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<document id="4D3F2DBAAB35CD0A281AF50E8FD1C82D" ID-DOI="10.11646/zootaxa.3946.4.6" ID-GBIF-Dataset="ff41f8d9-3726-4d32-9cef-c70d7709fb0e" ID-ISSN="1175-5326" ID-Zenodo-Dep="244907" ID-ZooBank="684012F3-13B1-48BE-8869-183D438B055B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461038875957" checkinUser="plazi" docAuthor="Izadizadeh, Mohammad, Talebi, Ali Asghar, Achterberg, Cornelis Van &amp; Rakhshani, Ehsan" docDate="2015" docId="03D3879DFFD8FFB9FF0DA0564CFAFCE2" docLanguage="en" docName="zt03946p582.pdf" docOrigin="Zootaxa 3946 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Helorus" docType="key" docVersion="8" lastPageNumber="578" masterDocId="FFEAFFE5FFD9FFB8FF9AA17E4966FFD9" masterDocTitle="First record of the family Heloridae (Hymenoptera: Proctotrupoidea) from Iran, with description of a new species" masterLastPageNumber="582" masterPageNumber="577" pageNumber="578" updateTime="1698575121966" updateUser="plazi">
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<mods:title id="E3DFABCA23F9C10C9C15C42840CB3E9A">First record of the family Heloridae (Hymenoptera: Proctotrupoidea) from Iran, with description of a new species</mods:title>
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<mods:name id="364EC3759F6CD1F062BD6C72F611417B" type="personal">
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<mods:namePart id="A9AA10BF79EB2552604F406C48ECD188">Izadizadeh, Mohammad</mods:namePart>
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<mods:namePart id="06F6FC0C09A9F974FFCEB5FBE8AB46AD">Talebi, Ali Asghar</mods:namePart>
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<mods:namePart id="9CB248DB57A0DAED629DD4C2CB0319CA">Achterberg, Cornelis Van</mods:namePart>
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<mods:roleTerm id="4D9172088C6AD8314F88B2E7530A80C0">Author</mods:roleTerm>
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<mods:namePart id="9F37D7B50F40BDD6F32F6834C12204B8">Rakhshani, Ehsan</mods:namePart>
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<mods:typeOfResource id="3939FABD5121C0785BFE5CF545980978">text</mods:typeOfResource>
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<mods:title id="0FF5D7E44DB2DA1588AC94D3BEBE4DB3">Zootaxa</mods:title>
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<mods:part id="1D44854332726BF9FD5E805828A1F7BD">
<mods:date id="A31CBBDC902C5920F888FEFA07F7EDF8">2015</mods:date>
<mods:detail id="7AEABAE9E3F1BE0104D81594A97A99C6" type="volume">
<mods:number id="141A69CFD19319903E3CFEB30B77A6B0">3946</mods:number>
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<mods:classification id="DB170E037155625678B39AE835367EC2">journal article</mods:classification>
<mods:identifier id="3DF7EF367DDFD3E9B8A99C80AD18D7AD" type="DOI">10.11646/zootaxa.3946.4.6</mods:identifier>
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<treatment id="03D3879DFFD8FFB9FF0DA0564CFAFCE2" ID-DOI="http://doi.org/10.5281/zenodo.6103182" ID-GBIF-Taxon="119609508" ID-Zenodo-Dep="6103182" LSID="urn:lsid:plazi:treatment:03D3879DFFD8FFB9FF0DA0564CFAFCE2" httpUri="http://treatment.plazi.org/id/03D3879DFFD8FFB9FF0DA0564CFAFCE2" lastPageNumber="578" pageId="1" pageNumber="578">
<subSubSection id="C3606500FFD8FFB9FF0DA0564A42FE98" box="[151,804,295,322]" pageId="1" pageNumber="578" type="nomenclature">
<paragraph id="8BC5368BFFD8FFB9FF0DA0564A42FE98" blockId="1.[151,804,295,322]" box="[151,804,295,322]" pageId="1" pageNumber="578">
<heading id="D08D81E7FFD8FFB9FF0DA0564A42FE98" bold="true" box="[151,804,295,322]" fontSize="11" level="1" pageId="1" pageNumber="578" reason="1">
<emphasis id="B90EEA99FFD8FFB9FF0DA0564A42FE98" bold="true" box="[151,804,295,322]" pageId="1" pageNumber="578">
Key to West Palaearctic species of the genus
<taxonomicName id="4C7A4D08FFD8FFB9FD59A0594A42FE98" ID-CoL="62VG6" ID-ENA="44345" box="[707,804,295,321]" class="Insecta" family="Heloridae" genus="Helorus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="578" phylum="Arthropoda" rank="genus">
<emphasis id="B90EEA99FFD8FFB9FD59A0594A42FE98" bold="true" box="[707,804,295,321]" italics="true" pageId="1" pageNumber="578">Helorus</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C3606500FFD8FFB9FF0DA0104CFAFCE2" pageId="1" pageNumber="578" type="key">
<keyStep id="308E2E2EFFD8FFB9FF0DA0104CFDFDCF" pageId="1" pageNumber="578">
<paragraph id="8BC5368BFFD8FFB9FF0DA0104CFDFE67" blockId="1.[151,1436,366,827]" pageId="1" pageNumber="578">
<keyLead id="308B95BEFFD8FFB9FF0DA0104CFDFE67" pageId="1" pageNumber="578">
1. First metasomal tergite slender subbasally; 2.7–3.3 × as long as its maximum width in dorsal view (in ♂
<taxonomicName id="4C7A4D08FFD8FFB9FB53A0104C27FE5A" box="[1225,1345,366,387]" class="Insecta" family="Heloridae" genus="Helorus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="578" phylum="Arthropoda" rank="species" species="striolatus">
<emphasis id="B90EEA99FFD8FFB9FB53A0104C27FE5A" box="[1225,1345,366,387]" italics="true" pageId="1" pageNumber="578">H. striolatus</emphasis>
</taxonomicName>
2.3–2.7), in lateral view relatively slender (
<figureCitation id="13412A0EFFD8FFB9FDAFA0F54B14FE79" box="[565,626,395,416]" captionStart="FIGURE 2" captionStartId="3.[151,250,965,987]" captionTargetBox="[250,1338,198,942]" captionTargetId="figure@3.[245,1341,195,943]" captionTargetPageId="3" captionText="FIGURE 2. Helorus alborzicus Izadizadeh, van Achterberg &amp; Talebi sp. nov. A. Dorsal aspect of head, B. dorsal aspect of mesosoma, C. Dorsal aspect of metasoma, D. Lateral aspect of metasoma." httpUri="https://zenodo.org/record/244909/files/figure.png" pageId="1" pageNumber="578">Fig. 2</figureCitation>
D, 4D); face and mesoscutum smooth, with inconspicuous sculpture and at most punctulate (e.g. marked with very small dots or punctures)..................................................... 2
</keyLead>
</paragraph>
<paragraph id="8BC5368BFFD8FFB9FF0DA0B84CFDFDCF" blockId="1.[151,1436,366,827]" pageId="1" pageNumber="578">
<keyLead id="308B95BEFFD8FFB9FF0DA0B84CFDFDCF" pageId="1" pageNumber="578">- First metasomal tergite wider subbasally, 1.8–2.3 × as long as its maximum width in dorsal view, in lateral view moderately high and mesoscutum distinctly finely punctate or coarsely rugose-punctate (e. g. marked with spots, holes or depressions)...................................................................................................... 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="308E2E2EFFD8FFB9FF0DA3604CF0FD52" pageId="1" pageNumber="578">
<paragraph id="8BC5368BFFD8FFB9FF0DA3604CFAFD89" blockId="1.[151,1436,366,827]" pageId="1" pageNumber="578">
<keyLead id="308B95BEFFD8FFB9FF0DA3604CFAFD89" pageId="1" pageNumber="578">
2. Pterostigma stout (
<figureCitation id="13412A0EFFD8FFB9FE0FA36048A8FDEA" box="[405,462,542,563]" captionStart="FIGURE 3" captionStartId="4.[151,250,1574,1596]" captionTargetBox="[195,1386,201,1527]" captionTargetId="figure@4.[194,1390,193,1532]" captionTargetPageId="4" captionText="FIGURE 3. Helorus ruficornis Foerster, 1856. A. Lateral aspect of male general habitus, B. Fore wing." httpUri="https://zenodo.org/record/244910/files/figure.png" pageId="1" pageNumber="578">Fig. 3</figureCitation>
B), 2.1–2.6 × as long as wide; mesopleuron with smooth area near anterior margin; antenna yellowish brown (
<figureCitation id="13412A0EFFD8FFB9FEB1A3454802FD89" box="[299,356,571,592]" captionStart="FIGURE 3" captionStartId="4.[151,250,1574,1596]" captionTargetBox="[195,1386,201,1527]" captionTargetId="figure@4.[194,1390,193,1532]" captionTargetPageId="4" captionText="FIGURE 3. Helorus ruficornis Foerster, 1856. A. Lateral aspect of male general habitus, B. Fore wing." httpUri="https://zenodo.org/record/244910/files/figure.png" pageId="1" pageNumber="578">Fig. 3</figureCitation>
A); fore and middle legs yellow or yellowish-brown (
<figureCitation id="13412A0EFFD8FFB9FCFFA3454AFBFD89" box="[869,925,571,592]" captionStart="FIGURE 3" captionStartId="4.[151,250,1574,1596]" captionTargetBox="[195,1386,201,1527]" captionTargetId="figure@4.[194,1390,193,1532]" captionTargetPageId="4" captionText="FIGURE 3. Helorus ruficornis Foerster, 1856. A. Lateral aspect of male general habitus, B. Fore wing." httpUri="https://zenodo.org/record/244910/files/figure.png" pageId="1" pageNumber="578">Fig. 3</figureCitation>
A)..................
<taxonomicName id="4C7A4D08FFD8FFB9FB01A3454CFAFD89" authority="Forster, 1856" authorityName="Forster" authorityYear="1856" box="[1179,1436,571,592]" class="Insecta" family="Heloridae" genus="Helorus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="578" phylum="Arthropoda" rank="species" species="ruficornis">
<emphasis id="B90EEA99FFD8FFB9FB01A3454C70FD89" box="[1179,1302,571,592]" italics="true" pageId="1" pageNumber="578">H. ruficornis</emphasis>
Förster, 1856
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BC5368BFFD8FFB9FF0DA3274CF0FD52" blockId="1.[151,1436,366,827]" pageId="1" pageNumber="578">
<keyLead id="308B95BEFFD8FFB9FF0DA3274CF0FD52" pageId="1" pageNumber="578">
- Pterostigma slender (
<figureCitation id="13412A0EFFD8FFB9FE36A3274884FDB7" box="[428,482,601,622]" captionStart="FIGURE 1" captionStartId="2.[151,250,1573,1595]" captionTargetBox="[252,1335,196,1545]" captionTargetId="figure@2.[247,1339,193,1552]" captionTargetPageId="2" captionText="FIGURE 1. Helorus alborzicus Izadizadeh, van Achterberg &amp; Talebi sp. nov. A. Lateral aspect of female general habitus, B. Fore wing." httpUri="https://zenodo.org/record/244908/files/figure.png" pageId="1" pageNumber="578">Fig. 1</figureCitation>
B), 2.9–3.6 × as long as wide; mesopleuron distinctly crenulate-rugose subanteriorly; antenna dark brown or black (
<figureCitation id="13412A0EFFD8FFB9FEE5A30848D3FD52" box="[383,437,630,651]" captionStart="FIGURE 1" captionStartId="2.[151,250,1573,1595]" captionTargetBox="[252,1335,196,1545]" captionTargetId="figure@2.[247,1339,193,1552]" captionTargetPageId="2" captionText="FIGURE 1. Helorus alborzicus Izadizadeh, van Achterberg &amp; Talebi sp. nov. A. Lateral aspect of female general habitus, B. Fore wing." httpUri="https://zenodo.org/record/244908/files/figure.png" pageId="1" pageNumber="578">Fig. 1</figureCitation>
A); coxae, trochanters and basal halves of femora of fore and middle legs black (
<figureCitation id="13412A0EFFD8FFB9FB24A3084D92FD52" box="[1214,1268,630,651]" captionStart="FIGURE 1" captionStartId="2.[151,250,1573,1595]" captionTargetBox="[252,1335,196,1545]" captionTargetId="figure@2.[247,1339,193,1552]" captionTargetPageId="2" captionText="FIGURE 1. Helorus alborzicus Izadizadeh, van Achterberg &amp; Talebi sp. nov. A. Lateral aspect of female general habitus, B. Fore wing." httpUri="https://zenodo.org/record/244908/files/figure.png" pageId="1" pageNumber="578">Fig. 1</figureCitation>
A)......... 3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="308E2E2EFFD8FFB9FF0DA3EA4CFAFCD8" pageId="1" pageNumber="578">
<paragraph id="8BC5368BFFD8FFB9FF0DA3EA4CFAFD1F" blockId="1.[151,1436,366,827]" pageId="1" pageNumber="578">
<keyLead id="308B95BEFFD8FFB9FF0DA3EA4CFAFD1F" pageId="1" pageNumber="578">
3. Frons laterally, vertex (
<figureCitation id="13412A0EFFD8FFB9FE25A3EA489EFD70" box="[447,504,660,681]" captionStart="FIGURE 2" captionStartId="3.[151,250,965,987]" captionTargetBox="[250,1338,198,942]" captionTargetId="figure@3.[245,1341,195,943]" captionTargetPageId="3" captionText="FIGURE 2. Helorus alborzicus Izadizadeh, van Achterberg &amp; Talebi sp. nov. A. Dorsal aspect of head, B. dorsal aspect of mesosoma, C. Dorsal aspect of metasoma, D. Lateral aspect of metasoma." httpUri="https://zenodo.org/record/244909/files/figure.png" pageId="1" pageNumber="578">Fig. 2</figureCitation>
A) and middle lobe of mesoscutum (
<figureCitation id="13412A0EFFD8FFB9FCCDA3EA4AF7FD70" box="[855,913,660,681]" captionStart="FIGURE 2" captionStartId="3.[151,250,965,987]" captionTargetBox="[250,1338,198,942]" captionTargetId="figure@3.[245,1341,195,943]" captionTargetPageId="3" captionText="FIGURE 2. Helorus alborzicus Izadizadeh, van Achterberg &amp; Talebi sp. nov. A. Dorsal aspect of head, B. dorsal aspect of mesosoma, C. Dorsal aspect of metasoma, D. Lateral aspect of metasoma." httpUri="https://zenodo.org/record/244909/files/figure.png" pageId="1" pageNumber="578">Fig. 2</figureCitation>
B) largely smooth; first metasomal tergite 3.2–3.5 × as long as wide in lateral view (
<figureCitation id="13412A0EFFD8FFB9FE6CA3CF4B56FD1F" box="[502,560,689,710]" captionStart="FIGURE 2" captionStartId="3.[151,250,965,987]" captionTargetBox="[250,1338,198,942]" captionTargetId="figure@3.[245,1341,195,943]" captionTargetPageId="3" captionText="FIGURE 2. Helorus alborzicus Izadizadeh, van Achterberg &amp; Talebi sp. nov. A. Dorsal aspect of head, B. dorsal aspect of mesosoma, C. Dorsal aspect of metasoma, D. Lateral aspect of metasoma." httpUri="https://zenodo.org/record/244909/files/figure.png" pageId="1" pageNumber="578">Fig. 2</figureCitation>
D).......................
<taxonomicName id="4C7A4D08FFD8FFB9FCFAA3CF4D37FD1F" authority="Izadizadeh" authorityName="Izadizadeh" box="[864,1105,689,710]" class="Insecta" family="Heloridae" genus="Helorus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="578" phylum="Arthropoda" rank="species" species="alborzicus">
<emphasis id="B90EEA99FFD8FFB9FCFAA3CF4A17FD1F" box="[864,881,689,710]" italics="true" pageId="1" pageNumber="578">H</emphasis>
.
<emphasis id="B90EEA99FFD8FFB9FCE7A3CF4A87FD1F" box="[893,993,689,710]" italics="true" pageId="1" pageNumber="578">alborzicus</emphasis>
Izadizadeh
</taxonomicName>
, van Achterberg &amp; Talebi
<emphasis id="B90EEA99FFD8FFB9FAD5A3CE4CFAFD1F" bold="true" box="[1359,1436,688,710]" pageId="1" pageNumber="578">sp. nov.</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BC5368BFFD8FFB9FF0DA3B04CFAFCD8" blockId="1.[151,1436,366,827]" pageId="1" pageNumber="578">
<keyLead id="308B95BEFFD8FFB9FF0DA3B04CFAFCD8" pageId="1" pageNumber="578">
- Frons laterally finely and more or less densely punctate and middle lobe of mesoscutum and vertex moderately shiny and finely punctulate; first metasomal tergite 2.3–3.0 × as long as wide in lateral view............
<taxonomicName id="4C7A4D08FFD8FFB9FB10A3924CFAFCD8" authority="Cameron, 1906" authorityName="Cameron" authorityYear="1906" box="[1162,1436,748,769]" class="Insecta" family="Heloridae" genus="Helorus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="578" phylum="Arthropoda" rank="species" species="striolatus">
<emphasis id="B90EEA99FFD8FFB9FB10A3924C64FCD8" box="[1162,1282,748,769]" italics="true" pageId="1" pageNumber="578">H. striolatus</emphasis>
<collectingCountry id="F36D761BFFD8FFB9FA92A3924C04FCD8" box="[1288,1378,748,769]" name="Cameroon" pageId="1" pageNumber="578">Cameron</collectingCountry>
, 1906
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="308E2E2EFFD8FFB9FF0DA2774CFAFCE2" pageId="1" pageNumber="578">
<paragraph id="8BC5368BFFD8FFB9FF0DA2774CFAFCC7" blockId="1.[151,1436,366,827]" box="[151,1436,777,798]" pageId="1" pageNumber="578">
<keyLead id="308B95BEFFD8FFB9FF0DA2774CFAFCC7" box="[151,1436,777,798]" pageId="1" pageNumber="578">
4. Face, vertex, temple and middle lobe of mesoscutum coarsely reticulate–rugose................
<taxonomicName id="4C7A4D08FFD8FFB9FB31A2774CFAFCC7" authority="Forster, 1856" authorityName="Forster" authorityYear="1856" box="[1195,1436,777,798]" class="Insecta" family="Heloridae" genus="Helorus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="578" phylum="Arthropoda" rank="species" species="nigripes">
<emphasis id="B90EEA99FFD8FFB9FB31A2774C70FCC7" box="[1195,1302,777,798]" italics="true" pageId="1" pageNumber="578">H. nigripes</emphasis>
Förster, 1856
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BC5368BFFD8FFB9FF0DA2584CFAFCE2" blockId="1.[151,1436,366,827]" box="[151,1436,806,827]" pageId="1" pageNumber="578">
<keyLead id="308B95BEFFD8FFB9FF0DA2584CFAFCE2" box="[151,1436,806,827]" pageId="1" pageNumber="578">
- Face, vertex, temple and middle lobe of mesoscutum moderately finely punctate............
<taxonomicName id="4C7A4D08FFD8FFB9FBE4A2584CFAFCE2" authority="Panzer, 1798" authorityName="Panzer" authorityYear="1798" box="[1150,1436,806,827]" class="Insecta" family="Heloridae" genus="Helorus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="578" phylum="Arthropoda" rank="species" species="anomalipes">
<emphasis id="B90EEA99FFD8FFB9FBE4A2584C6CFCE2" box="[1150,1290,806,827]" italics="true" pageId="1" pageNumber="578">H. anomalipes</emphasis>
(Panzer, 1798)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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