<documentid="1EE728D9B91871687615FABDB957454B"ID-DOI="10.5281/zenodo.13245398"ID-ISSN="2345-7600"ID-Zenodo-Dep="13245398"IM.bibliography_approvedBy="felipe"IM.illustrations_approvedBy="felipe"IM.materialsCitations_approvedBy="felipe"IM.metadata_approvedBy="felipe"IM.taxonomicNames_approvedBy="felipe"IM.treatmentCitations_approvedBy="felipe"IM.treatments_approvedBy="felipe"checkinTime="1722970648523"checkinUser="felipe"docAuthor="Marin, Ivan"docDate="2006"docId="03BFCE58FFEDFFBC15EBFBFF1B347F68"docLanguage="en"docName="RafflesBZool.54.2.321-340.pdf"docOrigin="Raffles Bulletin of Zoology 54 (2)"docStyle="DocumentStyle:C6443723295B2E4C6F22B9B23F08089C.5:RafflesBZool.2006-2013.journal_article"docStyleId="C6443723295B2E4C6F22B9B23F08089C"docStyleName="RafflesBZool.2006-2013.journal_article"docStyleVersion="5"docTitle="Crinotonia anastasiae Marin, 2006, new species"docType="treatment"docVersion="2"lastPageNumber="329"masterDocId="FF86B620FFECFFB11642FFBE19437D50"masterDocTitle="Description Of Crinotonia Anastasiae, New Genus, New Species, A New Crinoid Associated Pontoniine Shrimp (Crustacea: Caridea) From Nha Trang Bay, Vietnam, With Inclusion Of Periclimenes Attenuatus Bruce, 1971, In The New Genus"masterLastPageNumber="340"masterPageNumber="321"pageNumber="322"updateTime="1722976815852"updateUser="ExternalLinkService"zenodo-license-document="CC-BY-4.0">
<mods:titleid="BF80586A0C5577C08A04101143E409EA">Description Of Crinotonia Anastasiae, New Genus, New Species, A New Crinoid Associated Pontoniine Shrimp (Crustacea: Caridea) From Nha Trang Bay, Vietnam, With Inclusion Of Periclimenes Attenuatus Bruce, 1971, In The New Genus</mods:title>
<figureCitationid="132D63CBFFEDFFB01279FBDF1DD87928"box="[1083,1179,1121,1144]"captionStart-0="Fig"captionStart-1="Fig"captionStart-2="Fig"captionStart-3="Fig"captionStart-4="Fig"captionStart-5="Fig"captionStartId-0="2.[136,171,2092,2113]"captionStartId-1="3.[113,148,2063,2084]"captionStartId-2="4.[136,171,2065,2086]"captionStartId-3="5.[113,148,2035,2056]"captionStartId-4="6.[136,171,2093,2114]"captionStartId-5="7.[113,148,2035,2056]"captionTargetBox-0="[147,1460,858,2034]"captionTargetBox-2="[142,1473,780,2004]"captionTargetBox-3="[107,1450,319,1902]"captionTargetBox-4="[136,1466,1220,1931]"captionTargetBox-5="[115,1452,701,1938]"captionTargetId-0="figure-170@2.[115,1537,817,2067]"captionTargetId-2="figure-39@4.[97,1524,731,2060]"captionTargetId-3="figure-86@5.[60,1483,282,1949]"captionTargetId-4="figure-446@6.[76,1524,1189,1964]"captionTargetId-5="figure-68@7.[77,1489,647,2010]"captionTargetPageId-0="2"captionTargetPageId-2="4"captionTargetPageId-3="5"captionTargetPageId-4="6"captionTargetPageId-5="7"captionText-0="Fig. 1. Crinotonia anastasiae, new genus, new species: a, holotype female; b, allotype male. Scale bar = 5 mm."captionText-1="Fig. 2. Crinotonia anastasiae, new genus, new species, holotype (a, c) and allotype (b, d-h): a, b, carapace; c, d, front of carapace, dorsal; e, f, antennules; g, disto-lateral part of basal segment of antennules; h, antenna. Scale bar: a-d = 2 mm; e-h = 4.8 mm; g = no scale bar."captionText-2="Fig. 3. Mouthparts of Crinotonia anastasiae, new genus, new species, allotype: a, mandibule; b, maxillula; c, maxilla; d, first maxilliped; e, second maxilliped; f, third maxilliped. Scale bar = 1 mm."captionText-3="Fig. 4. Crinotonia anastasiae, new genus, new species, allotype: a, first pereiopod; b, c, chela of first pereiopod; d, major second pereiopod; e, minor second pereiopod; f, chela of minor second pereiopod; g, third pereiopod; h, same, ventral margin of propodus; i, dactylus of third pereiopod; j, same, distal part. Scale bar: a, e, g = 1 mm; b, c, f, i = 0.5 mm; d = 2 mm; j =no scale bar."captionText-4="Fig. 5. Crinotonia anastasiae, new genus, new species, allotype: a-d, dactylus of major second pereiopod. Scale bar = 2 mm."captionText-5="Fig. 6. Crinotonia anastasiae, new genus, new species, holotype (e), allotype (a-d, f): a, distal margin of sixth segment and telson, lateral view; b, telson, dorsal; c, uropods; d, e, distal part of telson; f, appendix interna of second male pleopods. Scale bar: a-c = 2 mm; d, e = 0.7 mm; f = 0.3 mm."figureDoi-0="http://doi.org/10.5281/zenodo.13245400"figureDoi-2="http://doi.org/10.5281/zenodo.13245402"figureDoi-3="http://doi.org/10.5281/zenodo.13245404"figureDoi-4="http://doi.org/10.5281/zenodo.13245406"figureDoi-5="http://doi.org/10.5281/zenodo.13245408"httpUri-0="https://zenodo.org/record/13245400/files/figure.png"httpUri-2="https://zenodo.org/record/13245402/files/figure.png"httpUri-3="https://zenodo.org/record/13245404/files/figure.png"httpUri-4="https://zenodo.org/record/13245406/files/figure.png"httpUri-5="https://zenodo.org/record/13245408/files/figure.png"pageId="1"pageNumber="322">Figs. 1-6</figureCitation>
female. Carapace swollen, smooth, with antennal and hepatic spines. Hepatic spine is distinctly smaller than antennal and directed horizontally. Rostrum short, straight, tapering and slightly compressed, with developed dorsal carina bearing four conspicuous teeth, without ventral teeth and ventral carina (
<figureCitationid="132D63CBFFEDFFB012A5F9941C107B11"box="[1255,1363,1578,1601]"captionStart="Fig"captionStartId="3.[113,148,2063,2084]"captionText="Fig. 2. Crinotonia anastasiae, new genus, new species, holotype (a, c) and allotype (b, d-h): a, b, carapace; c, d, front of carapace, dorsal; e, f, antennules; g, disto-lateral part of basal segment of antennules; h, antenna. Scale bar: a-d = 2 mm; e-h = 4.8 mm; g = no scale bar."pageId="1"pageNumber="322">Fig. 2a, b</figureCitation>
); lateral lamina feebly developed, continuous with orbital margin posteriorly (
<figureCitationid="132D63CBFFEDFFB015EBF9D41D4F7BD1"box="[937,1036,1642,1665]"captionStart="Fig"captionStartId="3.[113,148,2063,2084]"captionText="Fig. 2. Crinotonia anastasiae, new genus, new species, holotype (a, c) and allotype (b, d-h): a, b, carapace; c, d, front of carapace, dorsal; e, f, antennules; g, disto-lateral part of basal segment of antennules; h, antenna. Scale bar: a-d = 2 mm; e-h = 4.8 mm; g = no scale bar."pageId="1"pageNumber="322">Fig. 2c, d</figureCitation>
Abdominal somites smooth; pleura of first to fifth abdominal somites rounded, postero-lateral angle of sixth abdominal somite is broad and pointed, postero-ventral angle projecting and rounded (
<figureCitationid="132D63CBFFEDFFB015FDF8F41D647A31"box="[959,1063,1866,1889]"captionStart="Fig"captionStartId="7.[113,148,2035,2056]"captionTargetBox="[115,1452,701,1938]"captionTargetId="figure-68@7.[77,1489,647,2010]"captionTargetPageId="7"captionText="Fig. 6. Crinotonia anastasiae, new genus, new species, holotype (e), allotype (a-d, f): a, distal margin of sixth segment and telson, lateral view; b, telson, dorsal; c, uropods; d, e, distal part of telson; f, appendix interna of second male pleopods. Scale bar: a-c = 2 mm; d, e = 0.7 mm; f = 0.3 mm."figureDoi="http://doi.org/10.5281/zenodo.13245408"httpUri="https://zenodo.org/record/13245408/files/figure.png"pageId="1"pageNumber="322">Fig. 6a, b</figureCitation>
<figureCitationid="132D63CBFFEDFFB015CCF8D41A9D7AD1"box="[910,990,1898,1921]"captionStart="Fig"captionStartId="7.[113,148,2035,2056]"captionTargetBox="[115,1452,701,1938]"captionTargetId="figure-68@7.[77,1489,647,2010]"captionTargetPageId="7"captionText="Fig. 6. Crinotonia anastasiae, new genus, new species, holotype (e), allotype (a-d, f): a, distal margin of sixth segment and telson, lateral view; b, telson, dorsal; c, uropods; d, e, distal part of telson; f, appendix interna of second male pleopods. Scale bar: a-c = 2 mm; d, e = 0.7 mm; f = 0.3 mm."figureDoi="http://doi.org/10.5281/zenodo.13245408"httpUri="https://zenodo.org/record/13245408/files/figure.png"pageId="1"pageNumber="322">Fig. 6b</figureCitation>
), converge gradually posteriorly, with two pairs of conspicuous dorsal submarginal spines situated at 0.4 and 0.7 of telson length; three pairs of posterior spines contain short lateral, long and slender intermediate and slender submediate spines which are about half of intermediate spine length (
<figureCitationid="132D63CBFFEDFFB0153DF7B41A8C7571"box="[895,975,2058,2081]"captionStart="Fig"captionStartId="7.[113,148,2035,2056]"captionTargetBox="[115,1452,701,1938]"captionTargetId="figure-68@7.[77,1489,647,2010]"captionTargetPageId="7"captionText="Fig. 6. Crinotonia anastasiae, new genus, new species, holotype (e), allotype (a-d, f): a, distal margin of sixth segment and telson, lateral view; b, telson, dorsal; c, uropods; d, e, distal part of telson; f, appendix interna of second male pleopods. Scale bar: a-c = 2 mm; d, e = 0.7 mm; f = 0.3 mm."figureDoi="http://doi.org/10.5281/zenodo.13245408"httpUri="https://zenodo.org/record/13245408/files/figure.png"pageId="1"pageNumber="322">Fig. 6e</figureCitation>
<paragraphid="8BA97F4EFFEEFFB316CAFF7F1B0F7DA8"blockId="2.[136,775,193,248]"pageId="2"pageNumber="323">Eyes large and well developed, with slightly projecting cornea. Central cornea with small papilla.</paragraph>
<figureCitationid="132D63CBFFEEFFB3174EFE9F18267C68"box="[268,357,289,312]"captionStart="Fig"captionStartId="3.[113,148,2063,2084]"captionText="Fig. 2. Crinotonia anastasiae, new genus, new species, holotype (a, c) and allotype (b, d-h): a, b, carapace; c, d, front of carapace, dorsal; e, f, antennules; g, disto-lateral part of basal segment of antennules; h, antenna. Scale bar: a-d = 2 mm; e-h = 4.8 mm; g = no scale bar."pageId="2"pageNumber="323">Fig. 2e,f</figureCitation>
) well developed; basal segment is about two times longer than wide, with lateral border bearing an acute distolateral tooth, distal medial lobe with acute medial tooth, small setae is presented between teeth; distinct robust ventral tooth is situated on the middle of medial border of basal segment. The stylocerite is well developed. The intermediate segment is slender, about 1.5 times as long as wide, with disto-lateral inner border produced distally. The distal segment is about two times as long as wide. The upper flagellum with rami fused for the two proximal segments, the shorter ramus consists of four segments with five groups of aesthetascs, and the longer free ramus consists of more than twenty segments. The lower flagellum is filiform with more than twenty segments.
<figureCitationid="132D63CBFFEEFFB315E9FF7F1AB47D88"box="[939,1015,193,216]"captionStart="Fig"captionStartId="3.[113,148,2063,2084]"captionText="Fig. 2. Crinotonia anastasiae, new genus, new species, holotype (a, c) and allotype (b, d-h): a, b, carapace; c, d, front of carapace, dorsal; e, f, antennules; g, disto-lateral part of basal segment of antennules; h, antenna. Scale bar: a-d = 2 mm; e-h = 4.8 mm; g = no scale bar."pageId="2"pageNumber="323">Fig. 2h</figureCitation>
) well developed stylocerite, basicerite with small disto-ventral tooth; carpocerite is slender, about four times as long as wide; flagellum is well developed, with the length equal to body length; scaphocerite is slender and exceeding the distal margin and intermediate antennular segment, about 3 times as long as maximum width, with well developed distolateral tooth.
<figureCitationid="132D63CBFFEEFFB31325FE7F1CFA7C88"box="[1383,1465,449,472]"captionStart="Fig"captionStartId="4.[136,171,2065,2086]"captionTargetBox="[142,1473,780,2004]"captionTargetId="figure-39@4.[97,1524,731,2060]"captionTargetPageId="4"captionText="Fig. 3. Mouthparts of Crinotonia anastasiae, new genus, new species, allotype: a, mandibule; b, maxillula; c, maxilla; d, first maxilliped; e, second maxilliped; f, third maxilliped. Scale bar = 1 mm."figureDoi="http://doi.org/10.5281/zenodo.13245402"httpUri="https://zenodo.org/record/13245402/files/figure.png"pageId="2"pageNumber="323">Fig. 3a</figureCitation>
<figureCitationid="132D63CBFFEEFFB315F9FDDF1D4A7F28"box="[955,1033,609,632]"captionStart="Fig"captionStartId="4.[136,171,2065,2086]"captionTargetBox="[142,1473,780,2004]"captionTargetId="figure-39@4.[97,1524,731,2060]"captionTargetPageId="4"captionText="Fig. 3. Mouthparts of Crinotonia anastasiae, new genus, new species, allotype: a, mandibule; b, maxillula; c, maxilla; d, first maxilliped; e, second maxilliped; f, third maxilliped. Scale bar = 1 mm."figureDoi="http://doi.org/10.5281/zenodo.13245402"httpUri="https://zenodo.org/record/13245402/files/figure.png"pageId="2"pageNumber="323">Fig. 3b</figureCitation>
) is normal, with well developed bilobed palp; the upper lacinia is slightly curved, flaring distally, with strong setae distally; the lower lacinia is slender, tapering distally.
<taxonomicNameid="4C1604CDFFEEFFB3168CF79218EB7511"authorityName="Marin"authorityYear="2006"box="[206,424,2092,2113]"class="Malacostraca"family="Palaemonidae"genus="Crinotonia"higherTaxonomySource="CoL,GBIF"kingdom="Animalia"order="Decapoda"pageId="2"pageNumber="323"phylum="Arthropoda"rank="species"species="anastasiae"status="new genus, new species">Crinotonia anastasiae</taxonomicName>
,
</emphasis>
<taxonomicNameLabelid="A2511E27FFEEFFB317F0F7921BDC7511"box="[434,671,2092,2113]"pageId="2"pageNumber="323"rank="species">new genus, new species</taxonomicNameLabel>
: a, holotype female; b, allotype male. Scale bar = 5 mm.
<taxonomicNameid="4C1604CDFFEFFFB216F5F7B118D37574"authorityName="Marin"authorityYear="2006"box="[183,400,2063,2084]"class="Malacostraca"family="Palaemonidae"genus="Crinotonia"higherTaxonomySource="CoL,GBIF"kingdom="Animalia"order="Decapoda"pageId="3"pageNumber="324"phylum="Arthropoda"rank="species"species="anastasiae"status="new genus, new species">Crinotonia anastasiae</taxonomicName>
,
</emphasis>
<taxonomicNameLabelid="A2511E27FFEFFFB217D8F7B11BC67574"box="[410,645,2063,2084]"pageId="3"pageNumber="324"rank="species">new genus, new species</taxonomicNameLabel>
, holotype (a, c) and allotype (b, d-h): a, b, carapace; c, d, front of carapace, dorsal; e, f, antennules; g, disto-lateral part of basal segment of antennules; h, antenna. Scale bar: a-d = 2 mm; e-h = 4.8 mm; g = no scale bar.
<figureCitationid="132D63CBFFE8FFB516A9FF7F187B7D88"box="[235,312,193,216]"captionStart="Fig"captionStartId="4.[136,171,2065,2086]"captionTargetBox="[142,1473,780,2004]"captionTargetId="figure-39@4.[97,1524,731,2060]"captionTargetPageId="4"captionText="Fig. 3. Mouthparts of Crinotonia anastasiae, new genus, new species, allotype: a, mandibule; b, maxillula; c, maxilla; d, first maxilliped; e, second maxilliped; f, third maxilliped. Scale bar = 1 mm."figureDoi="http://doi.org/10.5281/zenodo.13245402"httpUri="https://zenodo.org/record/13245402/files/figure.png"pageId="4"pageNumber="325">Fig. 3c</figureCitation>
) with well developed simple palp; endites are well developed, basal endite feebly bilobed, distal lobes subequal, furnished with stiff, elongated setae; scaphognathite well developed, with plumose setae.
<figureCitationid="132D63CBFFE8FFB51706FEDF18D07C28"box="[324,403,353,376]"captionStart="Fig"captionStartId="4.[136,171,2065,2086]"captionTargetBox="[142,1473,780,2004]"captionTargetId="figure-39@4.[97,1524,731,2060]"captionTargetPageId="4"captionText="Fig. 3. Mouthparts of Crinotonia anastasiae, new genus, new species, allotype: a, mandibule; b, maxillula; c, maxilla; d, first maxilliped; e, second maxilliped; f, third maxilliped. Scale bar = 1 mm."figureDoi="http://doi.org/10.5281/zenodo.13245402"httpUri="https://zenodo.org/record/13245402/files/figure.png"pageId="4"pageNumber="325">Fig. 3d</figureCitation>
) with distinct and well developed endites, basal endite is significantly larger, with setae along lateral margin; exopod well developed with large caridean lobe bearing plumose setae; epipod is triangular in shape, well developed.
<figureCitationid="132D63CBFFE8FFB51727FD9F18F77F68"box="[357,436,545,568]"captionStart="Fig"captionStartId="4.[136,171,2065,2086]"captionTargetBox="[142,1473,780,2004]"captionTargetId="figure-39@4.[97,1524,731,2060]"captionTargetPageId="4"captionText="Fig. 3. Mouthparts of Crinotonia anastasiae, new genus, new species, allotype: a, mandibule; b, maxillula; c, maxilla; d, first maxilliped; e, second maxilliped; f, third maxilliped. Scale bar = 1 mm."figureDoi="http://doi.org/10.5281/zenodo.13245402"httpUri="https://zenodo.org/record/13245402/files/figure.png"pageId="4"pageNumber="325">Fig. 3e</figureCitation>
) with exopod well developed; distolateral margin of propodus broadly rounded, with slender setae; dactylus about three times longer than broad, with numerous spines along distal margin; epipod subrectangular; podobranch absent.
<figureCitationid="132D63CBFFE8FFB51249FF7F1D157D88"box="[1035,1110,193,216]"captionStart="Fig"captionStartId="4.[136,171,2065,2086]"captionTargetBox="[142,1473,780,2004]"captionTargetId="figure-39@4.[97,1524,731,2060]"captionTargetPageId="4"captionText="Fig. 3. Mouthparts of Crinotonia anastasiae, new genus, new species, allotype: a, mandibule; b, maxillula; c, maxilla; d, first maxilliped; e, second maxilliped; f, third maxilliped. Scale bar = 1 mm."figureDoi="http://doi.org/10.5281/zenodo.13245402"httpUri="https://zenodo.org/record/13245402/files/figure.png"pageId="4"pageNumber="325">Fig. 3f</figureCitation>
) with slender segments and well developed exopod exceeding the length of ischiomerus; ischiomerus is about 3.5 times as long as maximum width, tapering significantly distally, the penultimate segment is slender, about five times as long as wide; the terminal segment is also slender, about seven times as long as wide, with sparse long setae along the border; epipod oval; arthrobranch absent.
<figureCitationid="132D63CBFFE8FFB515ACFE7F1D7B7C88"box="[1006,1080,449,472]"captionStart="Fig"captionStartId="5.[113,148,2035,2056]"captionTargetBox="[107,1450,319,1902]"captionTargetId="figure-86@5.[60,1483,282,1949]"captionTargetPageId="5"captionText="Fig. 4. Crinotonia anastasiae, new genus, new species, allotype: a, first pereiopod; b, c, chela of first pereiopod; d, major second pereiopod; e, minor second pereiopod; f, chela of minor second pereiopod; g, third pereiopod; h, same, ventral margin of propodus; i, dactylus of third pereiopod; j, same, distal part. Scale bar: a, e, g = 1 mm; b, c, f, i = 0.5 mm; d = 2 mm; j =no scale bar."figureDoi="http://doi.org/10.5281/zenodo.13245404"httpUri="https://zenodo.org/record/13245404/files/figure.png"pageId="4"pageNumber="325">Fig. 4a</figureCitation>
) slender, with robust chela and broad, swollen fingers; carpus is slender, significantly flaring distally, palm is equal in width and length, slightly compressed laterally; fixed finger is robust with numerous terminal setae, with acute tip and feebly developed cutting edge; dactylus with acute tip and, subcylindrical, without conspicuous cutting edge (
<figureCitationid="132D63CBFFE8FFB51221FD3F1D847FC8"box="[1123,1223,641,664]"captionStart="Fig"captionStartId="5.[113,148,2035,2056]"captionTargetBox="[107,1450,319,1902]"captionTargetId="figure-86@5.[60,1483,282,1949]"captionTargetPageId="5"captionText="Fig. 4. Crinotonia anastasiae, new genus, new species, allotype: a, first pereiopod; b, c, chela of first pereiopod; d, major second pereiopod; e, minor second pereiopod; f, chela of minor second pereiopod; g, third pereiopod; h, same, ventral margin of propodus; i, dactylus of third pereiopod; j, same, distal part. Scale bar: a, e, g = 1 mm; b, c, f, i = 0.5 mm; d = 2 mm; j =no scale bar."figureDoi="http://doi.org/10.5281/zenodo.13245404"httpUri="https://zenodo.org/record/13245404/files/figure.png"pageId="4"pageNumber="325">Fig. 4b,c</figureCitation>
<taxonomicNameid="4C1604CDFFE8FFB5171DF7AF1B7B7576"authorityName="Marin"authorityYear="2006"box="[351,568,2065,2086]"class="Malacostraca"family="Palaemonidae"genus="Crinotonia"higherTaxonomySource="CoL,GBIF"kingdom="Animalia"order="Decapoda"pageId="4"pageNumber="325"phylum="Arthropoda"rank="species"species="anastasiae"status="new genus, new species">Crinotonia anastasiae</taxonomicName>
,
</emphasis>
<taxonomicNameLabelid="A2511E27FFE8FFB51401F7AF1A6D7576"box="[579,814,2065,2086]"pageId="4"pageNumber="325"rank="species">new genus, new species</taxonomicNameLabel>
, allotype: a, mandibule; b, maxillula; c, maxilla; d, first maxilliped; e, second maxilliped; f, third maxilliped. Scale bar = 1 mm.
<taxonomicNameid="4C1604CDFFE9FFB416F6F84A18C87559"authorityName="Marin"authorityYear="2006"box="[180,395,2036,2057]"class="Malacostraca"family="Palaemonidae"genus="Crinotonia"higherTaxonomySource="CoL,GBIF"kingdom="Animalia"order="Decapoda"pageId="5"pageNumber="326"phylum="Arthropoda"rank="species"species="anastasiae"status="new genus, new species">Crinotonia anastasiae</taxonomicName>
,
</emphasis>
<taxonomicNameLabelid="A2511E27FFE9FFB417D7F84D1B387558"box="[405,635,2035,2056]"pageId="5"pageNumber="326"rank="species">new genus, new species</taxonomicNameLabel>
, allotype: a, first pereiopod; b, c, chela of first pereiopod; d, major second pereiopod; e, minor second pereiopod; f, chela of minor second pereiopod; g, third pereiopod; h, same, ventral margin of propodus; i, dactylus of third pereiopod; j, same, distal part. Scale bar: a, e, g = 1 mm; b, c, f, i = 0.5 mm; d = 2 mm; j =no scale bar.
<figureCitationid="132D63CBFFEAFFB71740FF5F18127DA8"box="[258,337,225,248]"captionStart="Fig"captionStartId="5.[113,148,2035,2056]"captionTargetBox="[107,1450,319,1902]"captionTargetId="figure-86@5.[60,1483,282,1949]"captionTargetPageId="5"captionText="Fig. 4. Crinotonia anastasiae, new genus, new species, allotype: a, first pereiopod; b, c, chela of first pereiopod; d, major second pereiopod; e, minor second pereiopod; f, chela of minor second pereiopod; g, third pereiopod; h, same, ventral margin of propodus; i, dactylus of third pereiopod; j, same, distal part. Scale bar: a, e, g = 1 mm; b, c, f, i = 0.5 mm; d = 2 mm; j =no scale bar."figureDoi="http://doi.org/10.5281/zenodo.13245404"httpUri="https://zenodo.org/record/13245404/files/figure.png"pageId="6"pageNumber="327">Fig. 4d</figureCitation>
) with robust segments; ischium is about two times as long as wide; merus is about 3.4 times as long as maximal width, with medial part swollen centrally; carpus is flared distally, as long as maximal width, palm is subcylindrical, about 2.5 times as long as maximal width; fingers with acute, curved tips; fixed fingers robust, with two teeth opposing into deep pits on the dactylus, with distinct cutting edge on distal third of the length; dactylus is robust, about 4.5 times as long as wide, with distinct dorso-lateral flange along the proximal two-third of the length, ventral margin with distinct protruding cutting edge in distal third of the length, with two mesial deep pits and one tooth in proximal third of the length (
<figureCitationid="132D63CBFFEAFFB71718FDDF18D87F28"box="[346,411,609,632]"captionStart="Fig"captionStartId="5.[113,148,2035,2056]"captionTargetBox="[107,1450,319,1902]"captionTargetId="figure-86@5.[60,1483,282,1949]"captionTargetPageId="5"captionText="Fig. 4. Crinotonia anastasiae, new genus, new species, allotype: a, first pereiopod; b, c, chela of first pereiopod; d, major second pereiopod; e, minor second pereiopod; f, chela of minor second pereiopod; g, third pereiopod; h, same, ventral margin of propodus; i, dactylus of third pereiopod; j, same, distal part. Scale bar: a, e, g = 1 mm; b, c, f, i = 0.5 mm; d = 2 mm; j =no scale bar."figureDoi="http://doi.org/10.5281/zenodo.13245404"httpUri="https://zenodo.org/record/13245404/files/figure.png"pageId="6"pageNumber="327">Fig. 4</figureCitation>
<figureCitationid="132D63CBFFEAFFB714DAFDDF1BA67F28"box="[664,741,609,632]"captionStart="Fig"captionStartId="5.[113,148,2035,2056]"captionTargetBox="[107,1450,319,1902]"captionTargetId="figure-86@5.[60,1483,282,1949]"captionTargetPageId="5"captionText="Fig. 4. Crinotonia anastasiae, new genus, new species, allotype: a, first pereiopod; b, c, chela of first pereiopod; d, major second pereiopod; e, minor second pereiopod; f, chela of minor second pereiopod; g, third pereiopod; h, same, ventral margin of propodus; i, dactylus of third pereiopod; j, same, distal part. Scale bar: a, e, g = 1 mm; b, c, f, i = 0.5 mm; d = 2 mm; j =no scale bar."figureDoi="http://doi.org/10.5281/zenodo.13245404"httpUri="https://zenodo.org/record/13245404/files/figure.png"pageId="6"pageNumber="327">Fig. 4e</figureCitation>
) is extremely slender, ischium is about eight times as long as wide; merus is about thirteen times as long as width; carpus is flared distally, about fifteen times as long as maximal width, palm is subcylindrical, about five times as long as width, slightly tapering distally, fingers are slender, subcylindrical, equal to palm length, about 2.5 times as long as proximal width, with pointed, curved tips (
<figureCitationid="132D63CBFFEAFFB717B2FCFF1B7D7E08"box="[496,574,833,856]"captionStart="Fig"captionStartId="5.[113,148,2035,2056]"captionTargetBox="[107,1450,319,1902]"captionTargetId="figure-86@5.[60,1483,282,1949]"captionTargetPageId="5"captionText="Fig. 4. Crinotonia anastasiae, new genus, new species, allotype: a, first pereiopod; b, c, chela of first pereiopod; d, major second pereiopod; e, minor second pereiopod; f, chela of minor second pereiopod; g, third pereiopod; h, same, ventral margin of propodus; i, dactylus of third pereiopod; j, same, distal part. Scale bar: a, e, g = 1 mm; b, c, f, i = 0.5 mm; d = 2 mm; j =no scale bar."figureDoi="http://doi.org/10.5281/zenodo.13245404"httpUri="https://zenodo.org/record/13245404/files/figure.png"pageId="6"pageNumber="327">Fig. 4f</figureCitation>
<figureCitationid="132D63CBFFEAFFB71707FC3F18D77EC8"box="[325,404,897,920]"captionStart="Fig"captionStartId="5.[113,148,2035,2056]"captionTargetBox="[107,1450,319,1902]"captionTargetId="figure-86@5.[60,1483,282,1949]"captionTargetPageId="5"captionText="Fig. 4. Crinotonia anastasiae, new genus, new species, allotype: a, first pereiopod; b, c, chela of first pereiopod; d, major second pereiopod; e, minor second pereiopod; f, chela of minor second pereiopod; g, third pereiopod; h, same, ventral margin of propodus; i, dactylus of third pereiopod; j, same, distal part. Scale bar: a, e, g = 1 mm; b, c, f, i = 0.5 mm; d = 2 mm; j =no scale bar."figureDoi="http://doi.org/10.5281/zenodo.13245404"httpUri="https://zenodo.org/record/13245404/files/figure.png"pageId="6"pageNumber="327">
) slender, with unarmed segments; ischium is about 3.5 times as long as wide; merus is about seven times as long as wide; carpus is about 3.5 times as long as maximum wide, tapering proximally; propodus is about 10.5 times as long as wide, with serrated ventral margin (
<figureCitationid="132D63CBFFEAFFB71509FF7F1AD97D88"box="[843,922,193,216]"captionStart="Fig"captionStartId="5.[113,148,2035,2056]"captionTargetBox="[107,1450,319,1902]"captionTargetId="figure-86@5.[60,1483,282,1949]"captionTargetPageId="5"captionText="Fig. 4. Crinotonia anastasiae, new genus, new species, allotype: a, first pereiopod; b, c, chela of first pereiopod; d, major second pereiopod; e, minor second pereiopod; f, chela of minor second pereiopod; g, third pereiopod; h, same, ventral margin of propodus; i, dactylus of third pereiopod; j, same, distal part. Scale bar: a, e, g = 1 mm; b, c, f, i = 0.5 mm; d = 2 mm; j =no scale bar."figureDoi="http://doi.org/10.5281/zenodo.13245404"httpUri="https://zenodo.org/record/13245404/files/figure.png"pageId="6"pageNumber="327">Fig. 4h</figureCitation>
<figureCitationid="132D63CBFFEAFFB71257FF7F1D187D88"box="[1045,1115,193,216]"captionStart="Fig"captionStartId="5.[113,148,2035,2056]"captionTargetBox="[107,1450,319,1902]"captionTargetId="figure-86@5.[60,1483,282,1949]"captionTargetPageId="5"captionText="Fig. 4. Crinotonia anastasiae, new genus, new species, allotype: a, first pereiopod; b, c, chela of first pereiopod; d, major second pereiopod; e, minor second pereiopod; f, chela of minor second pereiopod; g, third pereiopod; h, same, ventral margin of propodus; i, dactylus of third pereiopod; j, same, distal part. Scale bar: a, e, g = 1 mm; b, c, f, i = 0.5 mm; d = 2 mm; j =no scale bar."figureDoi="http://doi.org/10.5281/zenodo.13245404"httpUri="https://zenodo.org/record/13245404/files/figure.png"pageId="6"pageNumber="327">Fig. 4i</figureCitation>
) slender, about 3 times as long as maximum width, significantly tapering distally, biunguiculate, with pointed slender curved unguis and preterminal triangular accessory tooth directed distally (
<figureCitationid="132D63CBFFEAFFB713D5FE9F1A187C08"captionStart="Fig"captionStartId="5.[113,148,2035,2056]"captionTargetBox="[107,1450,319,1902]"captionTargetId="figure-86@5.[60,1483,282,1949]"captionTargetPageId="5"captionText="Fig. 4. Crinotonia anastasiae, new genus, new species, allotype: a, first pereiopod; b, c, chela of first pereiopod; d, major second pereiopod; e, minor second pereiopod; f, chela of minor second pereiopod; g, third pereiopod; h, same, ventral margin of propodus; i, dactylus of third pereiopod; j, same, distal part. Scale bar: a, e, g = 1 mm; b, c, f, i = 0.5 mm; d = 2 mm; j =no scale bar."figureDoi="http://doi.org/10.5281/zenodo.13245404"httpUri="https://zenodo.org/record/13245404/files/figure.png"pageId="6"pageNumber="327">Fig. 4j</figureCitation>
<figureCitationid="132D63CBFFEAFFB715AEFE7F1D767C88"box="[1004,1077,449,472]"captionStart="Fig"captionStartId="7.[113,148,2035,2056]"captionTargetBox="[115,1452,701,1938]"captionTargetId="figure-68@7.[77,1489,647,2010]"captionTargetPageId="7"captionText="Fig. 6. Crinotonia anastasiae, new genus, new species, holotype (e), allotype (a-d, f): a, distal margin of sixth segment and telson, lateral view; b, telson, dorsal; c, uropods; d, e, distal part of telson; f, appendix interna of second male pleopods. Scale bar: a-c = 2 mm; d, e = 0.7 mm; f = 0.3 mm."figureDoi="http://doi.org/10.5281/zenodo.13245408"httpUri="https://zenodo.org/record/13245408/files/figure.png"pageId="6"pageNumber="327">Fig. 6f</figureCitation>
<figureCitationid="132D63CBFFEAFFB712EEFE7F1DB77C88"box="[1196,1268,449,472]"captionStart="Fig"captionStartId="7.[113,148,2035,2056]"captionTargetBox="[115,1452,701,1938]"captionTargetId="figure-68@7.[77,1489,647,2010]"captionTargetPageId="7"captionText="Fig. 6. Crinotonia anastasiae, new genus, new species, holotype (e), allotype (a-d, f): a, distal margin of sixth segment and telson, lateral view; b, telson, dorsal; c, uropods; d, e, distal part of telson; f, appendix interna of second male pleopods. Scale bar: a-c = 2 mm; d, e = 0.7 mm; f = 0.3 mm."figureDoi="http://doi.org/10.5281/zenodo.13245408"httpUri="https://zenodo.org/record/13245408/files/figure.png"pageId="6"pageNumber="327">Fig. 6c</figureCitation>
<figureCitationid="132D63CBFFEAFFB715F7FD3F1D627FC8"box="[949,1057,641,664]"captionStart="Fig"captionStartId="7.[113,148,2035,2056]"captionTargetBox="[115,1452,701,1938]"captionTargetId="figure-68@7.[77,1489,647,2010]"captionTargetPageId="7"captionText="Fig. 6. Crinotonia anastasiae, new genus, new species, holotype (e), allotype (a-d, f): a, distal margin of sixth segment and telson, lateral view; b, telson, dorsal; c, uropods; d, e, distal part of telson; f, appendix interna of second male pleopods. Scale bar: a-c = 2 mm; d, e = 0.7 mm; f = 0.3 mm."figureDoi="http://doi.org/10.5281/zenodo.13245408"httpUri="https://zenodo.org/record/13245408/files/figure.png"pageId="6"pageNumber="327">Figs. 6f, g</figureCitation>
<figureCitationid="132D63CBFFEAFFB71235FD1F1D877FE8"box="[1143,1220,673,696]"captionStart="Fig"captionStartId="7.[113,148,2035,2056]"captionTargetBox="[115,1452,701,1938]"captionTargetId="figure-68@7.[77,1489,647,2010]"captionTargetPageId="7"captionText="Fig. 6. Crinotonia anastasiae, new genus, new species, holotype (e), allotype (a-d, f): a, distal margin of sixth segment and telson, lateral view; b, telson, dorsal; c, uropods; d, e, distal part of telson; f, appendix interna of second male pleopods. Scale bar: a-c = 2 mm; d, e = 0.7 mm; f = 0.3 mm."figureDoi="http://doi.org/10.5281/zenodo.13245408"httpUri="https://zenodo.org/record/13245408/files/figure.png"pageId="6"pageNumber="327">Fig. 6h</figureCitation>
<taxonomicNameid="4C1604CDFFEAFFB7168CF79318EB7512"authorityName="Marin"authorityYear="2006"box="[206,424,2093,2114]"class="Malacostraca"family="Palaemonidae"genus="Crinotonia"higherTaxonomySource="CoL,GBIF"kingdom="Animalia"order="Decapoda"pageId="6"pageNumber="327"phylum="Arthropoda"rank="species"species="anastasiae"status="new genus, new species">Crinotonia anastasiae</taxonomicName>
,
</emphasis>
<taxonomicNameLabelid="A2511E27FFEAFFB717F0F7931BDD7512"box="[434,670,2093,2114]"pageId="6"pageNumber="327"rank="species">new genus, new species</taxonomicNameLabel>
, allotype: a-d, dactylus of major second pereiopod. Scale bar = 2 mm.
but clearly differs in having a more robust body and appendages (segments of antennules and antenna (scaphocerite), second pereiopods, especially the palm of the major cheliped, and the walking legs), the more developed rostral carina, especially in the female, the form of distal part of dactylus of third pereiopod and the presence of several denticles along its ventral border.
<emphasisid="B962A35CFFEBFFB61633FE5F18E97CA9"bold="true"box="[113,426,481,505]"italics="true"pageId="7"pageNumber="328">Host and associated fauna.</emphasis>
<bibRefCitationid="EF8702BFFFEBFFB61278FF7E1D847D88"author="Bruce, A"box="[1082,1223,192,216]"pageId="7"pageNumber="328"pagination="91 - 99"refId="ref5961"refString="Bruce, A. J., 1971. Periclimenes attenuatus sp. nov. a new commensal shrimp from Duke of York Islands. Pacific Science, 25: 91 - 99."type="journal article"year="1971">Bruce, 1971</bibRefCitation>
<figureCitationid="132D63CBFFEBFFB61276FF5F1DE07DA8"box="[1076,1187,225,248]"captionStart-0="Fig"captionStart-1="Fig"captionStart-2="Fig"captionStart-3="Fig"captionStartId-0="8.[136,171,2091,2112]"captionStartId-1="9.[113,148,2034,2055]"captionStartId-2="10.[136,171,2006,2027]"captionStartId-3="11.[113,148,887,908]"captionTargetBox-0="[140,1470,1419,2040]"captionTargetBox-3="[125,1436,196,834]"captionTargetId-0="figure-8@8.[100,1522,1382,2068]"captionTargetId-3="figure-28@11.[84,1494,163,868]"captionTargetPageId-0="8"captionTargetPageId-3="11"captionText-0="Fig. 7. Crinotonia attenuatus (Bruce), new combination: a, ovigerous female (PCL 3.8 mm); b, male (PCL 2.3 mm). Scale bar = 5 mm."captionText-1="Fig. 8. Crinotonia attenuatus (Bruce), new combination: a-d, carapace; e, f, front of carapace, dorsal; g, h, antennule; i, j, disto-lateral part of basal segment of antennule (left and right); k, antenna. a, male (PCL 3.2 mm); b, g, k, female (PCL 3.8 mm); c, e, male (PCL 3.2 mm); d, f, i, female (PCL 4.2 mm). Scale bar: a-f = 2 mm; g, h = 4.3 mm; k = 2.4 mm; i, j = no scale bar."captionText-2="Fig. 9. Crinotonia attenuatus (Bruce), new combination, male (PCL 3.2 mm): a, first pereiopod; b, major second pereiopod; c, fingers of major second pereiopod; d, minor second pereiopod; e, chela of minor second pereiopod; f, third pereiopod; g, dactylus of third pereiopod; h, same, distal part; i, distal part of sixth abdominal segment; j, telson. Scale bar: a, b = 1.1 mm; d, f = 1 mm; e = 0.7 mm; c, i, j = 0.7 mm; g = 0.4 mm; h = 0.2 mm."captionText-3="Fig. 10. Crinotonia attenuatus (Bruce), new combination, male (PCL 3.2 mm). Scale bar = 5 mm."figureDoi-0="http://doi.org/10.5281/zenodo.13245410"figureDoi-3="http://doi.org/10.5281/zenodo.13245412"httpUri-0="https://zenodo.org/record/13245410/files/figure.png"httpUri-3="https://zenodo.org/record/13245412/files/figure.png"pageId="7"pageNumber="328">Figs. 7-10</figureCitation>
<bibRefCitationid="EF8702BFFFEBFFB6125BFE9F1D827C66"author="Bruce, A"box="[1049,1217,289,310]"pageId="7"pageNumber="328"pagination="91 - 99"refId="ref5961"refString="Bruce, A. J., 1971. Periclimenes attenuatus sp. nov. a new commensal shrimp from Duke of York Islands. Pacific Science, 25: 91 - 99."type="journal article"year="1971">Bruce, 1971: 533</bibRefCitation>
<figureCitationid="132D63CBFFEBFFB61288FE9F1C637C66"box="[1226,1312,289,310]"captionStart-0="Fig"captionStart-1="Fig"captionStart-2="Fig"captionStart-3="Fig"captionStart-4="Fig"captionStartId-0="2.[136,171,2092,2113]"captionStartId-1="3.[113,148,2063,2084]"captionStartId-2="4.[136,171,2065,2086]"captionStartId-3="5.[113,148,2035,2056]"captionStartId-4="6.[136,171,2093,2114]"captionTargetBox-0="[147,1460,858,2034]"captionTargetBox-2="[142,1473,780,2004]"captionTargetBox-3="[107,1450,319,1902]"captionTargetBox-4="[136,1466,1220,1931]"captionTargetId-0="figure-170@2.[115,1537,817,2067]"captionTargetId-2="figure-39@4.[97,1524,731,2060]"captionTargetId-3="figure-86@5.[60,1483,282,1949]"captionTargetId-4="figure-446@6.[76,1524,1189,1964]"captionTargetPageId-0="2"captionTargetPageId-2="4"captionTargetPageId-3="5"captionTargetPageId-4="6"captionText-0="Fig. 1. Crinotonia anastasiae, new genus, new species: a, holotype female; b, allotype male. Scale bar = 5 mm."captionText-1="Fig. 2. Crinotonia anastasiae, new genus, new species, holotype (a, c) and allotype (b, d-h): a, b, carapace; c, d, front of carapace, dorsal; e, f, antennules; g, disto-lateral part of basal segment of antennules; h, antenna. Scale bar: a-d = 2 mm; e-h = 4.8 mm; g = no scale bar."captionText-2="Fig. 3. Mouthparts of Crinotonia anastasiae, new genus, new species, allotype: a, mandibule; b, maxillula; c, maxilla; d, first maxilliped; e, second maxilliped; f, third maxilliped. Scale bar = 1 mm."captionText-3="Fig. 4. Crinotonia anastasiae, new genus, new species, allotype: a, first pereiopod; b, c, chela of first pereiopod; d, major second pereiopod; e, minor second pereiopod; f, chela of minor second pereiopod; g, third pereiopod; h, same, ventral margin of propodus; i, dactylus of third pereiopod; j, same, distal part. Scale bar: a, e, g = 1 mm; b, c, f, i = 0.5 mm; d = 2 mm; j =no scale bar."captionText-4="Fig. 5. Crinotonia anastasiae, new genus, new species, allotype: a-d, dactylus of major second pereiopod. Scale bar = 2 mm."figureDoi-0="http://doi.org/10.5281/zenodo.13245400"figureDoi-2="http://doi.org/10.5281/zenodo.13245402"figureDoi-3="http://doi.org/10.5281/zenodo.13245404"figureDoi-4="http://doi.org/10.5281/zenodo.13245406"httpUri-0="https://zenodo.org/record/13245400/files/figure.png"httpUri-2="https://zenodo.org/record/13245402/files/figure.png"httpUri-3="https://zenodo.org/record/13245404/files/figure.png"httpUri-4="https://zenodo.org/record/13245406/files/figure.png"pageId="7"pageNumber="328">Figs. 1-5</figureCitation>
<bibRefCitationid="EF8702BFFFEBFFB6158CFEC91D037CDC"author="Bruce, A"box="[974,1088,375,396]"pageId="7"pageNumber="328"pagination="195 - 218"refId="ref6153"refString="Bruce, A. J., 1983 b. The pontoniine fauna of Australia. Australian Museum Memoirs, 18: 195 - 218."type="journal article"year="1983">1983b: 206</bibRefCitation>
<bibRefCitationid="EF8702BFFFEBFFB615F6FE2A1DC37CF9"author="De Grave, S"box="[948,1152,404,425]"pageId="7"pageNumber="328"pagination="119 - 148"refId="ref7284"refString="De Grave, S., 2000. Caridean shrimps (Crustacea, Decapoda) from Hansa Bay, Papua New Guinea: Palaemonidae and Gnathophyllidae. Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, Biologie, 70: 119 - 148."type="journal article"year="2000">De Grave, 2000: 134</bibRefCitation>
<bibRefCitationid="EF8702BFFFEBFFB612C8FE2A1C4D7CF9"author="Li, X"box="[1162,1294,404,425]"pageId="7"pageNumber="328"pagination="1 - 407"refId="ref8024"refString="Li, X., 2000. Catalog of the Genera and Species of Pontoniinae Kingsley, 1878. Xueyuan Press, Beijing. Pp. i-iv, 1 - 319, Figs. 1 - 407."type="book chapter"year="2000">Li, 2000: 159</bibRefCitation>
<figureCitationid="132D63CBFFEBFFB61535FE0F1AF97C96"box="[887,954,433,454]"captionStart="Fig"captionStartId="2.[136,171,2092,2113]"captionTargetBox="[147,1460,858,2034]"captionTargetId="figure-170@2.[115,1537,817,2067]"captionTargetPageId="2"captionText="Fig. 1. Crinotonia anastasiae, new genus, new species: a, holotype female; b, allotype male. Scale bar = 5 mm."figureDoi="http://doi.org/10.5281/zenodo.13245400"httpUri="https://zenodo.org/record/13245400/files/figure.png"pageId="7"pageNumber="328">Figs. 1</figureCitation>
<figureCitationid="132D63CBFFEBFFB61585FE0F1A967C96"box="[967,981,433,454]"captionStart="Fig"captionStartId="3.[113,148,2063,2084]"captionText="Fig. 2. Crinotonia anastasiae, new genus, new species, holotype (a, c) and allotype (b, d-h): a, b, carapace; c, d, front of carapace, dorsal; e, f, antennules; g, disto-lateral part of basal segment of antennules; h, antenna. Scale bar: a-d = 2 mm; e-h = 4.8 mm; g = no scale bar."pageId="7"pageNumber="328">2</figureCitation>
<taxonomicNameid="4C1604CDFFEBFFB616F5F84A18D37559"authorityName="Marin"authorityYear="2006"box="[183,400,2036,2057]"class="Malacostraca"family="Palaemonidae"genus="Crinotonia"higherTaxonomySource="CoL,GBIF"kingdom="Animalia"order="Decapoda"pageId="7"pageNumber="328"phylum="Arthropoda"rank="species"species="anastasiae"status="new genus, new species">Crinotonia anastasiae</taxonomicName>
,
</emphasis>
<taxonomicNameLabelid="A2511E27FFEBFFB617D9F84D1BC57558"box="[411,646,2035,2056]"pageId="7"pageNumber="328"rank="species">new genus, new species</taxonomicNameLabel>
, holotype (e), allotype (a-d, f): a, distal margin of sixth segment and telson, lateral view; b, telson, dorsal; c, uropods; d, e, distal part of telson; f, appendix interna of second male pleopods. Scale bar: a-c = 2 mm; d, e = 0.7 mm; f = 0.3 mm.
Body is slender. Carapace smooth, swollen, with antennal and small hepatic spines. Rostrum (
<figureCitationid="132D63CBFFE4FFB914DEFECA1B997CDB"box="[668,730,372,395]"captionStart="Fig"captionStartId="9.[113,148,2034,2055]"captionText="Fig. 8. Crinotonia attenuatus (Bruce), new combination: a-d, carapace; e, f, front of carapace, dorsal; g, h, antennule; i, j, disto-lateral part of basal segment of antennule (left and right); k, antenna. a, male (PCL 3.2 mm); b, g, k, female (PCL 3.8 mm); c, e, male (PCL 3.2 mm); d, f, i, female (PCL 4.2 mm). Scale bar: a-f = 2 mm; g, h = 4.3 mm; k = 2.4 mm; i, j = no scale bar."pageId="8"pageNumber="329">Fig. 8</figureCitation>
a-d) short, tapering, slightly upturned, with three-four dorsal teeth only, dorsal lamina is feebly developed, ventral teeth absent, lateral lamina feebly developed. Pterygostomial margin bluntly angular. Abdominal somites smooth; pleura of first to fifth abdominal somites rounded, postero-lateral angle of sixth abdominal somite pointed and postero-ventral angle projecting and rounded (
<figureCitationid="132D63CBFFE4FFB917C9FDEA18907F3B"box="[395,467,596,619]"captionStart="Fig"captionStartId="10.[136,171,2006,2027]"captionText="Fig. 9. Crinotonia attenuatus (Bruce), new combination, male (PCL 3.2 mm): a, first pereiopod; b, major second pereiopod; c, fingers of major second pereiopod; d, minor second pereiopod; e, chela of minor second pereiopod; f, third pereiopod; g, dactylus of third pereiopod; h, same, distal part; i, distal part of sixth abdominal segment; j, telson. Scale bar: a, b = 1.1 mm; d, f = 1 mm; e = 0.7 mm; c, i, j = 0.7 mm; g = 0.4 mm; h = 0.2 mm."pageId="8"pageNumber="329">Fig. 9i</figureCitation>
). Telson about three times as long as wide, with two pairs of dorsal and three pairs of posterior spines (
<figureCitationid="132D63CBFFE4FFB91706FD2A18D37FFB"box="[324,400,660,683]"captionStart="Fig"captionStartId="10.[136,171,2006,2027]"captionText="Fig. 9. Crinotonia attenuatus (Bruce), new combination, male (PCL 3.2 mm): a, first pereiopod; b, major second pereiopod; c, fingers of major second pereiopod; d, minor second pereiopod; e, chela of minor second pereiopod; f, third pereiopod; g, dactylus of third pereiopod; h, same, distal part; i, distal part of sixth abdominal segment; j, telson. Scale bar: a, b = 1.1 mm; d, f = 1 mm; e = 0.7 mm; c, i, j = 0.7 mm; g = 0.4 mm; h = 0.2 mm."pageId="8"pageNumber="329">Fig. 9j</figureCitation>
). Eyes slightly projecting distally, with corneal papillae. Antennules with large disto-lateral tooth and small teeth on distal medial lobe on proximal segment, robust ventral tooth in middle of medial border of basal segment (
<figureCitationid="132D63CBFFE4FFB916B3FCAA18157E7B"box="[241,342,788,811]"captionStart="Fig"captionStartId="9.[113,148,2034,2055]"captionText="Fig. 8. Crinotonia attenuatus (Bruce), new combination: a-d, carapace; e, f, front of carapace, dorsal; g, h, antennule; i, j, disto-lateral part of basal segment of antennule (left and right); k, antenna. a, male (PCL 3.2 mm); b, g, k, female (PCL 3.8 mm); c, e, male (PCL 3.2 mm); d, f, i, female (PCL 4.2 mm). Scale bar: a-f = 2 mm; g, h = 4.3 mm; k = 2.4 mm; i, j = no scale bar."pageId="8"pageNumber="329">Fig. 8h,g</figureCitation>
). Antenna (
<figureCitationid="132D63CBFFE4FFB91795FCAA1B657E7B"box="[471,550,788,811]"captionStart="Fig"captionStartId="9.[113,148,2034,2055]"captionText="Fig. 8. Crinotonia attenuatus (Bruce), new combination: a-d, carapace; e, f, front of carapace, dorsal; g, h, antennule; i, j, disto-lateral part of basal segment of antennule (left and right); k, antenna. a, male (PCL 3.2 mm); b, g, k, female (PCL 3.8 mm); c, e, male (PCL 3.2 mm); d, f, i, female (PCL 4.2 mm). Scale bar: a-f = 2 mm; g, h = 4.3 mm; k = 2.4 mm; i, j = no scale bar."pageId="8"pageNumber="329">Fig. 8k</figureCitation>
) with slender, about 4.5 times as long as maximum width. Second maxilliped without podobranch. First pereiopod (
<figureCitationid="132D63CBFFE4FFB9145DFCEA1B2E7E3B"box="[543,621,852,875]"captionStart="Fig"captionStartId="10.[136,171,2006,2027]"captionText="Fig. 9. Crinotonia attenuatus (Bruce), new combination, male (PCL 3.2 mm): a, first pereiopod; b, major second pereiopod; c, fingers of major second pereiopod; d, minor second pereiopod; e, chela of minor second pereiopod; f, third pereiopod; g, dactylus of third pereiopod; h, same, distal part; i, distal part of sixth abdominal segment; j, telson. Scale bar: a, b = 1.1 mm; d, f = 1 mm; e = 0.7 mm; c, i, j = 0.7 mm; g = 0.4 mm; h = 0.2 mm."pageId="8"pageNumber="329">Fig. 9a</figureCitation>
) slender, with robust chela and broad, swollen fingers bearing numerous setae. Major second pereiopod (
<figureCitationid="132D63CBFFE4FFB91446FC2A1B187EFB"box="[516,603,916,939]"captionStart="Fig"captionStartId="10.[136,171,2006,2027]"captionText="Fig. 9. Crinotonia attenuatus (Bruce), new combination, male (PCL 3.2 mm): a, first pereiopod; b, major second pereiopod; c, fingers of major second pereiopod; d, minor second pereiopod; e, chela of minor second pereiopod; f, third pereiopod; g, dactylus of third pereiopod; h, same, distal part; i, distal part of sixth abdominal segment; j, telson. Scale bar: a, b = 1.1 mm; d, f = 1 mm; e = 0.7 mm; c, i, j = 0.7 mm; g = 0.4 mm; h = 0.2 mm."pageId="8"pageNumber="329">Fig. 9b</figureCitation>
) with slender, subcylindrical palm about 4 times as long as wide; minor pereiopod (
<figureCitationid="132D63CBFFE4FFB916BDFC6A18087EBB"box="[255,331,980,1003]"captionStart="Fig"captionStartId="10.[136,171,2006,2027]"captionText="Fig. 9. Crinotonia attenuatus (Bruce), new combination, male (PCL 3.2 mm): a, first pereiopod; b, major second pereiopod; c, fingers of major second pereiopod; d, minor second pereiopod; e, chela of minor second pereiopod; f, third pereiopod; g, dactylus of third pereiopod; h, same, distal part; i, distal part of sixth abdominal segment; j, telson. Scale bar: a, b = 1.1 mm; d, f = 1 mm; e = 0.7 mm; c, i, j = 0.7 mm; g = 0.4 mm; h = 0.2 mm."pageId="8"pageNumber="329">Fig. 9d</figureCitation>
) is extremely slender, with simple slender fingers. Third pereiopod (
<figureCitationid="132D63CBFFE4FFB917E3FC4A18AE795B"box="[417,493,1012,1035]"captionStart="Fig"captionStartId="10.[136,171,2006,2027]"captionText="Fig. 9. Crinotonia attenuatus (Bruce), new combination, male (PCL 3.2 mm): a, first pereiopod; b, major second pereiopod; c, fingers of major second pereiopod; d, minor second pereiopod; e, chela of minor second pereiopod; f, third pereiopod; g, dactylus of third pereiopod; h, same, distal part; i, distal part of sixth abdominal segment; j, telson. Scale bar: a, b = 1.1 mm; d, f = 1 mm; e = 0.7 mm; c, i, j = 0.7 mm; g = 0.4 mm; h = 0.2 mm."pageId="8"pageNumber="329">Fig. 9f</figureCitation>
) with unarmed segments; ischium is about 5.5 times as long as wide; merus is about thirteen times as long as wide; carpus is about seven times as long as maximum wide, tapering proximally; propodus is about 11.5 times as long as wide, with serrated ventral margin; dactylus (
<figureCitationid="132D63CBFFE4FFB916B0FB2A180079FB"box="[242,323,1172,1195]"captionStart="Fig"captionStartId="10.[136,171,2006,2027]"captionText="Fig. 9. Crinotonia attenuatus (Bruce), new combination, male (PCL 3.2 mm): a, first pereiopod; b, major second pereiopod; c, fingers of major second pereiopod; d, minor second pereiopod; e, chela of minor second pereiopod; f, third pereiopod; g, dactylus of third pereiopod; h, same, distal part; i, distal part of sixth abdominal segment; j, telson. Scale bar: a, b = 1.1 mm; d, f = 1 mm; e = 0.7 mm; c, i, j = 0.7 mm; g = 0.4 mm; h = 0.2 mm."pageId="8"pageNumber="329">
) is slender, about three times as long as maximal width, significantly tapering distally, preterminal accessory tooth is triangular and situated perpendicularly to ventral margin (
<figureCitationid="132D63CBFFE4FFB91777FB4A18C4785B"box="[309,391,1268,1291]"captionStart="Fig"captionStartId="10.[136,171,2006,2027]"captionText="Fig. 9. Crinotonia attenuatus (Bruce), new combination, male (PCL 3.2 mm): a, first pereiopod; b, major second pereiopod; c, fingers of major second pereiopod; d, minor second pereiopod; e, chela of minor second pereiopod; f, third pereiopod; g, dactylus of third pereiopod; h, same, distal part; i, distal part of sixth abdominal segment; j, telson. Scale bar: a, b = 1.1 mm; d, f = 1 mm; e = 0.7 mm; c, i, j = 0.7 mm; g = 0.4 mm; h = 0.2 mm."pageId="8"pageNumber="329">Fig. 9h</figureCitation>
<bibRefCitationid="EF8702BFFFE4FFB9132DFF7E1ACF7DA7"author="Bruce, A"pageId="8"pageNumber="329"pagination="91 - 99"refId="ref5961"refString="Bruce, A. J., 1971. Periclimenes attenuatus sp. nov. a new commensal shrimp from Duke of York Islands. Pacific Science, 25: 91 - 99."type="journal article"year="1971">Bruce’s (1971)</bibRefCitation>
original description. However, Li (2004) showed podobranch in material from
. The rostral formula in Vietnamese material is 3-4/0 and similar to Bruce’s material. Li (2004) reported the presence of tiny sub-apical ventral tooth on rostrum and a “podobranch” on the second maxilliped and absence of epipod on the first maxilliped. During the material dissection I also received the same situation when epipod of first maxilliped was dissected together with second maxilliped attaching to it anteriorly. It looks like “podobranch” of second maxilliped while first maxilliped missing its epipod.
) shows the presence of a large disto-lateral tooth and three (large and two small) mesial teeth on the disto-medial border of basal segment of the antennules (
<figureCitationid="132D63CBFFE4FFB9128FFD061C557F9F"box="[1229,1302,696,719]"captionStart="Fig"captionStartId="9.[113,148,2034,2055]"captionText="Fig. 8. Crinotonia attenuatus (Bruce), new combination: a-d, carapace; e, f, front of carapace, dorsal; g, h, antennule; i, j, disto-lateral part of basal segment of antennule (left and right); k, antenna. a, male (PCL 3.2 mm); b, g, k, female (PCL 3.8 mm); c, e, male (PCL 3.2 mm); d, f, i, female (PCL 4.2 mm). Scale bar: a-f = 2 mm; g, h = 4.3 mm; k = 2.4 mm; i, j = no scale bar."pageId="8"pageNumber="329">Fig. 8i</figureCitation>
from the Mediterranean Sea includes a well developed rostrum bearing robust dorsal and ventral teeth, second maxilliped with podobranch, small arthrobranch on third maxilliped, slender first pereiopod with simple fingers; similar second pereiopods with chelae not borne in vertical plane, movable finger not ventral, finger without socket and plunger closure, third pereiopod with ventral margin of propodus smooth, with disto-ventral spines, dactylus distinctly biunguiculate (
<bibRefCitationid="EF8702BFFFE1FFBC163BFEDF18777C28"author="Holthuis, L"box="[121,308,353,376]"pageId="13"pageNumber="334"pagination="1 - 254"refId="ref7881"refString="Holthuis, L. B., 1952. The Decapoda of the Siboga Expedition, Part XI: The Palaemonidae collected by the Siboga and Snellius Expeditions, with remarks on other species, part II: Subfamily Pontoniinae. Siboga-Expeditie, 39 a (10): 1 - 254, 110 Figs."type="journal article"year="1952">Holthuis, 1952</bibRefCitation>
