treatments-xml/data/03/D7/87/03D787EE8F223569FF13FADDFDFB5C3E.xml

<document id="73426E427C82E209F53357CCA4712E31" ID="10.11646/zootaxa.4243.3.2SLASH10507" ID-CLB-Dataset="36267" ID-DOI="10.11646/zootaxa.4243.3.2" ID-GBIF-Dataset="934dbc23-a710-49b6-8f09-4a842f4258f6" ID-ISSN="1175-5326" ID-Zenodo-Dep="400114" ID-ZooBank="6771949E-C3B2-4A3F-ABFD-FA3D90454E9A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1489653541299" checkinUser="plazi" docAuthor="Loughman, Zachary J., Welsh, Stuart A. &amp; Thoma, Roger F." docDate="2017" docId="03D787EE8F223569FF13FADDFDFB5C3E" docLanguage="en" docName="zootaxa.4243.3.2SLASH10507.pdf" docOrigin="Zootaxa 4243 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cambarus (Cambarus) appalachiensis Loughman, Welsh &amp; Thoma, 2017, new species" docType="treatment" docVersion="9" lastPageNumber="446" masterDocId="FFEEFF968F213567FF84FF81FF895E26" masterDocTitle="Cambarus (C.) appalachiensis, a new species of crayfish (Decapoda: Cambaridae) from the New River Basin of Virginia and West Virginia, USA" masterLastPageNumber="454" masterPageNumber="432" pageNumber="435" updateTime="1698438004752" updateUser="ExternalLinkService">
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<mods:title id="18299BB1A5624BAC34C1F79BC6E515B3">Cambarus (C.) appalachiensis, a new species of crayfish (Decapoda: Cambaridae) from the New River Basin of Virginia and West Virginia, USA</mods:title>
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<mods:namePart id="E3A7D62F702F8DBE4C3E7F7C8A95F275">Loughman, Zachary J.</mods:namePart>
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<subSubSection id="C36465738F223564FF13FADDFD725B50" box="[151,763,1372,1399]" pageId="3" pageNumber="435" type="nomenclature">
<paragraph id="8BC136F88F223564FF13FADDFD725B50" blockId="3.[151,763,1372,1399]" box="[151,763,1372,1399]" pageId="3" pageNumber="435">
<heading id="D08981948F223564FF13FADDFD725B50" bold="true" box="[151,763,1372,1399]" fontSize="11" level="1" pageId="3" pageNumber="435" reason="1">
<emphasis id="B90AEAEA8F223564FF13FADDFD725B50" bold="true" box="[151,763,1372,1399]" pageId="3" pageNumber="435">
<taxonomicName id="4C7E4D7B8F223564FF13FADDFDEE5B50" ID-CoL="7RTVV" box="[151,615,1372,1399]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="435" phylum="Arthropoda" rank="species" species="appalachiensis" status="sp. nov." subGenus="Cambarus">
<emphasis id="B90AEAEA8F223564FF13FADDFE9D5B50" bold="true" box="[151,276,1372,1398]" italics="true" pageId="3" pageNumber="435">Cambarus</emphasis>
(
<emphasis id="B90AEAEA8F223564FEA1FADDFE2B5B50" bold="true" box="[293,418,1372,1398]" italics="true" pageId="3" pageNumber="435">Cambarus</emphasis>
) 
<emphasis id="B90AEAEA8F223564FE30FADDFDEE5B50" bold="true" box="[436,615,1372,1398]" italics="true" pageId="3" pageNumber="435">appalachiensis</emphasis>
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<subSubSection id="C36465738F223563FF13FA25FB3F5F96" lastPageId="4" lastPageNumber="436" pageId="3" pageNumber="435" type="diagnosis">
<paragraph id="8BC136F88F223563FF13FA25FB3F5F96" blockId="3.[151,1437,1444,2015]" lastBlockId="4.[151,1437,151,1875]" lastPageId="4" lastPageNumber="436" pageId="3" pageNumber="435">
<emphasis id="B90AEAEA8F223564FF13FA25FE985B9D" bold="true" box="[151,273,1444,1469]" pageId="3" pageNumber="435">Diagnosis.</emphasis>
Body and eyes pigmented. Rostrum broad, moderately excavated, and deflected anteriorly, margins thickened, parallel though rarely sub-parallel; converging to acumen abruptly at terminance of rostral margins. Acumen distinctly triangular with prominent dorsally deflected spiniform tubercle at terminus. Areola 2.2–5.1 (= 3.4, 
<emphasis id="B90AEAEA8F223564FF4EF993FF51580F" box="[202,216,1554,1577]" italics="true" pageId="3" pageNumber="435">n</emphasis>
= 98, 
<emphasis id="B90AEAEA8F223564FEA1F990FEBD580C" bold="true" box="[293,308,1553,1578]" pageId="3" pageNumber="435">σ</emphasis>
= 0.6) times as long as wide with 5–8 (usually 7) punctations across narrowest point. No cervical spines; 4–7 ( = 6.1, SE = 0.8) tubercles present. Mandibular, branchiostegal, and orbital regions of carapace with well-developed tubercles. Postorbital ridges short; pronounced, spiniform, dorsally deflected cephalic tubercle present in juveniles and subadults; adult postorbital ridge terminating in reduced to prominent dorsally deflected cephalic tubercle. Suborbital angle acute. Total carapace length (TCL) 1.8–2.1 ( = 1.9, 
<emphasis id="B90AEAEA8F223564FB0AF925FB15589D" box="[1166,1180,1700,1723]" italics="true" pageId="3" pageNumber="435">n</emphasis>
= 98, 
<emphasis id="B90AEAEA8F223564FB6DF925FA81589D" box="[1257,1288,1700,1723]" italics="true" pageId="3" pageNumber="435">SE</emphasis>
= 0.1) times longer than width. Form I and II males possessing hook on ischium of third pereopod only; hook gently curved at apex, overarching basioischial joint in form I males, not reaching basioischial joint in form II males; hooks not opposed by tubercle on basis. Mesial surface of chela nearly always possesses a single mesial row of tubercles; but rarely (3% of individuals; n = 98) an incomplete second row of tubercles also present; mesial most row with 6–10 ( = 7.9, 
<emphasis id="B90AEAEA8F223564FE83F8D8FE9C5956" box="[263,277,1881,1904]" italics="true" pageId="3" pageNumber="435">n</emphasis>
= 98, 
<emphasis id="B90AEAEA8F223564FEE0F8D8FE0A5956" box="[356,387,1881,1904]" italics="true" pageId="3" pageNumber="435">SE</emphasis>
= 0.5) tubercles, second incomplete row when present with 2–4 (= 2.2, 
<emphasis id="B90AEAEA8F223564FB64F8D8FB675956" box="[1248,1262,1881,1904]" italics="true" pageId="3" pageNumber="435">n</emphasis>
= 3, 
<emphasis id="B90AEAEA8F223564FAAAF8D8FAC45956" box="[1326,1357,1881,1904]" italics="true" pageId="3" pageNumber="435">SE</emphasis>
= 0.6) tubercles. Ventral surface of chelae with 1–3 ( = 1.9, 
<emphasis id="B90AEAEA8F223564FC99F8FEFCA259B0" box="[797,811,1919,1942]" italics="true" pageId="3" pageNumber="435">n</emphasis>
= 98, 
<emphasis id="B90AEAEA8F223564FCFAF8FEFC1459B0" box="[894,925,1919,1942]" italics="true" pageId="3" pageNumber="435">SE</emphasis>
= 0.7) subpalmar tubercles near the dactyl/ propodus junction. Dorsomedian ridge of fixed finger of propodus pronounced. Lateral impression at base of fixed finger shallow (61% of individuals) or absent (39% of individuals). Dactyl and fixed finger each with sharp corneous tip. Form I male palm length 64.3–77.1% ( = 69.9%, 
<emphasis id="B90AEAEA8F253563FCF8FF18FC035E96" box="[892,906,153,176]" italics="true" pageId="4" pageNumber="436">n</emphasis>
= 32, 
<emphasis id="B90AEAEA8F253563FC53FF18FC7F5E96" box="[983,1014,153,176]" italics="true" pageId="4" pageNumber="436">SE</emphasis>
= 2.8%) of palm width, form I male palm length 27.1–31.3% ( = 29.6%, 
<emphasis id="B90AEAEA8F253563FDD6FF3EFDE95EF0" box="[594,608,191,214]" italics="true" pageId="4" pageNumber="436">n</emphasis>
= 32, 
<emphasis id="B90AEAEA8F253563FD35FF3EFD595EF0" box="[689,720,191,214]" italics="true" pageId="4" pageNumber="436">SE</emphasis>
= 1.2%) of total propodus length; female dactyl length 55.2– 64.1% ( = 61.5%, 
<emphasis id="B90AEAEA8F253563FEFDFF65FE0E5EDD" box="[377,391,228,251]" italics="true" pageId="4" pageNumber="436">n</emphasis>
= 35, 
<emphasis id="B90AEAEA8F253563FE55FF65FE795EDD" box="[465,496,228,251]" italics="true" pageId="4" pageNumber="436">SE</emphasis>
= 2.2%) of total propodus length. First pleopod of form I male with short terminal elements. Central projection not tapering distally; recurved&gt;90° to main shaft, with distinct subapical notch. Mesial process directed 90° to shaft, bent cephalolaterally; inflated proximally, tapering to a point distinctly caudal to slighty cephalic to terminance of central projection and not projecting beyond the edge of the margin of gonopod shaft. Annulus ventralis immovable; distinctly asymmetrical caudally; cephalic portion with median trough leading to strongly sculptured central fossa; exaggerated “S” bend in sinus terminating at caudal edge.
</paragraph>
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<subSubSection id="C36465738F253560FF43FE3DFCDA581A" lastPageId="7" lastPageNumber="439" pageId="4" pageNumber="436" type="description">
<paragraph id="8BC136F88F253563FF43FE3DFB285815" blockId="4.[151,1437,151,1875]" pageId="4" pageNumber="436">
<emphasis id="B90AEAEA8F253563FF43FE3DFD355FF3" bold="true" box="[199,700,444,469]" pageId="4" pageNumber="436">Description of holotypic male, form I.</emphasis>
(
<figureCitation id="13452A7D8F253563FD51FE3DFCAB5FF2" box="[725,802,444,469]" captionStart="FIGURE 2" captionStartId="6.[151,250,1877,1899]" captionTargetBox="[172,1406,196,1842]" captionTargetId="figure@6.[172,1406,196,1842]" captionTargetPageId="6" captionText="FIGURE 2. Cambarus appalachiensis, new species: (A.) Lateral view of carapace; (B.) Lateral and (C.) mesial view of form I gonopod; (D.) Lateral and (E.) mesial view of form II gonopod; (F.) Annulus ventralis; (G.) Dorsal view of carapace; (H.) Right antennal scale; (I.) Dorsal view of right carpus and chelae; (J.) epistome. A-C and G-I from holotype; F from allotype; D- E from morphotypes." httpUri="https://zenodo.org/record/400116/files/figure.png" pageId="4" pageNumber="436">Fig. 2</figureCitation>
A–C, G–J; 
<tableCitation id="C6FC03438F253563FC4CFE3DFBAD5FF2" box="[968,1060,444,469]" captionStart="TABLE 1" captionStartId="5.[151,239,152,175]" captionTargetBox="[158,1278,202,1299]" captionTargetPageId="5" captionText="TABLE 1. Morphological measurements (mm) of Cambarus appalachiensis, new species." httpUri="http://table.plazi.org/id/DF0166708F243562FF13FF19FBD75E89" pageId="4" pageNumber="436" tableUuid="DF0166708F243562FF13FF19FBD75E89">Table 1</tableCitation>
). Body somewhat compressed dorsoventrally (
<figureCitation id="13452A7D8F253563FEC9FE5EFE1C5FDE" box="[333,405,479,504]" captionStart="FIGURE 2" captionStartId="6.[151,250,1877,1899]" captionTargetBox="[172,1406,196,1842]" captionTargetId="figure@6.[172,1406,196,1842]" captionTargetPageId="6" captionText="FIGURE 2. Cambarus appalachiensis, new species: (A.) Lateral view of carapace; (B.) Lateral and (C.) mesial view of form I gonopod; (D.) Lateral and (E.) mesial view of form II gonopod; (F.) Annulus ventralis; (G.) Dorsal view of carapace; (H.) Right antennal scale; (I.) Dorsal view of right carpus and chelae; (J.) epistome. A-C and G-I from holotype; F from allotype; D- E from morphotypes." httpUri="https://zenodo.org/record/400116/files/figure.png" pageId="4" pageNumber="436">Fig. 2</figureCitation>
A); thoracic section of carapace slightly wider than abdomen. Carapace depth less than carapace width at caudodorsal margin of cervical groove. Total carapace length 
<quantity id="4C869B1D8F253563FBB2FD85FB295C3A" box="[1078,1184,516,540]" metricMagnitude="-2" metricUnit="m" metricValue="3.81" pageId="4" pageNumber="436" unit="mm" value="38.1">38.1 mm</quantity>
; post orbital carapace length 
<quantity id="4C869B1D8F253563FF61FDA9FEC45C19" box="[229,333,552,576]" metricMagnitude="-2" metricUnit="m" metricValue="3.21" pageId="4" pageNumber="436" unit="mm" value="32.1">32.1 mm</quantity>
. Areola 2.8 times longer than wide, with 7 punctations across narrowest part (
<figureCitation id="13452A7D8F253563FB42FDA6FA835C66" box="[1222,1290,551,576]" captionStart="FIGURE 2" captionStartId="6.[151,250,1877,1899]" captionTargetBox="[172,1406,196,1842]" captionTargetId="figure@6.[172,1406,196,1842]" captionTargetPageId="6" captionText="FIGURE 2. Cambarus appalachiensis, new species: (A.) Lateral view of carapace; (B.) Lateral and (C.) mesial view of form I gonopod; (D.) Lateral and (E.) mesial view of form II gonopod; (F.) Annulus ventralis; (G.) Dorsal view of carapace; (H.) Right antennal scale; (I.) Dorsal view of right carpus and chelae; (J.) epistome. A-C and G-I from holotype; F from allotype; D- E from morphotypes." httpUri="https://zenodo.org/record/400116/files/figure.png" pageId="4" pageNumber="436">Fig. 2</figureCitation>
G); length of areola 34.9% of TCL (41.4% of postorbital carapace length (PCL). Rostrum weakly excavated, more so anteriorly than posteriorly; margins thickened and parallel; acumen margins noticeably thinner than rostral margin forming distinct 90 0 break from rostrum margins, continuous to base of acumen; floor of rostrum with numerous punctations. Rostrum 1.7 times longer than wide. Acumen distinctly triangular, ending in dorsally deflected corneous tip (
<figureCitation id="13452A7D8F253563FEB2FD5FFEF55CD1" box="[310,380,734,759]" captionStart="FIGURE 2" captionStartId="6.[151,250,1877,1899]" captionTargetBox="[172,1406,196,1842]" captionTargetId="figure@6.[172,1406,196,1842]" captionTargetPageId="6" captionText="FIGURE 2. Cambarus appalachiensis, new species: (A.) Lateral view of carapace; (B.) Lateral and (C.) mesial view of form I gonopod; (D.) Lateral and (E.) mesial view of form II gonopod; (F.) Annulus ventralis; (G.) Dorsal view of carapace; (H.) Right antennal scale; (I.) Dorsal view of right carpus and chelae; (J.) epistome. A-C and G-I from holotype; F from allotype; D- E from morphotypes." httpUri="https://zenodo.org/record/400116/files/figure.png" pageId="4" pageNumber="436">Fig. 2</figureCitation>
A). Postorbital ridges well developed, terminating in dorsally deflected spiniform tubercles. Suborbital angle acute, with tubercle (
<figureCitation id="13452A7D8F253563FDD5FC83FD1E5D3D" box="[593,663,770,795]" captionStart="FIGURE 2" captionStartId="6.[151,250,1877,1899]" captionTargetBox="[172,1406,196,1842]" captionTargetId="figure@6.[172,1406,196,1842]" captionTargetPageId="6" captionText="FIGURE 2. Cambarus appalachiensis, new species: (A.) Lateral view of carapace; (B.) Lateral and (C.) mesial view of form I gonopod; (D.) Lateral and (E.) mesial view of form II gonopod; (F.) Annulus ventralis; (G.) Dorsal view of carapace; (H.) Right antennal scale; (I.) Dorsal view of right carpus and chelae; (J.) epistome. A-C and G-I from holotype; F from allotype; D- E from morphotypes." httpUri="https://zenodo.org/record/400116/files/figure.png" pageId="4" pageNumber="436">Fig. 2</figureCitation>
A). Cervical spine absent; several weakly to moderately developed tubercles present. Mandibular, branchiostegal, and orbital regions of carapace punctated with well-developed tubercles; greatest tubercle density in hepatic region. Abdomen slightly longer than carapace, pleura rounded cephaloventrally, angled caudoventrally. Cephalic section of telson with 2 large spines in each caudolateral corner. Proximal podomere of uropod with distal spine on mesial lobe; mesial ramus of uropod with median ridge ending distally in distomedian spine not overreaching margin of ramus; laterodistal spine pronounced. Distal margin of proximal segment of lateral ramus of right uropod having 14 immovable, small spines and 1 lateral, large, movable spine. Cephalomedian lobe of epistome subtriangular, zygoma moderately arched (
<figureCitation id="13452A7D8F253563FBBDFC7FFBF45A31" box="[1081,1149,1022,1047]" captionStart="FIGURE 2" captionStartId="6.[151,250,1877,1899]" captionTargetBox="[172,1406,196,1842]" captionTargetId="figure@6.[172,1406,196,1842]" captionTargetPageId="6" captionText="FIGURE 2. Cambarus appalachiensis, new species: (A.) Lateral view of carapace; (B.) Lateral and (C.) mesial view of form I gonopod; (D.) Lateral and (E.) mesial view of form II gonopod; (F.) Annulus ventralis; (G.) Dorsal view of carapace; (H.) Right antennal scale; (I.) Dorsal view of right carpus and chelae; (J.) epistome. A-C and G-I from holotype; F from allotype; D- E from morphotypes." httpUri="https://zenodo.org/record/400116/files/figure.png" pageId="4" pageNumber="436">Fig. 2</figureCitation>
J); cephalolateral margins thickened, forming sharp angle at junction with endostyle; main body possessing prominent cephalomedian fovea. Antennal scale broadest anteriorly; lateral margin thickened, terminating in large corneous spine; (
<figureCitation id="13452A7D8F253563FB6AFBC7FABB5A79" box="[1262,1330,1094,1119]" captionStart="FIGURE 2" captionStartId="6.[151,250,1877,1899]" captionTargetBox="[172,1406,196,1842]" captionTargetId="figure@6.[172,1406,196,1842]" captionTargetPageId="6" captionText="FIGURE 2. Cambarus appalachiensis, new species: (A.) Lateral view of carapace; (B.) Lateral and (C.) mesial view of form I gonopod; (D.) Lateral and (E.) mesial view of form II gonopod; (F.) Annulus ventralis; (G.) Dorsal view of carapace; (H.) Right antennal scale; (I.) Dorsal view of right carpus and chelae; (J.) epistome. A-C and G-I from holotype; F from allotype; D- E from morphotypes." httpUri="https://zenodo.org/record/400116/files/figure.png" pageId="4" pageNumber="436">Fig. 2</figureCitation>
H). Right antennal scale 
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long, 
<quantity id="4C869B1D8F253563FE58FBEBFDBA5AA4" box="[476,563,1130,1155]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="4" pageNumber="436" unit="mm" value="2.5">2.5 mm</quantity>
wide. Tip of right antenna reaching middle of telson when appressed. Mesial surface of right chela with one well-formed row of 8 tubercles (
<figureCitation id="13452A7D8F253563FCE4FB0FFC2B5A81" box="[864,930,1166,1191]" captionStart="FIGURE 3" captionStartId="7.[151,250,943,965]" captionTargetBox="[151,1436,193,922]" captionTargetId="figure@7.[151,1436,193,922]" captionTargetPageId="7" captionText="FIGURE 3. Cambarus appalachiensis morphotype in life." httpUri="https://zenodo.org/record/400117/files/figure.png" pageId="4" pageNumber="436">Fig. 3</figureCitation>
K). Palm length 68.1 % of palm width; depth of palm 
<quantity id="4C869B1D8F253563FE87FB33FEEA5AEC" box="[259,355,1202,1227]" metricMagnitude="-3" metricUnit="m" metricValue="9.6" pageId="4" pageNumber="436" unit="mm" value="9.6">9.6 mm</quantity>
. Ventral surface of palm lacking subpalmar tubercles. Dorsal longitudinal ridge of dactyl developed and possessing moderate sized tubercles (
<figureCitation id="13452A7D8F253563FC9FFB57FCE15AC9" box="[795,872,1238,1263]" captionStart="FIGURE 2" captionStartId="6.[151,250,1877,1899]" captionTargetBox="[172,1406,196,1842]" captionTargetId="figure@6.[172,1406,196,1842]" captionTargetPageId="6" captionText="FIGURE 2. Cambarus appalachiensis, new species: (A.) Lateral view of carapace; (B.) Lateral and (C.) mesial view of form I gonopod; (D.) Lateral and (E.) mesial view of form II gonopod; (F.) Annulus ventralis; (G.) Dorsal view of carapace; (H.) Right antennal scale; (I.) Dorsal view of right carpus and chelae; (J.) epistome. A-C and G-I from holotype; F from allotype; D- E from morphotypes." httpUri="https://zenodo.org/record/400116/files/figure.png" pageId="4" pageNumber="436">
Fig. 
<date id="FFC010388F253563FCDCFB56FCE15AC9" box="[856,872,1239,1263]" pageId="4" pageNumber="436">2</date>
</figureCitation>
I); dactyl terminating in large corneous spine. Dorsomedian ridge of fixed finger of propodus weakly developed. Weakly developed lateral impression at the junction of fixed finger and palm; numerous setiferous punctations present. Dactyl and fixed finger with sharp, corneous tip. All measurements and counts from right chela. Carpus with prominent dorsal furrow (
<figureCitation id="13452A7D8F253563FA84FAC3FACD5B7D" box="[1280,1348,1346,1371]" captionStart="FIGURE 2" captionStartId="6.[151,250,1877,1899]" captionTargetBox="[172,1406,196,1842]" captionTargetId="figure@6.[172,1406,196,1842]" captionTargetPageId="6" captionText="FIGURE 2. Cambarus appalachiensis, new species: (A.) Lateral view of carapace; (B.) Lateral and (C.) mesial view of form I gonopod; (D.) Lateral and (E.) mesial view of form II gonopod; (F.) Annulus ventralis; (G.) Dorsal view of carapace; (H.) Right antennal scale; (I.) Dorsal view of right carpus and chelae; (J.) epistome. A-C and G-I from holotype; F from allotype; D- E from morphotypes." httpUri="https://zenodo.org/record/400116/files/figure.png" pageId="4" pageNumber="436">
Fig. 
<date id="FFC010388F253563FAB1FAC2FACD5B7D" box="[1333,1348,1347,1371]" pageId="4" pageNumber="436">2</date>
</figureCitation>
I) and 2 weak dorsomesial tubercles; rest of surface with setiferous punctations; mesial margin with large, procurved spine at about midlength, and one reduced proximal tubercle. Distodorsal surface of merus with 9 spiniform tubercles; ventrolateral ridge with 2 small spines and one large, corneous distal spine; ventromesial ridge with two welldeveloped spines. Carapace depth less than width. Hook on ischium of third pereopod only; hook gently curved at apex, overarching basioischial joint, not opposed by tubercle on basis. Form I gonopod as described in diagnosis (
<figureCitation id="13452A7D8F253563FF1BF99BFF6A5815" box="[159,227,1562,1587]" captionStart="FIGURE 2" captionStartId="6.[151,250,1877,1899]" captionTargetBox="[172,1406,196,1842]" captionTargetId="figure@6.[172,1406,196,1842]" captionTargetPageId="6" captionText="FIGURE 2. Cambarus appalachiensis, new species: (A.) Lateral view of carapace; (B.) Lateral and (C.) mesial view of form I gonopod; (D.) Lateral and (E.) mesial view of form II gonopod; (F.) Annulus ventralis; (G.) Dorsal view of carapace; (H.) Right antennal scale; (I.) Dorsal view of right carpus and chelae; (J.) epistome. A-C and G-I from holotype; F from allotype; D- E from morphotypes." httpUri="https://zenodo.org/record/400116/files/figure.png" pageId="4" pageNumber="436">Fig. 2</figureCitation>
B–C); tip reaching anterior margin of fourth caudomesial boss when abdomen flexed.
</paragraph>
<paragraph id="8BC136F88F253563FF43F9BFFB8E5975" blockId="4.[151,1437,151,1875]" pageId="4" pageNumber="436">
<emphasis id="B90AEAEA8F253563FF43F9BFFDB55871" bold="true" box="[199,572,1598,1623]" pageId="4" pageNumber="436">Description of allotypic female.</emphasis>
(
<figureCitation id="13452A7D8F253563FDCFF9BFFD075871" box="[587,654,1598,1623]" captionStart="FIGURE 2" captionStartId="6.[151,250,1877,1899]" captionTargetBox="[172,1406,196,1842]" captionTargetId="figure@6.[172,1406,196,1842]" captionTargetPageId="6" captionText="FIGURE 2. Cambarus appalachiensis, new species: (A.) Lateral view of carapace; (B.) Lateral and (C.) mesial view of form I gonopod; (D.) Lateral and (E.) mesial view of form II gonopod; (F.) Annulus ventralis; (G.) Dorsal view of carapace; (H.) Right antennal scale; (I.) Dorsal view of right carpus and chelae; (J.) epistome. A-C and G-I from holotype; F from allotype; D- E from morphotypes." httpUri="https://zenodo.org/record/400116/files/figure.png" pageId="4" pageNumber="436">Fig. 2</figureCitation>
F, 
<tableCitation id="C6FC03438F253563FD2EF9BFFD7E5871" box="[682,759,1598,1623]" captionStart="TABLE 1" captionStartId="5.[151,239,152,175]" captionTargetBox="[158,1278,202,1299]" captionTargetPageId="5" captionText="TABLE 1. Morphological measurements (mm) of Cambarus appalachiensis, new species." httpUri="http://table.plazi.org/id/DF0166708F243562FF13FF19FBD75E89" pageId="4" pageNumber="436" tableUuid="DF0166708F243562FF13FF19FBD75E89">Table 1</tableCitation>
).––Differing from 
<typeStatus id="54C5885A8F253563FC52F9BFFBB15871" box="[982,1080,1598,1623]" pageId="4" pageNumber="436" type="holotype">holotype</typeStatus>
in following respects; carapace height less than carapace width (16.9 and 
<quantity id="4C869B1D8F253563FDFBF9E2FD6E585C" box="[639,743,1635,1659]" metricMagnitude="-2" metricUnit="m" metricValue="2.11" pageId="4" pageNumber="436" unit="mm" value="21.1">21.1 mm</quantity>
, respectively); TCL 
<quantity id="4C869B1D8F253563FC52F9E3FBB6585C" box="[982,1087,1634,1659]" metricMagnitude="-2" metricUnit="m" metricValue="3.96" pageId="4" pageNumber="436" unit="mm" value="39.6">39.6 mm</quantity>
, PCL 
<quantity id="4C869B1D8F253563FB0EF9E3FB7A585C" box="[1162,1267,1634,1659]" metricMagnitude="-2" metricUnit="m" metricValue="3.35" pageId="4" pageNumber="436" unit="mm" value="33.5">33.5 mm</quantity>
. Areola length 35.1% of TCL (41.4% of PCL), 3.0 times as long as wide. Posterior portion of rostrum more excavated than anterior portion; rostrum 1.7 times longer than wide. Abdomen length 
<quantity id="4C869B1D8F253563FC49F92AFBB158E4" box="[973,1080,1707,1731]" metricMagnitude="-2" metricUnit="m" metricValue="4.11" pageId="4" pageNumber="436" unit="mm" value="41.1">41.1 mm</quantity>
. Mesial surface of chelae with single row of 8 tubercles. Palm length (10.0 mm) 62.5% of palm width (16.0 mm); depth of palm 
<quantity id="4C869B1D8F253563FA94F94EFAE258C1" box="[1296,1387,1743,1767]" metricMagnitude="-3" metricUnit="m" metricValue="9.2" pageId="4" pageNumber="436" unit="mm" value="9.2">9.2 mm</quantity>
. All measurements and counts from right chela. Antennal scale 
<quantity id="4C869B1D8F253563FCD7F973FC27592C" box="[851,942,1778,1803]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="4" pageNumber="436" unit="mm" value="5.7">5.7 mm</quantity>
long, 
<quantity id="4C869B1D8F253563FC7AF972FBD0592C" box="[1022,1113,1779,1803]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="4" pageNumber="436" unit="mm" value="2.3">2.3 mm</quantity>
wide. Annulus ventralis as described in diagnosis (
<figureCitation id="13452A7D8F253563FE27F897FE615909" box="[419,488,1814,1839]" captionStart="FIGURE 2" captionStartId="6.[151,250,1877,1899]" captionTargetBox="[172,1406,196,1842]" captionTargetId="figure@6.[172,1406,196,1842]" captionTargetPageId="6" captionText="FIGURE 2. Cambarus appalachiensis, new species: (A.) Lateral view of carapace; (B.) Lateral and (C.) mesial view of form I gonopod; (D.) Lateral and (E.) mesial view of form II gonopod; (F.) Annulus ventralis; (G.) Dorsal view of carapace; (H.) Right antennal scale; (I.) Dorsal view of right carpus and chelae; (J.) epistome. A-C and G-I from holotype; F from allotype; D- E from morphotypes." httpUri="https://zenodo.org/record/400116/files/figure.png" pageId="4" pageNumber="436">Fig. 2</figureCitation>
F); width of postannular sclerite half total width of annulus ventralis; first pleopods uniramous, reaching central region of annulus ventralis when abdomen flexed.
</paragraph>
<caption id="DF0166708F243562FF13FF19FBD75E89" ID-Table-UUID="DF0166708F243562FF13FF19FBD75E89" box="[151,1118,151,175]" httpUri="http://table.plazi.org/id/DF0166708F243562FF13FF19FBD75E89" pageId="5" pageNumber="437" targetBox="[158,1278,202,1299]" targetIsTable="true" targetPageId="5">
<paragraph id="8BC136F88F243562FF13FF19FBD75E89" blockId="5.[151,1118,151,175]" box="[151,1118,151,175]" pageId="5" pageNumber="437">
<emphasis id="B90AEAEA8F243562FF13FF19FE835E89" bold="true" box="[151,266,152,175]" pageId="5" pageNumber="437">TABLE 1.</emphasis>
Morphological measurements (mm) of 
<taxonomicName id="4C7E4D7B8F243562FD30FF16FC405E89" box="[692,969,151,175]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="437" phylum="Arthropoda" rank="species" species="appalachiensis" status="sp. nov.">
<emphasis id="B90AEAEA8F243562FD30FF16FC405E89" box="[692,969,151,175]" italics="true" pageId="5" pageNumber="437">Cambarus appalachiensis</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A23957918F243562FC52FF19FBD35E89" box="[982,1114,152,175]" pageId="5" pageNumber="437" rank="species">new species</taxonomicNameLabel>
.
</paragraph>
</caption>
<paragraph id="8BC136F88F243562FD1DFF4BFB285B35" pageId="5" pageNumber="437">
<table id="F97EC4588F24CA98FF1AFF4BFB775B35" box="[158,1278,202,1299]" gridcols="4" gridrows="24" pageId="5" pageNumber="437">
<tr id="354E34BA8F24CA98FF1AFF4BFB775EC6" box="[158,1278,202,224]" gridrow="0" pageId="5" pageNumber="437" rowspan-0="1">
<th id="769F5DC68F24CA98FD1DFF4BFD7E5EC6" box="[665,759,202,224]" gridcol="1" gridrow="0" pageId="5" pageNumber="437">Holotype</th>
<th id="769F5DC68F24CA98FC09FF4BFC6F5EC6" box="[909,998,202,224]" gridcol="2" gridrow="0" pageId="5" pageNumber="437">Allotype</th>
<th id="769F5DC68F24CA98FB05FF4BFB775EC6" box="[1153,1278,202,224]" gridcol="3" gridrow="0" pageId="5" pageNumber="437">Morphotype</th>
</tr>
<tr id="354E34BA8F24CA98FF1AFF75FB775F2C" box="[158,1278,244,266]" gridrow="1" pageId="5" pageNumber="437" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="769F5DC68F24CA98FF1AFF75FE4B5F2C" box="[158,450,244,266]" gridcol="0" gridrow="1" pageId="5" pageNumber="437">Carapace</th>
</tr>
<tr id="354E34BA8F24CA98FF1AFE9CFB775F15" box="[158,1278,285,307]" gridrow="2" pageId="5" pageNumber="437">
<th id="769F5DC68F24CA98FF1AFE9CFE4B5F15" box="[158,450,285,307]" gridcol="0" gridrow="2" pageId="5" pageNumber="437">Total carapace length</th>
<td id="769F5DC68F24CA98FD1DFE9CFD7E5F15" box="[665,759,285,307]" gridcol="1" gridrow="2" pageId="5" pageNumber="437">38.1</td>
<td id="769F5DC68F24CA98FC09FE9CFC6F5F15" box="[909,998,285,307]" gridcol="2" gridrow="2" pageId="5" pageNumber="437">39.6</td>
<td id="769F5DC68F24CA98FB05FE9CFB775F15" box="[1153,1278,285,307]" gridcol="3" gridrow="2" pageId="5" pageNumber="437">44.0</td>
</tr>
<tr id="354E34BA8F24CA98FF1AFEC7FB775F7A" box="[158,1278,326,348]" gridrow="3" pageId="5" pageNumber="437">
<th id="769F5DC68F24CA98FF1AFEC7FE4B5F7A" box="[158,450,326,348]" gridcol="0" gridrow="3" pageId="5" pageNumber="437">Postorbital length</th>
<td id="769F5DC68F24CA98FD1DFEC7FD7E5F7A" box="[665,759,326,348]" gridcol="1" gridrow="3" pageId="5" pageNumber="437">32.1</td>
<td id="769F5DC68F24CA98FC09FEC7FC6F5F7A" box="[909,998,326,348]" gridcol="2" gridrow="3" pageId="5" pageNumber="437">33.5</td>
<td id="769F5DC68F24CA98FB05FEC7FB775F7A" box="[1153,1278,326,348]" gridcol="3" gridrow="3" pageId="5" pageNumber="437">37.8</td>
</tr>
<tr id="354E34BA8F24CA98FF1AFEF1FB775FA0" box="[158,1278,368,390]" gridrow="4" pageId="5" pageNumber="437">
<th id="769F5DC68F24CA98FF1AFEF1FE4B5FA0" box="[158,450,368,390]" gridcol="0" gridrow="4" pageId="5" pageNumber="437">Length cephalic section</th>
<td id="769F5DC68F24CA98FD1DFEF1FD7E5FA0" box="[665,759,368,390]" gridcol="1" gridrow="4" pageId="5" pageNumber="437">24.8</td>
<td id="769F5DC68F24CA98FC09FEF1FC6F5FA0" box="[909,998,368,390]" gridcol="2" gridrow="4" pageId="5" pageNumber="437">25.7</td>
<td id="769F5DC68F24CA98FB05FEF1FB775FA0" box="[1153,1278,368,390]" gridcol="3" gridrow="4" pageId="5" pageNumber="437">27.8</td>
</tr>
<tr id="354E34BA8F24CA98FF1AFE18FB775F89" box="[158,1278,409,431]" gridrow="5" pageId="5" pageNumber="437">
<th id="769F5DC68F24CA98FF1AFE18FE4B5F89" box="[158,450,409,431]" gridcol="0" gridrow="5" pageId="5" pageNumber="437">Width</th>
<td id="769F5DC68F24CA98FD1DFE18FD7E5F89" box="[665,759,409,431]" gridcol="1" gridrow="5" pageId="5" pageNumber="437">20.2</td>
<td id="769F5DC68F24CA98FC09FE18FC6F5F89" box="[909,998,409,431]" gridcol="2" gridrow="5" pageId="5" pageNumber="437">21.1</td>
<td id="769F5DC68F24CA98FB05FE18FB775F89" box="[1153,1278,409,431]" gridcol="3" gridrow="5" pageId="5" pageNumber="437">4.3</td>
</tr>
<tr id="354E34BA8F24CA98FF1AFE43FB775FFE" box="[158,1278,450,472]" gridrow="6" pageId="5" pageNumber="437">
<th id="769F5DC68F24CA98FF1AFE43FE4B5FFE" box="[158,450,450,472]" gridcol="0" gridrow="6" pageId="5" pageNumber="437">Depth</th>
<td id="769F5DC68F24CA98FD1DFE43FD7E5FFE" box="[665,759,450,472]" gridcol="1" gridrow="6" pageId="5" pageNumber="437">16.2</td>
<td id="769F5DC68F24CA98FC09FE43FC6F5FFE" box="[909,998,450,472]" gridcol="2" gridrow="6" pageId="5" pageNumber="437">16.9</td>
<td id="769F5DC68F24CA98FB05FE43FB775FFE" box="[1153,1278,450,472]" gridcol="3" gridrow="6" pageId="5" pageNumber="437">19.0</td>
</tr>
<tr id="354E34BA8F24CA98FF1AFE6DFB775C24" box="[158,1278,492,514]" gridrow="7" pageId="5" pageNumber="437">
<th id="769F5DC68F24CA98FF1AFE6DFE4B5C24" box="[158,450,492,514]" gridcol="0" gridrow="7" pageId="5" pageNumber="437">Length rostrum</th>
<td id="769F5DC68F24CA98FD1DFE6DFD7E5C24" box="[665,759,492,514]" gridcol="1" gridrow="7" pageId="5" pageNumber="437">8.7</td>
<td id="769F5DC68F24CA98FC09FE6DFC6F5C24" box="[909,998,492,514]" gridcol="2" gridrow="7" pageId="5" pageNumber="437">9.1</td>
<td id="769F5DC68F24CA98FB05FE6DFB775C24" box="[1153,1278,492,514]" gridcol="3" gridrow="7" pageId="5" pageNumber="437">9.1</td>
</tr>
<tr id="354E34BA8F24CA98FF1AFD94FB775C0D" box="[158,1278,533,555]" gridrow="8" pageId="5" pageNumber="437">
<th id="769F5DC68F24CA98FF1AFD94FE4B5C0D" box="[158,450,533,555]" gridcol="0" gridrow="8" pageId="5" pageNumber="437">Length acumen</th>
<td id="769F5DC68F24CA98FD1DFD94FD7E5C0D" box="[665,759,533,555]" gridcol="1" gridrow="8" pageId="5" pageNumber="437">2.6</td>
<td id="769F5DC68F24CA98FC09FD94FC6F5C0D" box="[909,998,533,555]" gridcol="2" gridrow="8" pageId="5" pageNumber="437">2.9</td>
<td id="769F5DC68F24CA98FB05FD94FB775C0D" box="[1153,1278,533,555]" gridcol="3" gridrow="8" pageId="5" pageNumber="437">2.8</td>
</tr>
<tr id="354E34BA8F24CA98FF1AFDBFFB775C72" box="[158,1278,574,596]" gridrow="9" pageId="5" pageNumber="437">
<th id="769F5DC68F24CA98FF1AFDBFFE4B5C72" box="[158,450,574,596]" gridcol="0" gridrow="9" pageId="5" pageNumber="437">Length areola</th>
<td id="769F5DC68F24CA98FD1DFDBFFD7E5C72" box="[665,759,574,596]" gridcol="1" gridrow="9" pageId="5" pageNumber="437">13.3</td>
<td id="769F5DC68F24CA98FC09FDBFFC6F5C72" box="[909,998,574,596]" gridcol="2" gridrow="9" pageId="5" pageNumber="437">13.9</td>
<td id="769F5DC68F24CA98FB05FDBFFB775C72" box="[1153,1278,574,596]" gridcol="3" gridrow="9" pageId="5" pageNumber="437">16.2</td>
</tr>
<tr id="354E34BA8F24CA98FF1AFDE9FB775C58" box="[158,1278,616,638]" gridrow="10" pageId="5" pageNumber="437">
<th id="769F5DC68F24CA98FF1AFDE9FE4B5C58" box="[158,450,616,638]" gridcol="0" gridrow="10" pageId="5" pageNumber="437">Width areola</th>
<td id="769F5DC68F24CA98FD1DFDE9FD7E5C58" box="[665,759,616,638]" gridcol="1" gridrow="10" pageId="5" pageNumber="437">4.4</td>
<td id="769F5DC68F24CA98FC09FDE9FC6F5C58" box="[909,998,616,638]" gridcol="2" gridrow="10" pageId="5" pageNumber="437">4.4</td>
<td id="769F5DC68F24CA98FB05FDE9FB775C58" box="[1153,1278,616,638]" gridcol="3" gridrow="10" pageId="5" pageNumber="437">4.5</td>
</tr>
<tr id="354E34BA8F24CA98FF1AFD3BFB775CF6" box="[158,1278,698,720]" gridrow="11" pageId="5" pageNumber="437" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="769F5DC68F24CA98FF1AFD3BFE4B5CF6" box="[158,450,698,720]" gridcol="0" gridrow="11" pageId="5" pageNumber="437">Abdomen</th>
</tr>
<tr id="354E34BA8F24CA98FF1AFD65FB775CDC" box="[158,1278,740,762]" gridrow="12" pageId="5" pageNumber="437">
<th id="769F5DC68F24CA98FF1AFD65FE4B5CDC" box="[158,450,740,762]" gridcol="0" gridrow="12" pageId="5" pageNumber="437">Width</th>
<td id="769F5DC68F24CA98FD1DFD65FD7E5CDC" box="[665,759,740,762]" gridcol="1" gridrow="12" pageId="5" pageNumber="437">16.4</td>
<td id="769F5DC68F24CA98FC09FD65FC6F5CDC" box="[909,998,740,762]" gridcol="2" gridrow="12" pageId="5" pageNumber="437">20.0</td>
<td id="769F5DC68F24CA98FB05FD65FB775CDC" box="[1153,1278,740,762]" gridcol="3" gridrow="12" pageId="5" pageNumber="437">19.7</td>
</tr>
<tr id="354E34BA8F24CA98FF1AFC8CFB775D05" box="[158,1278,781,803]" gridrow="13" pageId="5" pageNumber="437">
<th id="769F5DC68F24CA98FF1AFC8CFE4B5D05" box="[158,450,781,803]" gridcol="0" gridrow="13" pageId="5" pageNumber="437">Length</th>
<td id="769F5DC68F24CA98FD1DFC8CFD7E5D05" box="[665,759,781,803]" gridcol="1" gridrow="13" pageId="5" pageNumber="437">38.8</td>
<td id="769F5DC68F24CA98FC09FC8CFC6F5D05" box="[909,998,781,803]" gridcol="2" gridrow="13" pageId="5" pageNumber="437">41.1</td>
<td id="769F5DC68F24CA98FB05FC8CFB775D05" box="[1153,1278,781,803]" gridcol="3" gridrow="13" pageId="5" pageNumber="437">43.0</td>
</tr>
<tr id="354E34BA8F24CA98FF1AFCE1FB775D50" box="[158,1278,864,886]" gridrow="14" pageId="5" pageNumber="437" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="769F5DC68F24CA98FF1AFCE1FE4B5D50" box="[158,450,864,886]" gridcol="0" gridrow="14" pageId="5" pageNumber="437">Cheliped</th>
</tr>
<tr id="354E34BA8F24CA98FF1AFC08FB775DB9" box="[158,1278,905,927]" gridrow="15" pageId="5" pageNumber="437">
<th id="769F5DC68F24CA98FF1AFC08FE4B5DB9" box="[158,450,905,927]" gridcol="0" gridrow="15" pageId="5" pageNumber="437">Length mesial margin palm</th>
<td id="769F5DC68F24CA98FD1DFC08FD7E5DB9" box="[665,759,905,927]" gridcol="1" gridrow="15" pageId="5" pageNumber="437">11.8</td>
<td id="769F5DC68F24CA98FC09FC08FC6F5DB9" box="[909,998,905,927]" gridcol="2" gridrow="15" pageId="5" pageNumber="437">10.0</td>
<td id="769F5DC68F24CA98FB05FC08FB775DB9" box="[1153,1278,905,927]" gridcol="3" gridrow="15" pageId="5" pageNumber="437">13.3</td>
</tr>
<tr id="354E34BA8F24CA98FF1AFC33FB775DEE" box="[158,1278,946,968]" gridrow="16" pageId="5" pageNumber="437">
<th id="769F5DC68F24CA98FF1AFC33FE4B5DEE" box="[158,450,946,968]" gridcol="0" gridrow="16" pageId="5" pageNumber="437">Width palm</th>
<td id="769F5DC68F24CA98FD1DFC33FD7E5DEE" box="[665,759,946,968]" gridcol="1" gridrow="16" pageId="5" pageNumber="437">17.3</td>
<td id="769F5DC68F24CA98FC09FC33FC6F5DEE" box="[909,998,946,968]" gridcol="2" gridrow="16" pageId="5" pageNumber="437">16.0</td>
<td id="769F5DC68F24CA98FB05FC33FB775DEE" box="[1153,1278,946,968]" gridcol="3" gridrow="16" pageId="5" pageNumber="437">20.3</td>
</tr>
<tr id="354E34BA8F24CA98FF1AFC5DFB775DD4" box="[158,1278,988,1010]" gridrow="17" pageId="5" pageNumber="437">
<th id="769F5DC68F24CA98FF1AFC5DFE4B5DD4" box="[158,450,988,1010]" gridcol="0" gridrow="17" pageId="5" pageNumber="437">Depth palm</th>
<td id="769F5DC68F24CA98FD1DFC5DFD7E5DD4" box="[665,759,988,1010]" gridcol="1" gridrow="17" pageId="5" pageNumber="437">16.2</td>
<td id="769F5DC68F24CA98FC09FC5DFC6F5DD4" box="[909,998,988,1010]" gridcol="2" gridrow="17" pageId="5" pageNumber="437">9.2</td>
<td id="769F5DC68F24CA98FB05FC5DFB775DD4" box="[1153,1278,988,1010]" gridcol="3" gridrow="17" pageId="5" pageNumber="437">11.2</td>
</tr>
<tr id="354E34BA8F24CA98FF1AFB84FB775A3D" box="[158,1278,1029,1051]" gridrow="18" pageId="5" pageNumber="437">
<th id="769F5DC68F24CA98FF1AFB84FE4B5A3D" box="[158,450,1029,1051]" gridcol="0" gridrow="18" pageId="5" pageNumber="437">Length dactyl</th>
<td id="769F5DC68F24CA98FD1DFB84FD7E5A3D" box="[665,759,1029,1051]" gridcol="1" gridrow="18" pageId="5" pageNumber="437">24.5</td>
<td id="769F5DC68F24CA98FC09FB84FC6F5A3D" box="[909,998,1029,1051]" gridcol="2" gridrow="18" pageId="5" pageNumber="437">20.0</td>
<td id="769F5DC68F24CA98FB05FB84FB775A3D" box="[1153,1278,1029,1051]" gridcol="3" gridrow="18" pageId="5" pageNumber="437">19.7</td>
</tr>
<tr id="354E34BA8F24CA98FF1AFBAFFB775A62" box="[158,1278,1070,1092]" gridrow="19" pageId="5" pageNumber="437">
<th id="769F5DC68F24CA98FF1AFBAFFE4B5A62" box="[158,450,1070,1092]" gridcol="0" gridrow="19" pageId="5" pageNumber="437">Length carpus</th>
<td id="769F5DC68F24CA98FD1DFBAFFD7E5A62" box="[665,759,1070,1092]" gridcol="1" gridrow="19" pageId="5" pageNumber="437">11.7</td>
<td id="769F5DC68F24CA98FC09FBAFFC6F5A62" box="[909,998,1070,1092]" gridcol="2" gridrow="19" pageId="5" pageNumber="437">12.2</td>
<td id="769F5DC68F24CA98FB05FBAFFB775A62" box="[1153,1278,1070,1092]" gridcol="3" gridrow="19" pageId="5" pageNumber="437">15.6</td>
</tr>
<tr id="354E34BA8F24CA98FF1AFBD9FB775A48" box="[158,1278,1112,1134]" gridrow="20" pageId="5" pageNumber="437">
<th id="769F5DC68F24CA98FF1AFBD9FE4B5A48" box="[158,450,1112,1134]" gridcol="0" gridrow="20" pageId="5" pageNumber="437">Width carpus</th>
<td id="769F5DC68F24CA98FD1DFBD9FD7E5A48" box="[665,759,1112,1134]" gridcol="1" gridrow="20" pageId="5" pageNumber="437">9.6</td>
<td id="769F5DC68F24CA98FC09FBD9FC6F5A48" box="[909,998,1112,1134]" gridcol="2" gridrow="20" pageId="5" pageNumber="437">8.7</td>
<td id="769F5DC68F24CA98FB05FBD9FB775A48" box="[1153,1278,1112,1134]" gridcol="3" gridrow="20" pageId="5" pageNumber="437">11.5</td>
</tr>
<tr id="354E34BA8F24CA98FF1AFB00FB775AB1" box="[158,1278,1153,1175]" gridrow="21" pageId="5" pageNumber="437">
<th id="769F5DC68F24CA98FF1AFB00FE4B5AB1" box="[158,450,1153,1175]" gridcol="0" gridrow="21" pageId="5" pageNumber="437">Length dorsal margin merus</th>
<td id="769F5DC68F24CA98FD1DFB00FD7E5AB1" box="[665,759,1153,1175]" gridcol="1" gridrow="21" pageId="5" pageNumber="437">10.5</td>
<td id="769F5DC68F24CA98FC09FB00FC6F5AB1" box="[909,998,1153,1175]" gridcol="2" gridrow="21" pageId="5" pageNumber="437">9.5</td>
<td id="769F5DC68F24CA98FB05FB00FB775AB1" box="[1153,1278,1153,1175]" gridcol="3" gridrow="21" pageId="5" pageNumber="437">9.9</td>
</tr>
<tr id="354E34BA8F24CA98FF1AFB2BFB775AE6" box="[158,1278,1194,1216]" gridrow="22" pageId="5" pageNumber="437">
<th id="769F5DC68F24CA98FF1AFB2BFE4B5AE6" box="[158,450,1194,1216]" gridcol="0" gridrow="22" pageId="5" pageNumber="437">Depth merus</th>
<td id="769F5DC68F24CA98FD1DFB2BFD7E5AE6" box="[665,759,1194,1216]" gridcol="1" gridrow="22" pageId="5" pageNumber="437">9.9</td>
<td id="769F5DC68F24CA98FC09FB2BFC6F5AE6" box="[909,998,1194,1216]" gridcol="2" gridrow="22" pageId="5" pageNumber="437">9.8</td>
<td id="769F5DC68F24CA98FB05FB2BFB775AE6" box="[1153,1278,1194,1216]" gridcol="3" gridrow="22" pageId="5" pageNumber="437">12.2</td>
</tr>
<tr id="354E34BA8F24CA98FF1AFB7CFB775B35" box="[158,1278,1277,1299]" gridrow="23" pageId="5" pageNumber="437" rowspan-2="1">
<th id="769F5DC68F24CA98FF1AFB7CFE4B5B35" box="[158,450,1277,1299]" gridcol="0" gridrow="23" pageId="5" pageNumber="437">Gonopod length</th>
<td id="769F5DC68F24CA98FD1DFB7CFD7E5B35" box="[665,759,1277,1299]" gridcol="1" gridrow="23" pageId="5" pageNumber="437">8.1</td>
<td id="769F5DC68F24CA98FB05FB7CFB775B35" box="[1153,1278,1277,1299]" gridcol="3" gridrow="23" pageId="5" pageNumber="437">9.9</td>
</tr>
</table>
</paragraph>
<paragraph id="8BC136F88F243562FF43FAD5FD6C58A9" blockId="5.[151,1437,1364,1823]" pageId="5" pageNumber="437">
<emphasis id="B90AEAEA8F243562FF43FAD5FD4F5B4B" bold="true" box="[199,710,1364,1389]" pageId="5" pageNumber="437">Description of morphotypic male, form II.</emphasis>
(
<figureCitation id="13452A7D8F243562FD53FAD5FC925B4A" box="[727,795,1364,1389]" captionStart="FIGURE 2" captionStartId="6.[151,250,1877,1899]" captionTargetBox="[172,1406,196,1842]" captionTargetId="figure@6.[172,1406,196,1842]" captionTargetPageId="6" captionText="FIGURE 2. Cambarus appalachiensis, new species: (A.) Lateral view of carapace; (B.) Lateral and (C.) mesial view of form I gonopod; (D.) Lateral and (E.) mesial view of form II gonopod; (F.) Annulus ventralis; (G.) Dorsal view of carapace; (H.) Right antennal scale; (I.) Dorsal view of right carpus and chelae; (J.) epistome. A-C and G-I from holotype; F from allotype; D- E from morphotypes." httpUri="https://zenodo.org/record/400116/files/figure.png" pageId="5" pageNumber="437">Fig. 2</figureCitation>
D–E, 
<tableCitation id="C6FC03438F243562FCDAFAD5FC3B5B4A" box="[862,946,1364,1389]" captionStart="TABLE 1" captionStartId="5.[151,239,152,175]" captionTargetBox="[158,1278,202,1299]" captionTargetPageId="5" captionText="TABLE 1. Morphological measurements (mm) of Cambarus appalachiensis, new species." httpUri="http://table.plazi.org/id/DF0166708F243562FF13FF19FBD75E89" pageId="5" pageNumber="437" tableUuid="DF0166708F243562FF13FF19FBD75E89">Table 1</tableCitation>
). Differing from 
<typeStatus id="54C5885A8F243562FBFFFAD5FB545B4B" box="[1147,1245,1364,1389]" pageId="5" pageNumber="437" type="holotype">holotype</typeStatus>
in the following respects: Carapace height less than carapace width (19.0 and 
<quantity id="4C869B1D8F243562FCCFFAF9FC395BA9" box="[843,944,1400,1424]" metricMagnitude="-2" metricUnit="m" metricValue="2.43" pageId="5" pageNumber="437" unit="mm" value="24.3">24.3 mm</quantity>
respectively); TCL 44.0 mm and PCL 
<quantity id="4C869B1D8F243562FAEFFAF9FF4F5B92" metricMagnitude="-2" metricUnit="m" metricValue="3.78" pageId="5" pageNumber="437" unit="mm" value="37.8">37.8 mm</quantity>
. Areola length 36.8% of TCL (42.8% of PCL), 3.6 times longer than wide. Rostrum margins parallel to base of acumen and thickened; rostrum ventrally deflected and excavated; rostrum 1.6 times as long as wide. Abdomen 43.0 mm long. Single mesial row of tubercles on palm of chela with 7 tubercles. Palm length (
<quantity id="4C869B1D8F243562FB3BFA65FAAF5BDA" box="[1215,1318,1508,1532]" metricMagnitude="-2" metricUnit="m" metricValue="1.33" pageId="5" pageNumber="437" unit="mm" value="13.3">13.3 mm</quantity>
) 65.5% of palm width (
<quantity id="4C869B1D8F243562FEA1F989FE065806" box="[293,399,1544,1568]" metricMagnitude="-2" metricUnit="m" metricValue="2.03" pageId="5" pageNumber="437" unit="mm" value="20.3">20.3 mm</quantity>
). All measurements and counts from right chela. Antennal scale 
<quantity id="4C869B1D8F243562FBE1F986FB325839" box="[1125,1211,1543,1568]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" pageId="5" pageNumber="437" unit="mm" value="6.6">6.6 mm</quantity>
long, 
<quantity id="4C869B1D8F243562FA86F989FAD15839" box="[1282,1368,1544,1568]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="5" pageNumber="437" unit="mm" value="2.9">2.9 mm</quantity>
wide. Gonopods reaching anterior margin of 4th pereopod caudomesial boss. Central projection curved 90° to shaft, with complete apex; rounded (
<figureCitation id="13452A7D8F243562FE36F9D3FE7D584D" box="[434,500,1618,1643]" captionStart="FIGURE 2" captionStartId="6.[151,250,1877,1899]" captionTargetBox="[172,1406,196,1842]" captionTargetId="figure@6.[172,1406,196,1842]" captionTargetPageId="6" captionText="FIGURE 2. Cambarus appalachiensis, new species: (A.) Lateral view of carapace; (B.) Lateral and (C.) mesial view of form I gonopod; (D.) Lateral and (E.) mesial view of form II gonopod; (F.) Annulus ventralis; (G.) Dorsal view of carapace; (H.) Right antennal scale; (I.) Dorsal view of right carpus and chelae; (J.) epistome. A-C and G-I from holotype; F from allotype; D- E from morphotypes." httpUri="https://zenodo.org/record/400116/files/figure.png" pageId="5" pageNumber="437">Fig. 2</figureCitation>
D–E). Mesial process tapered, bulbous, directed caudolaterally. Hook on ischium of third pereopod small, not reaching basioischial joint.
</paragraph>
<paragraph id="8BC136F88F243562FF43F91AFB8B5939" blockId="5.[151,1437,1364,1823]" pageId="5" pageNumber="437">
<emphasis id="B90AEAEA8F243562FF43F91AFF745895" bold="true" box="[199,253,1691,1715]" italics="true" pageId="5" pageNumber="437">Size.</emphasis>
Form I male (n = 32) TCL ranges in size from 
<quantity id="4C869B1D8F243562FC91F91BFC315894" box="[789,952,1690,1715]" metricMagnitude="-2" metricUnit="m" metricValue="3.935" metricValueMax="4.86" metricValueMin="3.01" pageId="5" pageNumber="437" unit="mm" value="39.35" valueMax="48.6" valueMin="30.1">30.1–48.6 mm</quantity>
(PCL 25.4–41.0 mm) with a mean TCL of 
<quantity id="4C869B1D8F243562FF13F93EFE8958F1" box="[151,256,1727,1751]" metricMagnitude="-2" metricUnit="m" metricValue="3.7399999999999998" pageId="5" pageNumber="437" unit="mm" value="37.4">37.4 mm</quantity>
. Form II male (n = 31) mean TCL is 
<quantity id="4C869B1D8F243562FD30F93FFC9358F1" box="[692,794,1726,1751]" metricMagnitude="-2" metricUnit="m" metricValue="3.66" pageId="5" pageNumber="437" unit="mm" value="36.6">36.6 mm</quantity>
and ranges in size from 
<quantity id="4C869B1D8F243562FBBDF93FFB5758F1" box="[1081,1246,1726,1751]" metricMagnitude="-2" metricUnit="m" metricValue="3.59" metricValueMax="4.6899999999999995" metricValueMin="2.4899999999999998" pageId="5" pageNumber="437" unit="mm" value="35.9" valueMax="46.9" valueMin="24.9">24.9–46.9 mm</quantity>
(PCL 20.7–40.0 mm). Female (n = 35) TCL mean is 37.0 mm and ranges from 11.1–45.0 mm (PCL 
<quantity id="4C869B1D8F243562FBE8F963FA8E58DD" box="[1132,1287,1762,1787]" metricMagnitude="-2" metricUnit="m" metricValue="2.33" metricValueMax="3.81" metricValueMin="0.85" pageId="5" pageNumber="437" unit="mm" value="23.3" valueMax="38.1" valueMin="8.5">8.5–38.1 mm</quantity>
). The largest specimen examined was a form I male with TCL of 
<quantity id="4C869B1D8F243562FD64F887FCCD5939" box="[736,836,1798,1823]" metricMagnitude="-2" metricUnit="m" metricValue="4.86" pageId="5" pageNumber="437" unit="mm" value="48.6">48.6 mm</quantity>
(PCL 41.0 mm).
</paragraph>
<caption id="DF0166708F273561FF13F8D4FEF959E1" httpUri="https://zenodo.org/record/400116/files/figure.png" pageId="6" pageNumber="438" targetBox="[172,1406,196,1842]" targetPageId="6">
<paragraph id="8BC136F88F273561FF13F8D4FEF959E1" blockId="6.[151,1436,1876,1991]" pageId="6" pageNumber="438">
<emphasis id="B90AEAEA8F273561FF13F8D4FD30594D" bold="true" box="[151,697,1876,1899]" pageId="6" pageNumber="438">
FIGURE 2. 
<taxonomicName id="4C7E4D7B8F273561FE9DF8D5FDAE594D" box="[281,551,1876,1899]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="438" phylum="Arthropoda" rank="species" species="appalachiensis" status="sp. nov.">
<emphasis id="B90AEAEA8F273561FE9DF8D5FDAE594D" bold="true" box="[281,551,1876,1899]" italics="true" pageId="6" pageNumber="438">Cambarus appalachiensis</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A23957918F273561FDB0F8D4FD3D594D" box="[564,692,1877,1899]" pageId="6" pageNumber="438" rank="species">new species</taxonomicNameLabel>
:
</emphasis>
(A.) Lateral view of carapace; (B.) Lateral and (C.) mesial view of form I gonopod; (D.) Lateral and (E.) mesial view of form II gonopod; (F.) Annulus ventralis; (G.) Dorsal view of carapace; (H.) Right antennal scale; (I.) Dorsal view of right carpus and chelae; (J.) epistome. A-C and G-I from holotype; F from allotype; D- E from morphotypes.
</paragraph>
</caption>
<caption id="DF0166708F263560FF13FC2EFD635DE3" box="[151,746,943,965]" httpUri="https://zenodo.org/record/400117/files/figure.png" pageId="7" pageNumber="439" targetBox="[151,1436,193,922]" targetPageId="7">
<paragraph id="8BC136F88F263560FF13FC2EFD635DE3" blockId="7.[151,746,943,965]" box="[151,746,943,965]" pageId="7" pageNumber="439">
<emphasis id="B90AEAEA8F263560FF13FC2EFE9B5DE3" bold="true" box="[151,274,943,965]" pageId="7" pageNumber="439">FIGURE 3.</emphasis>
<taxonomicName id="4C7E4D7B8F263560FE9DFC2EFDA85DE3" box="[281,545,943,965]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="439" phylum="Arthropoda" rank="species" species="appalachiensis">
<emphasis id="B90AEAEA8F263560FE9DFC2EFDA85DE3" box="[281,545,943,965]" italics="true" pageId="7" pageNumber="439">Cambarus appalachiensis</emphasis>
</taxonomicName>
morphotype in life.
</paragraph>
</caption>
<paragraph id="8BC136F88F263560FF43FC6DFCDA581A" blockId="7.[151,1437,1004,2026]" pageId="7" pageNumber="439">
<emphasis id="B90AEAEA8F263560FF43FC6DFDB45A23" box="[199,573,1004,1029]" italics="true" pageId="7" pageNumber="439">
<emphasis id="B90AEAEA8F263560FF43FC6DFE875A23" bold="true" box="[199,270,1004,1029]" italics="true" pageId="7" pageNumber="439">Color.</emphasis>
<taxonomicName id="4C7E4D7B8F263560FE93FC6CFDB45A23" box="[279,573,1005,1029]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="439" phylum="Arthropoda" rank="species" species="appalachiensis">Cambarus appalachiensis</taxonomicName>
</emphasis>
(
<figureCitation id="13452A7D8F263560FDCAFC6DFD1B5A23" box="[590,658,1004,1029]" captionStart="FIGURE 3" captionStartId="7.[151,250,943,965]" captionTargetBox="[151,1436,193,922]" captionTargetId="figure@7.[151,1436,193,922]" captionTargetPageId="7" captionText="FIGURE 3. Cambarus appalachiensis morphotype in life." httpUri="https://zenodo.org/record/400117/files/figure.png" pageId="7" pageNumber="439">Fig. 3</figureCitation>
) carapace ground color is chestnut brown to pink to orange-brown; posterior margin of carapace dark brown to black. Hepatic and antennal region of carapace punctuated with cream, olive, or tan tubercles. Postorbital ridge orange, red-brown, to light. Rostrum margins and acumen cream to orange, orange-brown, red-brown or tan. Cephalic section of carapace immediately anterior to and including cervical groove is black, forming weak saddle; mandibular abductor scars mottled, ranging from light-brown, brown, to dark-brown. Lateral margin of antennal scale olive to light brown; body of antennal scale brown to cream. Antennal flagellum and antennules green-brown, with olivaceous hue; dorsal surface of lamellae tan to brown; ventral surface light-green to olivaceous. Dorsal surface of chelae variable ranging from green, green-brown, olivaceous, orange, light-brown, or brown; occasionally with mesial row of tubercles cream, tan or orange. Denticles on opposable surfaces of fingers yellow, white, or tan. Ventral surface of chelae cream or tan. Dorsal surface of carpus light green, green-brown, or olivaceous; region adjacent to and including furrow green-brown to green; carpus spine orange, orange brown or cream. Merus green-brown, grey, or olivaceous brown. Podomeres of pereopods light blue, cream, or grey-blue; joints of pereopod podomeres pink. Dorsal and dorsolateral surface of abdomen same colors as carapace; tergal margins brown to olivaceous brown; abdomen lacking dorsal stripe. Uropods same colors as abdomen. Ventral surface of abdomen and carapace cream. Dorsal ridge of form I gonopod central projection amber; body of central projection, gonopod, and mesial process tan. Form II gonopod and all associated processes cream. Cephalic portion of annulus ventralis pink to pink-cream; ridge of fossa pink; caudal region of annulus ventralis ranges from pink to cream colored.
</paragraph>
</subSubSection>
<subSubSection id="C36465738F26356EFF43F9C5FDF45B9D" lastPageId="9" lastPageNumber="441" pageId="7" pageNumber="439" type="materials_examined">
<paragraph id="8BC136F88F263560FF43F9C5FCF1597F" blockId="7.[151,1437,1004,2026]" pageId="7" pageNumber="439">
<materialsCitation id="3B163CA58F263560FF43F9C5FC0458A7" ID-GBIF-Occurrence="1440156654" county="Summers County" location="Tom-Honaker Road" municipality="Pipestem Creek" pageId="7" pageNumber="439" specimenCount="1" stateProvince="West Virginia" typeStatus="holotype">
<emphasis id="B90AEAEA8F263560FF43F9C5FEE1587B" bold="true" box="[199,360,1604,1629]" pageId="7" pageNumber="439">
<typeStatus id="54C5885A8F263560FF43F9C5FE8B587B" box="[199,258,1604,1629]" pageId="7" pageNumber="439">Type</typeStatus>
locality.
</emphasis>
<collectingMunicipality id="6BA5AC828F263560FEF5F9C5FDAD587B" box="[369,548,1604,1629]" pageId="7" pageNumber="439">Pipestem Creek</collectingMunicipality>
at intersection of 
<location id="8EA160238F263560FD7CF9C5FC50587B" LSID="urn:lsid:plazi:treatment:03D787EE8F223569FF13FADDFDFB5C3E:8EA160238F263560FD7CF9C5FC50587B" box="[760,985,1604,1629]" county="Summers County" municipality="Pipestem Creek" name="Tom-Honaker Road" pageId="7" pageNumber="439" stateProvince="West Virginia">Tom-Honaker Road</location>
(CR 20-3) and 
<location id="8EA160238F263560FB16F9C4FA9F587B" LSID="urn:lsid:plazi:treatment:03D787EE8F223569FF13FADDFDFB5C3E:8EA160238F263560FB16F9C4FA9F587B" box="[1170,1302,1605,1629]" county="Summers County" municipality="Pipestem Creek" name="State Route" pageId="7" pageNumber="439" stateProvince="West Virginia">State Route</location>
20, 
<quantity id="4C869B1D8F263560FAC8F9C4FA15587B" box="[1356,1436,1604,1629]" metricMagnitude="3" metricUnit="m" metricValue="3.3" pageId="7" pageNumber="439" unit="km" value="3.3">3.3 km</quantity>
(
<quantity id="4C869B1D8F263560FF1AF9E8FF7058A7" box="[158,249,1640,1665]" metricMagnitude="3" metricUnit="m" metricValue="3.28306176" pageId="7" pageNumber="439" unit="mi" value="2.04">2.04 mi</quantity>
) north-east of 
<location id="8EA160238F263560FE20F9E9FD8758A7" LSID="urn:lsid:plazi:treatment:03D787EE8F223569FF13FADDFDFB5C3E:8EA160238F263560FE20F9E9FD8758A7" box="[420,526,1640,1665]" county="Summers County" municipality="Pipestem Creek" name="Pipestem" pageId="7" pageNumber="439" stateProvince="West Virginia">Pipestem</location>
, 
<collectingCounty id="62A04E748F263560FD9EF9E8FD6858A7" box="[538,737,1641,1665]" pageId="7" pageNumber="439">Summers County</collectingCounty>
, 
<collectingRegion id="49BAF81A8F263560FD6AF9E8FC0458A7" box="[750,909,1640,1665]" country="United States of America" name="West Virginia" pageId="7" pageNumber="439">West Virginia</collectingRegion>
</materialsCitation>
. 
<materialsCitation id="3B163CA58F263560FC1DF9E9FCF1597F" ID-GBIF-Occurrence="1440156659" collectingDate="2011-10" collectorName="Z. J. Loughman. At" elevation="7" location="Pipestem Creek" pageId="7" pageNumber="439" specimenCount="2" stateProvince="West Virginia" typeStatus="holotype">
The 
<typeStatus id="54C5885A8F263560FC4AF9E8FC7758A7" box="[974,1022,1641,1665]" pageId="7" pageNumber="439">type</typeStatus>
series was collected on 
<date id="FFC010388F263560FA9CF9E8FA1158A7" box="[1304,1432,1640,1665]" pageId="7" pageNumber="439" value="2011-10">
<collectingDate id="EF84E9D08F263560FA9CF9E8FA1158A7" box="[1304,1432,1640,1665]" pageId="7" pageNumber="439" value="2011-10">11 October</collectingDate>
</date>
, 2016 by 
<collectorName id="268B532E8F263560FF7BF90FFE6A5883" box="[255,483,1677,1702]" pageId="7" pageNumber="439">Z. J. Loughman. At</collectorName>
this site (
<figureCitation id="13452A7D8F263560FDDDF90CFD295883" box="[601,672,1677,1702]" captionStart="FIGURE 4" captionStartId="8.[151,250,1809,1831]" captionTargetBox="[340,1247,193,1788]" captionTargetId="figure@8.[340,1247,193,1788]" captionTargetPageId="8" captionText="FIGURE 4. Cambarus appalachiensis type locality. Pipestem Creek at intersection of Tom-Honaker Road (CR 20 - 3) and State Route 20, 3.3 km (2.04 mi) north-east of Pipestem, Summers County, West Virginia— 11 October, 2016. (A.) Run downstream of Tom-Honaker Road bridge where holotype and allotype were collected; (B.) pool immediately upstream of Tom-Honaker Road bridge." httpUri="https://zenodo.org/record/400118/files/figure.png" pageId="7" pageNumber="439">Fig. 4</figureCitation>
), 
<location id="8EA160238F263560FD3DF90CFCE55880" LSID="urn:lsid:plazi:treatment:03D787EE8F223569FF13FADDFDFB5C3E:8EA160238F263560FD3DF90CFCE55880" box="[697,876,1677,1702]" name="Pipestem Creek" pageId="7" pageNumber="439" stateProvince="West Virginia">Pipestem Creek</location>
is 
<quantity id="4C869B1D8F263560FC17F90CFC685883" box="[915,993,1677,1701]" metricMagnitude="0" metricUnit="m" metricValue="7.0" metricValueMax="10.0" metricValueMin="4.0" pageId="7" pageNumber="439" unit="m" value="7.0" valueMax="10.0" valueMin="4.0">
<elevation id="0053D1CB8F263560FC17F90CFC685883" box="[915,993,1677,1701]" metricMagnitude="0" metricUnit="m" metricValue="7.0" metricValueMax="10.0" metricValueMin="4.0" pageId="7" pageNumber="439" unit="m" value="7.0" valueMax="10.0" valueMin="4.0">4–10m</elevation>
</quantity>
wide, and consists of a series of high gradient riffles and glides, followed by moderate to low gradient runs with occasional pools. Substrates utilized by 
<taxonomicName id="4C7E4D7B8F263560FF13F954FEE958CB" box="[151,352,1749,1773]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="439" phylum="Arthropoda" rank="species" species="appalachiensis">
<emphasis id="B90AEAEA8F263560FF13F954FEE958CB" box="[151,352,1749,1773]" italics="true" pageId="7" pageNumber="439">C. appalachiensis</emphasis>
</taxonomicName>
included large slab-rock, boulders, and large cobbles, as well as leaf packs. Riparian conditions ranged from residential yards to intact mesophytic forest bordered by SR 20. Water depth ranged from 0.2–1.0 m deep. Here, 
<taxonomicName id="4C7E4D7B8F263560FEA1F89CFE7B5913" box="[293,498,1821,1845]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="439" phylum="Arthropoda" rank="species" species="appalachiensis">
<emphasis id="B90AEAEA8F263560FEA1F89CFE7B5913" box="[293,498,1821,1845]" italics="true" pageId="7" pageNumber="439">C. appalachiensis</emphasis>
</taxonomicName>
was both abundant and the only crayfish species observed. The 
<typeStatus id="54C5885A8F263560FB6FF89FFA925910" box="[1259,1307,1822,1846]" pageId="7" pageNumber="439">type</typeStatus>
series was collected (n=3), along with several additional specimens (n = 15).
</materialsCitation>
</paragraph>
<paragraph id="8BC136F88F263560FF43F8E4FAEC59CC" blockId="7.[151,1437,1004,2026]" pageId="7" pageNumber="439">
<materialsCitation id="3B163CA58F263560FF43F8E4FAAC5987" ID-GBIF-Occurrence="1440156656" collectionCode="NCSM" collectorName="Raleigh, NC" location="Science" pageId="7" pageNumber="439" specimenCode="NCSM 1251657, 1251658" specimenCount="3" stateProvince="North Carolina" typeStatus="holotype">
<emphasis id="B90AEAEA8F263560FF43F8E4FE315958" bold="true" box="[199,440,1893,1918]" pageId="7" pageNumber="439">
Disposition of 
<typeStatus id="54C5885A8F263560FEF7F8E4FE3A5958" box="[371,435,1893,1918]" pageId="7" pageNumber="439">types</typeStatus>
.
</emphasis>
The 
<typeStatus id="54C5885A8F263560FE76F8E4FDDE5958" box="[498,599,1893,1918]" pageId="7" pageNumber="439" type="holotype">holotype</typeStatus>
, 
<typeStatus id="54C5885A8F263560FDE6F8E4FD365958" box="[610,703,1893,1918]" pageId="7" pageNumber="439" type="allotype">allotype</typeStatus>
, and morphotype are deposited in the 
<collectingRegion id="49BAF81A8F263560FBEFF8E4FA9C5958" box="[1131,1301,1893,1918]" country="United States of America" name="North Carolina" pageId="7" pageNumber="439">North Carolina</collectingRegion>
Museum of 
<location id="8EA160238F263560FF13F809FF675987" LSID="urn:lsid:plazi:treatment:03D787EE8F223569FF13FADDFDFB5C3E:8EA160238F263560FF13F809FF675987" box="[151,238,1928,1953]" name="Science" pageId="7" pageNumber="439" stateProvince="North Carolina">Science</location>
(
<collectionCode id="ED6FAE3D8F263560FF7AF808FEDA5987" box="[254,339,1929,1953]" country="USA" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="7" pageNumber="439">NCSM</collectionCode>
), 
<collectorName id="268B532E8F263560FEE2F809FE7E5987" box="[358,503,1928,1953]" pageId="7" pageNumber="439">Raleigh, NC</collectorName>
. (catalogue numbers 
<specimenCode id="DBD89E838F263560FD63F808FB9A5987" box="[743,1043,1928,1953]" pageId="7" pageNumber="439">
<collectionCode id="ED6FAE3D8F263560FD63F808FCBF5987" box="[743,822,1929,1953]" country="USA" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="7" pageNumber="439">NCSM</collectionCode>
1251657, 1251658
</specimenCode>
, 1251659, respectively)
</materialsCitation>
. 
<materialsCitation id="3B163CA58F263560FAABF808FAE859CC" ID-GBIF-Occurrence="1440156657" collectionCode="MII, WLU, USNM, OSUMC" country="United States of America" location="Zoology" municipality="West Liberty University Astacology Collection" pageId="7" pageNumber="439" specimenCode="WLU 3003, USNM 1422178, OSUMC 9916" specimenCount="1" stateProvince="Ohio" typeStatus="paratype">
<typeStatus id="54C5885A8F263560FAABF808FA155987" box="[1327,1436,1929,1953]" pageId="7" pageNumber="439" type="paratype">Paratypes</typeStatus>
consisting of 1 M1, 1 
<collectionCode id="ED6FAE3D8F263560FE0DF82FFE3F59E3" box="[393,438,1966,1989]" country="Ireland" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13666" name="Museum of Irish Industry" pageId="7" pageNumber="439">MII</collectionCode>
, and 1 F are deposited in the 
<collectingMunicipality id="6BA5AC828F263560FD7DF82FFB7659E0" box="[761,1279,1965,1990]" pageId="7" pageNumber="439">West Liberty University Astacology Collection</collectingMunicipality>
(
<specimenCode id="DBD89E838F263560FA8AF82CFA0659E3" box="[1294,1423,1965,1989]" collectionCode="WLU" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15095" name="Wilfrid Laurier University" pageId="7" pageNumber="439">
<collectionCode id="ED6FAE3D8F263560FA8AF82CFAC659E3" box="[1294,1359,1965,1989]" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15095" name="Wilfrid Laurier University" pageId="7" pageNumber="439">WLU</collectionCode>
3003
</specimenCode>
), the 
<collectingCountry id="F36976688F263560FF3BF850FED859CF" box="[191,337,2001,2026]" name="United States of America" pageId="7" pageNumber="439">United States</collectingCountry>
National Museum (
<specimenCode id="DBD89E838F263560FDAFF850FD6C59CF" box="[555,741,2001,2025]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="7" pageNumber="439">
<collectionCode id="ED6FAE3D8F263560FDAFF850FDF559CF" box="[555,636,2001,2025]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="7" pageNumber="439">USNM</collectionCode>
1422178
</specimenCode>
) and the 
<collectingRegion id="49BAF81A8F263560FCCDF850FC0959CC" box="[841,896,2001,2026]" country="United States of America" name="Ohio" pageId="7" pageNumber="439">Ohio</collectingRegion>
State Museum of 
<location id="8EA160238F263560FBCCF850FB2C59CC" LSID="urn:lsid:plazi:treatment:03D787EE8F223569FF13FADDFDFB5C3E:8EA160238F263560FBCCF850FB2C59CC" box="[1096,1189,2001,2026]" country="United States of America" municipality="West Liberty University Astacology Collection" name="Zoology" pageId="7" pageNumber="439" stateProvince="Ohio">Zoology</location>
(
<specimenCode id="DBD89E838F263560FB37F850FAD059CC" box="[1203,1369,2001,2026]" collectionCode="OSUMC" pageId="7" pageNumber="439">
<collectionCode id="ED6FAE3D8F263560FB37F850FA9E59CF" box="[1203,1303,2001,2025]" pageId="7" pageNumber="439">OSUMC</collectionCode>
9916
</specimenCode>
)
</materialsCitation>
.
</paragraph>
<caption id="DF0166708F29356FFF13F890FE2759A5" httpUri="https://zenodo.org/record/400118/files/figure.png" pageId="8" pageNumber="440" targetBox="[340,1247,193,1788]" targetPageId="8">
<paragraph id="8BC136F88F29356FFF13F890FE2759A5" blockId="8.[151,1435,1809,1923]" pageId="8" pageNumber="440">
<emphasis id="B90AEAEA8F29356FFF13F890FD4B5901" bold="true" box="[151,706,1809,1832]" pageId="8" pageNumber="440">
FIGURE 4. 
<taxonomicName id="4C7E4D7B8F29356FFE9AF890FDA7590E" box="[286,558,1809,1832]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="440" phylum="Arthropoda" rank="species" species="appalachiensis">
<emphasis id="B90AEAEA8F29356FFE9AF890FDA7590E" bold="true" box="[286,558,1809,1832]" italics="true" pageId="8" pageNumber="440">Cambarus appalachiensis</emphasis>
</taxonomicName>
type locality.
</emphasis>
Pipestem Creek at intersection of Tom-Honaker Road (CR 20-3) and State Route 20, 3.3 km (2.04 mi) north-east of Pipestem, Summers County, West Virginia—11 October, 2016. (A.) Run downstream of Tom-Honaker Road bridge where holotype and allotype were collected; (B.) pool immediately upstream of Tom-Honaker Road bridge.
</paragraph>
</caption>
<caption id="DF0166708F28356EFF13FB30FD2A5AE1" box="[151,675,1201,1223]" httpUri="https://zenodo.org/record/400119/files/figure.png" pageId="9" pageNumber="441" targetBox="[151,1436,193,1180]" targetPageId="9">
<paragraph id="8BC136F88F28356EFF13FB30FD2A5AE1" blockId="9.[151,675,1201,1223]" box="[151,675,1201,1223]" pageId="9" pageNumber="441">
<emphasis id="B90AEAEA8F28356EFF13FB30FE9B5AE1" bold="true" box="[151,274,1201,1223]" pageId="9" pageNumber="441">FIGURE 5.</emphasis>
<taxonomicName id="4C7E4D7B8F28356EFE9DFB30FDA85AE1" box="[281,545,1201,1223]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="441" phylum="Arthropoda" rank="species" species="appalachiensis">
<emphasis id="B90AEAEA8F28356EFE9DFB30FDA85AE1" box="[281,545,1201,1223]" italics="true" pageId="9" pageNumber="441">Cambarus appalachiensis</emphasis>
</taxonomicName>
distribution.
</paragraph>
</caption>
<paragraph id="8BC136F88F28356EFF43FB6FFDF45B9D" blockId="9.[151,1436,1262,2016]" pageId="9" pageNumber="441">
<emphasis id="B90AEAEA8F28356EFF43FB6FFDEA5B21" bold="true" box="[199,611,1262,1287]" pageId="9" pageNumber="441">Range and specimens examined.</emphasis>
<taxonomicName id="4C7E4D7B8F28356EFDF7FB6FFC1E5B20" box="[627,919,1262,1286]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="441" phylum="Arthropoda" rank="species" species="appalachiensis">
<emphasis id="B90AEAEA8F28356EFDF7FB6FFC1E5B20" box="[627,919,1262,1286]" italics="true" pageId="9" pageNumber="441">Cambarus appalachiensis</emphasis>
</taxonomicName>
(
<figureCitation id="13452A7D8F28356EFC2BFB6FFC735B21" box="[943,1018,1262,1287]" captionStart="FIGURE 5" captionStartId="9.[151,250,1201,1223]" captionTargetBox="[151,1436,193,1180]" captionTargetId="figure@9.[151,1436,193,1180]" captionTargetPageId="9" captionText="FIGURE 5. Cambarus appalachiensis distribution." httpUri="https://zenodo.org/record/400119/files/figure.png" pageId="9" pageNumber="441">Fig. 5</figureCitation>
) occurs in the Ridge and Valley, Appalachian Plateau, and Allegheny Mountain physiographic provinces in the upper New River watershed in the middle New River basin in 
<collectingRegion id="49BAF81A8F28356EFE4DFAB7FDAD5B69" box="[457,548,1334,1359]" country="United States of America" name="Virginia" pageId="9" pageNumber="441">Virginia</collectingRegion>
and 
<collectingRegion id="49BAF81A8F28356EFDDEFAB6FD7F5B69" box="[602,758,1334,1359]" country="United States of America" name="West Virginia" pageId="9" pageNumber="441">West Virginia</collectingRegion>
, and the lower New, upper Kanawha, Gauley and Greenbrier river basins in 
<collectingRegion id="49BAF81A8F28356EFEC5FADAFE555B55" box="[321,476,1370,1395]" country="United States of America" name="West Virginia" pageId="9" pageNumber="441">West Virginia</collectingRegion>
(
<figureCitation id="13452A7D8F28356EFE6AFADBFDBD5B55" box="[494,564,1370,1395]" captionStart="FIGURE 5" captionStartId="9.[151,250,1201,1223]" captionTargetBox="[151,1436,193,1180]" captionTargetId="figure@9.[151,1436,193,1180]" captionTargetPageId="9" captionText="FIGURE 5. Cambarus appalachiensis distribution." httpUri="https://zenodo.org/record/400119/files/figure.png" pageId="9" pageNumber="441">Fig. 5</figureCitation>
). The southern/upstream portion of the range of 
<taxonomicName id="4C7E4D7B8F28356EFBE5FADAFAA55B54" box="[1121,1324,1370,1394]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="441" phylum="Arthropoda" rank="species" species="appalachiensis">
<emphasis id="B90AEAEA8F28356EFBE5FADAFAA55B54" box="[1121,1324,1370,1394]" italics="true" pageId="9" pageNumber="441">C. appalachiensis</emphasis>
</taxonomicName>
occurs in the Middle New River mainstem. Robust populations currently persist in the New River, as well as Peak Creek and Little River and their associated tributaries.
</paragraph>
</subSubSection>
<subSubSection id="C36465738F28356EFF43FA47FD99592D" pageId="9" pageNumber="441" type="reference_group">
<paragraph id="8BC136F88F28356EFF43FA47FD99592D" blockId="9.[151,1436,1262,2016]" pageId="9" pageNumber="441">
<taxonomicName id="4C7E4D7B8F28356EFF43FA47FE785BF8" ID-CoL="7RTVV" box="[199,497,1478,1502]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="441" phylum="Arthropoda" rank="species" species="appalachiensis">
<emphasis id="B90AEAEA8F28356EFF43FA47FE785BF8" box="[199,497,1478,1502]" italics="true" pageId="9" pageNumber="441">Cambarus appalachiensis</emphasis>
</taxonomicName>
is the dominant tertiary burrower in the middle New River basin, with large populations occurring in the New River mainstem and the Bluestone River, as well as all major New River tributaries. Greenbrier watershed populations are limited to the lower reaches of the basin, with the farthest upstream populations reported in Wolf Creek, Monroe County. Within the lower New River, 
<taxonomicName id="4C7E4D7B8F28356EFB54F9AAFA155864" ID-CoL="7RTVV" box="[1232,1436,1578,1602]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="441" phylum="Arthropoda" rank="species" species="appalachiensis">
<emphasis id="B90AEAEA8F28356EFB54F9AAFA155864" box="[1232,1436,1578,1602]" italics="true" pageId="9" pageNumber="441">C. appalachiensis</emphasis>
</taxonomicName>
occurs within the mainstem as well as all large tributaries. Gauley River drainage populations are stable and robust in the mainstem of the Gauley River, as well as the Williams, Cherry, Cranberry and Meadow Rivers and all associated tributaries, where 
<taxonomicName id="4C7E4D7B8F28356EFE78F90EFD465880" ID-CoL="7RTVV" box="[508,719,1678,1702]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="441" phylum="Arthropoda" rank="species" species="appalachiensis">
<emphasis id="B90AEAEA8F28356EFE78F90EFD465880" box="[508,719,1678,1702]" italics="true" pageId="9" pageNumber="441">C. appalachiensis</emphasis>
</taxonomicName>
is the dominant 
<taxonomicName id="4C7E4D7B8F28356EFC30F90FFBA05880" ID-CoL="7NRQP" box="[948,1065,1678,1702]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="441" phylum="Arthropoda" rank="genus">
<emphasis id="B90AEAEA8F28356EFC30F90FFBA05880" box="[948,1065,1678,1702]" italics="true" pageId="9" pageNumber="441">Cambarus</emphasis>
</taxonomicName>
species. Streams immediately downstream of Kanawha Falls in the upper Kanawha River are the only waterbodies within the basin that maintain 
<taxonomicName id="4C7E4D7B8F28356EFF13F950FEEC58CF" ID-CoL="7RTVV" box="[151,357,1745,1769]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="441" phylum="Arthropoda" rank="species" species="appalachiensis">
<emphasis id="B90AEAEA8F28356EFF13F950FEEC58CF" box="[151,357,1745,1769]" italics="true" pageId="9" pageNumber="441">C. appalachiensis</emphasis>
</taxonomicName>
. Downstream of Kanawha falls 
<taxonomicName id="4C7E4D7B8F28356EFD6FF950FC3358CF" ID-CoL="7RTVV" box="[747,954,1745,1769]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="441" phylum="Arthropoda" rank="species" species="appalachiensis">
<emphasis id="B90AEAEA8F28356EFD6FF950FC3358CF" box="[747,954,1745,1769]" italics="true" pageId="9" pageNumber="441">C. appalachiensis</emphasis>
</taxonomicName>
is replaced by its ecological equivalent, 
<taxonomicName id="4C7E4D7B8F28356EFF13F973FD85592D" ID-CoL="7RV4R" authority="Girard, 1852" authorityName="Girard" authorityYear="1852" box="[151,524,1778,1803]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="441" phylum="Arthropoda" rank="species" species="robustus">
<emphasis id="B90AEAEA8F28356EFF13F973FEFB592C" box="[151,370,1778,1802]" italics="true" pageId="9" pageNumber="441">Cambarus robustus</emphasis>
<bibRefCitation id="EFEF4B098F28356EFEFDF973FD85592D" author="Girard" box="[377,524,1778,1803]" pageId="9" pageNumber="441" refString="Girard, C. (1852) A revision of the North American Astaci, with observations on their habits and geographic distribution. Proceedings of the Academy of Natural Sciences of Philadelphia, 6, 87 - 91." type="journal article" year="1852">Girard, 1852</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C36465738F28356CFF43F895FC7558D9" lastPageId="11" lastPageNumber="443" pageId="9" pageNumber="441" type="materials_examined">
<paragraph id="8BC136F88F28356EFF43F895FE395952" blockId="9.[151,1436,1262,2016]" pageId="9" pageNumber="441">
Material examined included 1,089 specimens from 189 collections housed at either The 
<collectingRegion id="49BAF81A8F28356EFB2CF895FB69590B" box="[1192,1248,1812,1837]" country="United States of America" name="Ohio" pageId="9" pageNumber="441">Ohio</collectingRegion>
State University Museum of Biological Diversity Collection or the West Liberty University Astacology Collection and are presented in Appendix 1.
</paragraph>
<paragraph id="8BC136F88F28356DFF43F8FEFB9D5F42" blockId="9.[151,1436,1262,2016]" lastBlockId="10.[151,1437,151,1184]" lastPageId="10" lastPageNumber="442" pageId="9" pageNumber="441">
<emphasis id="B90AEAEA8F28356EFF43F8FEFDB059BE" bold="true" box="[199,569,1919,1944]" pageId="9" pageNumber="441">Habitat and life history notes.</emphasis>
The within-basin abundance and distribution of 
<taxonomicName id="4C7E4D7B8F28356EFBF0F801FA1559BE" box="[1140,1436,1920,1944]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="441" phylum="Arthropoda" rank="species" species="appalachiensis">
<emphasis id="B90AEAEA8F28356EFBF0F801FA1559BE" box="[1140,1436,1920,1944]" italics="true" pageId="9" pageNumber="441">Cambarus appalachiensis</emphasis>
</taxonomicName>
differs among drainage basins. In both the upper and middle New River, 
<taxonomicName id="4C7E4D7B8F28356EFC5AF825FB21599A" box="[990,1192,1956,1980]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="441" phylum="Arthropoda" rank="species" species="appalachiensis">
<emphasis id="B90AEAEA8F28356EFC5AF825FB21599A" box="[990,1192,1956,1980]" italics="true" pageId="9" pageNumber="441">C. appalachiensis</emphasis>
</taxonomicName>
is abundant in larger ordered streams, as well as the New River mainstem. As the New River increases in order, 
<taxonomicName id="4C7E4D7B8F28356EFB4BF849FA1559C6" box="[1231,1436,1992,2016]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="441" phylum="Arthropoda" rank="species" species="appalachiensis">
<emphasis id="B90AEAEA8F28356EFB4BF849FA1559C6" box="[1231,1436,1992,2016]" italics="true" pageId="9" pageNumber="441">C. appalachiensis</emphasis>
</taxonomicName>
occurrence decreases in the mainstem, and is limited primarily to larger ordered tributaries. Throughout the Gauley River system 
<taxonomicName id="4C7E4D7B8F2B356DFEB0FF3DFE765EF2" box="[308,511,188,212]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="442" phylum="Arthropoda" rank="species" species="appalachiensis">
<emphasis id="B90AEAEA8F2B356DFEB0FF3DFE765EF2" box="[308,511,188,212]" italics="true" pageId="10" pageNumber="442">C. appalachiensis</emphasis>
</taxonomicName>
is the dominant tertiary burrowing crayfish and occurs in all stream orders from small headwater systems through the Gauley River mainstem. In all situations, 
<taxonomicName id="4C7E4D7B8F2B356DFB8FFF61FB5D5EDE" box="[1035,1236,224,248]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="442" phylum="Arthropoda" rank="species" species="appalachiensis">
<emphasis id="B90AEAEA8F2B356DFB8FFF61FB5D5EDE" box="[1035,1236,224,248]" italics="true" pageId="10" pageNumber="442">C. appalachiensis</emphasis>
</taxonomicName>
occurs in streams with substrates of mixed cobble and gravels with isolated large boulders and slabs, which are the species preferred habitat (RFT and ZJL personal observation). 
<taxonomicName id="4C7E4D7B8F2B356DFD12FEA9FC335F66" box="[662,954,296,320]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="442" phylum="Arthropoda" rank="species" species="appalachiensis">
<emphasis id="B90AEAEA8F2B356DFD12FEA9FC335F66" box="[662,954,296,320]" italics="true" pageId="10" pageNumber="442">Cambarus appalachiensis</emphasis>
</taxonomicName>
also occurs in a myriad of gradients, from slack water eddies and slow runs, to rapids and high gradient step-pool streams.
</paragraph>
<paragraph id="8BC136F88F2B356DFF43FEEEFEF85C81" blockId="10.[151,1437,151,1184]" pageId="10" pageNumber="442">
<materialsCitation id="3B163CA58F2B356DFF43FEEEFD325C46" ID-GBIF-Occurrence="1440156658" collectorName="Based &amp; Appalachia &amp; Jezerinac &amp; Foltz &amp; Form I &amp; over Form II &amp; Population &amp; Zachary J. Loughman &amp; The &amp; June &amp; Form I." location="Pipestem Creek" pageId="10" pageNumber="442" specimenCount="1" stateProvince="West Virginia" typeStatus="holotype">
<collectorName id="268B532E8F2B356DFF43FEEEFE825FAE" box="[199,267,367,392]" pageId="10" pageNumber="442">Based</collectorName>
on collections examined to complete the description, 
<taxonomicName id="4C7E4D7B8F2B356DFCFEFEF1FBCD5FAE" box="[890,1092,368,392]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="442" phylum="Arthropoda" rank="species" species="appalachiensis">
<emphasis id="B90AEAEA8F2B356DFCFEFEF1FBCD5FAE" box="[890,1092,368,392]" italics="true" pageId="10" pageNumber="442">C. appalachiensis</emphasis>
</taxonomicName>
appears to have a life history similar to that of other larger tertiary burrowing 
<taxonomicName id="4C7E4D7B8F2B356DFD35FE15FCAF5F8A" box="[689,806,404,428]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="442" phylum="Arthropoda" rank="genus">
<emphasis id="B90AEAEA8F2B356DFD35FE15FCAF5F8A" box="[689,806,404,428]" italics="true" pageId="10" pageNumber="442">Cambarus</emphasis>
</taxonomicName>
species in 
<collectorName id="268B532E8F2B356DFC25FE15FBA95F8B" box="[929,1056,404,429]" pageId="10" pageNumber="442">Appalachia</collectorName>
(
<bibRefCitation id="EFEF4B098F2B356DFBAAFE15FA915F8B" author="Jezerinac" box="[1070,1304,404,429]" pageId="10" pageNumber="442" refString="Jezerinac, R. F., Stocker, G. W. &amp; Tarter, D. C. (1995) The crayfishes (Decapoda: Cambaridae) of West Virginia. Bulletin of the Ohio Biological Survey, 10, 1 - 193." type="journal article" year="1995">
<collectorName id="268B532E8F2B356DFBAAFE15FB115F8B" box="[1070,1176,404,429]" pageId="10" pageNumber="442">Jezerinac</collectorName>
<emphasis id="B90AEAEA8F2B356DFB1BFE14FB5E5F8A" box="[1183,1239,404,428]" italics="true" pageId="10" pageNumber="442">et al.</emphasis>
1995
</bibRefCitation>
; 
<bibRefCitation id="EFEF4B098F2B356DFAA0FE15FF5D5FF6" author="Foltz" pageId="10" pageNumber="442" refString="Foltz, D. A., Nolan IV., E. J., McGill, K. T. &amp; Loughman, Z. J. (2016) Life history of the coal fields crayfish Cambarus theepiensis Loughman, Foltz, Garrison and Welsh 2013 (Decapoda: Astacoidea: Cambaridae) in southwestern West Virginia, USA. Journal of Crustacean Biology, 36, 628 - 636." type="journal article" year="2016">
<collectorName id="268B532E8F2B356DFAA0FE15FAD75F8B" box="[1316,1374,404,429]" pageId="10" pageNumber="442">Foltz</collectorName>
<emphasis id="B90AEAEA8F2B356DFAE0FE14FA155F8A" box="[1380,1436,404,428]" italics="true" pageId="10" pageNumber="442">et al.</emphasis>
2016
</bibRefCitation>
). 
<collectorName id="268B532E8F2B356DFF62FE39FEBB5FE9" box="[230,306,440,463]" pageId="10" pageNumber="442">Form I</collectorName>
males dominate 
<collectorName id="268B532E8F2B356DFE6BFE39FDF75FE9" box="[495,638,440,463]" pageId="10" pageNumber="442">over Form II</collectorName>
males during autumn, winter and early spring months. 
<collectorName id="268B532E8F2B356DFB62FE36FAE95FF6" box="[1254,1376,439,464]" pageId="10" pageNumber="442">Population</collectorName>
wide molts occur in 
<collectingRegion id="49BAF81A8F2B356DFEC6FE5CFE555FD3" box="[322,476,476,501]" country="United States of America" name="West Virginia" pageId="10" pageNumber="442">West Virginia</collectingRegion>
beginning in mid-May, and are complete by mid-June (
<collectorName id="268B532E8F2B356DFBDCFE5DFB135FD2" box="[1112,1178,476,500]" name="Zachary J. Loughman" pageId="10" pageNumber="442">Z.J.L.</collectorName>
personal observation). 
<collectorName id="268B532E8F2B356DFF13FE7EFF4A5C3E" box="[151,195,511,536]" pageId="10" pageNumber="442">The</collectorName>
majority of adult males are form II from the latter part of 
<collectorName id="268B532E8F2B356DFC1FFD81FC475C31" box="[923,974,512,535]" pageId="10" pageNumber="442">June</collectorName>
through mid-September, when another population wide molt occurs, at which time the majority of males are 
<collectorName id="268B532E8F2B356DFC6CFDA4FBC85C1A" box="[1000,1089,549,572]" pageId="10" pageNumber="442">Form I.</collectorName>
This was evident during the collection of the 
<typeStatus id="54C5885A8F2B356DFEDAFDC9FE075C46" box="[350,398,584,608]" pageId="10" pageNumber="442">type</typeStatus>
series in 
<location id="8EA160238F2B356DFD80FDC6FD325C46" LSID="urn:lsid:plazi:treatment:03D787EE8F223569FF13FADDFDFB5C3E:8EA160238F2B356DFD80FDC6FD325C46" box="[516,699,583,608]" name="Pipestem Creek" pageId="10" pageNumber="442" stateProvince="West Virginia">Pipestem Creek</location>
</materialsCitation>
, 
<materialsCitation id="3B163CA58F2B356DFD4DFDC9FEF85C81" ID-GBIF-Occurrence="1440156655" collectingDate="2016-10-11" collectorName="Form II" location="Fifty-one" pageId="10" pageNumber="442" specimenCount="1" stateProvince="Summer County West Virginia" typeStatus="holotype">
<collectingRegion id="49BAF81A8F2B356DFD4DFDC9FBB95C46" box="[713,1072,583,608]" inferredBySuffix="true" pageId="10" pageNumber="442">
Summer County 
<collectingRegion id="49BAF81A8F2B356DFC14FDC9FBB95C46" box="[912,1072,583,608]" country="United States of America" name="West Virginia" pageId="10" pageNumber="442">West Virginia</collectingRegion>
</collectingRegion>
. Not all animals captured were retained, though basic biological data was ascertained. 
<location id="8EA160238F2B356DFC89FDEDFCFC5CA3" LSID="urn:lsid:plazi:treatment:03D787EE8F223569FF13FADDFDFB5C3E:8EA160238F2B356DFC89FDEDFCFC5CA3" box="[781,885,620,645]" name="Fifty-one" pageId="10" pageNumber="442" stateProvince="Summer County West Virginia">Fifty-one</location>
males were collected on 
<date id="FFC010388F2B356DFB1FFDEDFAD75CA3" box="[1179,1374,620,645]" pageId="10" pageNumber="442" value="2016-10-11">
<collectingDate id="EF84E9D08F2B356DFB1FFDEDFAD75CA3" box="[1179,1374,620,645]" pageId="10" pageNumber="442" value="2016-10-11">11 October 2016</collectingDate>
</date>
; only three were 
<collectorName id="268B532E8F2B356DFE97FD11FEE45C81" box="[275,365,656,679]" pageId="10" pageNumber="442">Form II</collectorName>
.
</materialsCitation>
</paragraph>
<paragraph id="8BC136F88F2B356DFF43FD35FE475DA6" blockId="10.[151,1437,151,1184]" pageId="10" pageNumber="442">
Mating likely occurs in the fall, winter and spring months. During the collection of the 
<typeStatus id="54C5885A8F2B356DFB38FD34FB655CEB" box="[1212,1260,693,717]" pageId="10" pageNumber="442">type</typeStatus>
series on three separate instances Form I males were collected alongside two to three females under large slabrock boulders. While mating was not observed, it was possible cohabitation in this scenario was associated with possible mating efforts. Active glare glands were not observed on any females collected in the 
<typeStatus id="54C5885A8F2B356DFBC5FCA1FBF85D1E" box="[1089,1137,800,824]" pageId="10" pageNumber="442">type</typeStatus>
series (n = 89). Females collected in early and mid-summer in both June and July in 
<collectingRegion id="49BAF81A8F2B356DFCC8FCC4FC6E5D7B" box="[844,999,836,861]" country="United States of America" name="West Virginia" pageId="10" pageNumber="442">West Virginia</collectingRegion>
frequently display active glare glands (ZJL personal observation).
</paragraph>
<paragraph id="8BC136F88F2B356DFF43FC0DFA105A86" blockId="10.[151,1437,151,1184]" pageId="10" pageNumber="442">
Ovigerous females as well as females carrying instars have been collected on several occasions in both 
<collectingRegion id="49BAF81A8F2B356DFF13FC2EFF7B5DEE" box="[151,242,943,968]" country="United States of America" name="Virginia" pageId="10" pageNumber="442">Virginia</collectingRegion>
and 
<collectingRegion id="49BAF81A8F2B356DFEACFC31FE495DEE" box="[296,448,943,968]" country="United States of America" name="West Virginia" pageId="10" pageNumber="442">West Virginia</collectingRegion>
(
<tableCitation id="C6FC03438F2B356DFE4BFC2EFDAF5DEE" box="[463,550,943,968]" captionStart="TABLE 2" captionStartId="10.[151,239,1228,1251]" captionTargetBox="[160,1399,1278,2014]" captionTargetPageId="10" captionText="TABLE 2. Cambarus appalachiensis pleopodal egg and juvenile counts." httpUri="http://table.plazi.org/id/DF0166708F2B356DFF13FB4DFC295AC5" pageId="10" pageNumber="442" tableUuid="DF0166708F2B356DFF13FB4DFC295AC5">Table 2</tableCitation>
). The earliest recorded date for females carrying eggs in this study was 
<date id="FFC010388F2B356DFAC2FC31FA115DEE" box="[1350,1432,943,968]" pageId="10" pageNumber="442" value="1924-07">24 July</date>
, 1989 with the latest occurrence of eggs on 
<date id="FFC010388F2B356DFD05FC55FD7F5DCA" box="[641,758,980,1004]" pageId="10" pageNumber="442" value="1919-08">19 August</date>
(
<tableCitation id="C6FC03438F2B356DFC82FC55FCD65DCA" box="[774,863,980,1005]" captionStart="TABLE 2" captionStartId="10.[151,239,1228,1251]" captionTargetBox="[160,1399,1278,2014]" captionTargetPageId="10" captionText="TABLE 2. Cambarus appalachiensis pleopodal egg and juvenile counts." httpUri="http://table.plazi.org/id/DF0166708F2B356DFF13FB4DFC295AC5" pageId="10" pageNumber="442" tableUuid="DF0166708F2B356DFF13FB4DFC295AC5">Table 2</tableCitation>
). The earliest recording of juveniles was 
<date id="FFC010388F2B356DFAB1FC55FA155DCA" box="[1333,1436,980,1004]" pageId="10" pageNumber="442">2 August</date>
when a female was collected with stage I juveniles. Females likely maintain juveniles throughout the fall, and possibly into the spring. During the collection of the 
<typeStatus id="54C5885A8F2B356DFD6DFB9CFC905A13" box="[745,793,1053,1077]" pageId="10" pageNumber="442">type</typeStatus>
series two females were collected maintaining 56 and 78 stage IV juveniles capable of living freely from their mothers (
<date id="FFC010388F2B356DFC00FBC1FB8E5A7E" box="[900,1031,1087,1112]" pageId="10" pageNumber="442" value="2011-10">11 October</date>
, 2016). Mean egg counts for nine females was 105.9 eggs; mean number of juveniles for 
<specimenCount id="9D78FD718F2B356DFCB6FBE5FC2A5A5B" box="[818,931,1124,1149]" pageId="10" pageNumber="442" type="female">6 females</specimenCount>
was 43.9 juveniles. The highest fecundity observed was 
<specimenCount id="9D78FD718F2B356DFEB1FB09FE425A86" box="[309,459,1159,1184]" pageId="10" pageNumber="442" type="juvenile">337 juveniles</specimenCount>
from a female collected in the Gauley River, Webster County, 
<collectingRegion id="49BAF81A8F2B356DFB08FB08FAAF5A86" box="[1164,1318,1159,1184]" country="United States of America" name="West Virginia" pageId="10" pageNumber="442">West Virginia</collectingRegion>
(
<tableCitation id="C6FC03438F2B356DFAB0FB06FA055A86" box="[1332,1420,1159,1184]" captionStart="TABLE 2" captionStartId="10.[151,239,1228,1251]" captionTargetBox="[160,1399,1278,2014]" captionTargetPageId="10" captionText="TABLE 2. Cambarus appalachiensis pleopodal egg and juvenile counts." httpUri="http://table.plazi.org/id/DF0166708F2B356DFF13FB4DFC295AC5" pageId="10" pageNumber="442" tableUuid="DF0166708F2B356DFF13FB4DFC295AC5">Table 2</tableCitation>
).
</paragraph>
<caption id="DF0166708F2B356DFF13FB4DFC295AC5" ID-Table-UUID="DF0166708F2B356DFF13FB4DFC295AC5" box="[151,928,1227,1251]" httpUri="http://table.plazi.org/id/DF0166708F2B356DFF13FB4DFC295AC5" pageId="10" pageNumber="442" targetBox="[160,1399,1278,2014]" targetIsTable="true" targetPageId="10">
<paragraph id="8BC136F88F2B356DFF13FB4DFC295AC5" blockId="10.[151,928,1227,1251]" box="[151,928,1227,1251]" pageId="10" pageNumber="442">
<emphasis id="B90AEAEA8F2B356DFF13FB4DFE805AC5" bold="true" box="[151,265,1228,1251]" pageId="10" pageNumber="442">TABLE 2.</emphasis>
<taxonomicName id="4C7E4D7B8F2B356DFE94FB4AFDAF5AC5" box="[272,550,1227,1251]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="442" phylum="Arthropoda" rank="species" species="appalachiensis">
<emphasis id="B90AEAEA8F2B356DFE94FB4AFDAF5AC5" box="[272,550,1227,1251]" italics="true" pageId="10" pageNumber="442">Cambarus appalachiensis</emphasis>
</taxonomicName>
pleopodal egg and juvenile counts.
</paragraph>
</caption>
<paragraph id="8BC136F88F2B356DFF24FB7FFDC559F8" pageId="10" pageNumber="442">
<table id="F97EC4588F2BCA98FF24FB7FFAFE59F8" box="[160,1399,1278,2014]" gridcols="8" gridrows="18" pageId="10" pageNumber="442">
<tr id="354E34BA8F2BCA98FF24FB7FFAFE5B32" box="[160,1399,1278,1300]" gridrow="0" pageId="10" pageNumber="442">
<th id="769F5DC68F2BCA98FF24FB7FFEF75B32" box="[160,382,1278,1300]" gridcol="0" gridrow="0" pageId="10" pageNumber="442">OSUMC #</th>
<th id="769F5DC68F2BCA98FE24FB7FFE6F5B32" box="[416,486,1278,1300]" gridcol="1" gridrow="0" pageId="10" pageNumber="442"># Eggs</th>
<th id="769F5DC68F2BCA98FD97FB7FFDE35B32" box="[531,618,1278,1300]" gridcol="2" gridrow="0" pageId="10" pageNumber="442"># Instars</th>
<th id="769F5DC68F2BCA98FD10FB7FFD725B32" box="[660,763,1278,1300]" gridcol="3" gridrow="0" pageId="10" pageNumber="442">Date</th>
<th id="769F5DC68F2BCA98FCB0FB7FFC285B32" box="[820,929,1278,1300]" gridcol="4" gridrow="0" pageId="10" pageNumber="442">County</th>
<th id="769F5DC68F2BCA98FC5EFB7FFBFA5B32" box="[986,1139,1278,1300]" gridcol="5" gridrow="0" pageId="10" pageNumber="442">Stream</th>
<th id="769F5DC68F2BCA98FB2CFB7FFA9C5B32" box="[1192,1301,1278,1300]" gridcol="6" gridrow="0" pageId="10" pageNumber="442">County</th>
<th id="769F5DC68F2BCA98FAC1FB7FFAFE5B32" box="[1349,1399,1278,1300]" gridcol="7" gridrow="0" pageId="10" pageNumber="442">State</th>
</tr>
<tr id="354E34BA8F2BCA98FF24FAA9FAFE5B18" box="[160,1399,1320,1342]" gridrow="1" pageId="10" pageNumber="442">
<th id="769F5DC68F2BCA98FF24FAA9FEF75B18" box="[160,382,1320,1342]" gridcol="0" gridrow="1" pageId="10" pageNumber="442">7647</th>
<td id="769F5DC68F2BCA98FE24FAA9FE6F5B18" box="[416,486,1320,1342]" gridcol="1" gridrow="1" pageId="10" pageNumber="442">159</td>
<td id="769F5DC68F2BCA98FD97FAA9FDE35B18" box="[531,618,1320,1342]" gridcol="2" gridrow="1" pageId="10" pageNumber="442">0</td>
<td id="769F5DC68F2BCA98FD10FAA9FD725B18" box="[660,763,1320,1342]" gridcol="3" gridrow="1" pageId="10" pageNumber="442">7/24/1989</td>
<td id="769F5DC68F2BCA98FCB0FAA9FC285B18" box="[820,929,1320,1342]" gridcol="4" gridrow="1" pageId="10" pageNumber="442">Monroe</td>
<td id="769F5DC68F2BCA98FC5EFAA9FBFA5B18" box="[986,1139,1320,1342]" gridcol="5" gridrow="1" pageId="10" pageNumber="442">Turkey Cr.</td>
<td id="769F5DC68F2BCA98FB2CFAA9FA9C5B18" box="[1192,1301,1320,1342]" gridcol="6" gridrow="1" pageId="10" pageNumber="442">Monroe</td>
<td id="769F5DC68F2BCA98FAC1FAA9FAFE5B18" box="[1349,1399,1320,1342]" gridcol="7" gridrow="1" pageId="10" pageNumber="442">WV</td>
</tr>
<tr id="354E34BA8F2BCA98FF24FAD3FAFE5B4E" box="[160,1399,1362,1384]" gridrow="2" pageId="10" pageNumber="442">
<th id="769F5DC68F2BCA98FF24FAD3FEF75B4E" box="[160,382,1362,1384]" gridcol="0" gridrow="2" pageId="10" pageNumber="442">7647</th>
<td id="769F5DC68F2BCA98FE24FAD3FE6F5B4E" box="[416,486,1362,1384]" gridcol="1" gridrow="2" pageId="10" pageNumber="442">117</td>
<td id="769F5DC68F2BCA98FD97FAD3FDE35B4E" box="[531,618,1362,1384]" gridcol="2" gridrow="2" pageId="10" pageNumber="442">0</td>
<td id="769F5DC68F2BCA98FD10FAD3FD725B4E" box="[660,763,1362,1384]" gridcol="3" gridrow="2" pageId="10" pageNumber="442">7/24/1989</td>
<td id="769F5DC68F2BCA98FCB0FAD3FC285B4E" box="[820,929,1362,1384]" gridcol="4" gridrow="2" pageId="10" pageNumber="442">Monroe</td>
<td id="769F5DC68F2BCA98FC5EFAD3FBFA5B4E" box="[986,1139,1362,1384]" gridcol="5" gridrow="2" pageId="10" pageNumber="442">Turkey Cr.</td>
<td id="769F5DC68F2BCA98FB2CFAD3FA9C5B4E" box="[1192,1301,1362,1384]" gridcol="6" gridrow="2" pageId="10" pageNumber="442">Monroe</td>
<td id="769F5DC68F2BCA98FAC1FAD3FAFE5B4E" box="[1349,1399,1362,1384]" gridcol="7" gridrow="2" pageId="10" pageNumber="442">WV</td>
</tr>
<tr id="354E34BA8F2BCA98FF24FAFDFAFE5BB4" box="[160,1399,1404,1426]" gridrow="3" pageId="10" pageNumber="442">
<th id="769F5DC68F2BCA98FF24FAFDFEF75BB4" box="[160,382,1404,1426]" gridcol="0" gridrow="3" pageId="10" pageNumber="442">7647</th>
<td id="769F5DC68F2BCA98FE24FAFDFE6F5BB4" box="[416,486,1404,1426]" gridcol="1" gridrow="3" pageId="10" pageNumber="442">98</td>
<td id="769F5DC68F2BCA98FD97FAFDFDE35BB4" box="[531,618,1404,1426]" gridcol="2" gridrow="3" pageId="10" pageNumber="442">0</td>
<td id="769F5DC68F2BCA98FD10FAFDFD725BB4" box="[660,763,1404,1426]" gridcol="3" gridrow="3" pageId="10" pageNumber="442">7/24/1989</td>
<td id="769F5DC68F2BCA98FCB0FAFDFC285BB4" box="[820,929,1404,1426]" gridcol="4" gridrow="3" pageId="10" pageNumber="442">Monroe</td>
<td id="769F5DC68F2BCA98FC5EFAFDFBFA5BB4" box="[986,1139,1404,1426]" gridcol="5" gridrow="3" pageId="10" pageNumber="442">Turkey Cr.</td>
<td id="769F5DC68F2BCA98FB2CFAFDFA9C5BB4" box="[1192,1301,1404,1426]" gridcol="6" gridrow="3" pageId="10" pageNumber="442">Monroe</td>
<td id="769F5DC68F2BCA98FAC1FAFDFAFE5BB4" box="[1349,1399,1404,1426]" gridcol="7" gridrow="3" pageId="10" pageNumber="442">WV</td>
</tr>
<tr id="354E34BA8F2BCA98FF24FA27FAFE5B9A" box="[160,1399,1446,1468]" gridrow="4" pageId="10" pageNumber="442">
<th id="769F5DC68F2BCA98FF24FA27FEF75B9A" box="[160,382,1446,1468]" gridcol="0" gridrow="4" pageId="10" pageNumber="442">6869</th>
<td id="769F5DC68F2BCA98FE24FA27FE6F5B9A" box="[416,486,1446,1468]" gridcol="1" gridrow="4" pageId="10" pageNumber="442">82</td>
<td id="769F5DC68F2BCA98FD97FA27FDE35B9A" box="[531,618,1446,1468]" gridcol="2" gridrow="4" pageId="10" pageNumber="442">0</td>
<td id="769F5DC68F2BCA98FD10FA27FD725B9A" box="[660,763,1446,1468]" gridcol="3" gridrow="4" pageId="10" pageNumber="442">8/2/2007</td>
<td id="769F5DC68F2BCA98FCB0FA27FC285B9A" box="[820,929,1446,1468]" gridcol="4" gridrow="4" pageId="10" pageNumber="442">Tazewell</td>
<td id="769F5DC68F2BCA98FC5EFA27FBFA5B9A" box="[986,1139,1446,1468]" gridcol="5" gridrow="4" pageId="10" pageNumber="442">Mud Fork</td>
<td id="769F5DC68F2BCA98FB2CFA27FA9C5B9A" box="[1192,1301,1446,1468]" gridcol="6" gridrow="4" pageId="10" pageNumber="442">Tazewell</td>
<td id="769F5DC68F2BCA98FAC1FA27FAFE5B9A" box="[1349,1399,1446,1468]" gridcol="7" gridrow="4" pageId="10" pageNumber="442">VA</td>
</tr>
<tr id="354E34BA8F2BCA98FF24FA51FAFE5BC0" box="[160,1399,1488,1510]" gridrow="5" pageId="10" pageNumber="442">
<th id="769F5DC68F2BCA98FF24FA51FEF75BC0" box="[160,382,1488,1510]" gridcol="0" gridrow="5" pageId="10" pageNumber="442">6874</th>
<td id="769F5DC68F2BCA98FE24FA51FE6F5BC0" box="[416,486,1488,1510]" gridcol="1" gridrow="5" pageId="10" pageNumber="442">0</td>
<td id="769F5DC68F2BCA98FD97FA51FDE35BC0" box="[531,618,1488,1510]" gridcol="2" gridrow="5" pageId="10" pageNumber="442">58</td>
<td id="769F5DC68F2BCA98FD10FA51FD725BC0" box="[660,763,1488,1510]" gridcol="3" gridrow="5" pageId="10" pageNumber="442">8/2/2007</td>
<td id="769F5DC68F2BCA98FCB0FA51FC285BC0" box="[820,929,1488,1510]" gridcol="4" gridrow="5" pageId="10" pageNumber="442">Tazewell</td>
<td id="769F5DC68F2BCA98FC5EFA51FBFA5BC0" box="[986,1139,1488,1510]" gridcol="5" gridrow="5" pageId="10" pageNumber="442">Bluestone R.</td>
<td id="769F5DC68F2BCA98FB2CFA51FA9C5BC0" box="[1192,1301,1488,1510]" gridcol="6" gridrow="5" pageId="10" pageNumber="442">Tazewell</td>
<td id="769F5DC68F2BCA98FAC1FA51FAFE5BC0" box="[1349,1399,1488,1510]" gridcol="7" gridrow="5" pageId="10" pageNumber="442">VA</td>
</tr>
<tr id="354E34BA8F2BCA98FF24FA7BFAFE5836" box="[160,1399,1530,1552]" gridrow="6" pageId="10" pageNumber="442">
<th id="769F5DC68F2BCA98FF24FA7BFEF75836" box="[160,382,1530,1552]" gridcol="0" gridrow="6" pageId="10" pageNumber="442">7123</th>
<td id="769F5DC68F2BCA98FE24FA7BFE6F5836" box="[416,486,1530,1552]" gridcol="1" gridrow="6" pageId="10" pageNumber="442">161</td>
<td id="769F5DC68F2BCA98FD97FA7BFDE35836" box="[531,618,1530,1552]" gridcol="2" gridrow="6" pageId="10" pageNumber="442">0</td>
<td id="769F5DC68F2BCA98FD10FA7BFD725836" box="[660,763,1530,1552]" gridcol="3" gridrow="6" pageId="10" pageNumber="442">8/18/1988</td>
<td id="769F5DC68F2BCA98FCB0FA7BFC285836" box="[820,929,1530,1552]" gridcol="4" gridrow="6" pageId="10" pageNumber="442">Monroe</td>
<td id="769F5DC68F2BCA98FC5EFA7BFBFA5836" box="[986,1139,1530,1552]" gridcol="5" gridrow="6" pageId="10" pageNumber="442">Wolf Cr.</td>
<td id="769F5DC68F2BCA98FB2CFA7BFA9C5836" box="[1192,1301,1530,1552]" gridcol="6" gridrow="6" pageId="10" pageNumber="442">Monroe</td>
<td id="769F5DC68F2BCA98FAC1FA7BFAFE5836" box="[1349,1399,1530,1552]" gridcol="7" gridrow="6" pageId="10" pageNumber="442">WV</td>
</tr>
<tr id="354E34BA8F2BCA98FF24F9A5FAFE581C" box="[160,1399,1572,1594]" gridrow="7" pageId="10" pageNumber="442">
<th id="769F5DC68F2BCA98FF24F9A5FEF7581C" box="[160,382,1572,1594]" gridcol="0" gridrow="7" pageId="10" pageNumber="442">7189</th>
<td id="769F5DC68F2BCA98FE24F9A5FE6F581C" box="[416,486,1572,1594]" gridcol="1" gridrow="7" pageId="10" pageNumber="442">118</td>
<td id="769F5DC68F2BCA98FD97F9A5FDE3581C" box="[531,618,1572,1594]" gridcol="2" gridrow="7" pageId="10" pageNumber="442">0</td>
<td id="769F5DC68F2BCA98FD10F9A5FD72581C" box="[660,763,1572,1594]" gridcol="3" gridrow="7" pageId="10" pageNumber="442">8/18/1988</td>
<td id="769F5DC68F2BCA98FCB0F9A5FC28581C" box="[820,929,1572,1594]" gridcol="4" gridrow="7" pageId="10" pageNumber="442">Monroe</td>
<td id="769F5DC68F2BCA98FC5EF9A5FBFA581C" box="[986,1139,1572,1594]" gridcol="5" gridrow="7" pageId="10" pageNumber="442">Indian Cr.</td>
<td id="769F5DC68F2BCA98FB2CF9A5FA9C581C" box="[1192,1301,1572,1594]" gridcol="6" gridrow="7" pageId="10" pageNumber="442">Monroe</td>
<td id="769F5DC68F2BCA98FAC1F9A5FAFE581C" box="[1349,1399,1572,1594]" gridcol="7" gridrow="7" pageId="10" pageNumber="442">WV</td>
</tr>
<tr id="354E34BA8F2BCA98FF24F9CFFAFE5842" box="[160,1399,1614,1636]" gridrow="8" pageId="10" pageNumber="442">
<th id="769F5DC68F2BCA98FF24F9CFFEF75842" box="[160,382,1614,1636]" gridcol="0" gridrow="8" pageId="10" pageNumber="442">7189</th>
<td id="769F5DC68F2BCA98FE24F9CFFE6F5842" box="[416,486,1614,1636]" gridcol="1" gridrow="8" pageId="10" pageNumber="442">88</td>
<td id="769F5DC68F2BCA98FD97F9CFFDE35842" box="[531,618,1614,1636]" gridcol="2" gridrow="8" pageId="10" pageNumber="442">0</td>
<td id="769F5DC68F2BCA98FD10F9CFFD725842" box="[660,763,1614,1636]" gridcol="3" gridrow="8" pageId="10" pageNumber="442">8/18/1988</td>
<td id="769F5DC68F2BCA98FCB0F9CFFC285842" box="[820,929,1614,1636]" gridcol="4" gridrow="8" pageId="10" pageNumber="442">Monroe</td>
<td id="769F5DC68F2BCA98FC5EF9CFFBFA5842" box="[986,1139,1614,1636]" gridcol="5" gridrow="8" pageId="10" pageNumber="442">Indian Cr.</td>
<td id="769F5DC68F2BCA98FB2CF9CFFA9C5842" box="[1192,1301,1614,1636]" gridcol="6" gridrow="8" pageId="10" pageNumber="442">Monroe</td>
<td id="769F5DC68F2BCA98FAC1F9CFFAFE5842" box="[1349,1399,1614,1636]" gridcol="7" gridrow="8" pageId="10" pageNumber="442">WV</td>
</tr>
<tr id="354E34BA8F2BCA98FF24F9F9FAFE58A8" box="[160,1399,1656,1678]" gridrow="9" pageId="10" pageNumber="442">
<th id="769F5DC68F2BCA98FF24F9F9FEF758A8" box="[160,382,1656,1678]" gridcol="0" gridrow="9" pageId="10" pageNumber="442">7189</th>
<td id="769F5DC68F2BCA98FE24F9F9FE6F58A8" box="[416,486,1656,1678]" gridcol="1" gridrow="9" pageId="10" pageNumber="442">15</td>
<td id="769F5DC68F2BCA98FD97F9F9FDE358A8" box="[531,618,1656,1678]" gridcol="2" gridrow="9" pageId="10" pageNumber="442">0</td>
<td id="769F5DC68F2BCA98FD10F9F9FD7258A8" box="[660,763,1656,1678]" gridcol="3" gridrow="9" pageId="10" pageNumber="442">8/18/1988</td>
<td id="769F5DC68F2BCA98FCB0F9F9FC2858A8" box="[820,929,1656,1678]" gridcol="4" gridrow="9" pageId="10" pageNumber="442">Monroe</td>
<td id="769F5DC68F2BCA98FC5EF9F9FBFA58A8" box="[986,1139,1656,1678]" gridcol="5" gridrow="9" pageId="10" pageNumber="442">Indian Cr.</td>
<td id="769F5DC68F2BCA98FB2CF9F9FA9C58A8" box="[1192,1301,1656,1678]" gridcol="6" gridrow="9" pageId="10" pageNumber="442">Monroe</td>
<td id="769F5DC68F2BCA98FAC1F9F9FAFE58A8" box="[1349,1399,1656,1678]" gridcol="7" gridrow="9" pageId="10" pageNumber="442">WV</td>
</tr>
<tr id="354E34BA8F2BCA98FF24F923FAFE589E" box="[160,1399,1698,1720]" gridrow="10" pageId="10" pageNumber="442">
<th id="769F5DC68F2BCA98FF24F923FEF7589E" box="[160,382,1698,1720]" gridcol="0" gridrow="10" pageId="10" pageNumber="442">7124</th>
<td id="769F5DC68F2BCA98FE24F923FE6F589E" box="[416,486,1698,1720]" gridcol="1" gridrow="10" pageId="10" pageNumber="442">115</td>
<td id="769F5DC68F2BCA98FD97F923FDE3589E" box="[531,618,1698,1720]" gridcol="2" gridrow="10" pageId="10" pageNumber="442">0</td>
<td id="769F5DC68F2BCA98FD10F923FD72589E" box="[660,763,1698,1720]" gridcol="3" gridrow="10" pageId="10" pageNumber="442">8/19/1988</td>
<td id="769F5DC68F2BCA98FCB0F923FC28589E" box="[820,929,1698,1720]" gridcol="4" gridrow="10" pageId="10" pageNumber="442">Summers</td>
<td id="769F5DC68F2BCA98FC5EF923FBFA589E" box="[986,1139,1698,1720]" gridcol="5" gridrow="10" pageId="10" pageNumber="442">Hungard Cr.</td>
<td id="769F5DC68F2BCA98FB2CF923FA9C589E" box="[1192,1301,1698,1720]" gridcol="6" gridrow="10" pageId="10" pageNumber="442">Summers</td>
<td id="769F5DC68F2BCA98FAC1F923FAFE589E" box="[1349,1399,1698,1720]" gridcol="7" gridrow="10" pageId="10" pageNumber="442">WV</td>
</tr>
<tr id="354E34BA8F2BCA98FF24F94DFAFE58C4" box="[160,1399,1740,1762]" gridrow="11" pageId="10" pageNumber="442">
<th id="769F5DC68F2BCA98FF24F94DFEF758C4" box="[160,382,1740,1762]" gridcol="0" gridrow="11" pageId="10" pageNumber="442">7276</th>
<td id="769F5DC68F2BCA98FE24F94DFE6F58C4" box="[416,486,1740,1762]" gridcol="1" gridrow="11" pageId="10" pageNumber="442">0</td>
<td id="769F5DC68F2BCA98FD97F94DFDE358C4" box="[531,618,1740,1762]" gridcol="2" gridrow="11" pageId="10" pageNumber="442">10</td>
<td id="769F5DC68F2BCA98FD10F94DFD7258C4" box="[660,763,1740,1762]" gridcol="3" gridrow="11" pageId="10" pageNumber="442">8/20/1988</td>
<td id="769F5DC68F2BCA98FCB0F94DFC2858C4" box="[820,929,1740,1762]" gridcol="4" gridrow="11" pageId="10" pageNumber="442">Greenbrier</td>
<td id="769F5DC68F2BCA98FC5EF94DFBFA58C4" box="[986,1139,1740,1762]" gridcol="5" gridrow="11" pageId="10" pageNumber="442">Little Clear Cr.</td>
<td id="769F5DC68F2BCA98FB2CF94DFA9C58C4" box="[1192,1301,1740,1762]" gridcol="6" gridrow="11" pageId="10" pageNumber="442">Greenbrier</td>
<td id="769F5DC68F2BCA98FAC1F94DFAFE58C4" box="[1349,1399,1740,1762]" gridcol="7" gridrow="11" pageId="10" pageNumber="442">WV</td>
</tr>
<tr id="354E34BA8F2BCA98FF24F977FAFE592A" box="[160,1399,1782,1804]" gridrow="12" pageId="10" pageNumber="442">
<th id="769F5DC68F2BCA98FF24F977FEF7592A" box="[160,382,1782,1804]" gridcol="0" gridrow="12" pageId="10" pageNumber="442">7191</th>
<td id="769F5DC68F2BCA98FE24F977FE6F592A" box="[416,486,1782,1804]" gridcol="1" gridrow="12" pageId="10" pageNumber="442">0</td>
<td id="769F5DC68F2BCA98FD97F977FDE3592A" box="[531,618,1782,1804]" gridcol="2" gridrow="12" pageId="10" pageNumber="442">337</td>
<td id="769F5DC68F2BCA98FD10F977FD72592A" box="[660,763,1782,1804]" gridcol="3" gridrow="12" pageId="10" pageNumber="442">8/22/1988</td>
<td id="769F5DC68F2BCA98FCB0F977FC28592A" box="[820,929,1782,1804]" gridcol="4" gridrow="12" pageId="10" pageNumber="442">Webster</td>
<td id="769F5DC68F2BCA98FC5EF977FBFA592A" box="[986,1139,1782,1804]" gridcol="5" gridrow="12" pageId="10" pageNumber="442">Gauley R.</td>
<td id="769F5DC68F2BCA98FB2CF977FA9C592A" box="[1192,1301,1782,1804]" gridcol="6" gridrow="12" pageId="10" pageNumber="442">Webster</td>
<td id="769F5DC68F2BCA98FAC1F977FAFE592A" box="[1349,1399,1782,1804]" gridcol="7" gridrow="12" pageId="10" pageNumber="442">WV</td>
</tr>
<tr id="354E34BA8F2BCA98FF24F8A1FAFE5910" box="[160,1399,1824,1846]" gridrow="13" pageId="10" pageNumber="442">
<th id="769F5DC68F2BCA98FF24F8A1FEF75910" box="[160,382,1824,1846]" gridcol="0" gridrow="13" pageId="10" pageNumber="442">7118</th>
<td id="769F5DC68F2BCA98FE24F8A1FE6F5910" box="[416,486,1824,1846]" gridcol="1" gridrow="13" pageId="10" pageNumber="442">0</td>
<td id="769F5DC68F2BCA98FD97F8A1FDE35910" box="[531,618,1824,1846]" gridcol="2" gridrow="13" pageId="10" pageNumber="442">18</td>
<td id="769F5DC68F2BCA98FD10F8A1FD725910" box="[660,763,1824,1846]" gridcol="3" gridrow="13" pageId="10" pageNumber="442">9/1/1989</td>
<td id="769F5DC68F2BCA98FCB0F8A1FC285910" box="[820,929,1824,1846]" gridcol="4" gridrow="13" pageId="10" pageNumber="442">McDowell</td>
<td id="769F5DC68F2BCA98FC5EF8A1FBFA5910" box="[986,1139,1824,1846]" gridcol="5" gridrow="13" pageId="10" pageNumber="442">Elkhorn Cr.</td>
<td id="769F5DC68F2BCA98FB2CF8A1FA9C5910" box="[1192,1301,1824,1846]" gridcol="6" gridrow="13" pageId="10" pageNumber="442">McDowell</td>
<td id="769F5DC68F2BCA98FAC1F8A1FAFE5910" box="[1349,1399,1824,1846]" gridcol="7" gridrow="13" pageId="10" pageNumber="442">WV</td>
</tr>
<tr id="354E34BA8F2BCA98FF24F8CBFAFE5946" box="[160,1399,1866,1888]" gridrow="14" pageId="10" pageNumber="442">
<th id="769F5DC68F2BCA98FF24F8CBFEF75946" box="[160,382,1866,1888]" gridcol="0" gridrow="14" pageId="10" pageNumber="442">7266</th>
<td id="769F5DC68F2BCA98FE24F8CBFE6F5946" box="[416,486,1866,1888]" gridcol="1" gridrow="14" pageId="10" pageNumber="442">0</td>
<td id="769F5DC68F2BCA98FD97F8CBFDE35946" box="[531,618,1866,1888]" gridcol="2" gridrow="14" pageId="10" pageNumber="442">54</td>
<td id="769F5DC68F2BCA98FD10F8CBFD725946" box="[660,763,1866,1888]" gridcol="3" gridrow="14" pageId="10" pageNumber="442">9/10/1988</td>
<td id="769F5DC68F2BCA98FCB0F8CBFC285946" box="[820,929,1866,1888]" gridcol="4" gridrow="14" pageId="10" pageNumber="442">Nicholas</td>
<td id="769F5DC68F2BCA98FC5EF8CBFBFA5946" box="[986,1139,1866,1888]" gridcol="5" gridrow="14" pageId="10" pageNumber="442">Cranberry R.</td>
<td id="769F5DC68F2BCA98FB2CF8CBFA9C5946" box="[1192,1301,1866,1888]" gridcol="6" gridrow="14" pageId="10" pageNumber="442">Nicholas</td>
<td id="769F5DC68F2BCA98FAC1F8CBFAFE5946" box="[1349,1399,1866,1888]" gridcol="7" gridrow="14" pageId="10" pageNumber="442">WV</td>
</tr>
<tr id="354E34BA8F2BCA98FF24F8F5FAFE59AC" box="[160,1399,1908,1930]" gridrow="15" pageId="10" pageNumber="442">
<th id="769F5DC68F2BCA98FF24F8F5FEF759AC" box="[160,382,1908,1930]" gridcol="0" gridrow="15" pageId="10" pageNumber="442">7543</th>
<td id="769F5DC68F2BCA98FE24F8F5FE6F59AC" box="[416,486,1908,1930]" gridcol="1" gridrow="15" pageId="10" pageNumber="442">0</td>
<td id="769F5DC68F2BCA98FD97F8F5FDE359AC" box="[531,618,1908,1930]" gridcol="2" gridrow="15" pageId="10" pageNumber="442">62</td>
<td id="769F5DC68F2BCA98FD10F8F5FD7259AC" box="[660,763,1908,1930]" gridcol="3" gridrow="15" pageId="10" pageNumber="442">9/10/1988</td>
<td id="769F5DC68F2BCA98FCB0F8F5FC2859AC" box="[820,929,1908,1930]" gridcol="4" gridrow="15" pageId="10" pageNumber="442">Nicholas</td>
<td id="769F5DC68F2BCA98FC5EF8F5FBFA59AC" box="[986,1139,1908,1930]" gridcol="5" gridrow="15" pageId="10" pageNumber="442">Cherry R.</td>
<td id="769F5DC68F2BCA98FB2CF8F5FA9C59AC" box="[1192,1301,1908,1930]" gridcol="6" gridrow="15" pageId="10" pageNumber="442">Nicholas</td>
<td id="769F5DC68F2BCA98FAC1F8F5FAFE59AC" box="[1349,1399,1908,1930]" gridcol="7" gridrow="15" pageId="10" pageNumber="442">WV</td>
</tr>
<tr id="354E34BA8F2BCA98FF24F81FFAFE5992" box="[160,1399,1950,1972]" gridrow="16" pageId="10" pageNumber="442" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
<th id="769F5DC68F2BCA98FF24F81FFEF75992" box="[160,382,1950,1972]" gridcol="0" gridrow="16" pageId="10" pageNumber="442">X egg/instar count =</th>
<td id="769F5DC68F2BCA98FE24F81FFE6F5992" box="[416,486,1950,1972]" gridcol="1" gridrow="16" pageId="10" pageNumber="442">105.9</td>
<td id="769F5DC68F2BCA98FD97F81FFDE35992" box="[531,618,1950,1972]" gridcol="2" gridrow="16" pageId="10" pageNumber="442">150.8</td>
</tr>
<tr id="354E34BA8F2BCA98FF24F849FAFE59F8" box="[160,1399,1992,2014]" gridrow="17" pageId="10" pageNumber="442" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
<th id="769F5DC68F2BCA98FF24F849FEF759F8" box="[160,382,1992,2014]" gridcol="0" gridrow="17" pageId="10" pageNumber="442">SD egg/instar count =</th>
<td id="769F5DC68F2BCA98FE24F849FE6F59F8" box="[416,486,1992,2014]" gridcol="1" gridrow="17" pageId="10" pageNumber="442">43.9</td>
<td id="769F5DC68F2BCA98FD97F849FDE359F8" box="[531,618,1992,2014]" gridcol="2" gridrow="17" pageId="10" pageNumber="442">123.0</td>
</tr>
</table>
</paragraph>
<caption id="DF0166708F2A356CFF13FA5DFD195836" httpUri="https://zenodo.org/record/400120/files/figure.png" pageId="11" pageNumber="443" targetBox="[151,1436,193,1478]" targetPageId="11">
<paragraph id="8BC136F88F2A356CFF13FA5DFD195836" blockId="11.[151,1436,1499,1552]" pageId="11" pageNumber="443">
<emphasis id="B90AEAEA8F2A356CFF13FA5DFBDF5BD4" bold="true" box="[151,1110,1499,1522]" pageId="11" pageNumber="443">
FIGURE 6. 
<taxonomicName id="4C7E4D7B8F2A356CFE9DFA5AFDAE5BD4" box="[281,551,1499,1522]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="443" phylum="Arthropoda" rank="species" species="appalachiensis">
<emphasis id="B90AEAEA8F2A356CFE9DFA5AFDAE5BD4" bold="true" box="[281,551,1499,1522]" italics="true" pageId="11" pageNumber="443">Cambarus appalachiensis</emphasis>
</taxonomicName>
chelae morphotypes: (A.) Elongate; (B.) Truncated.
</emphasis>
Over 90% of 
<taxonomicName id="4C7E4D7B8F2A356CFB62FA5DFA155BD4" box="[1254,1436,1500,1522]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="443" phylum="Arthropoda" rank="species" species="appalachiensis">
<emphasis id="B90AEAEA8F2A356CFB62FA5DFA155BD4" box="[1254,1436,1500,1522]" italics="true" pageId="11" pageNumber="443">C. appalachiensis</emphasis>
</taxonomicName>
examined maintained the elongated morphotypes.
</paragraph>
</caption>
<paragraph id="8BC136F88F2A356CFF43F9D7FC7558D9" blockId="11.[151,1436,1622,1935]" pageId="11" pageNumber="443">
<emphasis id="B90AEAEA8F2A356CFF43F9D7FE355849" bold="true" box="[199,444,1622,1647]" pageId="11" pageNumber="443">Conservation status.</emphasis>
<taxonomicName id="4C7E4D7B8F2A356CFE40F9D7FD605848" box="[452,745,1622,1646]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="443" phylum="Arthropoda" rank="species" species="appalachiensis">
<emphasis id="B90AEAEA8F2A356CFE40F9D7FD605848" box="[452,745,1622,1646]" italics="true" pageId="11" pageNumber="443">Cambarus appalachiensis</emphasis>
</taxonomicName>
should be listed as currently stable (CS) using the American Fisheries Society criteria (
<bibRefCitation id="EFEF4B098F2A356CFE4EF9FBFD2C58B5" author="Taylor" box="[458,677,1658,1683]" pageId="11" pageNumber="443" refString="Taylor, C. A., Schuster, G. A., Cooper, J. E., Distefano, R. J., Eversole, A. G., Hamr, P., Hobbs H. H. III, Robison, H. W., Skelton, C. E. &amp; Thoma, R. F. (2007) Reassessment of the conservation status of crayfishes of the United States and Canada after 10 + years of increased awareness. Fisheries, 32, 372 - 389." type="journal article" year="2007">
Taylor 
<emphasis id="B90AEAEA8F2A356CFDA4F9FAFDD758B4" box="[544,606,1658,1682]" italics="true" pageId="11" pageNumber="443">et al.</emphasis>
2007
</bibRefCitation>
), and assigned a G4/G5 ranking using the 
<bibRefCitation id="EFEF4B098F2A356CFB22F9FAFAC358B5" author="Master" box="[1190,1354,1659,1683]" pageId="11" pageNumber="443" refString="Master, L. (1991) Assessing threats and setting priorities for conservation. Conservation Biology, 5, 559 - 563. https: // doi. org / 10.1111 / j. 1523 - 1739.1991. tb 00370. x" type="journal article" year="1991">Master (1991)</bibRefCitation>
global conservation criteria for conservation listing because of its broad range and stable status. 
<taxonomicName id="4C7E4D7B8F2A356CFBFDF91FFA155890" box="[1145,1436,1694,1718]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="443" phylum="Arthropoda" rank="species" species="appalachiensis">
<emphasis id="B90AEAEA8F2A356CFBFDF91FFA155890" box="[1145,1436,1694,1718]" italics="true" pageId="11" pageNumber="443">Cambarus appalachiensis</emphasis>
</taxonomicName>
should be listed as least concern (LC) using the International Union for the Conservation of Nature (
<collectionCode id="ED6FAE3D8F2A356CFA88F942FADB58FD" box="[1292,1362,1731,1755]" pageId="11" pageNumber="443">IUCN</collectionCode>
2001) criteria because of its broad distribution (ZJL and RFT personal observation).
</paragraph>
</subSubSection>
<subSubSection id="C36465738F2A356AFF43F88BFDD25DD8" lastPageId="13" lastPageNumber="445" pageId="11" pageNumber="443" type="description">
<paragraph id="8BC136F88F2A356CFF43F88BFEA559A9" blockId="11.[151,1436,1622,1935]" pageId="11" pageNumber="443">
<emphasis id="B90AEAEA8F2A356CFF43F88BFE3D5905" bold="true" box="[199,436,1802,1827]" pageId="11" pageNumber="443">Crayfish associates.</emphasis>
<taxonomicName id="4C7E4D7B8F2A356CFE39F88BFD6A5904" box="[445,739,1802,1826]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="443" phylum="Arthropoda" rank="species" species="appalachiensis">
<emphasis id="B90AEAEA8F2A356CFE39F88BFD6A5904" box="[445,739,1802,1826]" italics="true" pageId="11" pageNumber="443">Cambarus appalachiensis</emphasis>
</taxonomicName>
has been collected with 
<taxonomicName id="4C7E4D7B8F2A356CFB86F88BFE915961" authority="Hay, 1914" authorityName="Hay" authorityYear="1914" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="443" phylum="Arthropoda" rank="species" species="carinirostris" subGenus="Cambarus">
<emphasis id="B90AEAEA8F2A356CFB86F88BFBFE5904" box="[1026,1143,1802,1826]" italics="true" pageId="11" pageNumber="443">Cambarus</emphasis>
(
<emphasis id="B90AEAEA8F2A356CFB0EF88BFB765904" box="[1162,1279,1802,1826]" italics="true" pageId="11" pageNumber="443">Cambarus</emphasis>
) 
<emphasis id="B90AEAEA8F2A356CFA95F88AFA155904" box="[1297,1436,1803,1826]" italics="true" pageId="11" pageNumber="443">carinirostris</emphasis>
<bibRefCitation id="EFEF4B098F2A356CFF13F8AEFE915961" author="Hay" box="[151,280,1839,1863]" pageId="11" pageNumber="443" refString="Hay, W. P. (1914) Cambarus bartonii carinirostris In: Faxon, W. (1914) Notes on the crayfishes in the United States National Museum and the Museum of Comparative Zoology with descriptions of new species and subspecies to which is appended a catalogue of the known species and subspecies. Memoirs of the Museum of Comparative Zoology, Harvard College, 40, 351 - 427." type="journal article" year="1914">Hay, 1914</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C7E4D7B8F2A356CFEABF8AFFC535961" authority="James 1966" authorityName="James" authorityYear="1966" box="[303,986,1838,1863]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="443" phylum="Arthropoda" rank="species" species="chasmodactylus" subGenus="Hiaticambarus">
<emphasis id="B90AEAEA8F2A356CFEABF8AFFE2D5960" box="[303,420,1838,1862]" italics="true" pageId="11" pageNumber="443">Cambarus</emphasis>
(
<emphasis id="B90AEAEA8F2A356CFE3BF8AFFDEE5960" box="[447,615,1838,1862]" italics="true" pageId="11" pageNumber="443">Hiaticambarus</emphasis>
) 
<emphasis id="B90AEAEA8F2A356CFD07F8AFFCBF5960" box="[643,822,1838,1862]" italics="true" pageId="11" pageNumber="443">chasmodactylus</emphasis>
<bibRefCitation id="EFEF4B098F2A356CFCCCF8AEFC535961" author="James" box="[840,986,1838,1863]" pageId="11" pageNumber="443" refString="James, H. A. (1966) Range and variations of the subspecies of Cambarus longulus (Decapoda, Astacidae). Proceedings of the United States National Museum, 119, 1 - 24." type="journal article" year="1966">James 1966</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C7E4D7B8F2A356CFC75F8AEFEB8594D" authority="Faxon,1884" authorityName="Faxon" authorityYear="1884" class="Malacostraca" family="Cambaridae" genus="Orconectes" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="443" phylum="Arthropoda" rank="species" species="sanbornii" subGenus="Crockerinus">
<emphasis id="B90AEAEA8F2A356CFC75F8AEFBE65960" box="[1009,1135,1839,1862]" italics="true" pageId="11" pageNumber="443">Orconectes</emphasis>
(
<emphasis id="B90AEAEA8F2A356CFB0FF8AFFA9D5960" box="[1163,1300,1838,1862]" italics="true" pageId="11" pageNumber="443">Crockerinus</emphasis>
) 
<emphasis id="B90AEAEA8F2A356CFAB5F8AFFA155960" box="[1329,1436,1838,1862]" italics="true" pageId="11" pageNumber="443">sanbornii</emphasis>
(
<bibRefCitation id="EFEF4B098F2A356CFF1BF8D2FEA0594D" author="Faxon" box="[159,297,1875,1899]" pageId="11" pageNumber="443" refString="Faxon, W. A. (1884) Descriptions of new species of Cambarus; to which is added a synonymical list of the known species of Cambarus and Astacus. Proceedings of the American Academy of Arts and Sciences, 20, 107 - 158. https: // doi. org / 10.2307 / 25138768" type="journal article" year="1884">Faxon,1884</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="4C7E4D7B8F2A356CFEB8F8D2FCF2594D" authority="Hagen, 1840" authorityName="Hagen" authorityYear="1840" box="[316,891,1874,1899]" class="Malacostraca" family="Cambaridae" genus="Orconectes" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="443" phylum="Arthropoda" rank="species" species="virilis" subGenus="Gremicambarus">
<emphasis id="B90AEAEA8F2A356CFEB8F8D2FE32594C" box="[316,443,1875,1898]" italics="true" pageId="11" pageNumber="443">Orconectes</emphasis>
(
<emphasis id="B90AEAEA8F2A356CFE4EF8D3FDF6594C" box="[458,639,1874,1898]" italics="true" pageId="11" pageNumber="443">Gremicambarus</emphasis>
) 
<emphasis id="B90AEAEA8F2A356CFD0BF8D3FD58594C" box="[655,721,1874,1898]" italics="true" pageId="11" pageNumber="443">virilis</emphasis>
(
<bibRefCitation id="EFEF4B098F2A356CFD5BF8D2FCFA594D" author="Hagen" box="[735,883,1875,1899]" pageId="11" pageNumber="443" refString="Hagen, H. A. (1840) Monograph of the North American Astacidae. Illustrated Catalogue of the Museum of Comparative Zoology. Welch, Bigelow &amp; co., University Press, Cambridge, 109 pp." type="book" year="1840">Hagen, 1840</bibRefCitation>
)
</taxonomicName>
, and 
<taxonomicName id="4C7E4D7B8F2A356CFC31F8D2FEA159A9" authority="Taylor, 2000" authorityName="Taylor" authorityYear="2000" class="Malacostraca" family="Cambaridae" genus="Orconectes" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="443" phylum="Arthropoda" rank="species" species="cristavarius" subGenus="Procericambarus">
<emphasis id="B90AEAEA8F2A356CFC31F8D2FBBD594C" box="[949,1076,1875,1898]" italics="true" pageId="11" pageNumber="443">Orconectes</emphasis>
(
<emphasis id="B90AEAEA8F2A356CFBC7F8D3FA8F594C" box="[1091,1286,1874,1898]" italics="true" pageId="11" pageNumber="443">Procericambarus</emphasis>
) 
<emphasis id="B90AEAEA8F2A356CFA92F8D2FA15594C" box="[1302,1436,1875,1898]" italics="true" pageId="11" pageNumber="443">cristavarius</emphasis>
<bibRefCitation id="EFEF4B098F2A356CFF13F8F7FEA159A9" author="Taylor" box="[151,296,1910,1935]" pageId="11" pageNumber="443" refString="Taylor, C. A. (2000) Systematic studies of the Orconectes juvenilis complex (Decapoda: Cambaridae), with descriptions of two new species. Journal of Crustacean Biology, 20, 132 - 152." type="journal article" year="2000">Taylor, 2000</bibRefCitation>
</taxonomicName>
.
</paragraph>
<caption id="DF0166708F2D356BFF13FA60FAEA5BD1" box="[151,1379,1505,1527]" httpUri="https://zenodo.org/record/400121/files/figure.png" pageId="12" pageNumber="444" targetBox="[151,1436,195,1480]" targetPageId="12">
<paragraph id="8BC136F88F2D356BFF13FA60FAEA5BD1" blockId="12.[151,1379,1505,1527]" box="[151,1379,1505,1527]" pageId="12" pageNumber="444">
<emphasis id="B90AEAEA8F2D356BFF13FA60FE9B5BD1" bold="true" box="[151,274,1505,1527]" pageId="12" pageNumber="444">FIGURE 7.</emphasis>
Differences in chelae structure between 
<taxonomicName id="4C7E4D7B8F2D356BFD2BFA60FC7C5BD1" authority="Top" authorityName="Top" box="[687,1013,1505,1527]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="444" phylum="Arthropoda" rank="species" species="appalachiensis">
<emphasis id="B90AEAEA8F2D356BFD2BFA60FC3E5BD1" box="[687,951,1505,1527]" italics="true" pageId="12" pageNumber="444">Cambarus appalachiensis</emphasis>
(Top)
</taxonomicName>
and 
<taxonomicName id="4C7E4D7B8F2D356BFBA2FA60FAD65BD1" authority="Bottom" authorityName="Bottom" box="[1062,1375,1505,1527]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="444" phylum="Arthropoda" rank="species" species="sciotensis">
<emphasis id="B90AEAEA8F2D356BFBA2FA60FB705BD1" box="[1062,1273,1505,1527]" italics="true" pageId="12" pageNumber="444">Cambarus sciotensis</emphasis>
(Bottom)
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BC136F88F2D356BFF43F99FFB425929" blockId="12.[151,1437,1566,2010]" pageId="12" pageNumber="444">
<emphasis id="B90AEAEA8F2D356BFF43F99FFEB65811" bold="true" box="[199,319,1566,1591]" pageId="12" pageNumber="444">Variation.</emphasis>
As in many 
<taxonomicName id="4C7E4D7B8F2D356BFE57F99FFDC15810" box="[467,584,1566,1590]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="444" phylum="Arthropoda" rank="genus">
<emphasis id="B90AEAEA8F2D356BFE57F99FFDC15810" box="[467,584,1566,1590]" italics="true" pageId="12" pageNumber="444">Cambarus</emphasis>
</taxonomicName>
species, 
<taxonomicName id="4C7E4D7B8F2D356BFD35F99EFCF25810" box="[689,891,1566,1590]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="444" phylum="Arthropoda" rank="species" species="appalachiensis">
<emphasis id="B90AEAEA8F2D356BFD35F99EFCF25810" box="[689,891,1566,1590]" italics="true" pageId="12" pageNumber="444">C. appalachiensis</emphasis>
</taxonomicName>
undergoes an ontogenic change in morphology, with juveniles and sub-adults being noticeably more spinose than adults. Specifically, juvenile postorbital ridges as well as acumen terminate in well pronounced spines, compared to adults whose postorbital ridges and acumens often terminate in either tubercles or spinose tubercles, rarely in defined spines. Juvenile/sub-adult rostra also are less broad and more lanceolate than adult rostra, which normally consist of parallel margins that are thickened, ending in strongly angled acumen, which are often absent in juveniles/sub-adults. Finally, chelae architecture is reduced in juvenile/subadult 
<taxonomicName id="4C7E4D7B8F2D356BFE5FF976FD2D5928" box="[475,676,1782,1806]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="444" phylum="Arthropoda" rank="species" species="appalachiensis">
<emphasis id="B90AEAEA8F2D356BFE5FF976FD2D5928" box="[475,676,1782,1806]" italics="true" pageId="12" pageNumber="444">C. appalachiensis</emphasis>
</taxonomicName>
and defined in adult animals as explained below.
</paragraph>
<paragraph id="8BC136F88F2D356AFF43F89AFD005F5E" blockId="12.[151,1437,1566,2010]" lastBlockId="13.[151,1436,151,2031]" lastPageId="13" lastPageNumber="445" pageId="12" pageNumber="444">
Two distinct chelae 
<typeStatus id="54C5885A8F2D356BFE22F89AFE685915" box="[422,481,1819,1843]" pageId="12" pageNumber="444">types</typeStatus>
exist in 
<taxonomicName id="4C7E4D7B8F2D356BFDBBF89AFC8E5914" box="[575,775,1818,1842]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="444" phylum="Arthropoda" rank="species" species="appalachiensis">
<emphasis id="B90AEAEA8F2D356BFDBBF89AFC8E5914" box="[575,775,1818,1842]" italics="true" pageId="12" pageNumber="444">C. appalachiensis</emphasis>
</taxonomicName>
, with both forms present in Form I and Form II individuals (
<figureCitation id="13452A7D8F2D356BFF1BF8BAFF6C5972" box="[159,229,1851,1876]" captionStart="FIGURE 6" captionStartId="11.[151,250,1500,1522]" captionTargetBox="[151,1436,193,1478]" captionTargetId="figure@11.[151,1436,193,1478]" captionTargetPageId="11" captionText="FIGURE 6. Cambarus appalachiensis chelae morphotypes: (A.) Elongate; (B.) Truncated. Over 90 % of C. appalachiensis examined maintained the elongated morphotypes." httpUri="https://zenodo.org/record/400120/files/figure.png" pageId="12" pageNumber="444">Fig. 6</figureCitation>
A–B). Elongated chelae (
<figureCitation id="13452A7D8F2D356BFD89F8BAFDDA5972" box="[525,595,1851,1876]" captionStart="FIGURE 6" captionStartId="11.[151,250,1500,1522]" captionTargetBox="[151,1436,193,1478]" captionTargetId="figure@11.[151,1436,193,1478]" captionTargetPageId="11" captionText="FIGURE 6. Cambarus appalachiensis chelae morphotypes: (A.) Elongate; (B.) Truncated. Over 90 % of C. appalachiensis examined maintained the elongated morphotypes." httpUri="https://zenodo.org/record/400120/files/figure.png" pageId="12" pageNumber="444">Fig. 6</figureCitation>
A) are far more frequently encountered than truncated chelae (
<figureCitation id="13452A7D8F2D356BFAB7F8BAFAF35972" box="[1331,1402,1851,1876]" captionStart="FIGURE 6" captionStartId="11.[151,250,1500,1522]" captionTargetBox="[151,1436,193,1478]" captionTargetId="figure@11.[151,1436,193,1478]" captionTargetPageId="11" captionText="FIGURE 6. Cambarus appalachiensis chelae morphotypes: (A.) Elongate; (B.) Truncated. Over 90 % of C. appalachiensis examined maintained the elongated morphotypes." httpUri="https://zenodo.org/record/400120/files/figure.png" pageId="12" pageNumber="444">Fig. 6</figureCitation>
B). Among adults, there is variation between animals in the Greenbrier, the lower, middle, and upper New River Basins, and the Gauley River basin. Chelae of adult 
<taxonomicName id="4C7E4D7B8F2D356BFD69F8FEFC3E59B0" box="[749,951,1918,1942]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="444" phylum="Arthropoda" rank="species" species="appalachiensis">
<emphasis id="B90AEAEA8F2D356BFD69F8FEFC3E59B0" box="[749,951,1918,1942]" italics="true" pageId="12" pageNumber="444">C. appalachiensis</emphasis>
</taxonomicName>
from the Greenbrier and New river basins lack well defined lateral impressions, lack extensive tuberculation on the dorsal surface of the dactyl, and often possess an extensive gape between the dactyl and fixed finger of the propodus when the dactyl is closed. Gauley River 
<taxonomicName id="4C7E4D7B8F2C356AFF5FFF19FE2D5E96" box="[219,420,152,176]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="445" phylum="Arthropoda" rank="species" species="appalachiensis">
<emphasis id="B90AEAEA8F2C356AFF5FFF19FE2D5E96" box="[219,420,152,176]" italics="true" pageId="13" pageNumber="445">C. appalachiensis</emphasis>
</taxonomicName>
have a more defined lateral impression, moderate tuberculation on the dorsal surface of the dactyl, and moderate to weak gape between the dactyl and propodus when the dactyl is closed. In addition to differences in chelae architecture, Greenbrier and New river animal’s rostra are normally broad, with steeply angled acumen. Gauley River 
<taxonomicName id="4C7E4D7B8F2C356AFD83FF7DFD5E5F32" box="[519,727,252,276]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="445" phylum="Arthropoda" rank="species" species="appalachiensis">
<emphasis id="B90AEAEA8F2C356AFD83FF7DFD5E5F32" box="[519,727,252,276]" italics="true" pageId="13" pageNumber="445">C. appalachiensis</emphasis>
</taxonomicName>
rostra can be less broad, occasionally approaching a sublanceolate condition, and can have less steeply angled acumen, though most animals observed possess rostra similar to those from other basins. The latter is an exception in the Gauley, but does occur with enough frequency to be mentioned (ZJL personal observation).
</paragraph>
<paragraph id="8BC136F88F2C356AFF43FE00FAD75C20" blockId="13.[151,1436,151,2031]" pageId="13" pageNumber="445">
<emphasis id="B90AEAEA8F2C356AFF43FE00FDCB5FBC" bold="true" box="[199,578,385,410]" pageId="13" pageNumber="445">Relationships and comparisons.</emphasis>
<taxonomicName id="4C7E4D7B8F2C356AFDCDFE00FCE45FBF" box="[585,877,385,409]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="445" phylum="Arthropoda" rank="species" species="appalachiensis">
<emphasis id="B90AEAEA8F2C356AFDCDFE00FCE45FBF" box="[585,877,385,409]" italics="true" pageId="13" pageNumber="445">Cambarus appalachiensis</emphasis>
</taxonomicName>
is placed in the subgenus 
<taxonomicName id="4C7E4D7B8F2C356AFB11FE00FA835FBF" box="[1173,1290,385,409]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="445" phylum="Arthropoda" rank="genus">
<emphasis id="B90AEAEA8F2C356AFB11FE00FA835FBF" box="[1173,1290,385,409]" italics="true" pageId="13" pageNumber="445">Cambarus</emphasis>
</taxonomicName>
based on (1) the presence of a subapical notch in the form I gonopod and (2) the lack of a well-developed mesial second tubercle row on the palm (
<bibRefCitation id="EFEF4B098F2C356AFEDEFE48FE6E5FC4" author="Hobbs" box="[346,487,457,482]" pageId="13" pageNumber="445" refString="Hobbs, H. H. Jr. (1969) On the distribution and phylogeny of the crayfish genus Cambarus. In: Holt, P. C. &amp; Hoffman, R. L, (Eds.), The Distributional History of the Biota of the Southern Appalachians. Part I: Invertebrates. Research Division Monograph 1. Virginia Polytechnic Institution, Blacksburg, Virginia. pp. 93 - 178." type="journal article" year="1969">Hobbs 1969</bibRefCitation>
). Among described members of the subgenus, 
<taxonomicName id="4C7E4D7B8F2C356AFC6AFE48FB3F5FC7" box="[1006,1206,457,481]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="445" phylum="Arthropoda" rank="species" species="appalachiensis">
<emphasis id="B90AEAEA8F2C356AFC6AFE48FB3F5FC7" box="[1006,1206,457,481]" italics="true" pageId="13" pageNumber="445">C. appalachiensis</emphasis>
</taxonomicName>
is most similar to 
<taxonomicName id="4C7E4D7B8F2C356AFA06FE48FE8D5C23" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="445" phylum="Arthropoda" rank="species" species="sciotensis">
<emphasis id="B90AEAEA8F2C356AFA06FE48FE8D5C23" italics="true" pageId="13" pageNumber="445">C. sciotensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7E4D7B8F2C356AFE96FE6FFE065C23" box="[274,399,493,517]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="445" phylum="Arthropoda" rank="species" species="hatfieldi">
<emphasis id="B90AEAEA8F2C356AFE96FE6FFE065C23" box="[274,399,493,517]" italics="true" pageId="13" pageNumber="445">C. hatfieldi</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C7E4D7B8F2C356AFE49FE6FFDD05C23" box="[461,601,493,517]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="445" phylum="Arthropoda" rank="species" species="angularis">
<emphasis id="B90AEAEA8F2C356AFE49FE6FFDD05C23" box="[461,601,493,517]" italics="true" pageId="13" pageNumber="445">C. angularis</emphasis>
</taxonomicName>
in overall body size and shape and thickening of the rostral margins.
</paragraph>
<paragraph id="8BC136F88F2C356AFF43FD90FDB65D00" blockId="13.[151,1436,151,2031]" pageId="13" pageNumber="445">
<taxonomicName id="4C7E4D7B8F2C356AFF43FD90FE655C0F" box="[199,492,529,553]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="445" phylum="Arthropoda" rank="species" species="appalachiensis">
<emphasis id="B90AEAEA8F2C356AFF43FD90FE655C0F" box="[199,492,529,553]" italics="true" pageId="13" pageNumber="445">Cambarus appalachiensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C7E4D7B8F2C356AFDA0FD90FD3A5C0F" box="[548,691,529,553]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="445" phylum="Arthropoda" rank="species" species="sciotensis">
<emphasis id="B90AEAEA8F2C356AFDA0FD90FD3A5C0F" box="[548,691,529,553]" italics="true" pageId="13" pageNumber="445">C. sciotensis</emphasis>
</taxonomicName>
were distinguished by morphometric ratios of palm length/width, and palm depth/length, as well as chelae architecture. Mean ± SE of palm length/width ratios of form I males, form II males, and females were 70.7% ± 0.49, 70.0% ± 0.40, and 70.6% ± 0.54 for 
<taxonomicName id="4C7E4D7B8F2C356AFB85FDDBFB435C57" box="[1025,1226,601,625]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="445" phylum="Arthropoda" rank="species" species="appalachiensis">
<emphasis id="B90AEAEA8F2C356AFB85FDDBFB435C57" box="[1025,1226,601,625]" italics="true" pageId="13" pageNumber="445">C. appalachiensis</emphasis>
</taxonomicName>
and 62.6% ± 0.65, 64.2% ± 0.90, and 63.8% ± 0.80 for 
<taxonomicName id="4C7E4D7B8F2C356AFDC6FDFFFD5B5CB3" box="[578,722,638,661]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="445" phylum="Arthropoda" rank="species" species="sciotensis">
<emphasis id="B90AEAEA8F2C356AFDC6FDFFFD5B5CB3" box="[578,722,638,661]" italics="true" pageId="13" pageNumber="445">C. sciotensis</emphasis>
</taxonomicName>
, respectively. Also, mean ± SE of palm depth/length ratios of form I males, form II males, and females were 83.7% ± 0.82, 85.6% ± 0.52, and 85.1% ± 0.61 for 
<taxonomicName id="4C7E4D7B8F2C356AFA06FD23FEB65CFB" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="445" phylum="Arthropoda" rank="species" species="appalachiensis">
<emphasis id="B90AEAEA8F2C356AFA06FD23FEB65CFB" italics="true" pageId="13" pageNumber="445">C. appalachiensis</emphasis>
</taxonomicName>
and 93.1% ± 1.1, 90.8% ± 1.2, and 92.7% ± 1.1 for 
<taxonomicName id="4C7E4D7B8F2C356AFC23FD47FBBE5CFB" box="[935,1079,710,733]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="445" phylum="Arthropoda" rank="species" species="sciotensis">
<emphasis id="B90AEAEA8F2C356AFC23FD47FBBE5CFB" box="[935,1079,710,733]" italics="true" pageId="13" pageNumber="445">C. sciotensis</emphasis>
</taxonomicName>
, respectively. The result of the previously mentioned ratios is 
<taxonomicName id="4C7E4D7B8F2C356AFE72FD6BFD485D27" box="[502,705,745,769]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="445" phylum="Arthropoda" rank="species" species="appalachiensis">
<emphasis id="B90AEAEA8F2C356AFE72FD6BFD485D27" box="[502,705,745,769]" italics="true" pageId="13" pageNumber="445">C. appalachiensis</emphasis>
</taxonomicName>
has a much more elongated chelae in profile than 
<taxonomicName id="4C7E4D7B8F2C356AFA82FD6BFA1C5D27" box="[1286,1429,746,769]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="445" phylum="Arthropoda" rank="species" species="sciotensis">
<emphasis id="B90AEAEA8F2C356AFA82FD6BFA1C5D27" box="[1286,1429,746,769]" italics="true" pageId="13" pageNumber="445">C. sciotensis</emphasis>
</taxonomicName>
, which possess a subrectangular chela.
</paragraph>
<paragraph id="8BC136F88F2C356AFF43FCB3FDD25DD8" blockId="13.[151,1436,151,2031]" pageId="13" pageNumber="445">
In addition to the previous ratios, 
<taxonomicName id="4C7E4D7B8F2C356AFDC0FCB3FC845D6F" box="[580,781,817,841]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="445" phylum="Arthropoda" rank="species" species="appalachiensis">
<emphasis id="B90AEAEA8F2C356AFDC0FCB3FC845D6F" box="[580,781,817,841]" italics="true" pageId="13" pageNumber="445">C. appalachiensis</emphasis>
</taxonomicName>
is readily differentiated from 
<taxonomicName id="4C7E4D7B8F2C356AFBE4FCB3FB675D6F" box="[1120,1262,818,841]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="445" phylum="Arthropoda" rank="species" species="sciotensis">
<emphasis id="B90AEAEA8F2C356AFBE4FCB3FB675D6F" box="[1120,1262,818,841]" italics="true" pageId="13" pageNumber="445">C. sciotensis</emphasis>
</taxonomicName>
by its lack of a second mesial row of tubercles, lack of a pronounced lateral impression, and lack of pronounced tubercles on the longitudinal ridge of the dactyl (
<figureCitation id="13452A7D8F2C356AFD86FCF8FDCF5DB4" box="[514,582,889,914]" captionStart="FIGURE 7" captionStartId="12.[151,250,1505,1527]" captionTargetBox="[151,1436,195,1480]" captionTargetId="figure@12.[151,1436,195,1480]" captionTargetPageId="12" captionText="FIGURE 7. Differences in chelae structure between Cambarus appalachiensis (Top) and Cambarus sciotensis (Bottom)." httpUri="https://zenodo.org/record/400121/files/figure.png" pageId="13" pageNumber="445">Fig. 7</figureCitation>
). 
<taxonomicName id="4C7E4D7B8F2C356AFDD8FCF8FCCC5DB7" box="[604,837,889,913]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="445" phylum="Arthropoda" rank="species" species="sciotensis">
<emphasis id="B90AEAEA8F2C356AFDD8FCF8FCCC5DB7" box="[604,837,889,913]" italics="true" pageId="13" pageNumber="445">Cambarus sciotensis</emphasis>
</taxonomicName>
has two mesial rows of tubercles, moderate to strong lateral impression, and pronounced tubercles on the dorsal longitudinal ridge of the dactyl (
<figureCitation id="13452A7D8F2C356AFB47FC1CFA835D90" box="[1219,1290,925,950]" captionStart="FIGURE 7" captionStartId="12.[151,250,1505,1527]" captionTargetBox="[151,1436,195,1480]" captionTargetId="figure@12.[151,1436,195,1480]" captionTargetPageId="12" captionText="FIGURE 7. Differences in chelae structure between Cambarus appalachiensis (Top) and Cambarus sciotensis (Bottom)." httpUri="https://zenodo.org/record/400121/files/figure.png" pageId="13" pageNumber="445">Fig. 7</figureCitation>
). Finally, 
<taxonomicName id="4C7E4D7B8F2C356AFA06FC1FFE8D5DFF" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="445" phylum="Arthropoda" rank="species" species="sciotensis">
<emphasis id="B90AEAEA8F2C356AFA06FC1FFE8D5DFF" italics="true" pageId="13" pageNumber="445">C. sciotensis</emphasis>
</taxonomicName>
postorbital ridge terminate in a straight tubercle or spine; 
<taxonomicName id="4C7E4D7B8F2C356AFC14FC43FB065DFF" authority="Post" authorityName="Post" box="[912,1167,961,985]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="445" phylum="Arthropoda" rank="species" species="appalachiensis">
<emphasis id="B90AEAEA8F2C356AFC14FC43FBD15DFF" box="[912,1112,961,985]" italics="true" pageId="13" pageNumber="445">C. appalachiensis</emphasis>
Post
</taxonomicName>
orbital ridges terminate in dorsally deflected spines or tubercles.
</paragraph>
</subSubSection>
<subSubSection id="C36465738F2C356AFF43FB88FEA15819" pageId="13" pageNumber="445" type="reference_group">
<paragraph id="8BC136F88F2C356AFF43FB88FD8A5B39" blockId="13.[151,1436,151,2031]" pageId="13" pageNumber="445">
<taxonomicName id="4C7E4D7B8F2C356AFF43FB88FE635A07" ID-CoL="7RTVV" box="[199,490,1033,1057]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="445" phylum="Arthropoda" rank="species" species="appalachiensis">
<emphasis id="B90AEAEA8F2C356AFF43FB88FE635A07" box="[199,490,1033,1057]" italics="true" pageId="13" pageNumber="445">Cambarus appalachiensis</emphasis>
</taxonomicName>
can be differentiated from both 
<taxonomicName id="4C7E4D7B8F2C356AFCD4FB88FAE45A04" ID-CoL="7RTVT" authority="Hobbs and Bouchard, 1994" authorityName="Hobbs and Bouchard" authorityYear="1994" box="[848,1389,1033,1058]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="445" phylum="Arthropoda" rank="species" species="angularis">
<emphasis id="B90AEAEA8F2C356AFCD4FB88FBBF5A07" box="[848,1078,1033,1057]" italics="true" pageId="13" pageNumber="445">Cambarus angularis</emphasis>
<bibRefCitation id="EFEF4B098F2C356AFBB8FB88FAE45A04" author="Hobbs" box="[1084,1389,1033,1058]" pageId="13" pageNumber="445" refString="Hobbs, H. H. Jr. &amp; Bouchard, R. W. (1994) Cambarus (Cambarus) angularis, a new crayfish (Decapoda: Cambaridae) from the Tennessee River basin of northeastern Tennessee and Virginia. Jeffersoniana, 5, 1 - 13." type="journal article" year="1994">Hobbs and Bouchard, 1994</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="4C7E4D7B8F2C356AFF13FBACFDF35A60" ID-CoL="7RTZ5" authority="Loughman et al. 2014" authorityName="Loughman et al." authorityYear="2014" box="[151,634,1069,1094]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="445" phylum="Arthropoda" rank="species" species="hatfieldi">
<emphasis id="B90AEAEA8F2C356AFF13FBACFEFB5A63" box="[151,370,1069,1093]" italics="true" pageId="13" pageNumber="445">Cambarus hatfieldi</emphasis>
<bibRefCitation id="EFEF4B098F2C356AFEF8FBACFDF35A60" author="Loughman" box="[380,634,1069,1094]" pageId="13" pageNumber="445" refString="Loughman, Z. J., Fagundo, R. A., Lau, E., Welsh, S. A. &amp; Thoma, R. F. (2014) Cambarus (C.) hatfieldi, a new species of crayfish (Decapoda: Cambaridae) from the Tug Fork River basin of Kentucky, Virginia, and West Virginia, USA. Zootaxa, 3750 (3), 223 - 236." type="journal article" year="2014">
Loughman 
<emphasis id="B90AEAEA8F2C356AFE7AFBAFFDB05A63" box="[510,569,1069,1093]" italics="true" pageId="13" pageNumber="445">et al.</emphasis>
2014
</bibRefCitation>
</taxonomicName>
by its more elongate chelae, compared to both the latter species subrectangular chelae. In addition to chelae shape, 
<taxonomicName id="4C7E4D7B8F2C356AFD54FBD3FC175A4F" ID-CoL="7RTVV" box="[720,926,1105,1129]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="445" phylum="Arthropoda" rank="species" species="appalachiensis">
<emphasis id="B90AEAEA8F2C356AFD54FBD3FC175A4F" box="[720,926,1105,1129]" italics="true" pageId="13" pageNumber="445">C. appalachiensis</emphasis>
</taxonomicName>
has less thickened rostral margins than 
<taxonomicName id="4C7E4D7B8F2C356AFA06FBD3FE8B5AAB" ID-CoL="7RTVT" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="445" phylum="Arthropoda" rank="species" species="angularis">
<emphasis id="B90AEAEA8F2C356AFA06FBD3FE8B5AAB" italics="true" pageId="13" pageNumber="445">C. angularis</emphasis>
</taxonomicName>
, and a more elongate rostrum than 
<taxonomicName id="4C7E4D7B8F2C356AFD21FBF7FCAF5AAB" ID-CoL="7RTZ5" box="[677,806,1141,1165]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="445" phylum="Arthropoda" rank="species" species="hatfieldi">
<emphasis id="B90AEAEA8F2C356AFD21FBF7FCAF5AAB" box="[677,806,1141,1165]" italics="true" pageId="13" pageNumber="445">C. hatfieldi</emphasis>
</taxonomicName>
. 
<taxonomicName id="4C7E4D7B8F2C356AFCB3FBF4FBD75AAB" ID-CoL="7RTVV" box="[823,1118,1141,1165]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="445" phylum="Arthropoda" rank="species" species="appalachiensis">
<emphasis id="B90AEAEA8F2C356AFCB3FBF4FBD75AAB" box="[823,1118,1141,1165]" italics="true" pageId="13" pageNumber="445">Cambarus appalachiensis</emphasis>
</taxonomicName>
central projection does not extend past the gonopod shaft; both 
<taxonomicName id="4C7E4D7B8F2C356AFDC1FB1BFD5C5A97" ID-CoL="7RTVT" box="[581,725,1177,1201]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="445" phylum="Arthropoda" rank="species" species="angularis">
<emphasis id="B90AEAEA8F2C356AFDC1FB1BFD5C5A97" box="[581,725,1177,1201]" italics="true" pageId="13" pageNumber="445">C. angularis</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C7E4D7B8F2C356AFC97FB1BFC1C5A97" ID-CoL="7RTZ5" box="[787,917,1177,1201]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="445" phylum="Arthropoda" rank="species" species="hatfieldi">
<emphasis id="B90AEAEA8F2C356AFC97FB1BFC1C5A97" box="[787,917,1177,1201]" italics="true" pageId="13" pageNumber="445">C. hatfieldi</emphasis>
</taxonomicName>
central projections typically extend past the gonopod shaft, and normally past the distal margin of the mesial process. 
<taxonomicName id="4C7E4D7B8F2C356AFC7EFB3CFAA85AF3" ID-CoL="7RTVV" box="[1018,1313,1213,1237]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="445" phylum="Arthropoda" rank="species" species="appalachiensis">
<emphasis id="B90AEAEA8F2C356AFC7EFB3CFAA85AF3" box="[1018,1313,1213,1237]" italics="true" pageId="13" pageNumber="445">Cambarus appalachiensis</emphasis>
</taxonomicName>
is also on average longer in total body length ( = 
<quantity id="4C869B1D8F2C356AFDDFFB63FD4B5ADC" box="[603,706,1250,1274]" metricMagnitude="-2" metricUnit="m" metricValue="7.44" pageId="13" pageNumber="445" unit="mm" value="74.4">74.4 mm</quantity>
; n = 98; SE ± 9.8%) than 
<taxonomicName id="4C7E4D7B8F2C356AFC65FB63FBD65ADC" ID-CoL="7RTZ5" box="[993,1119,1250,1274]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="445" phylum="Arthropoda" rank="species" species="hatfieldi">
<emphasis id="B90AEAEA8F2C356AFC65FB63FBD65ADC" box="[993,1119,1250,1274]" italics="true" pageId="13" pageNumber="445">C. hatfieldi</emphasis>
</taxonomicName>
( = 
<quantity id="4C869B1D8F2C356AFB1AFB63FA8D5ADC" box="[1182,1284,1250,1275]" metricMagnitude="-2" metricUnit="m" metricValue="6.76" pageId="13" pageNumber="445" unit="mm" value="67.6">67.6 mm</quantity>
; n = 52; SE ± 
<quantity id="4C869B1D8F2C356AFF13FA86FF795B38" box="[151,240,1287,1311]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="13" pageNumber="445" unit="mm" value="7.3">7.3 mm</quantity>
; 
<bibRefCitation id="EFEF4B098F2C356AFF7FFA87FE715B39" author="Loughman" box="[251,504,1286,1311]" pageId="13" pageNumber="445" refString="Loughman, Z. J., Fagundo, R. A., Lau, E., Welsh, S. A. &amp; Thoma, R. F. (2014) Cambarus (C.) hatfieldi, a new species of crayfish (Decapoda: Cambaridae) from the Tug Fork River basin of Kentucky, Virginia, and West Virginia, USA. Zootaxa, 3750 (3), 223 - 236." type="journal article" year="2014">
Loughman 
<emphasis id="B90AEAEA8F2C356AFEF8FA86FE3D5B38" box="[380,436,1286,1310]" italics="true" pageId="13" pageNumber="445">et al.</emphasis>
2014
</bibRefCitation>
).
</paragraph>
<paragraph id="8BC136F88F2C356AFF43FAAAFEA15819" blockId="13.[151,1436,151,2031]" pageId="13" pageNumber="445">
Four additional 
<taxonomicName id="4C7E4D7B8F2C356AFEFDFAAAFE675B65" ID-CoL="7NRQP" box="[377,494,1323,1347]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="445" phylum="Arthropoda" rank="genus">
<emphasis id="B90AEAEA8F2C356AFEFDFAAAFE675B65" box="[377,494,1323,1347]" italics="true" pageId="13" pageNumber="445">Cambarus</emphasis>
</taxonomicName>
species occur in the greater New River system that can easily be differentiated from 
<taxonomicName id="4C7E4D7B8F2C356AFF13FACEFED65B40" ID-CoL="7RTVV" box="[151,351,1358,1382]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="445" phylum="Arthropoda" rank="species" species="appalachiensis">
<emphasis id="B90AEAEA8F2C356AFF13FACEFED65B40" box="[151,351,1358,1382]" italics="true" pageId="13" pageNumber="445">C. appalachiensis</emphasis>
</taxonomicName>
. Both 
<taxonomicName id="4C7E4D7B8F2C356AFE2DFACFFCE15B41" ID-CoL="7RV57" authority="Loughman et al. 2011" authorityName="Loughman et al." authorityYear="2011" box="[425,872,1358,1383]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="445" phylum="Arthropoda" rank="species" species="smilax">
<emphasis id="B90AEAEA8F2C356AFE2DFACFFDE45B40" box="[425,621,1358,1382]" italics="true" pageId="13" pageNumber="445">Cambarus smilax</emphasis>
<bibRefCitation id="EFEF4B098F2C356AFDF0FACFFCE15B41" author="Loughman" box="[628,872,1358,1383]" pageId="13" pageNumber="445" refString="Loughman, Z. J., Simon, T. P. &amp; Welsh, S. A. (2011) Cambarus (Puncticambarus) smilax, a new species of crayfish (Crustacea: Decapoda: Cambaridae) from the Greenbrier River basin of West Virginia. Proceedings of the Biological Society of Washington, 124, 99 - 111." type="journal article" year="2011">
Loughman 
<emphasis id="B90AEAEA8F2C356AFD70FACEFCA25B40" box="[756,811,1358,1382]" italics="true" pageId="13" pageNumber="445">et al.</emphasis>
2011
</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="4C7E4D7B8F2C356AFC19FACFFAB95B41" ID-CoL="7RV4R" authority="Girard, 1852" authorityName="Girard" authorityYear="1852" box="[925,1328,1358,1383]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="445" phylum="Arthropoda" rank="species" species="robustus">
<emphasis id="B90AEAEA8F2C356AFC19FACFFB9B5B40" box="[925,1042,1358,1382]" italics="true" pageId="13" pageNumber="445">Cambarus</emphasis>
cf. 
<emphasis id="B90AEAEA8F2C356AFBB8FACFFB125B40" box="[1084,1179,1358,1382]" italics="true" pageId="13" pageNumber="445">robustus</emphasis>
<bibRefCitation id="EFEF4B098F2C356AFB26FACFFAB95B41" author="Girard" box="[1186,1328,1358,1383]" pageId="13" pageNumber="445" refString="Girard, C. (1852) A revision of the North American Astaci, with observations on their habits and geographic distribution. Proceedings of the Academy of Natural Sciences of Philadelphia, 6, 87 - 91." type="journal article" year="1852">Girard, 1852</bibRefCitation>
</taxonomicName>
have two mesial rows of tubercles as well as lanceolate rostra, compared to 
<emphasis id="B90AEAEA8F2C356AFCF2FAF2FBCE5BAD" box="[886,1095,1395,1419]" italics="true" pageId="13" pageNumber="445">C. appalachiensis’</emphasis>
single row of mesial tubercles and broad rostra. 
<taxonomicName id="4C7E4D7B8F2C356AFED8FA17FDA95B88" ID-CoL="7RV57" box="[348,544,1430,1454]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="445" phylum="Arthropoda" rank="species" species="smilax">
<emphasis id="B90AEAEA8F2C356AFED8FA17FDA95B88" box="[348,544,1430,1454]" italics="true" pageId="13" pageNumber="445">Cambarus smilax</emphasis>
</taxonomicName>
does not appear to be sympatric with 
<taxonomicName id="4C7E4D7B8F2C356AFC48FA16FB1C5B88" ID-CoL="7RTVV" box="[972,1173,1430,1454]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="445" phylum="Arthropoda" rank="species" species="appalachiensis">
<emphasis id="B90AEAEA8F2C356AFC48FA16FB1C5B88" box="[972,1173,1430,1454]" italics="true" pageId="13" pageNumber="445">C. appalachiensis</emphasis>
</taxonomicName>
, and is endemic to mid and headwater reaches of the Greenbrier River (
<bibRefCitation id="EFEF4B098F2C356AFD29FA3BFC2D5BF5" author="Loughman" box="[685,932,1466,1491]" pageId="13" pageNumber="445" refString="Loughman, Z. J., Simon, T. P. &amp; Welsh, S. A. (2011) Cambarus (Puncticambarus) smilax, a new species of crayfish (Crustacea: Decapoda: Cambaridae) from the Greenbrier River basin of West Virginia. Proceedings of the Biological Society of Washington, 124, 99 - 111." type="journal article" year="2011">
Loughman 
<emphasis id="B90AEAEA8F2C356AFCABFA3AFCEE5BF5" box="[815,871,1467,1491]" italics="true" pageId="13" pageNumber="445">et al.</emphasis>
2011
</bibRefCitation>
<emphasis id="B90AEAEA8F2C356AFC20FA3AFC3C5BF4" box="[932,949,1467,1490]" italics="true" pageId="13" pageNumber="445">).</emphasis>
<taxonomicName id="4C7E4D7B8F2C356AFC3FFA3AFB305BF5" ID-CoL="7RV4R" box="[955,1209,1466,1491]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="445" phylum="Arthropoda" rank="species" species="robustus">
<emphasis id="B90AEAEA8F2C356AFC3FFA3AFBB95BF5" box="[955,1072,1467,1491]" italics="true" pageId="13" pageNumber="445">Cambarus</emphasis>
cf. 
<emphasis id="B90AEAEA8F2C356AFBDEFA3AFB305BF5" box="[1114,1209,1467,1491]" italics="true" pageId="13" pageNumber="445">robustus</emphasis>
</taxonomicName>
, like 
<taxonomicName id="4C7E4D7B8F2C356AFB7CFA3AFAE85BF5" ID-CoL="7RV57" box="[1272,1377,1467,1491]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="445" phylum="Arthropoda" rank="species" species="smilax">
<emphasis id="B90AEAEA8F2C356AFB7CFA3AFAE85BF5" box="[1272,1377,1467,1491]" italics="true" pageId="13" pageNumber="445">C. smilax</emphasis>
</taxonomicName>
, also is a headwater species and endemic to the upper New River system of 
<collectingRegion id="49BAF81A8F2C356AFC3AFA5FFB905BD1" box="[958,1049,1502,1527]" country="United States of America" name="Virginia" pageId="13" pageNumber="445">Virginia</collectingRegion>
and 
<collectingRegion id="49BAF81A8F2C356AFBD7FA5FFB765BD1" box="[1107,1279,1502,1527]" country="United States of America" name="North Carolina" pageId="13" pageNumber="445">North Carolina</collectingRegion>
, and does not occur in larger ordered streams typically utilized by 
<taxonomicName id="4C7E4D7B8F2C356AFD6FF982FC3C583D" ID-CoL="7RTVV" box="[747,949,1539,1563]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="445" phylum="Arthropoda" rank="species" species="appalachiensis">
<emphasis id="B90AEAEA8F2C356AFD6FF982FC3C583D" box="[747,949,1539,1563]" italics="true" pageId="13" pageNumber="445">C. appalachiensis</emphasis>
</taxonomicName>
. Only two sites have been found to harbor both species.
</paragraph>
</subSubSection>
<subSubSection id="C36465738F2C3569FF43F9CAFCC15F3B" lastPageId="14" lastPageNumber="446" pageId="13" pageNumber="445" type="description">
<paragraph id="8BC136F88F2C356AFF43F9CAFCDB5981" blockId="13.[151,1436,151,2031]" pageId="13" pageNumber="445">
<taxonomicName id="4C7E4D7B8F2C356AFF43F9CAFE7C5845" box="[199,501,1611,1635]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="445" phylum="Arthropoda" rank="species" species="chasmodactylus">
<emphasis id="B90AEAEA8F2C356AFF43F9CAFE7C5845" box="[199,501,1611,1635]" italics="true" pageId="13" pageNumber="445">Cambarus chasmodactylus</emphasis>
</taxonomicName>
is the third 
<taxonomicName id="4C7E4D7B8F2C356AFDFCF9CAFD645845" box="[632,749,1611,1635]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="445" phylum="Arthropoda" rank="genus">
<emphasis id="B90AEAEA8F2C356AFDFCF9CAFD645845" box="[632,749,1611,1635]" italics="true" pageId="13" pageNumber="445">Cambarus</emphasis>
</taxonomicName>
species in the New River system, and does on occasion occur in sympatry with 
<taxonomicName id="4C7E4D7B8F2C356AFEDFF9EEFDAD58A0" box="[347,548,1646,1670]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="445" phylum="Arthropoda" rank="species" species="appalachiensis">
<emphasis id="B90AEAEA8F2C356AFEDFF9EEFDAD58A0" box="[347,548,1646,1670]" italics="true" pageId="13" pageNumber="445">C. appalachiensis</emphasis>
</taxonomicName>
. 
<taxonomicName id="4C7E4D7B8F2C356AFDB6F9EFFCDC58A0" box="[562,853,1646,1670]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="445" phylum="Arthropoda" rank="species" species="appalachiensis">
<emphasis id="B90AEAEA8F2C356AFDB6F9EFFCDC58A0" box="[562,853,1646,1670]" italics="true" pageId="13" pageNumber="445">Cambarus appalachiensis</emphasis>
</taxonomicName>
is readily differentiated from 
<taxonomicName id="4C7E4D7B8F2C356AFB21F9EEFAF058A0" box="[1189,1401,1646,1670]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="445" phylum="Arthropoda" rank="species" species="chasmodactylus">
<emphasis id="B90AEAEA8F2C356AFB21F9EEFAF058A0" box="[1189,1401,1646,1670]" italics="true" pageId="13" pageNumber="445">C. chasmodactylus</emphasis>
</taxonomicName>
by its elongate chelae, broad rostrum, and brown coloration compared to 
<emphasis id="B90AEAEA8F2C356AFC2EF912FB0E588D" box="[938,1159,1683,1707]" italics="true" pageId="13" pageNumber="445">C. chasmodactylus’</emphasis>
broad chelae, lanceolate rostrum, and blue, green, turquoise coloration. 
<taxonomicName id="4C7E4D7B8F2C356AFD2FF937FC5358E8" box="[683,986,1718,1742]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="445" phylum="Arthropoda" rank="species" species="chasmodactylus">
<emphasis id="B90AEAEA8F2C356AFD2FF937FC5358E8" box="[683,986,1718,1742]" italics="true" pageId="13" pageNumber="445">Cambarus chasmodactylus</emphasis>
</taxonomicName>
also has an extremely large gape in the chelae, which is substantially larger than the chelae gape observed in 
<taxonomicName id="4C7E4D7B8F2C356AFC3CF95AFB0A58D5" box="[952,1155,1755,1779]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="445" phylum="Arthropoda" rank="species" species="appalachiensis">
<emphasis id="B90AEAEA8F2C356AFC3CF95AFB0A58D5" box="[952,1155,1755,1779]" italics="true" pageId="13" pageNumber="445">C. appalachiensis</emphasis>
</taxonomicName>
. 
<taxonomicName id="4C7E4D7B8F2C356AFB17F95AFA1558D4" box="[1171,1436,1755,1779]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="445" phylum="Arthropoda" rank="species" species="carinirostris">
<emphasis id="B90AEAEA8F2C356AFB17F95AFA1558D4" box="[1171,1436,1755,1779]" italics="true" pageId="13" pageNumber="445">Cambarus carinirostris</emphasis>
</taxonomicName>
occurs in the northern New River system and functions as the basins secondary burrowing species. As such, it occurs primarily in headwater streams, and normally does not occur in sympatry with 
<taxonomicName id="4C7E4D7B8F2C356AFBD6F8A2FA93591D" box="[1106,1306,1827,1851]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="445" phylum="Arthropoda" rank="species" species="appalachiensis">
<emphasis id="B90AEAEA8F2C356AFBD6F8A2FA93591D" box="[1106,1306,1827,1851]" italics="true" pageId="13" pageNumber="445">C. appalachiensis</emphasis>
</taxonomicName>
. 
<taxonomicName id="4C7E4D7B8F2C356AFAA3F8A2FEAB5978" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="445" phylum="Arthropoda" rank="species" species="carinirostris">
<emphasis id="B90AEAEA8F2C356AFAA3F8A2FEAB5978" italics="true" pageId="13" pageNumber="445">Cambarus carinirostris</emphasis>
</taxonomicName>
has a broad rostrum like 
<taxonomicName id="4C7E4D7B8F2C356AFDC8F8C6FC9E5978" box="[588,791,1862,1886]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="445" phylum="Arthropoda" rank="species" species="appalachiensis">
<emphasis id="B90AEAEA8F2C356AFDC8F8C6FC9E5978" box="[588,791,1862,1886]" italics="true" pageId="13" pageNumber="445">C. appalachiensis</emphasis>
</taxonomicName>
, but can be differentiated from 
<taxonomicName id="4C7E4D7B8F2C356AFB0CF8C6FADA5978" box="[1160,1363,1862,1886]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="445" phylum="Arthropoda" rank="species" species="appalachiensis">
<emphasis id="B90AEAEA8F2C356AFB0CF8C6FADA5978" box="[1160,1363,1862,1886]" italics="true" pageId="13" pageNumber="445">C. appalachiensis</emphasis>
</taxonomicName>
by its subrectangular chelae which maintain a single row of appressed tubercles, compared to 
<taxonomicName id="4C7E4D7B8F2C356AFBE9F8EAFABF59A5" box="[1133,1334,1899,1923]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="445" phylum="Arthropoda" rank="species" species="appalachiensis">
<emphasis id="B90AEAEA8F2C356AFBE9F8EAFABF59A5" box="[1133,1334,1899,1923]" italics="true" pageId="13" pageNumber="445">C. appalachiensis</emphasis>
</taxonomicName>
elongate chelae which possess a defined single mesial row of tubercles.
</paragraph>
<paragraph id="8BC136F88F2C3569FF43F833FCC15F3B" blockId="13.[151,1436,151,2031]" lastBlockId="14.[151,1436,151,536]" lastPageId="14" lastPageNumber="446" pageId="13" pageNumber="445">
The upper Guyandotte River system is geographically proximate to the upper and middle New River system and is occupied by 
<taxonomicName id="4C7E4D7B8F2C356AFEF6F856FD9959C8" box="[370,528,2006,2030]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="445" phylum="Arthropoda" rank="species" species="theepiensis">
<emphasis id="B90AEAEA8F2C356AFEF6F856FD9959C8" box="[370,528,2006,2030]" italics="true" pageId="13" pageNumber="445">C. theepiensis</emphasis>
</taxonomicName>
. 
<taxonomicName id="4C7E4D7B8F2C356AFD9BF857FCCA59C8" box="[543,835,2006,2030]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="445" phylum="Arthropoda" rank="species" species="appalachiensis">
<emphasis id="B90AEAEA8F2C356AFD9BF857FCCA59C8" box="[543,835,2006,2030]" italics="true" pageId="13" pageNumber="445">Cambarus appalachiensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C7E4D7B8F2C356AFCFFF856FB9059C8" box="[891,1049,2006,2030]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="445" phylum="Arthropoda" rank="species" species="theepiensis">
<emphasis id="B90AEAEA8F2C356AFCFFF856FB9059C8" box="[891,1049,2006,2030]" italics="true" pageId="13" pageNumber="445">C. theepiensis</emphasis>
</taxonomicName>
have similar broad rostra, but can be differentiated from each other by the number of tubercles rows on the mesial margin of the palm as well as subpalmar tubercles. 
<taxonomicName id="4C7E4D7B8F2F3569FE00FF3DFDF25EF2" box="[388,635,188,212]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="446" phylum="Arthropoda" rank="species" species="theepiensis">
<emphasis id="B90AEAEA8F2F3569FE00FF3DFDF25EF2" box="[388,635,188,212]" italics="true" pageId="14" pageNumber="446">Cambarus theepiensis</emphasis>
</taxonomicName>
has two rows of tubercles on the mesial margin of the chelae, as well as 2–4 subpalmar tubercles (
<bibRefCitation id="EFEF4B098F2F3569FE45FF5EFD435EDE" author="Loughman" box="[449,714,223,248]" pageId="14" pageNumber="446" refString="Loughman, Z. J., Foltz, D. A., Garrison, N. L. &amp; Welsh, S. A. (2013) Cambarus (P.) theepiensis, a new species of crayfish (Decapoda: Cambaridae) from the coalfields region of eastern Kentucky and southwestern West Virginia, USA. Zootaxa, 3641 (1), 63 - 73." type="journal article" year="2013">
Loughman 
<emphasis id="B90AEAEA8F2F3569FDC3FF60FD0A5EDE" box="[583,643,224,248]" italics="true" pageId="14" pageNumber="446">et al.</emphasis>
2013
</bibRefCitation>
). 
<taxonomicName id="4C7E4D7B8F2F3569FD64FF61FB8E5EDE" box="[736,1031,224,248]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="446" phylum="Arthropoda" rank="species" species="appalachiensis">
<emphasis id="B90AEAEA8F2F3569FD64FF61FB8E5EDE" box="[736,1031,224,248]" italics="true" pageId="14" pageNumber="446">Cambarus appalachiensis</emphasis>
</taxonomicName>
has a single row of tubercles, and normally lacks or possesses only a single subpalmar tubercle.
</paragraph>
</subSubSection>
<subSubSection id="C36465738F2F3569FF43FEA6FDFB5C3E" pageId="14" pageNumber="446" type="etymology">
<paragraph id="8BC136F88F2F3569FF43FEA6FD8D5FD2" blockId="14.[151,1436,151,536]" pageId="14" pageNumber="446">
<emphasis id="B90AEAEA8F2F3569FF43FEA6FEC75F66" bold="true" box="[199,334,295,320]" pageId="14" pageNumber="446">Etymology.</emphasis>
<taxonomicName id="4C7E4D7B8F2F3569FEDCFEA9FDF65F66" box="[344,639,296,320]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="446" phylum="Arthropoda" rank="species" species="appalachiensis">
<emphasis id="B90AEAEA8F2F3569FEDCFEA9FDF65F66" box="[344,639,296,320]" italics="true" pageId="14" pageNumber="446">Cambarus appalachiensis</emphasis>
</taxonomicName>
is named after the Appalachian Mountains where 
<taxonomicName id="4C7E4D7B8F2F3569FB54FEA9FA155F66" box="[1232,1436,296,320]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="446" phylum="Arthropoda" rank="species" species="appalachiensis">
<emphasis id="B90AEAEA8F2F3569FB54FEA9FA155F66" box="[1232,1436,296,320]" italics="true" pageId="14" pageNumber="446">C. appalachiensis</emphasis>
</taxonomicName>
entire range occurs. The common name Conhaway Crayfish is in reference to Conhaway River, which was the name of both New and Bluestone Rivers in the 1700’s, and was the name Christopher Gist referred to the New River by in his field journal. It is theorized that Conhaway was the Shawnee name for the New River. Gist’s expeditions throughout the New River basin in 
<date id="FFC010388F2F3569FD39FE36FCE05FF6" box="[701,873,439,464]" pageId="14" pageNumber="446" value="1750" valueMax="1751">1750 and 1751</date>
were among the first expeditions throughout the New River valley by Europeans.
</paragraph>
<paragraph id="8BC136F88F2F3569FF43FE7EFDFB5C3E" blockId="14.[151,1436,151,536]" box="[199,626,511,536]" pageId="14" pageNumber="446">
<emphasis id="B90AEAEA8F2F3569FF43FE7EFE0C5C3E" bold="true" box="[199,389,511,536]" pageId="14" pageNumber="446">Common name.</emphasis>
Conhaway Crayfish.
</paragraph>
</subSubSection>
</treatment>
</document>