<documentid="1BE712D0D8CF4D2C067D82DBA940AAF5"ID-DOI="10.11646/zootaxa.5155.1.4"ID-ISSN="1175-5326"ID-Zenodo-Dep="6669176"ID-ZooBank="6E9C5E63-48B6-4D29-B98D-3DA2755A5AB8"IM.bibliography_approvedBy="juliana"IM.materialsCitations_approvedBy="juliana"IM.metadata_approvedBy="juliana"IM.taxonomicNames_approvedBy="felipe"IM.treatments_approvedBy="juliana"checkinTime="1655713213397"checkinUser="plazi"docAuthor="Golovatch, Sergei I., Bouzan, Rodrigo S., Gallo, Jéssica S. & Bichuette, Maria E."docDate="2022"docId="03AA0A59BB4AF170FF487677FAECF822"docLanguage="en"docName="zootaxa.5155.1.4.pdf"docOrigin="Zootaxa 5155 (1)"docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article"docStyleId="647186512141C8FC8976D5BCC54AEB7D"docStyleName="Zootaxa.2013-.journal_article"docStyleVersion="9"docTitle="Rotundotergum elevatum Golovatch, Bouzan & Gallo 2022, sp. nov."docType="treatment"docVersion="5"lastPageNumber="101"masterDocId="FF937221BB41F17EFFDF7046FFAAFFCF"masterDocTitle="A new genus and two new species of the millipede family Chelodesmidae from Bahia state, northeastern Brazil, including a likely troglobiont (Diplopoda, Polydesmida)"masterLastPageNumber="104"masterPageNumber="87"pageNumber="98"updateTime="1731957614320"updateUser="juliana"zenodo-license-document="CLOSED">
<mods:titleid="AA4B2F5A76A04AA83A8B15FE95D4C29E">A new genus and two new species of the millipede family Chelodesmidae from Bahia state, northeastern Brazil, including a likely troglobiont (Diplopoda, Polydesmida)</mods:title>
<mods:affiliationid="2177619E1B7E0D25A1CEB9F27BC094DE">Institute for Problems of Ecology and Evolution, Russian Academy of Sciences, Moscow, Russia.</mods:affiliation>
<mods:affiliationid="DC912671CA342C5ED473D7E758880A4F">Laboratório de Coleções Zoológicas, Instituto Butantan, Avenida Vital Brasil, 1500, 05503 - 090, São Paulo, Brasil</mods:affiliation>
<mods:affiliationid="219FE0AAC395ABC813F971D64C797EA7">Laboratório de Estudos Subterrâneos, Universidade Federal de São Carlos (UFSCar), Rodovia Washington Luis, km 235, 13565 - 905, CP 676. São Carlos, São Paulo, Brasil</mods:affiliation>
<mods:affiliationid="75BEFDFE5C27D81F57C644C960CC0B09">Laboratório de Estudos Subterrâneos, Universidade Federal de São Carlos (UFSCar), Rodovia Washington Luis, km 235, 13565 - 905, CP 676. São Carlos, São Paulo, Brasil</mods:affiliation>
<taxonomicNameid="4C03C0CCBB4AF175FF487677FC87F984"ID-CoL="CHG3D"authority="Golovatch, Bouzan & Gallo"authorityName="Golovatch, Bouzan & Gallo"authorityYear="2022"box="[151,813,1585,1612]"class="Diplopoda"family="Chelodesmidae"genus="Rotundotergum"kingdom="Animalia"order="Polydesmida"pageId="11"pageNumber="98"phylum="Arthropoda"rank="species"species="elevatum"status="sp. nov.">
<figureCitationid="1338A7CABB4AF175FF487610FEADF9BF"box="[151,263,1622,1648]"captionStart-0="FIGURE 8"captionStart-1="FIGURE 9"captionStart-2="FIGURE 10"captionStartId-0="12.[151,250,1740,1764]"captionStartId-1="13.[151,250,1928,1952]"captionStartId-2="14.[151,250,1753,1777]"captionTargetBox-0="[160,1420,496,1717]"captionTargetBox-1="[151,1436,921,1904]"captionTargetBox-2="[151,1436,489,1729]"captionTargetId-0="figure-164@12.[151,1436,489,1717]"captionTargetId-1="figure-361@13.[151,1436,921,1904]"captionTargetId-2="figure-201@14.[151,1436,489,1729]"captionTargetPageId-0="12"captionTargetPageId-1="13"captionTargetPageId-2="14"captionText-0="FIGURE 8. Rotundotergum elevatum sp. nov., male paratype. A-C. Habitus, dorsal, lateral and ventral views, respectively. Photographs by K.V. Makarov, taken not to scale."captionText-1="FIGURE 9. Rotundotergum elevatum sp. nov., male paratype. A. Incisura lateralis (I) beneath antenna, sublateral view. B. Body ring 7 with gonopods removed to show gonopod aperture, ventral view. C & D. Body ring 7 with intact gonopods, ventral and lateral views, respectively. Abbreviation: I, Incisura lateralis. Photographs by K.V. Makarov, taken not to scale."captionText-2="FIGURE 10. Rotundotergum elevatum sp. nov., male paratype. A & B. Both gonopods, anterior and posterior views, respectively. C-F. Right gonopod, mesal, lateral, ventrocaudal and caudal views, respectively. Abbreviations: A, acropodite; c, cannula; Cx, gonocoxa; AP, acropodital process; lo, lateral lobe; Pf, prefemoral region; PfP, prefemoral process; sl, solenomere; st, sternal rudiment; Te, telopodite. Photographs by K.V. Makarov, taken not to scale."figureDoi-0="http://doi.org/10.5281/zenodo.6669211"figureDoi-1="http://doi.org/10.5281/zenodo.6669213"figureDoi-2="http://doi.org/10.5281/zenodo.6669215"httpUri-0="https://zenodo.org/record/6669211/files/figure.png"httpUri-1="https://zenodo.org/record/6669213/files/figure.png"httpUri-2="https://zenodo.org/record/6669215/files/figure.png"pageId="11"pageNumber="98">Figs 8–10</figureCitation>
<materialsCitationid="3B6BB112BB4AF175FE5776D8FD86F8CF"collectingDate="2011-11-02"collectionCode="R"collectorName="M. E. Bichuette & J. E. Gallao & C. S. Fernandes & D. R. Pedroso"country="Brazil"elevation="580"latitude="-12.374333"location="Gruta Sumidouro do Joao Baio cave"longLatPrecision="1"longitude="-44.89214"municipality="Sao Desiderio Municipality"pageId="11"pageNumber="98"specimenCode="LES0027945"specimenCount="1"specimenCount-male="1"stateProvince="Bahia State"typeStatus="holotype">
<specimenCodeid="DBA51334BB4AF175FDE176D8FD72F976"box="[574,728,1694,1721]"collectionCode="LES"country="United Kingdom"httpUri="http://biocol.org/urn:lsid:biocol.org:col:13487"lsid="urn:lsid:biocol.org:col:13487"name="Leeds Museums and Galleries"pageId="11"pageNumber="98"type="Museum">LES0027945</specimenCode>
<locationid="8EDCED94BB4AF175FACE76D8FE1DF913"LSID="urn:lsid:plazi:treatment:03AA0A59BB4AF170FF487677FAECF822:8EDCED94BB4AF175FACE76D8FE1DF913"country="Brazil"latitude="-12.374333"longLatPrecision="1"longitude="-44.89214"municipality="Sao Desiderio Municipality"name="Gruta Sumidouro do Joao Baio cave"pageId="11"pageNumber="98"stateProvince="Bahia State">Gruta Sumidouro do João Baio cave</location>
<specimenCodeid="DBA51334BB4AF175FDA2774CFCBCF8EA"box="[637,790,1802,1829]"collectionCode="LES"country="United Kingdom"httpUri="http://biocol.org/urn:lsid:biocol.org:col:13487"lsid="urn:lsid:biocol.org:col:13487"name="Leeds Museums and Galleries"pageId="11"pageNumber="98"type="Museum">LES0027946</specimenCode>
), adults with lighter to yellowish antennae, legs, lateral calluses on paraterga, and tip of epiproct (
<figureCitationid="1338A7CABB4DF172FE3570FDFD9BFF1A"box="[490,561,187,213]"captionStart="FIGURE 8"captionStartId="12.[151,250,1740,1764]"captionTargetBox="[160,1420,496,1717]"captionTargetId="figure-164@12.[151,1436,489,1717]"captionTargetPageId="12"captionText="FIGURE 8. Rotundotergum elevatum sp. nov., male paratype. A-C. Habitus, dorsal, lateral and ventral views, respectively. Photographs by K.V. Makarov, taken not to scale."figureDoi="http://doi.org/10.5281/zenodo.6669211"httpUri="https://zenodo.org/record/6669211/files/figure.png"pageId="12"pageNumber="99">Figs 8</figureCitation>
,
<figureCitationid="1338A7CABB4DF172FDE170FDFD2EFF1A"box="[574,644,187,213]"captionStart="FIGURE 9"captionStartId="13.[151,250,1928,1952]"captionTargetBox="[151,1436,921,1904]"captionTargetId="figure-361@13.[151,1436,921,1904]"captionTargetPageId="13"captionText="FIGURE 9. Rotundotergum elevatum sp. nov., male paratype. A. Incisura lateralis (I) beneath antenna, sublateral view. B. Body ring 7 with gonopods removed to show gonopod aperture, ventral view. C & D. Body ring 7 with intact gonopods, ventral and lateral views, respectively. Abbreviation: I, Incisura lateralis. Photographs by K.V. Makarov, taken not to scale."figureDoi="http://doi.org/10.5281/zenodo.6669213"httpUri="https://zenodo.org/record/6669213/files/figure.png"pageId="12"pageNumber="99">9A, B</figureCitation>
<figureCitationid="1338A7CABB4DF172FD507099FD7DFF36"box="[655,727,223,249]"captionStart="FIGURE 8"captionStartId="12.[151,250,1740,1764]"captionTargetBox="[160,1420,496,1717]"captionTargetId="figure-164@12.[151,1436,489,1717]"captionTargetPageId="12"captionText="FIGURE 8. Rotundotergum elevatum sp. nov., male paratype. A-C. Habitus, dorsal, lateral and ventral views, respectively. Photographs by K.V. Makarov, taken not to scale."figureDoi="http://doi.org/10.5281/zenodo.6669211"httpUri="https://zenodo.org/record/6669211/files/figure.png"pageId="12"pageNumber="99">Fig. 8</figureCitation>
). Tegument mainly poorly shining, texture mostly leathery to finely striolate. Head poorly setose over clypeolabral region, vertex bare, epicranial suture evident (
<figureCitationid="1338A7CABB4DF172FB237145FAFFFED2"box="[1276,1365,259,285]"captionStart="FIGURE 9"captionStartId="13.[151,250,1928,1952]"captionTargetBox="[151,1436,921,1904]"captionTargetId="figure-361@13.[151,1436,921,1904]"captionTargetPageId="13"captionText="FIGURE 9. Rotundotergum elevatum sp. nov., male paratype. A. Incisura lateralis (I) beneath antenna, sublateral view. B. Body ring 7 with gonopods removed to show gonopod aperture, ventral view. C & D. Body ring 7 with intact gonopods, ventral and lateral views, respectively. Abbreviation: I, Incisura lateralis. Photographs by K.V. Makarov, taken not to scale."figureDoi="http://doi.org/10.5281/zenodo.6669213"httpUri="https://zenodo.org/record/6669213/files/figure.png"pageId="12"pageNumber="99">Fig. 9C</figureCitation>
); isthmus between antennae about as wide as diameter of antennal socket (
<figureCitationid="1338A7CABB4DF172FC4A7161FC46FE8E"box="[917,1004,295,321]"captionStart="FIGURE 8"captionStartId="12.[151,250,1740,1764]"captionTargetBox="[160,1420,496,1717]"captionTargetId="figure-164@12.[151,1436,489,1717]"captionTargetPageId="12"captionText="FIGURE 8. Rotundotergum elevatum sp. nov., male paratype. A-C. Habitus, dorsal, lateral and ventral views, respectively. Photographs by K.V. Makarov, taken not to scale."figureDoi="http://doi.org/10.5281/zenodo.6669211"httpUri="https://zenodo.org/record/6669211/files/figure.png"pageId="12"pageNumber="99">Fig. 8C</figureCitation>
). Antennae relatively long and slender, poorly clavate due to only a slightly enlarged antennomere 6,
projecting past ring 3 dorsally (male) or 2 (subadult female); in length, antennomeres 2 = 5 = 6> 3 = 4> 1> 7 (
<figureCitationid="1338A7CABB4DF172FC787129FC41FE46"box="[935,1003,367,393]"captionStart="FIGURE 8"captionStartId="12.[151,250,1740,1764]"captionTargetBox="[160,1420,496,1717]"captionTargetId="figure-164@12.[151,1436,489,1717]"captionTargetPageId="12"captionText="FIGURE 8. Rotundotergum elevatum sp. nov., male paratype. A-C. Habitus, dorsal, lateral and ventral views, respectively. Photographs by K.V. Makarov, taken not to scale."figureDoi="http://doi.org/10.5281/zenodo.6669211"httpUri="https://zenodo.org/record/6669211/files/figure.png"pageId="12"pageNumber="99">Fig. 8</figureCitation>
). Genae subquadrate laterally (
<figureCitationid="1338A7CABB4DF172FA967129FA27FE46"box="[1353,1421,367,393]"captionStart="FIGURE 8"captionStartId="12.[151,250,1740,1764]"captionTargetBox="[160,1420,496,1717]"captionTargetId="figure-164@12.[151,1436,489,1717]"captionTargetPageId="12"captionText="FIGURE 8. Rotundotergum elevatum sp. nov., male paratype. A-C. Habitus, dorsal, lateral and ventral views, respectively. Photographs by K.V. Makarov, taken not to scale."figureDoi="http://doi.org/10.5281/zenodo.6669211"httpUri="https://zenodo.org/record/6669211/files/figure.png"pageId="12"pageNumber="99">Fig. 8</figureCitation>
), incisura lateralis small and round, located beneath antenna (
<figureCitationid="1338A7CABB4DF172FCF071D5FC0EFE62"box="[815,932,403,429]"captionStart="FIGURE 9"captionStartId="13.[151,250,1928,1952]"captionTargetBox="[151,1436,921,1904]"captionTargetId="figure-361@13.[151,1436,921,1904]"captionTargetPageId="13"captionText="FIGURE 9. Rotundotergum elevatum sp. nov., male paratype. A. Incisura lateralis (I) beneath antenna, sublateral view. B. Body ring 7 with gonopods removed to show gonopod aperture, ventral view. C & D. Body ring 7 with intact gonopods, ventral and lateral views, respectively. Abbreviation: I, Incisura lateralis. Photographs by K.V. Makarov, taken not to scale."figureDoi="http://doi.org/10.5281/zenodo.6669213"httpUri="https://zenodo.org/record/6669213/files/figure.png"pageId="12"pageNumber="99">
<taxonomicNameid="4C03C0CCBB4DF172FEC1768AFDB4F92B"authority="Golovatch & Bouzan & Gallo & Bichuette, 2022"authorityName="Golovatch & Bouzan & Gallo & Bichuette"authorityYear="2022"box="[286,542,1740,1764]"class="Diplopoda"family="Chelodesmidae"genus="Rotundotergum"kingdom="Animalia"order="Polydesmida"pageId="12"pageNumber="99"phylum="Arthropoda"rank="species"species="elevatum"status="sp. nov.">
In width, head <ring 5=16 <collum <2=3=4 (male), body gradually tapering towards telson on rings 16–19 (
<figureCitationid="1338A7CABB4DF172FF7F771CFF4CF8BB"box="[160,230,1882,1908]"captionStart="FIGURE 8"captionStartId="12.[151,250,1740,1764]"captionTargetBox="[160,1420,496,1717]"captionTargetId="figure-164@12.[151,1436,489,1717]"captionTargetPageId="12"captionText="FIGURE 8. Rotundotergum elevatum sp. nov., male paratype. A-C. Habitus, dorsal, lateral and ventral views, respectively. Photographs by K.V. Makarov, taken not to scale."figureDoi="http://doi.org/10.5281/zenodo.6669211"httpUri="https://zenodo.org/record/6669211/files/figure.png"pageId="12"pageNumber="99">Fig. 8</figureCitation>
). Dorsum mostly slightly convex, generally smooth, metaterga largely very delicately striolate and rather densely microtuberculate, only caudal row of microtuberculations in front of rear tergal margin being regular (
<figureCitationid="1338A7CABB4DF172FAB07738FF0DF873"captionStart="FIGURE 8"captionStartId="12.[151,250,1740,1764]"captionTargetBox="[160,1420,496,1717]"captionTargetId="figure-164@12.[151,1436,489,1717]"captionTargetPageId="12"captionText="FIGURE 8. Rotundotergum elevatum sp. nov., male paratype. A-C. Habitus, dorsal, lateral and ventral views, respectively. Photographs by K.V. Makarov, taken not to scale."figureDoi="http://doi.org/10.5281/zenodo.6669211"httpUri="https://zenodo.org/record/6669211/files/figure.png"pageId="12"pageNumber="99">Fig. 8</figureCitation>
). Surface below paraterga more clearly striolate. Paraterga very strongly developed, mostly clearly upturned and reaching the level of dorsum, smaller, lower and subhorizontal in subadult females, in adult males set high, mostly at about upper ¼ height of metazonae (
<figureCitationid="1338A7CABB4CF173FDB970D1FD1BFF7E"box="[614,689,151,177]"captionStart="FIGURE 8"captionStartId="12.[151,250,1740,1764]"captionTargetBox="[160,1420,496,1717]"captionTargetId="figure-164@12.[151,1436,489,1717]"captionTargetPageId="12"captionText="FIGURE 8. Rotundotergum elevatum sp. nov., male paratype. A-C. Habitus, dorsal, lateral and ventral views, respectively. Photographs by K.V. Makarov, taken not to scale."figureDoi="http://doi.org/10.5281/zenodo.6669211"httpUri="https://zenodo.org/record/6669211/files/figure.png"pageId="13"pageNumber="100">Figs 8</figureCitation>
&
<figureCitationid="1338A7CABB4CF173FD0370D1FCABFF7E"box="[732,769,151,177]"captionStart="FIGURE 9"captionStartId="13.[151,250,1928,1952]"captionTargetBox="[151,1436,921,1904]"captionTargetId="figure-361@13.[151,1436,921,1904]"captionTargetPageId="13"captionText="FIGURE 9. Rotundotergum elevatum sp. nov., male paratype. A. Incisura lateralis (I) beneath antenna, sublateral view. B. Body ring 7 with gonopods removed to show gonopod aperture, ventral view. C & D. Body ring 7 with intact gonopods, ventral and lateral views, respectively. Abbreviation: I, Incisura lateralis. Photographs by K.V. Makarov, taken not to scale."figureDoi="http://doi.org/10.5281/zenodo.6669213"httpUri="https://zenodo.org/record/6669213/files/figure.png"pageId="13"pageNumber="100">9D</figureCitation>
), collum crescent-shaped, regularly convex at a narrowly rimmed anterior margin, slightly concave in the middle at a faintly rimmed caudal margin; following metaterga shoulder-shaped at a clearly rimmed anterior margin, metaterga 2–5 each with a very small and sharp denticle drawn laterally at a clearly angular anterolateral corner, metaterga 2–4 each with a very flat and broad lateral callus; following metaterga smooth at lateral margin, each with similarly and clearly rounded lateral margins and both more narrowly rounded anterolateral and caudolateral corners, calluses being particularly thick and prominent on pore-bearing rings, and interrupted anteriorly by a small incision behind anterolateral corner; caudal margins of paraterga narrowly rimmed much like anterior margin (
<figureCitationid="1338A7CABB4CF173FCDC71D5FCE0FE62"box="[771,842,403,429]"captionStart="FIGURE 8"captionStartId="12.[151,250,1740,1764]"captionTargetBox="[160,1420,496,1717]"captionTargetId="figure-164@12.[151,1436,489,1717]"captionTargetPageId="12"captionText="FIGURE 8. Rotundotergum elevatum sp. nov., male paratype. A-C. Habitus, dorsal, lateral and ventral views, respectively. Photographs by K.V. Makarov, taken not to scale."figureDoi="http://doi.org/10.5281/zenodo.6669211"httpUri="https://zenodo.org/record/6669211/files/figure.png"pageId="13"pageNumber="100">Figs 8</figureCitation>
&
<figureCitationid="1338A7CABB4CF173FCB071D5FC1CFE62"box="[879,950,403,429]"captionStart="FIGURE 9"captionStartId="13.[151,250,1928,1952]"captionTargetBox="[151,1436,921,1904]"captionTargetId="figure-361@13.[151,1436,921,1904]"captionTargetPageId="13"captionText="FIGURE 9. Rotundotergum elevatum sp. nov., male paratype. A. Incisura lateralis (I) beneath antenna, sublateral view. B. Body ring 7 with gonopods removed to show gonopod aperture, ventral view. C & D. Body ring 7 with intact gonopods, ventral and lateral views, respectively. Abbreviation: I, Incisura lateralis. Photographs by K.V. Makarov, taken not to scale."figureDoi="http://doi.org/10.5281/zenodo.6669213"httpUri="https://zenodo.org/record/6669213/files/figure.png"pageId="13"pageNumber="100">9B–D</figureCitation>
). Caudal corners of paraterga almost sharp, beak-shaped and distinctly drawn past rear tergal margin only in rings 17–19 (
<figureCitationid="1338A7CABB4CF173FC2171F1FBE8FE1E"box="[1022,1090,439,465]"captionStart="FIGURE 8"captionStartId="12.[151,250,1740,1764]"captionTargetBox="[160,1420,496,1717]"captionTargetId="figure-164@12.[151,1436,489,1717]"captionTargetPageId="12"captionText="FIGURE 8. Rotundotergum elevatum sp. nov., male paratype. A-C. Habitus, dorsal, lateral and ventral views, respectively. Photographs by K.V. Makarov, taken not to scale."figureDoi="http://doi.org/10.5281/zenodo.6669211"httpUri="https://zenodo.org/record/6669211/files/figure.png"pageId="13"pageNumber="100">Fig. 8</figureCitation>
). Pore formula normal (5, 7, 9, 10, 12, 13, 15–19); ozopores small, located inside ovoid pits considerably before caudal corners of paraterga (
<figureCitationid="1338A7CABB4CF173FAB5719DFF0FFDD6"captionStart="FIGURE 8"captionStartId="12.[151,250,1740,1764]"captionTargetBox="[160,1420,496,1717]"captionTargetId="figure-164@12.[151,1436,489,1717]"captionTargetPageId="12"captionText="FIGURE 8. Rotundotergum elevatum sp. nov., male paratype. A-C. Habitus, dorsal, lateral and ventral views, respectively. Photographs by K.V. Makarov, taken not to scale."figureDoi="http://doi.org/10.5281/zenodo.6669211"httpUri="https://zenodo.org/record/6669211/files/figure.png"pageId="13"pageNumber="100">Figs 8</figureCitation>
&
<figureCitationid="1338A7CABB4CF173FF1271B9FF58FDD6"box="[205,242,511,537]"captionStart="FIGURE 9"captionStartId="13.[151,250,1928,1952]"captionTargetBox="[151,1436,921,1904]"captionTargetId="figure-361@13.[151,1436,921,1904]"captionTargetPageId="13"captionText="FIGURE 9. Rotundotergum elevatum sp. nov., male paratype. A. Incisura lateralis (I) beneath antenna, sublateral view. B. Body ring 7 with gonopods removed to show gonopod aperture, ventral view. C & D. Body ring 7 with intact gonopods, ventral and lateral views, respectively. Abbreviation: I, Incisura lateralis. Photographs by K.V. Makarov, taken not to scale."figureDoi="http://doi.org/10.5281/zenodo.6669213"httpUri="https://zenodo.org/record/6669213/files/figure.png"pageId="13"pageNumber="100">9D</figureCitation>
). Tergal setae absent, no setation pattern traceable on metaterga. A weak transverse impression traceable on most metaterga. Stricture between pro- and metazonae distinct, but rather narrow, shallow and smooth (male). Limbus conspicuously brush-like (
<figureCitationid="1338A7CABB4CF173FDC07201FDD2FDAE"box="[543,632,583,609]"captionStart="FIGURE 9"captionStartId="13.[151,250,1928,1952]"captionTargetBox="[151,1436,921,1904]"captionTargetId="figure-361@13.[151,1436,921,1904]"captionTargetPageId="13"captionText="FIGURE 9. Rotundotergum elevatum sp. nov., male paratype. A. Incisura lateralis (I) beneath antenna, sublateral view. B. Body ring 7 with gonopods removed to show gonopod aperture, ventral view. C & D. Body ring 7 with intact gonopods, ventral and lateral views, respectively. Abbreviation: I, Incisura lateralis. Photographs by K.V. Makarov, taken not to scale."figureDoi="http://doi.org/10.5281/zenodo.6669213"httpUri="https://zenodo.org/record/6669213/files/figure.png"pageId="13"pageNumber="100">Fig. 9B</figureCitation>
). Pleurosternal carinae increasingly low, arcuate, longitudinal ridges or bulges traceable from ring 2 until ring 6 (male). A poor axial line sometimes traceable on metaterga of adults (
<figureCitationid="1338A7CABB4CF173FAB0722DFF0DFD66"captionStart="FIGURE 8"captionStartId="12.[151,250,1740,1764]"captionTargetBox="[160,1420,496,1717]"captionTargetId="figure-164@12.[151,1436,489,1717]"captionTargetPageId="12"captionText="FIGURE 8. Rotundotergum elevatum sp. nov., male paratype. A-C. Habitus, dorsal, lateral and ventral views, respectively. Photographs by K.V. Makarov, taken not to scale."figureDoi="http://doi.org/10.5281/zenodo.6669211"httpUri="https://zenodo.org/record/6669211/files/figure.png"pageId="13"pageNumber="100">Fig. 8</figureCitation>
). Epiproct rather long, conical, slightly curved caudally, tip rounded; pre-apical papillae very small (
<figureCitationid="1338A7CABB4CF173FAC972C9FAF1FD66"box="[1302,1371,655,681]"captionStart="FIGURE 8"captionStartId="12.[151,250,1740,1764]"captionTargetBox="[160,1420,496,1717]"captionTargetId="figure-164@12.[151,1436,489,1717]"captionTargetPageId="12"captionText="FIGURE 8. Rotundotergum elevatum sp. nov., male paratype. A-C. Habitus, dorsal, lateral and ventral views, respectively. Photographs by K.V. Makarov, taken not to scale."figureDoi="http://doi.org/10.5281/zenodo.6669211"httpUri="https://zenodo.org/record/6669211/files/figure.png"pageId="13"pageNumber="100">Fig. 8</figureCitation>
). Hypoproct subtriangular, sharpened at tip; setigerous papillae at caudal corners very small and well separated; sides faintly convex (
<figureCitationid="1338A7CABB4CF173FE987291FE0AFD3E"box="[327,416,727,753]"captionStart="FIGURE 8"captionStartId="12.[151,250,1740,1764]"captionTargetBox="[160,1420,496,1717]"captionTargetId="figure-164@12.[151,1436,489,1717]"captionTargetPageId="12"captionText="FIGURE 8. Rotundotergum elevatum sp. nov., male paratype. A-C. Habitus, dorsal, lateral and ventral views, respectively. Photographs by K.V. Makarov, taken not to scale."figureDoi="http://doi.org/10.5281/zenodo.6669211"httpUri="https://zenodo.org/record/6669211/files/figure.png"pageId="13"pageNumber="100">Fig. 8C</figureCitation>
Sterna broad, without notable modifications even before ring 7, nearly glabrous, cross-impressions weak (
<figureCitationid="1338A7CABB4CF173FAB072BDFF11FCF6"captionStart="FIGURE 8"captionStartId="12.[151,250,1740,1764]"captionTargetBox="[160,1420,496,1717]"captionTargetId="figure-164@12.[151,1436,489,1717]"captionTargetPageId="12"captionText="FIGURE 8. Rotundotergum elevatum sp. nov., male paratype. A-C. Habitus, dorsal, lateral and ventral views, respectively. Photographs by K.V. Makarov, taken not to scale."figureDoi="http://doi.org/10.5281/zenodo.6669211"httpUri="https://zenodo.org/record/6669211/files/figure.png"pageId="13"pageNumber="100">Fig. 8C</figureCitation>
). Legs long and slender (
<figureCitationid="1338A7CABB4CF173FE097359FDB0FCF6"box="[470,538,799,825]"captionStart="FIGURE 8"captionStartId="12.[151,250,1740,1764]"captionTargetBox="[160,1420,496,1717]"captionTargetId="figure-164@12.[151,1436,489,1717]"captionTargetPageId="12"captionText="FIGURE 8. Rotundotergum elevatum sp. nov., male paratype. A-C. Habitus, dorsal, lateral and ventral views, respectively. Photographs by K.V. Makarov, taken not to scale."figureDoi="http://doi.org/10.5281/zenodo.6669211"httpUri="https://zenodo.org/record/6669211/files/figure.png"pageId="13"pageNumber="100">Fig. 8</figureCitation>
), 1.6–1.7 times as long as mid-body height, considerably shorter in subadult females; male legs without tibial chelae; in length, femur >> tarsus> tibia> prefemur = postfemur> coxa; claw short and simple (
<figureCitationid="1338A7CABB4CF173FEFE7321FECFFC4E"box="[289,357,871,897]"captionStart="FIGURE 8"captionStartId="12.[151,250,1740,1764]"captionTargetBox="[160,1420,496,1717]"captionTargetId="figure-164@12.[151,1436,489,1717]"captionTargetPageId="12"captionText="FIGURE 8. Rotundotergum elevatum sp. nov., male paratype. A-C. Habitus, dorsal, lateral and ventral views, respectively. Photographs by K.V. Makarov, taken not to scale."figureDoi="http://doi.org/10.5281/zenodo.6669211"httpUri="https://zenodo.org/record/6669211/files/figure.png"pageId="13"pageNumber="100">Fig. 8</figureCitation>
). Male gonopores located on distinct, distomesal, coniform gonapophyses (
<figureCitationid="1338A7CABB4CF173FB6C7321FAA6FC4E"box="[1203,1292,871,897]"captionStart="FIGURE 8"captionStartId="12.[151,250,1740,1764]"captionTargetBox="[160,1420,496,1717]"captionTargetId="figure-164@12.[151,1436,489,1717]"captionTargetPageId="12"captionText="FIGURE 8. Rotundotergum elevatum sp. nov., male paratype. A-C. Habitus, dorsal, lateral and ventral views, respectively. Photographs by K.V. Makarov, taken not to scale."figureDoi="http://doi.org/10.5281/zenodo.6669211"httpUri="https://zenodo.org/record/6669211/files/figure.png"pageId="13"pageNumber="100">Fig. 8C</figureCitation>
<taxonomicNameid="4C03C0CCBB4CF173FEC877CEFDBEF86F"authority="Golovatch & Bouzan & Gallo & Bichuette, 2022"authorityName="Golovatch & Bouzan & Gallo & Bichuette"authorityYear="2022"box="[279,532,1928,1952]"class="Diplopoda"family="Chelodesmidae"genus="Rotundotergum"kingdom="Animalia"order="Polydesmida"pageId="13"pageNumber="100"phylum="Arthropoda"rank="species"species="elevatum"status="sp. nov.">
Gonopodal aperture subcordiform, much broader than following sterna, depressed caudal shelf with a few small and transverse crests (
<figureCitationid="1338A7CABB4FF170FE4B70FDFE45FF1A"box="[404,495,187,213]"captionStart="FIGURE 9"captionStartId="13.[151,250,1928,1952]"captionTargetBox="[151,1436,921,1904]"captionTargetId="figure-361@13.[151,1436,921,1904]"captionTargetPageId="13"captionText="FIGURE 9. Rotundotergum elevatum sp. nov., male paratype. A. Incisura lateralis (I) beneath antenna, sublateral view. B. Body ring 7 with gonopods removed to show gonopod aperture, ventral view. C & D. Body ring 7 with intact gonopods, ventral and lateral views, respectively. Abbreviation: I, Incisura lateralis. Photographs by K.V. Makarov, taken not to scale."figureDoi="http://doi.org/10.5281/zenodo.6669213"httpUri="https://zenodo.org/record/6669213/files/figure.png"pageId="14"pageNumber="101">Fig. 9B</figureCitation>
). Gonopods (
<figureCitationid="1338A7CABB4FF170FD5670FDFCA3FF1A"box="[649,777,187,213]"captionStart="FIGURE 9"captionStartId="13.[151,250,1928,1952]"captionTargetBox="[151,1436,921,1904]"captionTargetId="figure-361@13.[151,1436,921,1904]"captionTargetPageId="13"captionText="FIGURE 9. Rotundotergum elevatum sp. nov., male paratype. A. Incisura lateralis (I) beneath antenna, sublateral view. B. Body ring 7 with gonopods removed to show gonopod aperture, ventral view. C & D. Body ring 7 with intact gonopods, ventral and lateral views, respectively. Abbreviation: I, Incisura lateralis. Photographs by K.V. Makarov, taken not to scale."figureDoi="http://doi.org/10.5281/zenodo.6669213"httpUri="https://zenodo.org/record/6669213/files/figure.png"pageId="14"pageNumber="101">Figs 9C, D</figureCitation>
&
<figureCitationid="1338A7CABB4FF170FCEE70FDFCE4FF1A"box="[817,846,187,213]"captionStart="FIGURE 10"captionStartId="14.[151,250,1753,1777]"captionTargetBox="[151,1436,489,1729]"captionTargetId="figure-201@14.[151,1436,489,1729]"captionTargetPageId="14"captionText="FIGURE 10. Rotundotergum elevatum sp. nov., male paratype. A & B. Both gonopods, anterior and posterior views, respectively. C-F. Right gonopod, mesal, lateral, ventrocaudal and caudal views, respectively. Abbreviations: A, acropodite; c, cannula; Cx, gonocoxa; AP, acropodital process; lo, lateral lobe; Pf, prefemoral region; PfP, prefemoral process; sl, solenomere; st, sternal rudiment; Te, telopodite. Photographs by K.V. Makarov, taken not to scale."figureDoi="http://doi.org/10.5281/zenodo.6669215"httpUri="https://zenodo.org/record/6669215/files/figure.png"pageId="14"pageNumber="101">10</figureCitation>
) stout and mostly robust, with evident sternal rudiments (
<figureCitationid="1338A7CABB4FF170FF657129FEA6FE46"box="[186,268,367,393]"captionStart="FIGURE 10"captionStartId="14.[151,250,1753,1777]"captionTargetBox="[151,1436,489,1729]"captionTargetId="figure-201@14.[151,1436,489,1729]"captionTargetPageId="14"captionText="FIGURE 10. Rotundotergum elevatum sp. nov., male paratype. A & B. Both gonopods, anterior and posterior views, respectively. C-F. Right gonopod, mesal, lateral, ventrocaudal and caudal views, respectively. Abbreviations: A, acropodite; c, cannula; Cx, gonocoxa; AP, acropodital process; lo, lateral lobe; Pf, prefemoral region; PfP, prefemoral process; sl, solenomere; st, sternal rudiment; Te, telopodite. Photographs by K.V. Makarov, taken not to scale."figureDoi="http://doi.org/10.5281/zenodo.6669215"httpUri="https://zenodo.org/record/6669215/files/figure.png"pageId="14"pageNumber="101">Fig. 10</figureCitation>
<figureCitationid="1338A7CABB4FF170FD0071F1FC90FE1E"box="[735,826,439,465]"captionStart="FIGURE 9"captionStartId="13.[151,250,1928,1952]"captionTargetBox="[151,1436,921,1904]"captionTargetId="figure-361@13.[151,1436,921,1904]"captionTargetPageId="13"captionText="FIGURE 9. Rotundotergum elevatum sp. nov., male paratype. A. Incisura lateralis (I) beneath antenna, sublateral view. B. Body ring 7 with gonopods removed to show gonopod aperture, ventral view. C & D. Body ring 7 with intact gonopods, ventral and lateral views, respectively. Abbreviation: I, Incisura lateralis. Photographs by K.V. Makarov, taken not to scale."figureDoi="http://doi.org/10.5281/zenodo.6669213"httpUri="https://zenodo.org/record/6669213/files/figure.png"pageId="14"pageNumber="101">Figs 9D</figureCitation>
&
<figureCitationid="1338A7CABB4FF170FC8171F1FCD7FE1E"box="[862,893,439,465]"captionStart="FIGURE 10"captionStartId="14.[151,250,1753,1777]"captionTargetBox="[151,1436,489,1729]"captionTargetId="figure-201@14.[151,1436,489,1729]"captionTargetPageId="14"captionText="FIGURE 10. Rotundotergum elevatum sp. nov., male paratype. A & B. Both gonopods, anterior and posterior views, respectively. C-F. Right gonopod, mesal, lateral, ventrocaudal and caudal views, respectively. Abbreviations: A, acropodite; c, cannula; Cx, gonocoxa; AP, acropodital process; lo, lateral lobe; Pf, prefemoral region; PfP, prefemoral process; sl, solenomere; st, sternal rudiment; Te, telopodite. Photographs by K.V. Makarov, taken not to scale."figureDoi="http://doi.org/10.5281/zenodo.6669215"httpUri="https://zenodo.org/record/6669215/files/figure.png"pageId="14"pageNumber="101">10</figureCitation>
<taxonomicNameid="4C03C0CCBB4FF170FEF7769FFD8DF93E"authority="Golovatch & Bouzan & Gallo & Bichuette, 2022"authorityName="Golovatch & Bouzan & Gallo & Bichuette"authorityYear="2022"box="[296,551,1753,1777]"class="Diplopoda"family="Chelodesmidae"genus="Rotundotergum"kingdom="Animalia"order="Polydesmida"pageId="14"pageNumber="101"phylum="Arthropoda"rank="species"species="elevatum"status="sp. nov.">
. Based on most or much of the body being clearly pigmented, up to red-brown, superficially this species is certainly not more than troglophilic. This only emphasizes the general impression that