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<document id="72DD8393F01BCB79C6AACC06B4EF44E9" ID-CLB-Dataset="115951" ID-DOI="10.1093/zoolinnean/zlab033" ID-GBIF-Dataset="75b10efe-a17d-4d10-b46e-094c6a7ebf01" ID-ISSN="0024-4082" ID-Zenodo-Dep="6967774" ID-ZooBank="BE2B3859-8F6A-4543-8A69-91840F82377B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1653991067759" checkinUser="plazi" docAuthor="Lin, Shou-Wang, Lopardo, Lara &amp; Uhl, Gabriele" docDate="2022" docId="03B787C7FF5476A7FF0ED02E4EF7F23D" docLanguage="en" docName="zlab033.pdf" docOrigin="Zoological Journal of the Linnean Society 195" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Oedothorax stylus Tanasevitch 2015, INCERTAE SEDIS" docType="treatment" docVersion="7" lastPageNumber="554" masterDocId="FF8EFFBFFFDD762DFFABD4584A34F51A" masterDocTitle="Evolution of nuptial-gift-related male prosomal structures: taxonomic revision and cladistic analysis of the genus Oedothorax (Araneae: Linyphiidae: Erigoninae)" masterLastPageNumber="584" masterPageNumber="417" pageNumber="554" updateTime="1699338434129" updateUser="ExternalLinkService">
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<mods:title id="C79F169EFCBBAE19E2580D5804270021">Evolution of nuptial-gift-related male prosomal structures: taxonomic revision and cladistic analysis of the genus Oedothorax (Araneae: Linyphiidae: Erigoninae)</mods:title>
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<mods:namePart id="AC76E87E2BDA85795376EE9B9FE9D87D">Lin, Shou-Wang</mods:namePart>
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<mods:namePart id="B7A5007CAE9D0C1953CDCDEC4B0EE4E8">Lopardo, Lara</mods:namePart>
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<mods:namePart id="24FC9CEE71D8C2D3E09A6FB1E9F45E40">Uhl, Gabriele</mods:namePart>
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<mods:date id="71CD443C1975450D9E9A0E4F909F9D7F">2022</mods:date>
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<treatment id="03B787C7FF5476A7FF0ED02E4EF7F23D" ID-DOI="http://doi.org/10.5281/zenodo.6968054" ID-GBIF-Taxon="198618792" ID-Zenodo-Dep="6968054" LSID="urn:lsid:plazi:treatment:03B787C7FF5476A7FF0ED02E4EF7F23D" httpUri="http://treatment.plazi.org/id/03B787C7FF5476A7FF0ED02E4EF7F23D" lastPageId="138" lastPageNumber="554" pageId="137" pageNumber="554">
<subSubSection id="C304655AFF5476A4FF0ED02E4BCCF1B6" pageId="137" pageNumber="554" type="nomenclature">
<paragraph id="8BA136D1FF5476A4FF0ED02E4BCCF1B6" blockId="137.[165,776,1142,1196]" pageId="137" pageNumber="554">
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<taxonomicName id="4C1E4D52FF5476A4FF0ED02E48A5F195" ID-CoL="48WVK" authority="TANASEVITCH, 2015 INCERTAE" authorityName="Tanasevitch" authorityYear="2015" box="[165,657,1142,1167]" class="Arachnida" family="Linyphiidae" genus="Oedothorax" kingdom="Animalia" order="Araneae" pageId="137" pageNumber="554" phylum="Arthropoda" rank="species" species="stylus" status="INCERTAE SEDIS">
<emphasis id="B96AEAC3FF5476A4FF0ED02E4B9CF194" box="[165,424,1142,1166]" italics="true" pageId="137" pageNumber="554">OEDOTHORAX STYLUS</emphasis>
<bibRefCitation id="EF8F4B20FF5476A4FE05D02E48A5F195" author="Tanasevitch AV" box="[430,657,1142,1167]" pageId="137" pageNumber="554" pagination="381 - 398" refId="ref95840" refString="Tanasevitch AV. 2015. Notes on the spider genus Oedothorax Bertkau, 1883 with description of eleven new species from India (Linyphiidae: Erigoninae). Revue suisse de Zoologie 122: 381 - 398." type="journal article" year="2015">TANASEVITCH, 2015</bibRefCitation>
</taxonomicName>
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<emphasis id="B96AEAC3FF5476A4FD33D0224BCCF1B6" italics="true" pageId="137" pageNumber="554">INCERTAE SEDIS</emphasis>
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</heading>
</paragraph>
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<subSubSection id="C304655AFF5476A4FEFFD0E3486DF1CE" box="[340,601,1211,1236]" pageId="137" pageNumber="554" type="description">
<paragraph id="8BA136D1FF5476A4FEFFD0E3486DF1CE" blockId="137.[340,601,1211,1236]" box="[340,601,1211,1236]" pageId="137" pageNumber="554">
(
<figureCitation id="13252A54FF5476A4FEF6D0E34B82F1C9" box="[349,438,1211,1236]" captionStart="Figure 7" captionStartId="13.[163,240,1315,1337]" captionTargetBox="[166,1440,198,1272]" captionTargetId="figure-25@13.[163,1443,195,1275]" captionTargetPageId="13" captionText="Figure 7. Male prosomal morphology, lateral.A, Emertongone montifera (Emerton, 1882) (Paquin &amp; Dupérré, 2003, fig. 1187). B, Shaanxinus mingchihensis Lin, 2019 (traced from photograph). C, Oedothorax paracymbialis Tanasevitch, 2015 (traced from photograph). D, Oe. kodaikanal Tanasevitch, 2015 (traced from photograph). E, Oe. cunur Tanasevitch, 2015 (traced from photograph). F, Oe. stylus Tanasevitch, 2015 (traced from photograph). G, Atypena cirrifrons (Heimer, 1984) (traced from photograph). H, A. formosana (Oi, 1977) (Oi, 1977, fig. 2). I, Oe. nazareti Scharff, 1989 (Scharff, 1989: fig. 7). J, Gongylidium rufipes (Linnaeus, 1758) (traced from photograph). K, Ummeliata insecticeps (Bösenberg &amp; Strand, 1906) (Tu &amp; Li 2004: fig. 8A). L, U. esyunini (Zhang, Zhang &amp; Yu, 2003) (Zhang et al., 2003, fig. 2A). M, Hylyphantes graminicola (Sundevall, 1830) (traced from photograph). N, Tmeticus tolli Kulczyński, 1908 (traced from photograph). O, Oedothorax gibbosus (Blackwall, 1841) [Roberts, 1987: fig. 21 (e)]. P, Oe. trilobatus (Banks, 1896) (Bishop &amp; Crosby, 1935: fig. 79). Q, Oe. gibbifer (Kulczyński, 1882) (Heimer &amp; Nentwig, 1991: fig.604.5).R, Oe.apicatus (Blackwall, 1850) [Roberts, 1987: fig.23 (d)].S, Oe. retusus (Westring, 1851) [Roberts, 1987: fig. 23 (c)]. T, Oe. paludigena Simon, 1926 [Millidge, 1975: fig. (5)]. U, Oe. meridionalis Tanasevitch, 1987 (Tanasevitch, 1987: fig. 107).V, Oe. agrestis (Blackwall, 1853) (Roberts, 1987: fig. 23B).W, Oe. fuscus (Blackwall, 1834) (Roberts, 1987: fig. 23A). X, Oe. tingitanus (Simon, 1884) (Denis, 1968: fig. 11)." figureDoi="http://doi.org/10.5281/zenodo.6967802" httpUri="https://zenodo.org/record/6967802/files/figure.png" pageId="137" pageNumber="554">FIGS 7F</figureCitation>
,
<figureCitation id="13252A54FF5476A4FE69D0E34BDAF1C9" box="[450,494,1211,1236]" captionStart="Figure 67" captionStartId="110.[144,224,1301,1323]" captionTargetBox="[147,1426,197,1260]" captionTargetId="figure-175@110.[145,1428,195,1261]" captionTargetPageId="110" captionText="Figure 67. Male chelicerae, stridulatory striae on lateral side. A, Jilinus hulongensis (Zhu &amp; Wen, 1980). B, Cornitibia simplicithorax (Tanasevitch, 1998). C, Emertongone montifera (Emerton, 1882). D, Oedothorax paracymbialis Tanasevitch, 2015. E, Oe. kodaikanal Tanasevitch, 2015. F, Oe. cunur Tanasevitch, 2015. G, Oe. stylus Tanasevitch, 2015. H, Atypena cirrifrons (Heimer, 1984). I, A. formosana (Oi, 1977). J, Oe. nazareti Scharff, 1989. K, Gongylidium rufipes (Linnaeus, 1758). L, Ummeliata insecticeps (Bösenberg &amp; Strand, 1906). M, U. esyunini (Zhang, Zhang &amp; Yu, 2003). N, Hylyphantes graminicola (Sundevall, 1830). O, Tmeticus tolli Kulczyński, 1908. Scale bars 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.6967988" httpUri="https://zenodo.org/record/6967988/files/figure.png" pageId="137" pageNumber="554">67G</figureCitation>
,
<figureCitation id="13252A54FF5476A4FE50D0E34813F1C9" box="[507,551,1211,1236]" captionStart="Figure 68" captionStartId="111.[163,245,1308,1330]" captionTargetBox="[164,1444,196,1268]" captionTargetId="figure-26@111.[163,1446,195,1269]" captionTargetPageId="111" captionText="Figure 68. Male opisthosoma, dorsal view. A, Jilinus hulongensis (Zhu &amp; Wen, 1980). B, Cornitibia simplicithorax (Tanasevitch, 1998). C, Emertongone montifera (Emerton, 1882). D, Oedothorax paracymbialis Tanasevitch, 2015. E, Oe. kodaikanal Tanasevitch, 2015. F, Oe. cunur Tanasevitch, 2015. G, Oe. stylus Tanasevitch, 2015. H, Atypena cirrifrons (Heimer, 1984). I, A. formosana (Oi, 1977). J, Oe. nazareti Scharff, 1989. K, Gongylidium rufipes (Linnaeus, 1758). L, Ummeliata insecticeps (Bösenberg &amp; Strand, 1906). M, U. esyunini (Zhang, Zhang &amp; Yu, 2003). N, Hylyphantes graminicola (Sundevall, 1830). O, Tmeticus tolli Kulczyński, 1908. Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6967990" httpUri="https://zenodo.org/record/6967990/files/figure.png" pageId="137" pageNumber="554">68G</figureCitation>
,
<figureCitation id="13252A54FF5476A4FD9FD0E34864F1CE" box="[564,592,1211,1236]" captionStart="Figure 80" captionStartId="138.[145,223,1612,1634]" captionTargetBox="[148,1425,202,1571]" captionTargetId="figure-106@138.[145,1428,195,1572]" captionTargetPageId="138" captionText="Figure 80. Oedothorax stylus Tanasevitch, 2015. AD, male right palp, images flipped horizontally. A, retrolateral view. B, prolateral view. C, dorsal view. D, ventral view. E, F, epigyne. E, ventrolateral view. F, external morphology. G, male posterior median spinnerets and posterior lateral spinnerets, dorsal view. H, female spinnerets. Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6968028" httpUri="https://zenodo.org/record/6968028/files/figure.png" pageId="137" pageNumber="554">80</figureCitation>
)
</paragraph>
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<treatmentCitation id="0ABF10C0FF5476A4FF08D0BC48BCF1E3" author="Tanasevitch AV" box="[163,648,1252,1274]" page="393" pageId="137" pageNumber="554" year="2015">
<taxonomicName id="4C1E4D52FF5476A4FF08D0BC48BCF1E3" ID-CoL="48WVK" authority="Tanasevitch, 2015: 393" authorityName="Tanasevitch" authorityPageNumber="393" authorityYear="2015" box="[163,648,1252,1274]" class="Arachnida" family="Linyphiidae" genus="Oedothorax" kingdom="Animalia" order="Araneae" pageId="137" pageNumber="554" phylum="Arthropoda" rank="species" species="stylus">
<emphasis id="B96AEAC3FF5476A4FF08D0BC4B40F1E3" box="[163,372,1252,1273]" italics="true" pageId="137" pageNumber="554">Oedothorax stylus</emphasis>
<bibRefCitation id="EF8F4B20FF5476A4FED0D0BC48BCF1E3" author="Tanasevitch AV" box="[379,648,1252,1274]" pageId="137" pageNumber="554" pagination="381 - 398" refId="ref95840" refString="Tanasevitch AV. 2015. Notes on the spider genus Oedothorax Bertkau, 1883 with description of eleven new species from India (Linyphiidae: Erigoninae). Revue suisse de Zoologie 122: 381 - 398." type="journal article" year="2015">Tanasevitch, 2015: 393</bibRefCitation>
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, figs 8385 (Dmf).
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</paragraph>
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<subSubSection id="C304655AFF5476A4FF08D16248E4F31D" pageId="137" pageNumber="554" type="materials_examined">
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<emphasis id="B96AEAC3FF5476A4FF08D1624B69F054" box="[163,349,1337,1359]" italics="true" pageId="137" pageNumber="554">Type material:</emphasis>
<materialsCitation id="3B763C8CFF5476A4FEC6D1614B1CF0B1" ID-GBIF-Occurrence="3880945391" collectingDate="1972-11-30" collectorName="C. Besuchet &amp; I. Lobl" country="India" elevation="900" location="Kaikatty" municipality="Nelliampathi Hills" pageId="137" pageNumber="554" specimenCount="1" specimenCount-male="1" stateProvince="Kerala" typeStatus="holotype">
<typeStatus id="54A58873FF5476A4FEC6D1614BD6F055" box="[365,482,1337,1359]" pageId="137" pageNumber="554" type="holotype">Holotype</typeStatus>
:
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<collectingCountry id="F3097641FF5476A4FE5AD161480AF055" box="[497,574,1337,1359]" name="India" pageId="137" pageNumber="554">India</collectingCountry>
:
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, NW of
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,
<location id="8EC1600AFF5476A4FE21D1004BC4F074" LSID="urn:lsid:plazi:treatment:03B787C7FF5476A7FF0ED02E4EF7F23D:8EC1600AFF5476A4FE21D1004BC4F074" box="[394,496,1368,1390]" country="India" municipality="Nelliampathi Hills" name="Kaikatty" pageId="137" pageNumber="554" stateProvince="Kerala">Kaikatty</location>
,
<quantity id="4CE69B34FF5476A4FE57D1004872F077" box="[508,582,1368,1389]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="137" pageNumber="554" unit="m" value="900.0">
<elevation id="0033D1E2FF5476A4FE57D1004872F077" box="[508,582,1368,1389]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="137" pageNumber="554" unit="m" value="900.0">900 m</elevation>
</quantity>
, sifting in forest, neara spring,
<specimenCount id="9D18FD58FF5476A4FEEED1214B61F091" box="[325,341,1401,1419]" pageId="137" pageNumber="554" type="male"></specimenCount>
<date id="FFA01011FF5476A4FEF6D12E4BE2F096" box="[349,470,1398,1420]" pageId="137" pageNumber="554" value="1972-11-30">
<collectingDate id="EFE4E9F9FF5476A4FEF6D12E4BE2F096" box="[349,470,1398,1420]" pageId="137" pageNumber="554" value="1972-11-30">30.xi.1972</collectingDate>
</date>
, leg.
<collectorName id="26EB5307FF5476A4FDB9D12F48AFF096" box="[530,667,1398,1420]" pageId="137" pageNumber="554">C. Besuchet</collectorName>
&amp;
<collectorName id="26EB5307FF5476A4FD14D12F493EF096" box="[703,778,1398,1420]" pageId="137" pageNumber="554">I. Löbl</collectorName>
(examined)
</materialsCitation>
.
<materialsCitation id="3B763C8CFF5476A4FE9FD1CE48E9F0F2" ID-GBIF-Occurrence="3880945336" country="India" county="Valparai" elevation="1250" location="Madras" municipality="Anaimalai Hills" pageId="137" pageNumber="554" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<typeStatus id="54A58873FF5476A4FE9FD1CE4B99F0B1" box="[308,429,1430,1451]" pageId="137" pageNumber="554" type="paratype">Paratypes</typeStatus>
:
<emphasis id="B96AEAC3FF5476A4FE12D1CD4832F0B1" bold="true" box="[441,518,1429,1451]" pageId="137" pageNumber="554">
<collectingCountry id="F3097641FF5476A4FE12D1CD4835F0B1" box="[441,513,1429,1451]" name="India" pageId="137" pageNumber="554">India</collectingCountry>
:
</emphasis>
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, collected together with the
<typeStatus id="54A58873FF5476A4FEB9D1EC4B4EF0D0" box="[274,378,1460,1482]" pageId="137" pageNumber="554" type="holotype">holotype</typeStatus>
;
<location id="8EC1600AFF5476A4FE2CD1EC4BD7F0D0" LSID="urn:lsid:plazi:treatment:03B787C7FF5476A7FF0ED02E4EF7F23D:8EC1600AFF5476A4FE2CD1EC4BD7F0D0" box="[391,483,1460,1482]" country="India" county="Valparai" municipality="Anaimalai Hills" name="Madras" pageId="137" pageNumber="554">Madras</location>
,
<collectingMunicipality id="6BC5ACABFF5476A4FE5BD1EC4887F0D0" box="[496,691,1460,1482]" pageId="137" pageNumber="554">Anaimalai Hills</collectingMunicipality>
,
<quantity id="4CE69B34FF5476A4FD6BD1EC493EF0D0" box="[704,778,1460,1482]" metricMagnitude="4" metricUnit="m" metricValue="1.8" pageId="137" pageNumber="554" unit="km" value="18.0">18 km</quantity>
north of
<collectingCounty id="62C04E5DFF5476A4FEA5D18A4B4CF0F2" box="[270,376,1490,1512]" pageId="137" pageNumber="554">Valparai</collectingCounty>
,
<quantity id="4CE69B34FF5476A4FE2DD18A4BD6F0F2" box="[390,482,1490,1512]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="137" pageNumber="554" unit="m" value="1250.0">
<elevation id="0033D1E2FF5476A4FE2DD18A4BD6F0F2" box="[390,482,1490,1512]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="137" pageNumber="554" unit="m" value="1250.0">1250 m</elevation>
</quantity>
, forest, sifting litter
</materialsCitation>
,
<specimenCount id="9D18FD58FF5476A4FD40D18B493EF0FD" box="[747,778,1491,1512]" pageId="137" pageNumber="554" type="male">1♂</specimenCount>
<date id="FFA01011FF5476A4FF08D1A94B2EF31D" box="[163,282,1521,1543]" pageId="137" pageNumber="554" value="1972-11-18">18.xi.1972</date>
, leg. C. Besuchet &amp; I. Löbl (examined).
</paragraph>
</subSubSection>
<subSubSection id="C304655AFF5476A4FF08D2704B7CF279" pageId="137" pageNumber="554" type="diagnosis">
<paragraph id="8BA136D1FF5476A4FF08D2704B2EF327" blockId="137.[163,282,1576,1597]" box="[163,282,1576,1597]" pageId="137" pageNumber="554">
<emphasis id="B96AEAC3FF5476A4FF08D2704B2EF327" box="[163,282,1576,1597]" italics="true" pageId="137" pageNumber="554">Diagnosis:</emphasis>
</paragraph>
<paragraph id="8BA136D1FF5476A4FF08D2064938F3CA" blockId="137.[163,780,1630,1744]" pageId="137" pageNumber="554">
<emphasis id="B96AEAC3FF5476A4FF08D2064AC5F369" box="[163,241,1630,1651]" italics="true" pageId="137" pageNumber="554">Males:</emphasis>
This species can be diagnosed by the scaly elevation representing palpal tibial prolateral apophysis, the exceptionally long and slender ventral radical process, and the lack of prosomal modification.
</paragraph>
<paragraph id="8BA136D1FF5476A4FF08D2A94B7CF279" blockId="137.[163,779,1777,1891]" pageId="137" pageNumber="554">
<emphasis id="B96AEAC3FF5476A4FF08D2A94B3EF21C" box="[163,266,1777,1798]" italics="true" pageId="137" pageNumber="554">Females:</emphasis>
The general appearance of this species is
<taxonomicName id="4C1E4D52FF5476A4FF08D3484B1EF23F" authorityName="Lin &amp; Lopardo &amp; Uhl" authorityYear="2022" box="[163,298,1808,1829]" class="Arachnida" family="Linyphiidae" genus="Oedothorax" kingdom="Animalia" order="Araneae" pageId="137" pageNumber="554" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAC3FF5476A4FF08D3484B1EF23F" box="[163,298,1808,1829]" italics="true" pageId="137" pageNumber="554">Oedothorax</emphasis>
</taxonomicName>
-like. It can be identified by the anterior position of the copulatory ducts opening to the spermathecae.
</paragraph>
</subSubSection>
<subSubSection id="C304655AFF5476A4FC90D49E499BF37B" pageId="137" pageNumber="554" type="description">
<paragraph id="8BA136D1FF5476A4FC90D49E49F7F5C1" blockId="137.[827,963,198,219]" box="[827,963,198,219]" pageId="137" pageNumber="554">
<emphasis id="B96AEAC3FF5476A4FC90D49E49F7F5C1" box="[827,963,198,219]" italics="true" pageId="137" pageNumber="554">Description:</emphasis>
</paragraph>
<paragraph id="8BA136D1FF5476A4FC90D55C4E9EF12C" blockId="137.[827,1444,260,1079]" pageId="137" pageNumber="554">
<emphasis id="B96AEAC3FF5476A4FC90D55C49C5F403" box="[827,1009,260,281]" italics="true" pageId="137" pageNumber="554">
Male (
<typeStatus id="54A58873FF5476A4FC2AD55C49D0F403" box="[897,996,260,281]" pageId="137" pageNumber="554" type="paratype">paratype</typeStatus>
):
</emphasis>
Total length: 1.70. Prosoma: 0.77 long, 0.67 wide, unmodified (
<figureCitation id="13252A54FF5476A4FB94D57A4EB9F422" box="[1087,1165,290,312]" captionStart="Figure 7" captionStartId="13.[163,240,1315,1337]" captionTargetBox="[166,1440,198,1272]" captionTargetId="figure-25@13.[163,1443,195,1275]" captionTargetPageId="13" captionText="Figure 7. Male prosomal morphology, lateral.A, Emertongone montifera (Emerton, 1882) (Paquin &amp; Dupérré, 2003, fig. 1187). B, Shaanxinus mingchihensis Lin, 2019 (traced from photograph). C, Oedothorax paracymbialis Tanasevitch, 2015 (traced from photograph). D, Oe. kodaikanal Tanasevitch, 2015 (traced from photograph). E, Oe. cunur Tanasevitch, 2015 (traced from photograph). F, Oe. stylus Tanasevitch, 2015 (traced from photograph). G, Atypena cirrifrons (Heimer, 1984) (traced from photograph). H, A. formosana (Oi, 1977) (Oi, 1977, fig. 2). I, Oe. nazareti Scharff, 1989 (Scharff, 1989: fig. 7). J, Gongylidium rufipes (Linnaeus, 1758) (traced from photograph). K, Ummeliata insecticeps (Bösenberg &amp; Strand, 1906) (Tu &amp; Li 2004: fig. 8A). L, U. esyunini (Zhang, Zhang &amp; Yu, 2003) (Zhang et al., 2003, fig. 2A). M, Hylyphantes graminicola (Sundevall, 1830) (traced from photograph). N, Tmeticus tolli Kulczyński, 1908 (traced from photograph). O, Oedothorax gibbosus (Blackwall, 1841) [Roberts, 1987: fig. 21 (e)]. P, Oe. trilobatus (Banks, 1896) (Bishop &amp; Crosby, 1935: fig. 79). Q, Oe. gibbifer (Kulczyński, 1882) (Heimer &amp; Nentwig, 1991: fig.604.5).R, Oe.apicatus (Blackwall, 1850) [Roberts, 1987: fig.23 (d)].S, Oe. retusus (Westring, 1851) [Roberts, 1987: fig. 23 (c)]. T, Oe. paludigena Simon, 1926 [Millidge, 1975: fig. (5)]. U, Oe. meridionalis Tanasevitch, 1987 (Tanasevitch, 1987: fig. 107).V, Oe. agrestis (Blackwall, 1853) (Roberts, 1987: fig. 23B).W, Oe. fuscus (Blackwall, 1834) (Roberts, 1987: fig. 23A). X, Oe. tingitanus (Simon, 1884) (Denis, 1968: fig. 11)." figureDoi="http://doi.org/10.5281/zenodo.6967802" httpUri="https://zenodo.org/record/6967802/files/figure.png" pageId="137" pageNumber="554">Fig. 7F</figureCitation>
). Eyes: AME-AME: 0.02, AME width: 0.05, AME-ALE: 0.02, ALE width: 0.08, ALE-PLE: 0.01, PLE width: 0.08, PLE-PME: 0.03, PME width: 0.08, PME-PME: 0.03. Clypeus: not hirsute, one sub-AME seta. Sternum: 0.48 long, 0.48 wide. Chelicerae: mastidia absent; stridulatory striae rows compressed proximally (
<figureCitation id="13252A54FF5476A4FBEED5824E94F4EA" box="[1093,1184,474,496]" captionStart="Figure 67" captionStartId="110.[144,224,1301,1323]" captionTargetBox="[147,1426,197,1260]" captionTargetId="figure-175@110.[145,1428,195,1261]" captionTargetPageId="110" captionText="Figure 67. Male chelicerae, stridulatory striae on lateral side. A, Jilinus hulongensis (Zhu &amp; Wen, 1980). B, Cornitibia simplicithorax (Tanasevitch, 1998). C, Emertongone montifera (Emerton, 1882). D, Oedothorax paracymbialis Tanasevitch, 2015. E, Oe. kodaikanal Tanasevitch, 2015. F, Oe. cunur Tanasevitch, 2015. G, Oe. stylus Tanasevitch, 2015. H, Atypena cirrifrons (Heimer, 1984). I, A. formosana (Oi, 1977). J, Oe. nazareti Scharff, 1989. K, Gongylidium rufipes (Linnaeus, 1758). L, Ummeliata insecticeps (Bösenberg &amp; Strand, 1906). M, U. esyunini (Zhang, Zhang &amp; Yu, 2003). N, Hylyphantes graminicola (Sundevall, 1830). O, Tmeticus tolli Kulczyński, 1908. Scale bars 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.6967988" httpUri="https://zenodo.org/record/6967988/files/figure.png" pageId="137" pageNumber="554">Fig. 67G</figureCitation>
). Legs: tibia chaetotaxy 2-2-1-1, dorsal proximal macroseta on tibia I 1.56 times diameter of tibia; Tm I: 0.54. All metatarsi with trichobothrium. Pedipalp: patella prolateral proximal vertical macrosetae absent; tibia with one prolateral, two retrolateral trichobothria; TPS absent; TPA short, stout, apical surface scaly; TRA absent (
<figureCitation id="13252A54FF5476A4FAA1D6CA4F5EF7BD" box="[1290,1386,658,680]" captionStart="Figure 80" captionStartId="138.[145,223,1612,1634]" captionTargetBox="[148,1425,202,1571]" captionTargetId="figure-106@138.[145,1428,195,1572]" captionTargetPageId="138" captionText="Figure 80. Oedothorax stylus Tanasevitch, 2015. AD, male right palp, images flipped horizontally. A, retrolateral view. B, prolateral view. C, dorsal view. D, ventral view. E, F, epigyne. E, ventrolateral view. F, external morphology. G, male posterior median spinnerets and posterior lateral spinnerets, dorsal view. H, female spinnerets. Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6968028" httpUri="https://zenodo.org/record/6968028/files/figure.png" pageId="137" pageNumber="554">Fig. 80C</figureCitation>
); PC median-sized, base not visible from dorsal view, distal setae close to distal clasp, distal clasp without striae, clasp directed apically (
<figureCitation id="13252A54FF5476A4FB96D6B64EACF619" box="[1085,1176,750,772]" captionStart="Figure 80" captionStartId="138.[145,223,1612,1634]" captionTargetBox="[148,1425,202,1571]" captionTargetId="figure-106@138.[145,1428,195,1572]" captionTargetPageId="138" captionText="Figure 80. Oedothorax stylus Tanasevitch, 2015. AD, male right palp, images flipped horizontally. A, retrolateral view. B, prolateral view. C, dorsal view. D, ventral view. E, F, epigyne. E, ventrolateral view. F, external morphology. G, male posterior median spinnerets and posterior lateral spinnerets, dorsal view. H, female spinnerets. Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6968028" httpUri="https://zenodo.org/record/6968028/files/figure.png" pageId="137" pageNumber="554">Fig. 80A</figureCitation>
); T without papillae; PT long, with median-sized papillae; TS absent (
<figureCitation id="13252A54FF5476A4FA9ED7554FA0F638" box="[1333,1428,781,803]" captionStart="Figure 80" captionStartId="138.[145,223,1612,1634]" captionTargetBox="[148,1425,202,1571]" captionTargetId="figure-106@138.[145,1428,195,1572]" captionTargetPageId="138" captionText="Figure 80. Oedothorax stylus Tanasevitch, 2015. AD, male right palp, images flipped horizontally. A, retrolateral view. B, prolateral view. C, dorsal view. D, ventral view. E, F, epigyne. E, ventrolateral view. F, external morphology. G, male posterior median spinnerets and posterior lateral spinnerets, dorsal view. H, female spinnerets. Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6968028" httpUri="https://zenodo.org/record/6968028/files/figure.png" pageId="137" pageNumber="554">Fig. 80D</figureCitation>
); MSA present; DSA tip slightly wavy, with a protuberance on dorsal side in front of MSA; EM flat, anterior margin with papillae, length equals ARP (
<figureCitation id="13252A54FF5476A4FB40D7314F66F664" box="[1259,1362,873,895]" captionStart="Figure 80" captionStartId="138.[145,223,1612,1634]" captionTargetBox="[148,1425,202,1571]" captionTargetId="figure-106@138.[145,1428,195,1572]" captionTargetPageId="138" captionText="Figure 80. Oedothorax stylus Tanasevitch, 2015. AD, male right palp, images flipped horizontally. A, retrolateral view. B, prolateral view. C, dorsal view. D, ventral view. E, F, epigyne. E, ventrolateral view. F, external morphology. G, male posterior median spinnerets and posterior lateral spinnerets, dorsal view. H, female spinnerets. Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6968028" httpUri="https://zenodo.org/record/6968028/files/figure.png" pageId="137" pageNumber="554">Fig. 80A</figureCitation>
); ARP pointed, weakly sclerotized; LER small, without striae, not extended dorsal to E; VRP long, straight, apically directed; TP tip pointed, narrow; E retrolaterally spiral, anterior margin at base wavy (
<figureCitation id="13252A54FF5476A4FB3CD7BC4EC1F6E3" box="[1175,1269,996,1018]" captionStart="Figure 80" captionStartId="138.[145,223,1612,1634]" captionTargetBox="[148,1425,202,1571]" captionTargetId="figure-106@138.[145,1428,195,1572]" captionTargetPageId="138" captionText="Figure 80. Oedothorax stylus Tanasevitch, 2015. AD, male right palp, images flipped horizontally. A, retrolateral view. B, prolateral view. C, dorsal view. D, ventral view. E, F, epigyne. E, ventrolateral view. F, external morphology. G, male posterior median spinnerets and posterior lateral spinnerets, dorsal view. H, female spinnerets. Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6968028" httpUri="https://zenodo.org/record/6968028/files/figure.png" pageId="137" pageNumber="554">Fig. 80B</figureCitation>
). Opisthosoma: dorsal pattern see
<figureCitation id="13252A54FF5476A4FBBDD05A4E4DF102" box="[1046,1145,1026,1048]" captionStart="Figure 68" captionStartId="111.[163,245,1308,1330]" captionTargetBox="[164,1444,196,1268]" captionTargetId="figure-26@111.[163,1446,195,1269]" captionTargetPageId="111" captionText="Figure 68. Male opisthosoma, dorsal view. A, Jilinus hulongensis (Zhu &amp; Wen, 1980). B, Cornitibia simplicithorax (Tanasevitch, 1998). C, Emertongone montifera (Emerton, 1882). D, Oedothorax paracymbialis Tanasevitch, 2015. E, Oe. kodaikanal Tanasevitch, 2015. F, Oe. cunur Tanasevitch, 2015. G, Oe. stylus Tanasevitch, 2015. H, Atypena cirrifrons (Heimer, 1984). I, A. formosana (Oi, 1977). J, Oe. nazareti Scharff, 1989. K, Gongylidium rufipes (Linnaeus, 1758). L, Ummeliata insecticeps (Bösenberg &amp; Strand, 1906). M, U. esyunini (Zhang, Zhang &amp; Yu, 2003). N, Hylyphantes graminicola (Sundevall, 1830). O, Tmeticus tolli Kulczyński, 1908. Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6967990" httpUri="https://zenodo.org/record/6967990/files/figure.png" pageId="137" pageNumber="554">Fig. 68G</figureCitation>
; PMS with mAP, two AC; PLS with triad, 3+ AC (
<figureCitation id="13252A54FF5476A4FB95D0794EAFF12C" box="[1086,1179,1057,1079]" captionStart="Figure 80" captionStartId="138.[145,223,1612,1634]" captionTargetBox="[148,1425,202,1571]" captionTargetId="figure-106@138.[145,1428,195,1572]" captionTargetPageId="138" captionText="Figure 80. Oedothorax stylus Tanasevitch, 2015. AD, male right palp, images flipped horizontally. A, retrolateral view. B, prolateral view. C, dorsal view. D, ventral view. E, F, epigyne. E, ventrolateral view. F, external morphology. G, male posterior median spinnerets and posterior lateral spinnerets, dorsal view. H, female spinnerets. Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6968028" httpUri="https://zenodo.org/record/6968028/files/figure.png" pageId="137" pageNumber="554">Fig. 80G</figureCitation>
).
</paragraph>
<paragraph id="8BA136D1FF5476A4FC90D0384946F18E" blockId="137.[827,1442,1120,1172]" pageId="137" pageNumber="554">
<emphasis id="B96AEAC3FF5476A4FC90D03849C4F16F" box="[827,1008,1120,1141]" italics="true" pageId="137" pageNumber="554">
Male (
<typeStatus id="54A58873FF5476A4FC28D03849D6F16F" box="[899,994,1120,1141]" pageId="137" pageNumber="554" type="holotype">holotype</typeStatus>
):
</emphasis>
Prosoma: 0.77 long, 0.63 wide. Tm I: 0.54.
</paragraph>
<paragraph id="8BA136D1FF5476A4FC90D0E5499BF37B" blockId="137.[827,1444,1213,1633]" pageId="137" pageNumber="554">
<emphasis id="B96AEAC3FF5476A4FC90D0E54E25F1C8" box="[827,1041,1213,1234]" italics="true" pageId="137" pageNumber="554">
Female (
<typeStatus id="54A58873FF5476A4FC36D0E54E37F1C8" box="[925,1027,1213,1234]" pageId="137" pageNumber="554" type="paratype">paratype</typeStatus>
):
</emphasis>
Total length: 1.96. Prosoma: 0.85 long, 0.66 wide. Eyes: AME-AME: 0.02, AME width: 0.05, AME-ALE: 0.02, ALE width: 0.09, ALE-PLE: 0, PLE width: 0.09, PLE-PME: 0.04, PME width: 0.08, PME-PME: 0.05. Clypeus: not hirsute, one sub- AME seta. Sternum: 0.53 long; 0.51 wide. Legs: tibia chaetotaxy 2-2-1-1, dorsal proximal macroseta on tibia I, II, III and IV 2.72, 2.82, 2.60 and 3.39 times diameter of tibia, respectively; Tm I: 0.54. All metatarsi with trichobothrium. Epigyne: Clade 13 characteristic morphology, CO anteriorly oriented (
<figureCitation id="13252A54FF5476A4FAA3D1A84FA6F31F" box="[1288,1426,1520,1542]" captionStart="Figure 80" captionStartId="138.[145,223,1612,1634]" captionTargetBox="[148,1425,202,1571]" captionTargetId="figure-106@138.[145,1428,195,1572]" captionTargetPageId="138" captionText="Figure 80. Oedothorax stylus Tanasevitch, 2015. AD, male right palp, images flipped horizontally. A, retrolateral view. B, prolateral view. C, dorsal view. D, ventral view. E, F, epigyne. E, ventrolateral view. F, external morphology. G, male posterior median spinnerets and posterior lateral spinnerets, dorsal view. H, female spinnerets. Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6968028" httpUri="https://zenodo.org/record/6968028/files/figure.png" pageId="137" pageNumber="554">Fig. 80E, F</figureCitation>
). Opisthosoma: dorsal pattern same as male; PMS with mAP, two AC, one CY; PLS with triad, two CY, 3+ AC (
<figureCitation id="13252A54FF5476A4FCE8D2134995F37B" box="[835,929,1611,1633]" captionStart="Figure 80" captionStartId="138.[145,223,1612,1634]" captionTargetBox="[148,1425,202,1571]" captionTargetId="figure-106@138.[145,1428,195,1572]" captionTargetPageId="138" captionText="Figure 80. Oedothorax stylus Tanasevitch, 2015. AD, male right palp, images flipped horizontally. A, retrolateral view. B, prolateral view. C, dorsal view. D, ventral view. E, F, epigyne. E, ventrolateral view. F, external morphology. G, male posterior median spinnerets and posterior lateral spinnerets, dorsal view. H, female spinnerets. Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6968028" httpUri="https://zenodo.org/record/6968028/files/figure.png" pageId="137" pageNumber="554">Fig. 80H</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C304655AFF5476A4FC90D2D24EC6F3A5" pageId="137" pageNumber="554" type="distribution">
<paragraph id="8BA136D1FF5476A4FC90D2D24EC6F3A5" blockId="137.[827,1442,1674,1727]" pageId="137" pageNumber="554">
<emphasis id="B96AEAC3FF5476A4FC90D2D249F9F385" box="[827,973,1674,1695]" italics="true" pageId="137" pageNumber="554">Distribution:</emphasis>
At present only known from
<collectingRegion id="49DAF833FF5476A4FA89D2D24F44F3BA" box="[1314,1392,1674,1696]" country="India" name="Kerala" pageId="137" pageNumber="554">Kerala</collectingRegion>
and Madras (currently
<collectingRegion id="49DAF833FF5476A4FBBAD2F14EAFF3A5" box="[1041,1179,1705,1727]" country="India" name="Tamil Nadu" pageId="137" pageNumber="554">Tamil Nadu</collectingRegion>
),
<collectingCountry id="F3097641FF5476A4FB05D2F14EDAF3A5" box="[1198,1262,1705,1727]" name="India" pageId="137" pageNumber="554">India</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C304655AFF5476A4FC90D2B04E00F3E4" box="[827,1076,1768,1790]" pageId="137" pageNumber="554" type="biology_ecology">
<paragraph id="8BA136D1FF5476A4FC90D2B04E00F3E4" blockId="137.[827,1076,1768,1790]" box="[827,1076,1768,1790]" pageId="137" pageNumber="554">
<emphasis id="B96AEAC3FF5476A4FC90D2B049AEF3E7" box="[827,922,1768,1789]" italics="true" pageId="137" pageNumber="554">Habitat:</emphasis>
Forest litter.
</paragraph>
</subSubSection>
<subSubSection id="C304655AFF5476A7FC90D37E4EF7F23D" lastPageId="138" lastPageNumber="555" pageId="137" pageNumber="554" type="discussion">
<paragraph id="8BA136D1FF5476A7FC90D37E4EF7F23D" blockId="137.[827,1443,1830,1883]" lastBlockId="138.[809,1425,1748,1831]" lastPageId="138" lastPageNumber="555" pageId="137" pageNumber="554">
<emphasis id="B96AEAC3FF5476A4FC90D37E4992F221" box="[827,934,1830,1851]" italics="true" pageId="137" pageNumber="554">Remarks:</emphasis>
According to the results of our phylogenetic analysis and the lack of the distinctive characteristics of
<emphasis id="B96AEAC3FF5776A7FF1DD28C4BB5F3F3" box="[182,385,1748,1769]" italics="true" pageId="138" pageNumber="555">
<taxonomicName id="4C1E4D52FF5776A7FF1DD28C4B78F3F3" authorityName="Lin &amp; Lopardo &amp; Uhl" authorityYear="2022" box="[182,332,1748,1769]" class="Arachnida" family="Linyphiidae" genus="Oedothorax" kingdom="Animalia" order="Araneae" pageId="138" pageNumber="555" phylum="Arthropoda" rank="genus" sensu="stricto">Oedothorax</taxonomicName>
<taxonomicNameLabel id="A25957B8FF5776A7FEF1D28C4BB5F3F3" box="[346,385,1748,1769]" pageId="138" pageNumber="555" sensu="stricto">s. s.</taxonomicNameLabel>
</emphasis>
(see general description of
<emphasis id="B96AEAC3FF5776A7FF3AD2AA4B77F212" box="[145,323,1778,1800]" italics="true" pageId="138" pageNumber="555">
<taxonomicName id="4C1E4D52FF5776A7FF3AD2AA4B2DF21D" authorityName="Lin &amp; Lopardo &amp; Uhl" authorityYear="2022" box="[145,281,1778,1799]" class="Arachnida" family="Linyphiidae" genus="Oedothorax" kingdom="Animalia" order="Araneae" pageId="138" pageNumber="555" phylum="Arthropoda" rank="genus" sensu="stricto">Oedothorax</taxonomicName>
<taxonomicNameLabel id="A25957B8FF5776A7FE8AD2AB4B77F212" box="[289,323,1779,1800]" pageId="138" pageNumber="555" sensu="stricto">s.s.</taxonomicNameLabel>
</emphasis>
), this species is more closely related to
<emphasis id="B96AEAC3FF5776A7FF1FD3494AEEF23C" box="[180,218,1809,1830]" italics="true" pageId="138" pageNumber="555">Oe.</emphasis>
<taxonomicName id="4C1E4D52FF5776A7FF43D34A4B19F23D" authorityName="Tanasevitch" authorityYear="2015" box="[232,301,1810,1831]" class="Arachnida" family="Linyphiidae" genus="Oedothorax" kingdom="Animalia" order="Araneae" pageId="138" pageNumber="555" phylum="Arthropoda" rank="species" species="cunur">
<emphasis id="B96AEAC3FF5776A7FF43D34A4B19F23D" box="[232,301,1810,1831]" italics="true" pageId="138" pageNumber="555">cunur</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1E4D52FF5776A7FEC1D3494BFEF23C" baseAuthorityName="Jocque" baseAuthorityYear="1983" box="[362,458,1809,1830]" class="Arachnida" family="Linyphiidae" genus="Atypena" kingdom="Animalia" order="Araneae" pageId="138" pageNumber="555" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAC3FF5776A7FEC1D3494BFEF23C" box="[362,458,1809,1830]" italics="true" pageId="138" pageNumber="555">Atypena</emphasis>
</taxonomicName>
which, as in this species, are distributed in the
<collectingRegion id="49DAF833FF5776A7FB8BD28C4EB4F3F0" box="[1056,1152,1748,1770]" country="Morocco" name="Oriental" pageId="138" pageNumber="555">Oriental</collectingRegion>
realm. We provisionally leave the taxonomic status of this species unchanged until further data become available.
</paragraph>
</subSubSection>
</treatment>
</document>