 2656. A, habitus; B, frontal view of carapace; C, opening of left efferent branchial channel, external view; D, chela of large cheliped, external view; E, left third maxilliped, external view." httpUri="https://zenodo.org/record/214843/files/figure.png" pageId="1" pageNumber="84">Fig. 1</figureCitation>
A) with straight, wide, deep cervical grooves, becoming indistinct towards margins of carapace; antero-lateral margins with wide deep notch behind outer orbital angle, notch fringed by papillae, with 5 papillae before cervical grooves; lateral margins with approximately 14 papilliform teeth, diminishing in size posteriorly; postfrontal lobes high, oval, delimited anteriorly by transverse depression; median groove shallow; front high, slightly excavated, upper border convex with slight middle depression in dorsal view, marked with row of papillae, lower margin sinuous in frontal view, conspicuously thickened; upper, lower orbital margins each fringed with tubercles (
<figureCitation id="133143CEA5452240FC57FACEA584FAD2" box="[931,996,1339,1363]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1966]" captionTargetBox="[228,1335,193,1925]" captionTargetId="figure@2.[223,1358,193,1927]" captionTargetPageId="2" captionText="FIGURE 1. Phallangothelphusa martensis new species, male holotype, 1 male, cl 17.3 mm, cb 30.1 mm, ICN-MHN-CR 2656. A, habitus; B, frontal view of carapace; C, opening of left efferent branchial channel, external view; D, chela of large cheliped, external view; E, left third maxilliped, external view." httpUri="https://zenodo.org/record/214843/files/figure.png" pageId="1" pageNumber="84">Fig. 1</figureCitation>
B); dorsal surface of carapace smooth, covered by small papillae, regions demarcated; third maxilliped with shallow depression on subdistal external margin of merus, exognath 0.9 times length of ischium (
<figureCitation id="133143CEA5452240FCFAFA76A52DFA1A" box="[782,845,1411,1435]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1966]" captionTargetBox="[228,1335,193,1925]" captionTargetId="figure@2.[223,1358,193,1927]" captionTargetPageId="2" captionText="FIGURE 1. Phallangothelphusa martensis new species, male holotype, 1 male, cl 17.3 mm, cb 30.1 mm, ICN-MHN-CR 2656. A, habitus; B, frontal view of carapace; C, opening of left efferent branchial channel, external view; D, chela of large cheliped, external view; E, left third maxilliped, external view." httpUri="https://zenodo.org/record/214843/files/figure.png" pageId="1" pageNumber="84">Fig. 1</figureCitation>
E); orifice of efferent branchial channel open, nearly ovate with recurved extension of lateral lobe of epistome (
<figureCitation id="133143CEA5452240FCD2FA52A505FA3E" box="[806,869,1447,1471]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1966]" captionTargetBox="[228,1335,193,1925]" captionTargetId="figure@2.[223,1358,193,1927]" captionTargetPageId="2" captionText="FIGURE 1. Phallangothelphusa martensis new species, male holotype, 1 male, cl 17.3 mm, cb 30.1 mm, ICN-MHN-CR 2656. A, habitus; B, frontal view of carapace; C, opening of left efferent branchial channel, external view; D, chela of large cheliped, external view; E, left third maxilliped, external view." httpUri="https://zenodo.org/record/214843/files/figure.png" pageId="1" pageNumber="84">Fig. 1</figureCitation>
C).
</paragraph>
<paragraph id="8BB55F4BA5452240FF33FA3EA797F882" blockId="1.[151,1437,535,2011]" pageId="1" pageNumber="84">
First pereopods heterochelous (
<figureCitation id="133143CEA5452240FDDDFA3EA40AFA62" box="[553,618,1483,1507]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1966]" captionTargetBox="[228,1335,193,1925]" captionTargetId="figure@2.[223,1358,193,1927]" captionTargetPageId="2" captionText="FIGURE 1. Phallangothelphusa martensis new species, male holotype, 1 male, cl 17.3 mm, cb 30.1 mm, ICN-MHN-CR 2656. A, habitus; B, frontal view of carapace; C, opening of left efferent branchial channel, external view; D, chela of large cheliped, external view; E, left third maxilliped, external view." httpUri="https://zenodo.org/record/214843/files/figure.png" pageId="1" pageNumber="84">Fig. 1</figureCitation>
A); right cheliped larger than left; merus with 3 crests as follows: upper crest with rows of tubercles of different sizes, internal lower crest with 2 tight rows of tubercles, diminishing in size proximally, external lower crest with row of papillae; carpus with blunt subdistal spine, follows by 6 rounded tubercles, decreasing in size proximally; palms of both chelipeds smooth, swollen, with row of papillae on external, internal margin; fingers of chelae with rows of tubercles, gaping when closed, tips crossing (
<figureCitation id="133143CEA5452240FB69F9AEA2BCF9F2" box="[1181,1244,1627,1651]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1966]" captionTargetBox="[228,1335,193,1925]" captionTargetId="figure@2.[223,1358,193,1927]" captionTargetPageId="2" captionText="FIGURE 1. Phallangothelphusa martensis new species, male holotype, 1 male, cl 17.3 mm, cb 30.1 mm, ICN-MHN-CR 2656. A, habitus; B, frontal view of carapace; C, opening of left efferent branchial channel, external view; D, chela of large cheliped, external view; E, left third maxilliped, external view." httpUri="https://zenodo.org/record/214843/files/figure.png" pageId="1" pageNumber="84">Fig. 1</figureCitation>
D). Walking legs (second to fifth pereopods) slender (
<figureCitation id="133143CEA5452240FDC5F98AA411F916" box="[561,625,1663,1687]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1966]" captionTargetBox="[228,1335,193,1925]" captionTargetId="figure@2.[223,1358,193,1927]" captionTargetPageId="2" captionText="FIGURE 1. Phallangothelphusa martensis new species, male holotype, 1 male, cl 17.3 mm, cb 30.1 mm, ICN-MHN-CR 2656. A, habitus; B, frontal view of carapace; C, opening of left efferent branchial channel, external view; D, chela of large cheliped, external view; E, left third maxilliped, external view." httpUri="https://zenodo.org/record/214843/files/figure.png" pageId="1" pageNumber="84">Fig. 1</figureCitation>
A), dactyli each about 1.5 times as long as propodi, with 5 longitudinal rows of large spines diminishing in size proximally, arrangement of spines on dactylus of left second pereopod as follows: anterolateral, anteroventral rows each with 8 spines, external row with 9 spines, postero-ventral, posterolateral rows each with 4 spines.
</paragraph>
<paragraph id="8BB55F4BA5452245FF33F8FAA78BFAAA" blockId="1.[151,1437,535,2011]" lastBlockId="4.[151,1436,1155,2010]" lastPageId="4" lastPageNumber="87" pageId="1" pageNumber="84">
First male gonopod with subdistal portion bent laterally; caudal lobe wide basally, distally, narrower near middle, hourglass-like, subdistal portion bent laterally, in caudal view; lateral side with subdistal constriction, prominent ridge running parallel to caudal lobe; mesial process formed by 2 projections (distal, proximal), separated by wide curvature between; distal projection triangular, semi-acute distally, in caudal view, with basal acute spine, directed downwards in distal view; proximal projection rectangular, recurved meso-caudally with notch on basal portion in caudal view (
<figureCitation id="133143CEA5452240FDBBF836A4FDF85A" box="[591,669,1987,2011]" captionStart="FIGURE 2" captionStartId="3.[151,250,1917,1940]" captionTargetBox="[208,1363,199,1874]" captionTargetId="figure@3.[208,1367,193,1877]" captionTargetPageId="3" captionText="FIGURE 2. Phallangothelphusa martensis new species, male holotype, ICN-MHN-CR 2656. A, left first gonopod, caudal view; B, same, lateral view; C, same, mesial view; D, same, cephalic view; E, apex distal view; F, second gonopod, caudal view; G, distal part of second gonopod. 1, distal projection of mesial process; 2, curvature; 3, proximal projection of mesial process; 4, lateral process." httpUri="https://zenodo.org/record/214844/files/figure.png" pageId="1" pageNumber="84">Figs. 2</figureCitation>
A, D, E); lateral process near semicircular, inwardly turned, parallel to axis of gonopod, approximately 60% of length, remaining 40% projected mesially, basal portion covered by rows of spinules, separated from proximal projection of mesial process by deep gap (
<figureCitation id="133143CEA5402245FBA4FB52A2F3FB3E" box="[1104,1171,1191,1215]" captionStart="FIGURE 2" captionStartId="3.[151,250,1917,1940]" captionTargetBox="[208,1363,199,1874]" captionTargetId="figure@3.[208,1367,193,1877]" captionTargetPageId="3" captionText="FIGURE 2. Phallangothelphusa martensis new species, male holotype, ICN-MHN-CR 2656. A, left first gonopod, caudal view; B, same, lateral view; C, same, mesial view; D, same, cephalic view; E, apex distal view; F, second gonopod, caudal view; G, distal part of second gonopod. 1, distal projection of mesial process; 2, curvature; 3, proximal projection of mesial process; 4, lateral process." httpUri="https://zenodo.org/record/214844/files/figure.png" pageId="4" pageNumber="87">Fig. 2</figureCitation>
C–E); lateral process as long as proximal projection of mesial process; apex outline oval in distal view (
<figureCitation id="133143CEA5402245FBD8FB3EA211FB62" box="[1068,1137,1227,1251]" captionStart="FIGURE 2" captionStartId="3.[151,250,1917,1940]" captionTargetBox="[208,1363,199,1874]" captionTargetId="figure@3.[208,1367,193,1877]" captionTargetPageId="3" captionText="FIGURE 2. Phallangothelphusa martensis new species, male holotype, ICN-MHN-CR 2656. A, left first gonopod, caudal view; B, same, lateral view; C, same, mesial view; D, same, cephalic view; E, apex distal view; F, second gonopod, caudal view; G, distal part of second gonopod. 1, distal projection of mesial process; 2, curvature; 3, proximal projection of mesial process; 4, lateral process." httpUri="https://zenodo.org/record/214844/files/figure.png" pageId="4" pageNumber="87">Fig. 2</figureCitation>
D, E); conspicuous caudal setae basally, laterally. Second male gonopod slightly sinuous with distal internal portion nearly flat, covered by rows of spinules (
<figureCitation id="133143CEA5402245FEAAFAE6A7CBFAAA" box="[350,427,1299,1323]" captionStart="FIGURE 2" captionStartId="3.[151,250,1917,1940]" captionTargetBox="[208,1363,199,1874]" captionTargetId="figure@3.[208,1367,193,1877]" captionTargetPageId="3" captionText="FIGURE 2. Phallangothelphusa martensis new species, male holotype, ICN-MHN-CR 2656. A, left first gonopod, caudal view; B, same, lateral view; C, same, mesial view; D, same, cephalic view; E, apex distal view; F, second gonopod, caudal view; G, distal part of second gonopod. 1, distal projection of mesial process; 2, curvature; 3, proximal projection of mesial process; 4, lateral process." httpUri="https://zenodo.org/record/214844/files/figure.png" pageId="4" pageNumber="87">Figs. 2</figureCitation>
F, G).
</paragraph>
<caption id="DF750FC3A5462243FF63F862A572F86B" httpUri="https://zenodo.org/record/214843/files/figure.png" pageId="2" pageNumber="85" targetBox="[228,1335,193,1925]" targetPageId="2">
<paragraph id="8BB55F4BA5462243FF63F862A572F86B" blockId="2.[151,1436,1943,2026]" pageId="2" pageNumber="85">
<emphasis id="B97E8359A5462243FF63F862A776F82D" bold="true" box="[151,278,1943,1966]" pageId="2" pageNumber="85">FIGURE 1.</emphasis>
<taxonomicName id="4C0A24C8A5462243FEEBF862A437F82C" box="[287,599,1943,1965]" class="Malacostraca" family="Pseudothelphusidae" genus="Phallangothelphusa" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="85" phylum="Arthropoda" rank="species" species="martensis" status="sp. nov.">
<emphasis id="B97E8359A5462243FEEBF862A437F82C" box="[287,599,1943,1965]" italics="true" pageId="2" pageNumber="85">Phallangothelphusa martensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D3E22A5462243FD95F86DA480F82F" box="[609,736,1943,1966]" pageId="2" pageNumber="85" rank="species">
<emphasis id="B97E8359A5462243FD95F86DA480F82F" bold="true" box="[609,736,1943,1966]" pageId="2" pageNumber="85">new species</emphasis>
</taxonomicNameLabel>
, male holotype, 1 male, cl 17.3 mm, cb 30.1 mm, ICN-MHN-CR 2656. A, habitus; B, frontal view of carapace; C, opening of left efferent branchial channel, external view; D, chela of large cheliped, external view; E, left third maxilliped, external view.
</paragraph>
</caption>
<caption id="DF750FC3A5472242FF63F888A7C2F871" httpUri="https://zenodo.org/record/214844/files/figure.png" pageId="3" pageNumber="86" targetBox="[208,1363,199,1874]" targetPageId="3">
<paragraph id="8BB55F4BA5472242FF63F888A7C2F871" blockId="3.[151,1436,1917,2032]" pageId="3" pageNumber="86">
<emphasis id="B97E8359A5472242FF63F888A775F812" bold="true" box="[151,277,1917,1940]" pageId="3" pageNumber="86">FIGURE 2.</emphasis>
<taxonomicName id="4C0A24C8A5472242FEEAF88BA435F815" box="[286,597,1918,1940]" class="Malacostraca" family="Pseudothelphusidae" genus="Phallangothelphusa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="86" phylum="Arthropoda" rank="species" species="martensis" status="sp. nov.">
<emphasis id="B97E8359A5472242FEEAF88BA435F815" box="[286,597,1918,1940]" italics="true" pageId="3" pageNumber="86">Phallangothelphusa martensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D3E22A5472242FDABF88BA4BDF815" box="[607,733,1917,1940]" pageId="3" pageNumber="86" rank="species">
<emphasis id="B97E8359A5472242FDABF88BA4BDF815" bold="true" box="[607,733,1917,1940]" pageId="3" pageNumber="86">new species</emphasis>
</taxonomicNameLabel>
, male holotype, ICN-MHN-CR 2656. A, left first gonopod, caudal view; B, same, lateral view; C, same, mesial view; D, same, cephalic view; E, apex distal view; F, second gonopod, caudal view; G, distal part of second gonopod. 1, distal projection of mesial process; 2, curvature; 3, proximal projection of mesial process; 4, lateral process.
</paragraph>
</caption>
<caption id="DF750FC3A5402245FF63FBF6A2FFFBD7" httpUri="https://zenodo.org/record/214845/files/figure.png" pageId="4" pageNumber="87" targetBox="[163,1430,196,987]" targetPageId="4">
<paragraph id="8BB55F4BA5402245FF63FBF6A2FFFBD7" blockId="4.[151,1436,1027,1110]" pageId="4" pageNumber="87">
<emphasis id="B97E8359A5402245FF63FBF6A772FB99" bold="true" box="[151,274,1027,1050]" pageId="4" pageNumber="87">FIGURE 3.</emphasis>
<taxonomicName id="4C0A24C8A5402245FEECFBF6A42DFB98" box="[280,589,1027,1049]" class="Malacostraca" family="Pseudothelphusidae" genus="Phallangothelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="87" phylum="Arthropoda" rank="species" species="martensis" status="sp. nov.">
<emphasis id="B97E8359A5402245FEECFBF6A42DFB98" box="[280,589,1027,1049]" italics="true" pageId="4" pageNumber="87">Phallangothelphusa martensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D3E22A5402245FDA6FBF1A4AEFB9B" box="[594,718,1027,1050]" pageId="4" pageNumber="87" rank="species">
<emphasis id="B97E8359A5402245FDA6FBF1A4AEFB9B" bold="true" box="[594,718,1027,1050]" pageId="4" pageNumber="87">new species</emphasis>
</taxonomicNameLabel>
, male holotype, ICN-MHN-CR 2656, left first gonopod: A, apex distal view;
<taxonomicName id="4C0A24C8A5402245FF2EFBD7A4C2FBB9" authority="Zimmer, 1912" authorityName="Zimmer" authorityYear="1912" box="[218,674,1058,1080]" class="Malacostraca" family="Pseudothelphusidae" genus="Phallangothelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="87" phylum="Arthropoda" rank="species" species="dispar">
<emphasis id="B97E8359A5402245FF2EFBD7A791FBB9" box="[218,497,1058,1080]" italics="true" pageId="4" pageNumber="87">Phallangothelphusa dispar</emphasis>
(Zimmer, 1912)
</taxonomicName>
, ICN-MHN-CR 0 732, left first gonopod: B, apex, distal view. 1, distal projection of mesial process; 2, curvature; 3, proximal projection of mesial process; 4, lateral process.
</paragraph>
</caption>
<paragraph id="8BB55F4BA5402245FF31FAC2A705FA5E" blockId="4.[151,1436,1155,2010]" pageId="4" pageNumber="87">
<emphasis id="B97E8359A5402245FF31FAC2A76EFAD1" bold="true" box="[197,270,1335,1360]" pageId="4" pageNumber="87">Color.</emphasis>
The freshly‒alcohol‒preserved
<typeStatus id="54B1E1E9A5402245FD7DFAC2A48BFACE" box="[649,747,1335,1359]" pageId="4" pageNumber="87" type="holotype">holotype</typeStatus>
is brown (near Burn Umber, 22) with light specks (near Cinnamon, 123A) on the posterior dorsal side of the carapace. The walking legs dorsally, ventrally brown (Burn Umber, 22). The chelae brown dorsally (near Mikado Brown, 121C), light brown ventrally (near Sayal Brown, 223C). The ventral surface of the carapace is brown (near Olive Brown, 28) with light specks (near Sayal Brown, 223C) posteriorly.
</paragraph>
<paragraph id="8BB55F4BA5402245FF31FA1EA5B5F9CA" blockId="4.[151,1436,1155,2010]" pageId="4" pageNumber="87">
<emphasis id="B97E8359A5402245FF31FA1EA748F985" bold="true" box="[197,296,1515,1540]" pageId="4" pageNumber="87">Habitat.</emphasis>
The specimens were collected by hand, under not submerged rocks in secondary forest streams (
<quantity id="4CF2F2AEA5402245FA84FA1EA3FCF982" box="[1392,1436,1515,1539]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="87" unit="m" value="2.0">2 m</quantity>
average width, depth between
<quantity id="4CF2F2AEA5402245FE09F9FAA426F9A6" box="[509,582,1551,1575]" metricMagnitude="-1" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="87" unit="cm" value="30.0">30 cm</quantity>
and
<quantity id="4CF2F2AEA5402245FD70F9FAA4BAF9A6" box="[644,730,1551,1575]" metricMagnitude="0" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="87" unit="m" value="1.5">1.50 m</quantity>
). The streams were surrounded by a gallery forest and were characterized by low turbidity water, sandy clay soil and pebble substrate.
</paragraph>
</subSubSection>
<subSubSection id="C3100CC0A5402245FF31F9ADA2A0F9EE" box="[197,1216,1623,1648]" pageId="4" pageNumber="87" type="etymology">
<paragraph id="8BB55F4BA5402245FF31F9ADA2A0F9EE" blockId="4.[151,1436,1155,2010]" box="[197,1216,1623,1648]" pageId="4" pageNumber="87">
<emphasis id="B97E8359A5402245FF31F9ADA72CF9F1" bold="true" box="[197,332,1624,1648]" pageId="4" pageNumber="87">Etymology.</emphasis>
The species is named after the
<typeStatus id="54B1E1E9A5402245FD5FF9ADA4BBF9F1" box="[683,731,1624,1648]" pageId="4" pageNumber="87">type</typeStatus>
locality, Municipio San Martín,
<collectingCountry id="F31D1FDBA5402245FBBEF9A2A2DCF9EE" box="[1098,1212,1623,1647]" name="Colombia" pageId="4" pageNumber="87">Colombia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3100CC0A5402244FF31F989A572FE4E" lastPageId="5" lastPageNumber="88" pageId="4" pageNumber="87" type="discussion">
<paragraph id="8BB55F4BA5402244FF31F989A731FE06" blockId="4.[151,1436,1155,2010]" lastBlockId="5.[151,1436,151,464]" lastPageId="5" lastPageNumber="88" pageId="4" pageNumber="87">
<emphasis id="B97E8359A5402245FF31F989A759F915" bold="true" box="[197,313,1660,1684]" pageId="4" pageNumber="87">Remarks.</emphasis>
This new species closely resembles
<taxonomicName id="4C0A24C8A5402245FD20F98EA2DEF912" authority="Zimmer, 1912" authorityName="Zimmer" authorityYear="1912" box="[724,1214,1659,1683]" class="Malacostraca" family="Pseudothelphusidae" genus="Phallangothelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="87" phylum="Arthropoda" rank="species" species="dispar">
<emphasis id="B97E8359A5402245FD20F98EA265F912" box="[724,1029,1659,1683]" italics="true" pageId="4" pageNumber="87">Phallangothelphusa dispar</emphasis>
(Zimmer, 1912)
</taxonomicName>
(see
<bibRefCitation id="EF9B22BAA5402245FB0DF98EA3F8F912" author="Campos" box="[1273,1432,1659,1683]" pageId="4" pageNumber="87" refString="Campos, M. R. (2005) Freshwater crabs from Colombia. A taxonomic and distributional study. Academia Colombiana de Ciencias Exactas Fisicas y Naturales. Col. Jorge Alvarez Lleras, 24, 363 pp." type="book" year="2005">Campos 2005</bibRefCitation>
: figs. 40A–H). The main distinguishing feature between the two species is the first male gonopod. The gonopod of
<taxonomicName id="4C0A24C8A5402245FF6DF931A741F95A" box="[153,289,1732,1755]" class="Malacostraca" family="Pseudothelphusidae" genus="Phallangothelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="87" phylum="Arthropoda" rank="species" species="martensis">
<emphasis id="B97E8359A5402245FF6DF931A741F95A" box="[153,289,1732,1755]" italics="true" pageId="4" pageNumber="87">P. martensis</emphasis>
</taxonomicName>
has a laterally bent subdistal portion (straight in
<emphasis id="B97E8359A5402245FCB1F931A5C4F95A" box="[837,932,1731,1755]" italics="true" pageId="4" pageNumber="87">P. d i s p a r</emphasis>
); the caudal lobe is hourglass-like, narrower near the middle portion, with subdistal portion laterally bent (straight, with slight subdistal constriction in
<taxonomicName id="4C0A24C8A5402245FA73F91DA681F8A2" class="Malacostraca" family="Pseudothelphusidae" genus="Phallangothelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="87" phylum="Arthropoda" rank="species" species="dispar">
<emphasis id="B97E8359A5402245FA73F91DA681F8A2" italics="true" pageId="4" pageNumber="87">P. dispar</emphasis>
</taxonomicName>
); the lateral side has a subdistal constriction and a prominent ridge running parallel to the caudal lobe (only a subdistal slight depression in
<taxonomicName id="4C0A24C8A5402245FE08F8C5A400F8C6" box="[508,608,1839,1863]" class="Malacostraca" family="Pseudothelphusidae" genus="Phallangothelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="87" phylum="Arthropoda" rank="species" species="dispar">
<emphasis id="B97E8359A5402245FE08F8C5A400F8C6" box="[508,608,1839,1863]" italics="true" pageId="4" pageNumber="87">P. dispar</emphasis>
</taxonomicName>
); the curvature between the distal and proximal projections of the mesial process in greater than 90° (
<figureCitation id="133143CEA5402245FE21F8A3A479F8EF" box="[469,537,1878,1902]" captionStart="FIGURE 3" captionStartId="4.[151,250,1027,1050]" captionTargetBox="[163,1430,196,987]" captionTargetId="figure@4.[151,1436,193,1006]" captionTargetPageId="4" captionText="FIGURE 3. Phallangothelphusa martensis new species, male holotype, ICN-MHN-CR 2656, left first gonopod: A, apex distal view; Phallangothelphusa dispar (Zimmer, 1912), ICN-MHN-CR 0 732, left first gonopod: B, apex, distal view. 1, distal projection of mesial process; 2, curvature; 3, proximal projection of mesial process; 4, lateral process." httpUri="https://zenodo.org/record/214845/files/figure.png" pageId="4" pageNumber="87">Fig. 3</figureCitation>
A) (nearly 60° in
<taxonomicName id="4C0A24C8A5402245FD2BF8A2A525F8EF" box="[735,837,1878,1902]" class="Malacostraca" family="Pseudothelphusidae" genus="Phallangothelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="87" phylum="Arthropoda" rank="species" species="dispar">
<emphasis id="B97E8359A5402245FD2BF8A2A525F8EF" box="[735,837,1878,1902]" italics="true" pageId="4" pageNumber="87">P. dispar</emphasis>
</taxonomicName>
;
<figureCitation id="133143CEA5402245FCA2F8A3A5FAF8EF" box="[854,922,1878,1902]" captionStart="FIGURE 3" captionStartId="4.[151,250,1027,1050]" captionTargetBox="[163,1430,196,987]" captionTargetId="figure@4.[151,1436,193,1006]" captionTargetPageId="4" captionText="FIGURE 3. Phallangothelphusa martensis new species, male holotype, ICN-MHN-CR 2656, left first gonopod: A, apex distal view; Phallangothelphusa dispar (Zimmer, 1912), ICN-MHN-CR 0 732, left first gonopod: B, apex, distal view. 1, distal projection of mesial process; 2, curvature; 3, proximal projection of mesial process; 4, lateral process." httpUri="https://zenodo.org/record/214845/files/figure.png" pageId="4" pageNumber="87">Fig. 3</figureCitation>
B); the inner margin of the curvature slightly overlaps the mesial surface (
<figureCitation id="133143CEA5402245FE14F88FA443F813" box="[480,547,1914,1938]" captionStart="FIGURE 3" captionStartId="4.[151,250,1027,1050]" captionTargetBox="[163,1430,196,987]" captionTargetId="figure@4.[151,1436,193,1006]" captionTargetPageId="4" captionText="FIGURE 3. Phallangothelphusa martensis new species, male holotype, ICN-MHN-CR 2656, left first gonopod: A, apex distal view; Phallangothelphusa dispar (Zimmer, 1912), ICN-MHN-CR 0 732, left first gonopod: B, apex, distal view. 1, distal projection of mesial process; 2, curvature; 3, proximal projection of mesial process; 4, lateral process." httpUri="https://zenodo.org/record/214845/files/figure.png" pageId="4" pageNumber="87">Fig. 3</figureCitation>
A) (extensively overlaps the mesial surface, ending in a wide base spine in
<taxonomicName id="4C0A24C8A5402245FA73F88EA681F837" class="Malacostraca" family="Pseudothelphusidae" genus="Phallangothelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="87" phylum="Arthropoda" rank="species" species="dispar">
<emphasis id="B97E8359A5402245FA73F88EA681F837" italics="true" pageId="4" pageNumber="87">P. dispar</emphasis>
</taxonomicName>
;
<figureCitation id="133143CEA5402245FF06F86BA756F837" box="[242,310,1950,1974]" captionStart="FIGURE 3" captionStartId="4.[151,250,1027,1050]" captionTargetBox="[163,1430,196,987]" captionTargetId="figure@4.[151,1436,193,1006]" captionTargetPageId="4" captionText="FIGURE 3. Phallangothelphusa martensis new species, male holotype, ICN-MHN-CR 2656, left first gonopod: A, apex distal view; Phallangothelphusa dispar (Zimmer, 1912), ICN-MHN-CR 0 732, left first gonopod: B, apex, distal view. 1, distal projection of mesial process; 2, curvature; 3, proximal projection of mesial process; 4, lateral process." httpUri="https://zenodo.org/record/214845/files/figure.png" pageId="4" pageNumber="87">Fig. 3</figureCitation>
B); the distal projection is triangular, semi-acute distally, in caudal view, with a basal acute spine directed downwards, in distal view (
<figureCitation id="133143CEA5402245FDC8F837A4E0F85B" box="[572,640,1986,2010]" captionStart="FIGURE 3" captionStartId="4.[151,250,1027,1050]" captionTargetBox="[163,1430,196,987]" captionTargetId="figure@4.[151,1436,193,1006]" captionTargetPageId="4" captionText="FIGURE 3. Phallangothelphusa martensis new species, male holotype, ICN-MHN-CR 2656, left first gonopod: A, apex distal view; Phallangothelphusa dispar (Zimmer, 1912), ICN-MHN-CR 0 732, left first gonopod: B, apex, distal view. 1, distal projection of mesial process; 2, curvature; 3, proximal projection of mesial process; 4, lateral process." httpUri="https://zenodo.org/record/214845/files/figure.png" pageId="4" pageNumber="87">Fig. 3</figureCitation>
A) (rounded and devoid of the basal spine in
<emphasis id="B97E8359A5402245FB66F836A293F85B" box="[1170,1267,1986,2010]" italics="true" pageId="4" pageNumber="87">P. d i s p a r</emphasis>
;
<figureCitation id="133143CEA5402245FAFCF837A32CF85B" box="[1288,1356,1986,2010]" captionStart="FIGURE 3" captionStartId="4.[151,250,1027,1050]" captionTargetBox="[163,1430,196,987]" captionTargetId="figure@4.[151,1436,193,1006]" captionTargetPageId="4" captionText="FIGURE 3. Phallangothelphusa martensis new species, male holotype, ICN-MHN-CR 2656, left first gonopod: A, apex distal view; Phallangothelphusa dispar (Zimmer, 1912), ICN-MHN-CR 0 732, left first gonopod: B, apex, distal view. 1, distal projection of mesial process; 2, curvature; 3, proximal projection of mesial process; 4, lateral process." httpUri="https://zenodo.org/record/214845/files/figure.png" pageId="4" pageNumber="87">Fig. 3</figureCitation>
B); the proximal projection has a notch on basal portion in caudal view (
<figureCitation id="133143CEA5412244FC7CFF62A5ACFF2E" box="[904,972,151,175]" captionStart="FIGURE 3" captionStartId="4.[151,250,1027,1050]" captionTargetBox="[163,1430,196,987]" captionTargetId="figure@4.[151,1436,193,1006]" captionTargetPageId="4" captionText="FIGURE 3. Phallangothelphusa martensis new species, male holotype, ICN-MHN-CR 2656, left first gonopod: A, apex distal view; Phallangothelphusa dispar (Zimmer, 1912), ICN-MHN-CR 0 732, left first gonopod: B, apex, distal view. 1, distal projection of mesial process; 2, curvature; 3, proximal projection of mesial process; 4, lateral process." httpUri="https://zenodo.org/record/214845/files/figure.png" pageId="5" pageNumber="88">Fig. 3</figureCitation>
A) (a slight depression in
<taxonomicName id="4C0A24C8A5412244FB0DFF6CA33EFF31" box="[1273,1374,152,176]" class="Malacostraca" family="Pseudothelphusidae" genus="Phallangothelphusa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="88" phylum="Arthropoda" rank="species" species="dispar">
<emphasis id="B97E8359A5412244FB0DFF6CA33EFF31" box="[1273,1374,152,176]" italics="true" pageId="5" pageNumber="88">P. dispar</emphasis>
</taxonomicName>
;
<figureCitation id="133143CEA5412244FA9BFF62A6C7FF55" captionStart="FIGURE 3" captionStartId="4.[151,250,1027,1050]" captionTargetBox="[163,1430,196,987]" captionTargetId="figure@4.[151,1436,193,1006]" captionTargetPageId="4" captionText="FIGURE 3. Phallangothelphusa martensis new species, male holotype, ICN-MHN-CR 2656, left first gonopod: A, apex distal view; Phallangothelphusa dispar (Zimmer, 1912), ICN-MHN-CR 0 732, left first gonopod: B, apex, distal view. 1, distal projection of mesial process; 2, curvature; 3, proximal projection of mesial process; 4, lateral process." httpUri="https://zenodo.org/record/214845/files/figure.png" pageId="5" pageNumber="88">Fig. 3</figureCitation>
B); the lateral process is inwardly turned and parallel to the axis of the gonopod, approximately 60% of the length, the remaining 40% projected mesially (
<figureCitation id="133143CEA5412244FD90FF2AA4C8FF76" box="[612,680,223,247]" captionStart="FIGURE 3" captionStartId="4.[151,250,1027,1050]" captionTargetBox="[163,1430,196,987]" captionTargetId="figure@4.[151,1436,193,1006]" captionTargetPageId="4" captionText="FIGURE 3. Phallangothelphusa martensis new species, male holotype, ICN-MHN-CR 2656, left first gonopod: A, apex distal view; Phallangothelphusa dispar (Zimmer, 1912), ICN-MHN-CR 0 732, left first gonopod: B, apex, distal view. 1, distal projection of mesial process; 2, curvature; 3, proximal projection of mesial process; 4, lateral process." httpUri="https://zenodo.org/record/214845/files/figure.png" pageId="5" pageNumber="88">Fig. 3</figureCitation>
A) (perpendicular to the axis of gonopod and projected mesially, covering the proximal projection of the mesial process in
<taxonomicName id="4C0A24C8A5412244FCE8FEF0A51FFE9D" box="[796,895,260,284]" class="Malacostraca" family="Pseudothelphusidae" genus="Phallangothelphusa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="88" phylum="Arthropoda" rank="species" species="dispar">
<emphasis id="B97E8359A5412244FCE8FEF0A51FFE9D" box="[796,895,260,284]" italics="true" pageId="5" pageNumber="88">P. dispar</emphasis>
</taxonomicName>
;
<figureCitation id="133143CEA5412244FC79FEF1A5AFFE9D" box="[909,975,260,284]" captionStart="FIGURE 3" captionStartId="4.[151,250,1027,1050]" captionTargetBox="[163,1430,196,987]" captionTargetId="figure@4.[151,1436,193,1006]" captionTargetPageId="4" captionText="FIGURE 3. Phallangothelphusa martensis new species, male holotype, ICN-MHN-CR 2656, left first gonopod: A, apex distal view; Phallangothelphusa dispar (Zimmer, 1912), ICN-MHN-CR 0 732, left first gonopod: B, apex, distal view. 1, distal projection of mesial process; 2, curvature; 3, proximal projection of mesial process; 4, lateral process." httpUri="https://zenodo.org/record/214845/files/figure.png" pageId="5" pageNumber="88">Fig. 3</figureCitation>
B). Another morphological character that can be used to differentiate the two species is the length of the exognath of the third maxilliped, which corresponds to 0.9 times the length of the ischium in
<taxonomicName id="4C0A24C8A5412244FD93FEB8A490FEE5" box="[615,752,333,356]" class="Malacostraca" family="Pseudothelphusidae" genus="Phallangothelphusa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="88" phylum="Arthropoda" rank="species" species="martensis">
<emphasis id="B97E8359A5412244FD93FEB8A490FEE5" box="[615,752,333,356]" italics="true" pageId="5" pageNumber="88">P. martensis</emphasis>
</taxonomicName>
(
<figureCitation id="133143CEA5412244FCF4FEB9A521FEE5" box="[768,833,332,356]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1966]" captionTargetBox="[228,1335,193,1925]" captionTargetId="figure@2.[223,1358,193,1927]" captionTargetPageId="2" captionText="FIGURE 1. Phallangothelphusa martensis new species, male holotype, 1 male, cl 17.3 mm, cb 30.1 mm, ICN-MHN-CR 2656. A, habitus; B, frontal view of carapace; C, opening of left efferent branchial channel, external view; D, chela of large cheliped, external view; E, left third maxilliped, external view." httpUri="https://zenodo.org/record/214843/files/figure.png" pageId="5" pageNumber="88">Fig. 1</figureCitation>
E), whereas it is only 0.8 times in
<taxonomicName id="4C0A24C8A5412244FB3FFEB8A34FFEE5" box="[1227,1327,332,356]" class="Malacostraca" family="Pseudothelphusidae" genus="Phallangothelphusa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="88" phylum="Arthropoda" rank="species" species="dispar">
<emphasis id="B97E8359A5412244FB3FFEB8A34FFEE5" box="[1227,1327,332,356]" italics="true" pageId="5" pageNumber="88">P. dispar</emphasis>
</taxonomicName>
(
<bibRefCitation id="EF9B22BAA5412244FACAFEB9A725FE06" author="Campos" pageId="5" pageNumber="88" refString="Campos, M. R. (2005) Freshwater crabs from Colombia. A taxonomic and distributional study. Academia Colombiana de Ciencias Exactas Fisicas y Naturales. Col. Jorge Alvarez Lleras, 24, 363 pp." type="book" year="2005">Campos 2005: fig. 40 E</bibRefCitation>
).
</paragraph>
<paragraph id="8BB55F4BA5412244FF33FE60A572FE4E" blockId="5.[151,1436,151,464]" pageId="5" pageNumber="88">In some of the specimens examined the length of the lateral process was shorter than the proximal projection of the mesial process and the gap between them was minor.</paragraph>
</subSubSection>
</treatment>
</document>
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