167 lines
19 KiB
XML
167 lines
19 KiB
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<document id="2D3C647FB311265451D88FCE2C60BF5E" ID-DOI="10.5281/zenodo.5318944" ID-ISSN="0374-1036" ID-Zenodo-Dep="5318944" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630132583789" checkinUser="felipe" docAuthor="Švihla, Vladimír" docDate="2009" docId="03D387E4EE36FFB9FE554754FE0AFA0D" docLanguage="en" docName="ActaEntMusNatPra.2009.49.1.191-216.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 49 (1)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Malthodes lysosensis Švihla 2009, sp. nov." docType="treatment" docVersion="5" lastPageNumber="207" masterDocId="FFEAFF9CEE26FFA9FF8B4575FF8EFFF0" masterDocTitle="New species of the subfamily Malthininae (Coleoptera: Cantharidae) from the western Palaearctic Region" masterLastPageNumber="216" masterPageNumber="191" pageNumber="207" updateTime="1699162932936" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:mods id="4BFBCA0675265BFC4E5B6D8B62FF854D" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="16AC2BDFEB18914987D68B9AE9E18A0D">
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<mods:title id="D0C6D705900485D56A1EE7F85D85E980">New species of the subfamily Malthininae (Coleoptera: Cantharidae) from the western Palaearctic Region</mods:title>
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</mods:titleInfo>
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<mods:name id="57C593C7FCFA95D7B6E772C72E4757A8" type="personal">
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<mods:roleTerm id="2CA4CC996BD1CEEA018CEF128325C8D7">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="58D4A39E00DFBE47C08F93A57843EE68">Švihla, Vladimír</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="E14D80F4921B7ACFD48760C4C725698A">text</mods:typeOfResource>
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<mods:relatedItem id="21EBF24E46D4F7264F4ED9F8AD6497AE" type="host">
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<mods:titleInfo id="D3CA50895C2E9583938E2CA237425AC6">
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<mods:title id="3C1678418B0C5C5FE960ECC3EF0036A8">Acta Entomologica Musei Nationalis Pragae</mods:title>
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</mods:titleInfo>
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<mods:part id="4139E18D16A5998E93AD4522D3C30EF4">
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<mods:date id="CB6EB14AB26EC08EE0E82A52E5491C60">2009</mods:date>
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<mods:detail id="BAF97AB4F50466F9AAF9C0754CB9EA8B" type="pubDate">
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<mods:number id="0B70CC393D3DE06F9B788BC80E5471BA">2009-06-30</mods:number>
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</mods:detail>
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<mods:detail id="70DD4EB2C58F21715292344C3AE4AA21" type="volume">
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<mods:number id="D8A5327E216B3EF040259D21CB144898">49</mods:number>
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</mods:detail>
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<mods:detail id="BD7EE2348750168E9804D05EAA19AEE2" type="issue">
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<mods:number id="2ADA76C8199969E54B68C881F85C7BE2">1</mods:number>
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</mods:detail>
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<mods:extent id="A19A3571D75BFD365AF370BD3A0F9267" unit="page">
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="DF1DA35CFB00D3C91AC17F5C54A580D3">journal article</mods:classification>
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<mods:identifier id="87D95F3A06A33DBEA2ADC89068BF291D" type="DOI">10.5281/zenodo.5318944</mods:identifier>
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<mods:identifier id="F122F941BDF31C464B6C1D5354B6FA18" type="ISSN">0374-1036</mods:identifier>
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<mods:identifier id="77C2BAAEB632429CDCF5FB59B02897BF" type="Zenodo-Dep">5318944</mods:identifier>
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</mods:mods>
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<treatment id="03D387E4EE36FFB9FE554754FE0AFA0D" ID-DOI="http://doi.org/10.5281/zenodo.5337888" ID-Zenodo-Dep="5337888" LSID="urn:lsid:plazi:treatment:03D387E4EE36FFB9FE554754FE0AFA0D" httpUri="http://treatment.plazi.org/id/03D387E4EE36FFB9FE554754FE0AFA0D" lastPageNumber="207" pageId="16" pageNumber="207">
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<subSubSection id="C3606579EE36FFB9FE554754FCAEFDCA" box="[478,800,545,570]" pageId="16" pageNumber="207" type="nomenclature">
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<paragraph id="8BC536F2EE36FFB9FE554754FCAEFDCA" blockId="16.[478,800,545,570]" box="[478,800,545,570]" pageId="16" pageNumber="207">
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<heading id="D08D819EEE36FFB9FE554754FCAEFDCA" bold="true" box="[478,800,545,570]" centered="true" fontSize="8" level="2" pageId="16" pageNumber="207" reason="3">
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<emphasis id="B90EEAE0EE36FFB9FE554754FCAEFDCA" bold="true" box="[478,800,545,570]" pageId="16" pageNumber="207">
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<taxonomicName id="4C7A4D71EE36FFB9FE554754FD4AFDCA" authority="Švihla, 2009" authorityName="Švihla" authorityYear="2009" box="[478,708,545,570]" class="Insecta" family="Cantharidae" genus="Malthodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="207" phylum="Arthropoda" rank="species" species="lysosensis" status="sp. nov.">
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<emphasis id="B90EEAE0EE36FFB9FE554754FD4AFDCA" bold="true" box="[478,708,545,570]" italics="true" pageId="16" pageNumber="207">Malthodes lysosensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23D579BEE36FFB9FD404757FCAEFDCA" box="[715,800,546,570]" pageId="16" pageNumber="207" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3606579EE36FFB9FDBE4735FD46FDA3" box="[565,712,576,595]" pageId="16" pageNumber="207" type="description">
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<paragraph id="8BC536F2EE36FFB9FDBE4735FD46FDA3" blockId="16.[565,712,576,595]" box="[565,712,576,595]" pageId="16" pageNumber="207">
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(
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<figureCitation id="13412A77EE36FFB9FDB74735FD0DFDA3" box="[572,643,576,595]" captionStart="Figs" captionStartId="7.[113,152,1542,1561]" captionTargetBox="[160,1042,220,1511]" captionTargetId="figure-12@7.[160,1042,220,1512]" captionTargetPageId="7" captionText="Figs. 13–16. Aedeagus, dorsal view. 13 – Malthodes lysosensis sp. nov. (holotype); 14 – M. anadyomenae sp. nov. (paratype); 15 – M. cyprogenius sp. nov. (holotype); 16 – M. andreasi sp. nov. (paratype). Scale bars = 0.1 mm. Arrows refer to diagnostic characters mentioned in descriptions." figureDoi="http://doi.org/10.5281/zenodo.5363037" httpUri="https://zenodo.org/record/5363037/files/figure.png" pageId="16" pageNumber="207">Figs. 13</figureCitation>
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,
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<figureCitation id="13412A77EE36FFB9FD074735FD4DFDA3" box="[652,707,576,595]" captionStart="Figs" captionStartId="15.[113,152,1435,1454]" captionTargetBox="[141,1088,219,1372]" captionTargetId="figure-12@15.[113,1091,219,1396]" captionTargetPageId="15" captionText="Figs. 28–33. 28–30 – Malthodes fritzlari sp. nov. (holotype): 28 – last abdominal segments of male, lateral view; 29 – last tergite of male, dorsal view; 30 – apical part of last sternite of male, ventro-caudal view. 31–33 – M. lysosensis sp. nov. (holotype): 31 – last abdominal segments of male, lateral view; 32 – apical part of last sternite of male, ventro-caudal view; 33 – last tergite of male, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5363043" httpUri="https://zenodo.org/record/5363043/files/figure.png" pageId="16" pageNumber="207">31–33</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3606579EE36FFB9FF1C471FFC0BFD3D" pageId="16" pageNumber="207" type="materials_examined">
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<paragraph id="8BC536F2EE36FFB9FF1C471FFC90FD8D" blockId="16.[151,1127,617,717]" box="[151,798,617,637]" pageId="16" pageNumber="207">
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<emphasis id="B90EEAE0EE36FFB9FF1C471FFE9EFD8D" bold="true" box="[151,272,617,637]" pageId="16" pageNumber="207">
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<typeStatus id="54C18850EE36FFB9FF1C471FFF4AFD8D" box="[151,196,618,637]" pageId="16" pageNumber="207">Type</typeStatus>
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locality.
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</emphasis>
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Northwestern
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<collectingCountry id="F36D7662EE36FFB9FE05471FFE40FD8D" box="[398,462,618,637]" name="Cyprus" pageId="16" pageNumber="207">Cyprus</collectingCountry>
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, Lysos, southwest of Polis,
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<quantity id="4C829B17EE36FFB9FD31471FFD7FFD8D" box="[698,753,618,637]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="16" pageNumber="207" unit="m" value="600.0">600 m</quantity>
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a.s.l.
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</paragraph>
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<paragraph id="8BC536F2EE36FFB9FF1C47F0FC0BFD3D" blockId="16.[151,1127,617,717]" pageId="16" pageNumber="207">
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<emphasis id="B90EEAE0EE36FFB9FF1C47F0FE90FD68" bold="true" box="[151,286,644,664]" pageId="16" pageNumber="207">Type material.</emphasis>
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<materialsCitation id="3B123CAFEE36FFB9FEAF47F0FEBBFD42" collectingDate="2006-04-16" collectionCode="NMEG" collectorName="Polis SW Lysos & F. Fritzlar" country="Cyprus" elevation="600" location="ZYPERN" pageId="16" pageNumber="207" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54C18850EE36FFB9FEAF47F0FEF6FD66" box="[292,376,645,664]" pageId="16" pageNumber="207" type="holotype">HOLOTYPE</typeStatus>
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: J, ‘
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<collectingCountry id="F36D7662EE36FFB9FE2F47F0FE73FD68" box="[420,509,645,664]" name="Cyprus" pageId="16" pageNumber="207">ZYPERN</collectingCountry>
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, NW,
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<collectorName id="268F5324EE36FFB9FDBD47F1FD4CFD68" box="[566,706,644,664]" pageId="16" pageNumber="207">Polis SW Lysos</collectorName>
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,
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<quantity id="4C829B17EE36FFB9FD4747F1FC88FD68" box="[716,774,644,664]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="16" pageNumber="207" unit="m" value="600.0">
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<elevation id="0057D1C1EE36FFB9FD4747F1FC88FD68" box="[716,774,644,664]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="16" pageNumber="207" unit="m" value="600.0">600 m</elevation>
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</quantity>
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,
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<date id="FFC41032EE36FFB9FC9A47F1FCE3FD67" box="[785,877,644,663]" pageId="16" pageNumber="207" value="2006-04-16">
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<collectingDate id="EF80E9DAEE36FFB9FC9A47F1FCE3FD67" box="[785,877,644,663]" pageId="16" pageNumber="207" value="2006-04-16">16.iv.2006</collectingDate>
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</date>
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, leg.
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<collectorName id="268F5324EE36FFB9FC1647F0FC7DFD67" box="[925,1011,644,664]" pageId="16" pageNumber="207">F. Fritzlar</collectorName>
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[white label, printed]’ (
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<collectionCode id="ED6BAE37EE36FFB9FF6047EAFEA1FD42" box="[235,303,671,690]" country="Germany" httpUri="http://grbio.org/cool/3082-3vhh" name="Naturkundesmuseum" pageId="16" pageNumber="207">NMEG</collectionCode>
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)
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</materialsCitation>
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.
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<typeStatus id="54C18850EE36FFB9FEB647D5FE1AFD41" box="[317,404,672,691]" pageId="16" pageNumber="207" type="paratype">PARATYPES</typeStatus>
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: 5 JJ
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<specimenCount id="9D7CFD7BEE36FFB9FE5847EAFE7CFD42" box="[467,498,665,690]" pageId="16" pageNumber="207" type="female">1 ♀</specimenCount>
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, ‘Cyprus,
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<collectingRegion id="49BEF810EE36FFB9FDC747EAFD32FD42" box="[588,700,671,690]" country="Cyprus" name="Pafos" pageId="16" pageNumber="207">Paphos distr.</collectingRegion>
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, Polis
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<quantity id="4C829B17EE36FFB9FD7847EAFCA6FD42" box="[755,808,671,690]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="16" pageNumber="207" unit="km" value="10.0">10 km</quantity>
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W, Aphrodite Bay footpath,
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<geoCoordinate id="EE4E5035EE36FFB9FB9647EAFBE9FD42" box="[1053,1127,671,690]" degrees="35" direction="north" minutes="04" orientation="latitude" pageId="16" pageNumber="207" precision="925" value="35.066666">35°04´N</geoCoordinate>
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<geoCoordinate id="EE4E5035EE36FFB9FF1C47CFFF6EFD3D" box="[151,224,698,717]" degrees="32" direction="east" minutes="18" orientation="longitude" pageId="16" pageNumber="207" precision="925" value="32.3">32°18´E</geoCoordinate>
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,
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<quantity id="4C829B17EE36FFB9FF6247CFFEAFFD3D" box="[233,289,698,717]" metricMagnitude="2" metricUnit="m" metricValue="1.58" pageId="16" pageNumber="207" unit="m" value="158.0">158 m</quantity>
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,
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<date id="FFC41032EE36FFB9FEA147CFFE0BFD3D" box="[298,389,698,717]" pageId="16" pageNumber="207" value="1999-04-21">21.iv.1999</date>
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, R. Constantin [lgt.] [white label, printed]’ (
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<collectionCode id="ED6BAE37EE36FFB9FD7647CFFCBDFD3D" box="[765,819,698,717]" pageId="16" pageNumber="207">RCSF</collectionCode>
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,
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<collectionCode id="ED6BAE37EE36FFB9FCB047CFFCF5FD3D" box="[827,891,698,717]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="16" pageNumber="207">NMPC</collectionCode>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3606579EE36FFB9FF1C4790FE1DFB0D" pageId="16" pageNumber="207" type="description">
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<paragraph id="8BC536F2EE36FFB9FF1C4790FE67FC8D" blockId="16.[151,1127,741,1533]" pageId="16" pageNumber="207">
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<emphasis id="B90EEAE0EE36FFB9FF1C4790FEACFD0D" bold="true" box="[151,290,741,765]" pageId="16" pageNumber="207">Description.</emphasis>
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Coloration. Head black, antennae sooty, mandibles sepia. Prothorax saffron yellow, anterior corners and cross-shaped median longitudinal stripe sepia; the stripe not reaching posterior margin of pronotum. Meso- and metasternum and legs sepia; ventral part of abdomen sepia, sternites narrowly bordered with yellow. Scutellum and elytra sepia, each elytron with yellow apical spot.
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</paragraph>
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<paragraph id="8BC536F2EE36FFB9FF3E46F0FE8BFB4D" blockId="16.[151,1127,741,1533]" pageId="16" pageNumber="207">
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Male. Head moderately elongate; eyes of medium size, slightly protruding; head across eyes approximately as wide as pronotum, temporal margins arcuately narrowing posteriorly; surface of head very finely punctate and yellow pubescent, lustrous. Antenna moderately exceeding abdominal apex. Pronotum slightly wider than long, anterior margin widely rounded, anterior corners obtuse, very slightly rounded, lateral margins moderately concave, posterior corners obtusely rounded, posterior margin protruding apicad, rounded; surface of pronotum sculptured and pubescent similarly to head, lustrous. Elytra finely rugulose-lacunose, finely yellow pubescent, semilustrous. Last abdominal segments as in
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<figureCitation id="13412A77EE36FFB9FCD34110FC55FB8D" box="[856,987,1125,1149]" captionStart="Figs" captionStartId="15.[113,152,1435,1454]" captionTargetBox="[141,1088,219,1372]" captionTargetId="figure-12@15.[113,1091,219,1396]" captionTargetPageId="15" captionText="Figs. 28–33. 28–30 – Malthodes fritzlari sp. nov. (holotype): 28 – last abdominal segments of male, lateral view; 29 – last tergite of male, dorsal view; 30 – apical part of last sternite of male, ventro-caudal view. 31–33 – M. lysosensis sp. nov. (holotype): 31 – last abdominal segments of male, lateral view; 32 – apical part of last sternite of male, ventro-caudal view; 33 – last tergite of male, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5363043" httpUri="https://zenodo.org/record/5363043/files/figure.png" pageId="16" pageNumber="207">Figs. 31–33</figureCitation>
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, last sternite ventrally in its apical half before bifurcation with median longitudinal groove. Aedeagus as in
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<figureCitation id="13412A77EE36FFB9FF3841D0FE8EFB4D" box="[179,256,1189,1213]" captionStart="Figs" captionStartId="7.[113,152,1542,1561]" captionTargetBox="[160,1042,220,1511]" captionTargetId="figure-12@7.[160,1042,220,1512]" captionTargetPageId="7" captionText="Figs. 13–16. Aedeagus, dorsal view. 13 – Malthodes lysosensis sp. nov. (holotype); 14 – M. anadyomenae sp. nov. (paratype); 15 – M. cyprogenius sp. nov. (holotype); 16 – M. andreasi sp. nov. (paratype). Scale bars = 0.1 mm. Arrows refer to diagnostic characters mentioned in descriptions." figureDoi="http://doi.org/10.5281/zenodo.5363037" httpUri="https://zenodo.org/record/5363037/files/figure.png" pageId="16" pageNumber="207">Fig. 13</figureCitation>
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.
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</paragraph>
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<paragraph id="8BC536F2EE36FFB9FF3E41B0FEFBFB2D" blockId="16.[151,1127,741,1533]" box="[181,373,1221,1245]" pageId="16" pageNumber="207">Female unknown.</paragraph>
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<paragraph id="8BC536F2EE36FFB9FF3E4190FE1DFB0D" blockId="16.[151,1127,741,1533]" box="[181,403,1248,1278]" pageId="16" pageNumber="207">
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Length (J):
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<quantity id="4C829B17EE36FFB9FEB14190FE01FB0D" box="[314,399,1253,1277]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="16" pageNumber="207" unit="mm" value="3.4">3.4 mm</quantity>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3606579EE36FFB9FF1C4070FE5AFA4D" pageId="16" pageNumber="207" type="diagnosis">
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<paragraph id="8BC536F2EE36FFB9FF1C4070FE5AFA4D" blockId="16.[151,1127,741,1533]" pageId="16" pageNumber="207">
|
|||
|
|
<emphasis id="B90EEAE0EE36FFB9FF1C4070FE1EFAED" bold="true" box="[151,400,1285,1309]" pageId="16" pageNumber="207">Differential diagnosis.</emphasis>
|
|||
|
|
<taxonomicName id="4C7A4D71EE36FFB9FE1E4072FDF8FAEE" authority="Švihla, 2009" authorityName="Švihla" authorityYear="2009" box="[405,630,1287,1310]" class="Insecta" family="Cantharidae" genus="Malthodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="207" phylum="Arthropoda" rank="species" species="lysosensis" status="sp. nov.">
|
|||
|
|
<emphasis id="B90EEAE0EE36FFB9FE1E4072FDF8FAEE" box="[405,630,1287,1310]" italics="true" pageId="16" pageNumber="207">Malthodes lysosensis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A23D579BEE36FFB9FDF74073FD43FAED" box="[636,717,1286,1309]" pageId="16" pageNumber="207" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
is similar to
|
|||
|
|
<taxonomicName id="4C7A4D71EE36FFB9FCD84072FB87FAEE" authority="Švihla, 2009" authorityName="Švihla" authorityYear="2009" box="[851,1033,1287,1310]" class="Insecta" family="Cantharidae" genus="Malthodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="207" phylum="Arthropoda" rank="species" species="anadyomenae" status="sp. nov.">
|
|||
|
|
<emphasis id="B90EEAE0EE36FFB9FCD84072FB87FAEE" box="[851,1033,1287,1310]" italics="true" pageId="16" pageNumber="207">M. anadyomenae</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A23D579BEE36FFB9FB844073FBEDFAED" box="[1039,1123,1286,1309]" pageId="16" pageNumber="207" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
, from which it differs by the less emarginate last sternite with more slender branches of the bifurcation, presence of the median longitudinal groove on the ventral side of the last sternite restricted to the apical half, shorter last tergite lacking laterally protruding inner portions, absence of protuberances at the base of the aedeagus, shorter laterophyses and less emarginate ventral portion of the phallus.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C3606579EE36FFB9FF1C40B0FDD2FA2D" box="[151,604,1477,1502]" pageId="16" pageNumber="207" type="etymology">
|
|||
|
|
<paragraph id="8BC536F2EE36FFB9FF1C40B0FDD2FA2D" blockId="16.[151,1127,741,1533]" box="[151,604,1477,1502]" pageId="16" pageNumber="207">
|
|||
|
|
<emphasis id="B90EEAE0EE36FFB9FF1C40B0FE9CFA2D" bold="true" box="[151,274,1477,1501]" pageId="16" pageNumber="207">Etymology</emphasis>
|
|||
|
|
. Named after the
|
|||
|
|
<typeStatus id="54C18850EE36FFB9FE5940B3FD8EFA2E" box="[466,512,1478,1502]" pageId="16" pageNumber="207">type</typeStatus>
|
|||
|
|
locality.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C3606579EE36FFB9FF1C4090FE0AFA0D" box="[151,388,1509,1533]" pageId="16" pageNumber="207" type="distribution">
|
|||
|
|
<paragraph id="8BC536F2EE36FFB9FF1C4090FE0AFA0D" blockId="16.[151,1127,741,1533]" box="[151,388,1509,1533]" pageId="16" pageNumber="207">
|
|||
|
|
<emphasis id="B90EEAE0EE36FFB9FF1C4090FEA4FA0D" bold="true" box="[151,298,1509,1533]" pageId="16" pageNumber="207">Distribution.</emphasis>
|
|||
|
|
<collectingCountry id="F36D7662EE36FFB9FEBB4090FE0EFA0D" box="[304,384,1509,1533]" name="Cyprus" pageId="16" pageNumber="207">Cyprus</collectingCountry>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|