treatments-xml/data/03/9C/87/039C87E5EF11FFB804FCF8AFFDACFE7F.xml

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<mods:title id="D4C3D7664159ED0560426B5A7B94503B">Pleurothallis hawkingii and Pleurothallis vide-vallis (Orchidaceae; Epidendroideae), two new species from Cordillera de Guanacaste in Costa Rica</mods:title>
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<mods:namePart id="9093392CEB967D26617BB7F4F281FF36">Karremans, Adam P.</mods:namePart>
<mods:affiliation id="D6E39C802EAF24D95C90428804A17F57">Naturalis Biodiversity Center, The Netherlands</mods:affiliation>
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<mods:namePart id="98F9A91A89C6A806FCBF626959CCBB7B">Jiménez, José Esteban</mods:namePart>
<mods:affiliation id="1B61EBCADFE017FC8937CA63F5D364D9">Herbario Luis A. Fournier Origgi, Universidad de Costa Rica, Apdo. 11501 - 2060, San José, Costa Rica</mods:affiliation>
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<taxonomicName id="4C354D70EF11FFBC04FCF8AFFD7CF8C9" ID-CoL="8X3DY" authority="Karremans &amp; J.E.Jimenez" authorityName="Karremans &amp; J. E. Jimenez" authorityYear="2018" box="[136,710,1876,1903]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="0" pageNumber="185" phylum="Tracheophyta" rank="species" species="hawkingii" status="sp. nov.">
<emphasis id="B941EAE1EF11FFBC04FCF8AFFE2DF8C9" bold="true" box="[136,407,1876,1903]" italics="true" pageId="0" pageNumber="185">Pleurothallis hawkingii</emphasis>
Karremans &amp; J.E.Jiménez
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, 
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<taxonomicNameLabel id="A272579AEF11FFBC06A5F8ADFC9CF8C9" box="[721,806,1878,1903]" pageId="0" pageNumber="185" rank="species">sp. nov.</taxonomicNameLabel>
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(
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)
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<paragraph id="8B8A36F3EF11FFBC04FCF887FD02F85B" blockId="0.[136,1451,1916,2045]" pageId="0" pageNumber="185">
<materialsCitation id="3B5D3CAEEF11FFBC04FCF887FD0EF85B" collectingDate="2006-02-02" collectionCode="USJ" collectorName="de Tierras Morenas" country="Costa Rica" elevation="950" latitude="10.603223" location="Rio Cabuyo" longLatPrecision="1" longitude="-85.00147" pageId="0" pageNumber="185" specimenCount="2" stateProvince="Guanacaste" typeStatus="holotype">
Type:— 
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. 
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: Tilarán, Tierras Morenas, desviación a la izquierda después del 
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, camino al Proyecto Geotérmico Tenorio y Cerro Jilguero, ca. 
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norte 
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, ladera sureste del Volcán Tenorio, 
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, 
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, bosque pluvial premontano, epífitas a orillas del camino y en bordes de potreros, 
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, 
<emphasis id="B941EAE1EF11FFBC04C9F813FEF8F85B" box="[189,322,2024,2045]" italics="true" pageId="0" pageNumber="185">Bogarín 2386</emphasis>
(
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: 
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!; 
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: JBL-spirit!)
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.
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<emphasis id="B941EAE1EF10FFBD04FCF8ADFF44F8CD" bold="true" box="[136,254,1878,1899]" pageId="1" pageNumber="186">FIGURE 1.</emphasis>
<taxonomicName id="4C354D70EF10FFBD0577F8ADFE5EF8CD" authorityName="Karremans &amp; J. E. Jimenez" authorityYear="2018" box="[259,484,1878,1899]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="186" phylum="Tracheophyta" rank="species" species="hawkingii">
<emphasis id="B941EAE1EF10FFBD0577F8ADFE5EF8CD" box="[259,484,1878,1899]" italics="true" pageId="1" pageNumber="186">Pleurothallis hawkingii</emphasis>
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. A. Habit. B. Inflorescence. C. Flower. D. Dissected perianth. E. Ovary, column and lip, lateral view. F. Anther cap and pollinarium. Based on type specimen, 
<emphasis id="B941EAE1EF10FFBD06D9F881FC88F829" box="[685,818,1914,1935]" italics="true" pageId="1" pageNumber="186">Bogarín 2386</emphasis>
(USJ, JBL-spirit). Lankester Composite Digital Plate (LCDP) by APK.
</paragraph>
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The fasciculate inflorescence with numerous simultaneously open flowers in 
<taxonomicName id="4C354D70EF13FFBE078FFF5BFABEFF1F" authorityName="Karremans &amp; J. E. Jimenez" authorityYear="2018" box="[1019,1284,160,185]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="187" phylum="Tracheophyta" rank="species" species="hawkingii">
<emphasis id="B941EAE1EF13FFBE078FFF5BFABEFF1F" box="[1019,1284,160,185]" italics="true" pageId="2" pageNumber="187">Pleurothallis hawkingii</emphasis>
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is reminiscent of 
<taxonomicName id="4C354D70EF13FFBE04DFFF3FFDA8FF7B" authority="Luer (1996: 70 - 72)" authorityName="Luer" authorityPageNumber="70 - 72" authorityYear="1996" box="[171,530,196,221]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="187" phylum="Tracheophyta" rank="species" species="bothros">
<emphasis id="B941EAE1EF13FFBE04DFFF3FFEA5FF7B" box="[171,287,196,221]" italics="true" pageId="2" pageNumber="187">P. bothros</emphasis>
<bibRefCitation id="EFA44B02EF13FFBE055EFF3FFDA8FF7B" author="Luer, C. A." box="[298,530,196,221]" pageId="2" pageNumber="187" pagination="54 - 113" refId="ref2731" refString="Luer, C. A. (1996) New species in the Pleurothallidinae (Orchidaceae) from Costa Rica. Lindleyana 11: 54 - 113." type="journal article" year="1996">Luer (1996: 70–72)</bibRefCitation>
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. It differs, however, in the non-spreading, pale yellow to whitish flowers with pinkish markings (vs. greenish), oblong synsepal (vs. ovate), obtuse petals (vs. acute) and lip horizontally disposed, panduriform, smaller, ca. 2.7 × 
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(vs. recurved, ovate, 3.5 × 
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).
</paragraph>
<paragraph id="8B8A36F3EF13FFBE04C9FECBFC62FD73" blockId="2.[136,1452,160,977]" pageId="2" pageNumber="187">
Epiphytic, erect herbs, up to 
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tall. Roots flexuous, thin, 0.5–1.0 mm diameter, densely spaced, appearing fasciculate. Ramicauls erect, thin, up to 
<quantity id="4CCD9B16EF13FFBE0631FEAFFD30FECB" box="[581,650,340,365]" metricMagnitude="-1" metricUnit="m" metricValue="2.5" pageId="2" pageNumber="187" unit="cm" value="25.0">25 cm</quantity>
long, covered by tubular sheaths close to the base, tightly adpressed, up to 
<quantity id="4CCD9B16EF13FFBE04D1FE83FF67FE37" box="[165,221,376,401]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="187" unit="cm" value="5.0">5 cm</quantity>
long. Leaves horizontal or nearly so, sessile, ovate, shortly acuminate, 8.0–9.5 × 3.0– 
<quantity id="4CCD9B16EF13FFBE00D2FE83FB4EFE37" box="[1190,1268,376,401]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="2" pageNumber="187" unit="cm" value="3.5">3.5 cm</quantity>
, deeply cordate. Inflorescence a fascicle of 3–9 simultaneously open flowers, subtended by a spathaceous bract 1.0– 
<quantity id="4CCD9B16EF13FFBE016CFE67FADDFE13" box="[1304,1383,412,437]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="187" unit="cm" value="1.5">1.5 cm</quantity>
long; pedicels terete, 
<quantity id="4CCD9B16EF13FFBE0542FE3BFE29FE7F" box="[310,403,448,473]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="2" pageNumber="187" unit="mm" value="5.5" valueMax="6.0" valueMin="5.0">5–6 mm</quantity>
long, pale green. Ovary clavate, 
<quantity id="4CCD9B16EF13FFBE0777FE3BFC30FE7F" box="[771,906,448,473]" metricMagnitude="-3" metricUnit="m" metricValue="3.35" metricValueMax="3.5" metricValueMin="3.2" pageId="2" pageNumber="187" unit="mm" value="3.35" valueMax="3.5" valueMin="3.2">3.2–3.5 mm</quantity>
long, green. Flowers non-spreading, pale yellow with pinkish markings to solid pink; dorsal sepal oblong, slightly concave, obtuse, 3(5)-veined, 7.0–7.3 × 
<quantity id="4CCD9B16EF13FFBE016BFE1FFA1DFE5B" box="[1311,1447,484,509]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="3.1" metricValueMin="2.9" pageId="2" pageNumber="187" unit="mm" value="3.0" valueMax="3.1" valueMin="2.9">2.9–3.1 mm</quantity>
, apiculate; lateral sepals connate in an oblong, slightly concave synsepal, obtuse, 6-veined, 6.5–7.0 × 
<quantity id="4CCD9B16EF13FFBE016EFDF3FA1DFD87" box="[1306,1447,520,545]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" metricValueMax="3.5" metricValueMin="3.3" pageId="2" pageNumber="187" unit="mm" value="3.4" valueMax="3.5" valueMin="3.3">3.3–3.5 mm</quantity>
; petals oblong, oblique, acute, 3.5–3.7 × 
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; lip panduriform, horizontally disposed, 2.6–2.8 × 
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, margins raised, apex obtuse; glenion a depression between elevated lateral margins, the disk ca. 
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long; column straight, transversely subrectangular, with a column foot, 1.5–1.7 × 
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, the anther and stigma apical; anther cap ovate, cucullate, acute at the base, bilobed apically, 2-celled, ca. 0.7 × 
<quantity id="4CCD9B16EF13FFBE07CDFD63FBAAFD17" box="[953,1040,664,689]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="187" unit="mm" value="0.5">0.5 mm</quantity>
; pollinia 2, narrowly ovate-pyriform, 
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long, connected to an elliptic viscidium. Fruits and seeds unknown.
</paragraph>
<paragraph id="8B8A36F3EF13FFBE04C9FD1BFD56FC77" blockId="2.[136,1452,160,977]" pageId="2" pageNumber="187">
Additional specimens examined:— 
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<collectingCountry id="F3227663EF13FFBE0633FD1BFD48FD5F" box="[583,754,736,761]" name="Costa Rica" pageId="2" pageNumber="187">COSTA RICA</collectingCountry>
. 
<collectingRegion id="49F1F811EF13FFBE0688FD1BFC38FD5F" box="[764,898,736,761]" country="Costa Rica" name="Guanacaste" pageId="2" pageNumber="187">Guanacaste</collectingRegion>
: 
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<collectingCounty id="62EB4E7FEF13FFBE07F9FD1BFC56FD5F" box="[909,1004,736,761]" pageId="2" pageNumber="187">La Cruz</collectingCounty>
</collectorName>
, 
<collectingMunicipality id="6BEEAC89EF13FFBE078CFD1BFB09FD5F" box="[1016,1203,736,761]" pageId="2" pageNumber="187">Parque Nacional</collectingMunicipality>
<collectingRegion id="49F1F811EF13FFBE00CFFD1BFA85FD5F" box="[1211,1343,736,761]" country="Costa Rica" name="Guanacaste" pageId="2" pageNumber="187">Guanacaste</collectingRegion>
, 
<location id="8EEA6028EF13FFBE013FFD1BFEE9FCBB" LSID="urn:lsid:plazi:treatment:039C87E5EF11FFB804FCF8AFFDACFE7F:8EEA6028EF13FFBE013FFD1BFEE9FCBB" country="Costa Rica" county="La Cruz" latitude="10.98" longLatPrecision="21" longitude="-85.475" municipality="Parque Nacional" name="Estacion Biologica Maritza" pageId="2" pageNumber="187" stateProvince="Guanacaste">Estación Biológica Maritza</location>
, 
<location id="8EEA6028EF13FFBE052FFCFFFE54FCBB" LSID="urn:lsid:plazi:treatment:039C87E5EF11FFB804FCF8AFFDACFE7F:8EEA6028EF13FFBE052FFCFFFE54FCBB" box="[347,494,772,797]" country="Costa Rica" county="La Cruz" latitude="10.98" longLatPrecision="21" longitude="-85.475" municipality="Parque Nacional" name="Volcan Orosi" pageId="2" pageNumber="187" stateProvince="Guanacaste">Volcán Orosí</location>
, cima del volcán entre el hito y la piedra, 
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</quantity>
, 
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, 
<geoCoordinate id="EE015034EF13FFBE00D4FCFFFA89FCBB" box="[1184,1331,772,797]" degrees="85" direction="west" minutes="28" orientation="longitude" pageId="2" pageNumber="187" precision="15" seconds="30" value="-85.475">85°28’30” W</geoCoordinate>
, 
<date id="FF8B1033EF13FFBE0148FCFFFF7AFCE7" pageId="2" pageNumber="187" value="2017-08-21">
<collectingDate id="EFCFE9DBEF13FFBE0148FCFFFF7AFCE7" pageId="2" pageNumber="187" value="2017-08-21">21Agosto 2017</collectingDate>
</date>
(fl), 
<emphasis id="B941EAE1EF13FFBE048AFCD3FD3BFCE7" box="[254,641,808,833]" italics="true" pageId="2" pageNumber="187">
<collectorName id="26C05325EF13FFBE048AFCD3FEE4FCE7" box="[254,350,808,833]" pageId="2" pageNumber="187">Morales</collectorName>
, 
<collectorName id="26C05325EF13FFBE0518FCD3FE72FCE7" box="[364,456,808,833]" pageId="2" pageNumber="187">Grayum</collectorName>
&amp; 
<collectorName id="26C05325EF13FFBE0585FCD3FD8BFCE7" box="[497,561,808,833]" pageId="2" pageNumber="187">Rojas</collectorName>
15521
</emphasis>
(INB-CR)
</materialsCitation>
; 
<materialsCitation id="3B5D3CAEEF13FFBE077AFCD3FD14FC2F" collectingDate="2017-08-21" collectorName="La Cruz &amp; Morales &amp; Grayum &amp; Rojas" country="Costa Rica" county="La Cruz" elevation="1300" latitude="10.98" location="Estacion Biologica Maritza" longLatPrecision="21" longitude="-85.475" municipality="Parque Nacional" pageId="2" pageNumber="187" specimenCount="1" stateProvince="Guanacaste">
<collectorName id="26C05325EF13FFBE077AFCD3FCD4FCE7" box="[782,878,808,833]" pageId="2" pageNumber="187">
<collectingCounty id="62EB4E7FEF13FFBE077AFCD3FCD4FCE7" box="[782,878,808,833]" pageId="2" pageNumber="187">La Cruz</collectingCounty>
</collectorName>
, 
<collectingMunicipality id="6BEEAC89EF13FFBE0708FCD3FB83FCE7" box="[892,1081,808,833]" pageId="2" pageNumber="187">Parque Nacional</collectingMunicipality>
<collectingRegion id="49F1F811EF13FFBE0036FCD3FB7CFCE7" box="[1090,1222,808,833]" country="Costa Rica" name="Guanacaste" pageId="2" pageNumber="187">Guanacaste</collectingRegion>
, 
<location id="8EEA6028EF13FFBE00A0FCD3FF58FCC3" LSID="urn:lsid:plazi:treatment:039C87E5EF11FFB804FCF8AFFDACFE7F:8EEA6028EF13FFBE00A0FCD3FF58FCC3" country="Costa Rica" county="La Cruz" latitude="10.98" longLatPrecision="21" longitude="-85.475" municipality="Parque Nacional" name="Estacion Biologica Maritza" pageId="2" pageNumber="187" stateProvince="Guanacaste">Estación Biológica Maritza</location>
, 
<location id="8EEA6028EF13FFBE0499FCB7FE38FCC3" LSID="urn:lsid:plazi:treatment:039C87E5EF11FFB804FCF8AFFDACFE7F:8EEA6028EF13FFBE0499FCB7FE38FCC3" box="[237,386,844,869]" country="Costa Rica" county="La Cruz" latitude="10.98" longLatPrecision="21" longitude="-85.475" municipality="Parque Nacional" name="Volcan Orosi" pageId="2" pageNumber="187" stateProvince="Guanacaste">Volcán Orosí</location>
, cima del volcán entre el hito y la piedra, 
<quantity id="4CCD9B16EF13FFBE072CFCB7FC15FCC3" box="[856,943,844,869]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="2" pageNumber="187" unit="m" value="1300.0">
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</quantity>
, 
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, 
<geoCoordinate id="EE015034EF13FFBE002DFCB7FB55FCC3" box="[1113,1263,844,869]" degrees="85" direction="west" minutes="28" orientation="longitude" pageId="2" pageNumber="187" precision="15" seconds="30" value="-85.475">85°28’30” W</geoCoordinate>
, 
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<collectingDate id="EFCFE9DBEF13FFBE008EFCB7FA16FCC3" box="[1274,1452,844,869]" pageId="2" pageNumber="187" value="2017-08-21">21 Agosto 2017</collectingDate>
</date>
(fl), 
<emphasis id="B941EAE1EF13FFBE04CEFC8BFD88FC2F" box="[186,562,880,905]" italics="true" pageId="2" pageNumber="187">
<collectorName id="26C05325EF13FFBE04CEFC8BFEA0FC2F" box="[186,282,880,905]" pageId="2" pageNumber="187">Morales</collectorName>
, 
<collectorName id="26C05325EF13FFBE0551FC8BFE3BFC2F" box="[293,385,880,905]" pageId="2" pageNumber="187">Grayum</collectorName>
&amp; 
<collectorName id="26C05325EF13FFBE05D1FC8BFE5FFC2F" box="[421,485,880,905]" pageId="2" pageNumber="187">Rojas</collectorName>
15523
</emphasis>
(INB-CR)
</materialsCitation>
; 
<materialsCitation id="3B5D3CAEEF13FFBE06CDFC8BFD52FC77" collectingDate="2017-08-23" collectorName="La Cruz &amp; Morales" country="Costa Rica" county="La Cruz" elevation="1300" latitude="10.975833" location="Estacion Biologica Maritza" longLatPrecision="21" longitude="-85.48167" municipality="Parque Nacional" pageId="2" pageNumber="187" specimenCount="1" stateProvince="Guanacaste">
<collectorName id="26C05325EF13FFBE06CDFC8BFCADFC2F" box="[697,791,880,905]" pageId="2" pageNumber="187">
<collectingCounty id="62EB4E7FEF13FFBE06CDFC8BFCADFC2F" box="[697,791,880,905]" pageId="2" pageNumber="187">La Cruz</collectingCounty>
</collectorName>
, 
<collectingMunicipality id="6BEEAC89EF13FFBE0756FC8BFC66FC2F" box="[802,988,880,905]" pageId="2" pageNumber="187">Parque Nacional</collectingMunicipality>
<collectingRegion id="49F1F811EF13FFBE0797FC8BFBDDFC2F" box="[995,1127,880,905]" country="Costa Rica" name="Guanacaste" pageId="2" pageNumber="187">Guanacaste</collectingRegion>
, 
<location id="8EEA6028EF13FFBE0006FC8BFA1DFC2F" LSID="urn:lsid:plazi:treatment:039C87E5EF11FFB804FCF8AFFDACFE7F:8EEA6028EF13FFBE0006FC8BFA1DFC2F" box="[1138,1447,880,905]" country="Costa Rica" county="La Cruz" latitude="10.975833" longLatPrecision="21" longitude="-85.48167" municipality="Parque Nacional" name="Estacion Biologica Maritza" pageId="2" pageNumber="187" stateProvince="Guanacaste">Estación Biológica Maritza</location>
, 
<location id="8EEA6028EF13FFBE04FCFC6FFEA7FC0B" LSID="urn:lsid:plazi:treatment:039C87E5EF11FFB804FCF8AFFDACFE7F:8EEA6028EF13FFBE04FCFC6FFEA7FC0B" box="[136,285,916,941]" country="Costa Rica" county="La Cruz" latitude="10.975833" longLatPrecision="21" longitude="-85.48167" municipality="Parque Nacional" name="Volcan Orosi" pageId="2" pageNumber="187" stateProvince="Guanacaste">Volcán Orosí</location>
, cima del volcán entre la base del campamento y sendero a 
<location id="8EEA6028EF13FFBE07C4FC6FFBB0FC0B" LSID="urn:lsid:plazi:treatment:039C87E5EF11FFB804FCF8AFFDACFE7F:8EEA6028EF13FFBE07C4FC6FFBB0FC0B" box="[944,1034,916,941]" country="Costa Rica" county="La Cruz" latitude="10.975833" longLatPrecision="21" longitude="-85.48167" municipality="Parque Nacional" name="Maritza" pageId="2" pageNumber="187" stateProvince="Guanacaste">Maritza</location>
, 
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<elevation id="0018D1C0EF13FFBE0060FC6FFBD0FC0B" box="[1044,1130,916,941]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="2" pageNumber="187" unit="m" value="1300.0">1300 m</elevation>
</quantity>
, 
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, 
<geoCoordinate id="EE015034EF13FFBE0166FC6FFA12FC0B" box="[1298,1448,916,941]" degrees="85" direction="west" minutes="28" orientation="longitude" pageId="2" pageNumber="187" precision="15" seconds="54" value="-85.48167">85°28’54” W</geoCoordinate>
, 
<date id="FF8B1033EF13FFBE04FCFC43FE83FC77" box="[136,313,952,977]" pageId="2" pageNumber="187" value="2017-08-23">
<collectingDate id="EFCFE9DBEF13FFBE04FCFC43FE83FC77" box="[136,313,952,977]" pageId="2" pageNumber="187" value="2017-08-23">23 Agosto 2017</collectingDate>
</date>
(fl), 
<emphasis id="B941EAE1EF13FFBE0506FC43FDA6FC77" box="[370,540,952,977]" italics="true" pageId="2" pageNumber="187">
<collectorName id="26C05325EF13FFBE0506FC43FE75FC77" box="[370,463,952,977]" pageId="2" pageNumber="187">Morales</collectorName>
15527
</emphasis>
(INB-CR; 
<figureCitation id="130E2A76EF13FFBE06EEFC43FD65FC77" box="[666,735,952,977]" captionStart="FIGURE 2" captionStartId="2.[136,229,1844,1865]" captionTargetBox="[151,1436,1001,1819]" captionTargetId="figure-393@2.[151,1436,1001,1819]" captionTargetPageId="2" captionText="FIGURE 2. Pleurothallis hawkingii, Morales 15527 (INB-CR).A. In situ on the Orosí Volcano. B. Plants of the same population showing colour variation. Photographs by J.F. Morales." figureDoi="http://doi.org/10.5281/zenodo.13707375" httpUri="https://zenodo.org/record/13707375/files/figure.png" pageId="2" pageNumber="187">Fig. 2</figureCitation>
)
</materialsCitation>
.
</paragraph>
<caption id="DF4A667BEF13FFBE04FCF8CFFDFFF8CB" ID-DOI="http://doi.org/10.5281/zenodo.13707375" ID-Zenodo-Dep="13707375" httpUri="https://zenodo.org/record/13707375/files/figure.png" pageId="2" pageNumber="187" startId="2.[136,229,1844,1865]" targetBox="[151,1436,1001,1819]" targetPageId="2" targetType="figure">
<paragraph id="8B8A36F3EF13FFBE04FCF8CFFDFFF8CB" blockId="2.[136,1451,1844,1901]" pageId="2" pageNumber="187">
<emphasis id="B941EAE1EF13FFBE04FCF8CFFF47F8EF" bold="true" box="[136,253,1844,1865]" pageId="2" pageNumber="187">FIGURE 2.</emphasis>
<taxonomicName id="4C354D70EF13FFBE0576F8CFFD84F8EF" authority=", Morales" authorityName="Morales" box="[258,574,1844,1865]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="187" phylum="Tracheophyta" rank="species" species="hawkingii">
<emphasis id="B941EAE1EF13FFBE0576F8CFFE59F8EF" box="[258,483,1844,1865]" italics="true" pageId="2" pageNumber="187">Pleurothallis hawkingii</emphasis>
, 
<emphasis id="B941EAE1EF13FFBE059AF8CFFD84F8EF" box="[494,574,1844,1865]" italics="true" pageId="2" pageNumber="187">Morales</emphasis>
</taxonomicName>
<emphasis id="B941EAE1EF13FFBE0637F8CFFDC5F8EF" box="[579,639,1844,1865]" italics="true" pageId="2" pageNumber="187">15527</emphasis>
(INB-CR). A. 
<emphasis id="B941EAE1EF13FFBE0778F8CFFCF2F8EF" box="[780,840,1844,1865]" italics="true" pageId="2" pageNumber="187">In situ</emphasis>
on the Orosí Volcano. B. Plants of the same population showing colour variation. Photographs by J.F. Morales.
</paragraph>
</caption>
<paragraph id="8B8A36F3EF13FFBE04C9F865FC33F87D" blockId="2.[136,1451,1950,2083]" pageId="2" pageNumber="187">
Eponomy:—Honouring the English theoretical physicist, cosmologist and author Stephen William Hawking, who passed away the day this manuscript was submitted, 
<date id="FF8B1033EF13FFBE06A2F839FC3FF87D" box="[726,901,1986,2011]" pageId="2" pageNumber="187" value="2018-03-14">14 March 2018</date>
.
</paragraph>
<paragraph id="8B8A36F3EF13FFBE04C9F81DFBD1F859" blockId="2.[136,1451,1950,2083]" box="[189,1131,2022,2047]" pageId="2" pageNumber="187">Phenology:—Flowering August in the field but January and February in cultivation.</paragraph>
<paragraph id="8B8A36F3EF13FFBE04C9F7F1FD62F785" blockId="2.[136,1451,1950,2083]" box="[189,728,2058,2083]" pageId="2" pageNumber="187">
Distribution:—Endemic to northern 
<collectingCountry id="F3227663EF13FFBE0622F7F1FD6EF785" box="[598,724,2058,2083]" name="Costa Rica" pageId="2" pageNumber="187">Costa Rica</collectingCountry>
.
</paragraph>
<caption id="DF4A667BEF12FFBF04FCF8ADFDA6F815" ID-DOI="http://doi.org/10.5281/zenodo.13707377" ID-Zenodo-Dep="13707377" httpUri="https://zenodo.org/record/13707377/files/figure.png" pageId="3" pageNumber="188" startId="3.[136,229,1878,1899]" targetBox="[151,1436,190,1853]" targetPageId="3" targetType="figure">
<paragraph id="8B8A36F3EF12FFBF04FCF8ADFDA6F815" blockId="3.[136,1453,1878,1971]" pageId="3" pageNumber="188">
<emphasis id="B941EAE1EF12FFBF04FCF8ADFF47F8CD" bold="true" box="[136,253,1878,1899]" pageId="3" pageNumber="188">FIGURE 3.</emphasis>
<taxonomicName id="4C354D70EF12FFBF0576F8ADFE52F8CD" authorityName="Karremans &amp; J. E. Jimenez" authorityYear="2018" box="[258,488,1878,1899]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="188" phylum="Tracheophyta" rank="species" species="vide-vallis">
<emphasis id="B941EAE1EF12FFBF0576F8ADFE52F8CD" box="[258,488,1878,1899]" italics="true" pageId="3" pageNumber="188">Pleurothallis vide-vallis</emphasis>
</taxonomicName>
. A. Habit. B. Inflorescence. C. Flower. D. Dissected perianth. E. Ovary, column and lip, lateral view. F. Column lateral and dorsal view. G. Anther cap and pollinarium. Based on type specimen 
<emphasis id="B941EAE1EF12FFBF079DF881FB71F829" box="[1001,1227,1914,1935]" italics="true" pageId="3" pageNumber="188">Calderón &amp; Jiménez 96</emphasis>
(JBL-spirit). Lankester Composite Digital Plate (LCDP) by APK.
</paragraph>
</caption>
<paragraph id="8B8A36F3EF15FFB804C9FF5BFDB3FF7B" blockId="4.[136,1452,160,473]" pageId="4" pageNumber="189">
Habitat and ecology:—The species is only known from the Orosí Volcano and the Tilarán area in Cordillera de 
<collectingRegion id="49F1F811EF15FFB804FCFF3FFEB3FF7B" box="[136,265,196,221]" country="Costa Rica" name="Guanacaste" pageId="4" pageNumber="189">Guanacaste</collectingRegion>
between 
<quantity id="4CCD9B16EF15FFB80500FF3FFDBEFF7B" box="[372,516,196,221]" metricMagnitude="3" metricUnit="m" metricValue="1.1" metricValueMax="1.3" metricValueMin="0.9" pageId="4" pageNumber="189" unit="m" value="1100.0" valueMax="1300.0" valueMin="900.0">900–1300 m</quantity>
.
</paragraph>
<paragraph id="8B8A36F3EF15FFB804C9FF13FDACFE7F" blockId="4.[136,1452,160,473]" pageId="4" pageNumber="189">
Notes:— 
<taxonomicName id="4C354D70EF15FFB80556FF13FD9CFEA7" box="[290,550,232,257]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="189" phylum="Tracheophyta" rank="species" species="hawkingii">
<emphasis id="B941EAE1EF15FFB80556FF13FD9CFEA7" box="[290,550,232,257]" italics="true" pageId="4" pageNumber="189">Pleurothallis hawkingii</emphasis>
</taxonomicName>
belongs to 
<emphasis id="B941EAE1EF15FFB806D5FF13FD0CFEA7" box="[673,694,232,257]" italics="true" pageId="4" pageNumber="189">
<collectionCode id="ED24AE36EF15FFB806D5FF13FD08FEA7" box="[673,690,232,257]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="4" pageNumber="189" type="Herbarium">P</collectionCode>
.
</emphasis>
sect. 
<emphasis id="B941EAE1EF15FFB80685FF13FB9DFEA7" box="[753,1063,232,257]" italics="true" pageId="4" pageNumber="189">Macrophyllae-Fasciculatae</emphasis>
(
<bibRefCitation id="EFA44B02EF15FFB80047FF13FB10FEA7" author="Luer C. A." box="[1075,1194,232,257]" pageId="4" pageNumber="189" pagination="1 - 311" refId="ref2852" refString="Luer C. A. (2005) Icones pleurothallidinarum XXVII. Dryadella and Acronia sect. Macrophyllaceae - Fasciculatae. Monographs in Systematic Botany 103: 1 - 311." type="journal article" year="2005">Luer 2005</bibRefCitation>
), and can be recognised by the fasciculate inflorescence of numerous simultaneously open flowers, a rare feature in this species-rich group. It shares the fasciculate inflorescence of several simultaneously open flowers with 
<taxonomicName id="4C354D70EF15FFB80040FECBFB1CFEEF" authorityName="Luer" authorityYear="1996" box="[1076,1190,304,329]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="189" phylum="Tracheophyta" rank="species" species="bothros">
<emphasis id="B941EAE1EF15FFB80040FECBFB1CFEEF" box="[1076,1190,304,329]" italics="true" pageId="4" pageNumber="189">P. bothros</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C354D70EF15FFB800AAFECBFAD5FEEF" authorityName="Karremans &amp; J. E. Jimenez" authorityYear="2018" box="[1246,1391,304,329]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="189" phylum="Tracheophyta" rank="species" species="vide-vallis">
<emphasis id="B941EAE1EF15FFB800AAFECBFAD5FEEF" box="[1246,1391,304,329]" italics="true" pageId="4" pageNumber="189">P. vide-vallis</emphasis>
</taxonomicName>
, also described in this paper. 
<taxonomicName id="4C354D70EF15FFB805ECFEAFFD1AFECB" box="[408,672,340,365]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="189" phylum="Tracheophyta" rank="species" species="hawkingii">
<emphasis id="B941EAE1EF15FFB805ECFEAFFD1AFECB" box="[408,672,340,365]" italics="true" pageId="4" pageNumber="189">Pleurothallis hawkingii</emphasis>
</taxonomicName>
can be distinguished from 
<taxonomicName id="4C354D70EF15FFB807ADFEAFFBD1FECB" authorityName="Karremans &amp; J. E. Jimenez" authorityYear="2018" box="[985,1131,340,365]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="189" phylum="Tracheophyta" rank="species" species="vide-vallis">
<emphasis id="B941EAE1EF15FFB807ADFEAFFBD1FECB" box="[985,1131,340,365]" italics="true" pageId="4" pageNumber="189">P. vide-vallis</emphasis>
</taxonomicName>
by the oblong synsepal (vs. elliptic), narrower oblong (vs. elliptic) petals, horizontal lip (vs. apically recurved) that is shorter and broader (2.6–2.8 × 
<quantity id="4CCD9B16EF15FFB804EAFE67FE9FFE13" box="[158,293,412,437]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" metricValueMax="1.8" metricValueMin="1.6" pageId="4" pageNumber="189" unit="mm" value="1.7" valueMax="1.8" valueMin="1.6">1.6–1.8 mm</quantity>
vs. 3.4–3.6 × 
<quantity id="4CCD9B16EF15FFB805B7FE67FDF1FE13" box="[451,587,412,437]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="4" pageNumber="189" unit="mm" value="1.4" valueMax="1.5" valueMin="1.3">1.3–1.5 mm</quantity>
) and has raised margins, lacking a callus higher at the base. It is also found at lower elevations than 
<taxonomicName id="4C354D70EF15FFB8050BFE3BFDB5FE7F" authorityName="Karremans &amp; J. E. Jimenez" authorityYear="2018" box="[383,527,448,473]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="189" phylum="Tracheophyta" rank="species" species="vide-vallis">
<emphasis id="B941EAE1EF15FFB8050BFE3BFDB5FE7F" box="[383,527,448,473]" italics="true" pageId="4" pageNumber="189">P. vide-vallis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>