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<mods:title id="DB10071A626F29B17DA7E93DEE16FD1E">A new classification of the tribe Hygrotini Portevin, 1929 (Coleoptera: Dytiscidae: Hydroporinae)</mods:title>
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<subSubSection id="C32F65530F35FF97FF1F62BDCA9550B6" box="[151,677,475,502]" pageId="22" pageNumber="521" type="nomenclature">
<paragraph id="8B8A36D80F35FF97FF1F62BDCA9550B6" blockId="22.[151,677,475,502]" box="[151,677,475,502]" pageId="22" pageNumber="521">
<heading id="D0C281B40F35FF97FF1F62BDCA9550B6" bold="true" box="[151,677,475,502]" fontSize="11" level="1" pageId="22" pageNumber="521" reason="1">
<emphasis id="B941EACA0F35FF97FF1F62BDCA9550B6" bold="true" box="[151,677,475,502]" pageId="22" pageNumber="521">
Subgenus 
<taxonomicName id="4C354D5B0F35FF97FE9F62BACA5350B6" ID-CoL="8357637c-3589-44f2-9358-bd2514039a1f" authority="Sharp, 1880 n" authorityName="Sharp" authorityYear="1880" box="[279,611,475,502]" class="Insecta" family="Dytiscidae" genus="Hyphoporus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="521" phylum="Arthropoda" rank="genus">
<emphasis id="B941EACA0F35FF97FE9F62BAC99C50B5" bold="true" box="[279,428,475,501]" italics="true" pageId="22" pageNumber="521">Hyphoporus</emphasis>
<bibRefCitation id="EFA44B290F35FF97FE3C62BDCA5350B6" author="Sharp" box="[436,611,476,502]" pageId="22" pageNumber="521" refString="Sharp, D. (1880) Avis preliminaire d'une nouvelle classification de la famille des Dytiscidae. Annales de la Societe Entomologique de Belgique, 23 (Comptes Rendus), cxlvii - cli." type="book chapter" year="1880" yearSuffix="n">Sharp, 1880 n</bibRefCitation>
</taxonomicName>
. stat.
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32F65530F35FF97FF1F6143C8CA53E2" pageId="22" pageNumber="521" type="materials_examined">
<paragraph id="8B8A36D80F35FF97FF1F6143CB035378" blockId="22.[151,819,546,568]" box="[151,819,546,568]" pageId="22" pageNumber="521">
<emphasis id="B941EACA0F35FF97FF1F6143C92C5378" bold="true" box="[151,284,546,568]" pageId="22" pageNumber="521">Type species</emphasis>
: 
<taxonomicName id="4C354D5B0F35FF97FEA26143CAAC5378" authority="Aube, 1838: 554" authorityName="Aube" authorityPageNumber="554" authorityYear="1838" box="[298,668,546,568]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="521" phylum="Arthropoda" rank="species" species="solieri">
<emphasis id="B941EACA0F35FF97FEA26143C9DB5378" box="[298,491,546,568]" italics="true" pageId="22" pageNumber="521">Hydroporus solieri</emphasis>
<bibRefCitation id="EFA44B290F35FF97FE796143CA5A5378" author="Aube" box="[497,618,546,568]" pageId="22" pageNumber="521" refString="Aube, C. (1838) Hydrocanthares et gyriniens. In: Dejean, P. F. M. A. Species general des coleopteres de la collection de M. le Comte Dejean. Vol. 6. Mequignon Pere et Fils, Paris, xvi + 804 pp." type="book" year="1838">Aubé, 1838</bibRefCitation>
: 554
</taxonomicName>
, by monotypy.
</paragraph>
<paragraph id="8B8A36D80F35FF97FF1F6104C8CA53E2" blockId="22.[151,1436,613,1322]" pageId="22" pageNumber="521">
<materialsCitation id="3B5D3C850F35FF97FF1F6104C8CA53E2" ID-GBIF-Occurrence="1608130917" location="Oriental" pageId="22" pageNumber="521" specimenCount="1" stateProvince="Oriental">
The subgenus 
<taxonomicName id="4C354D5B0F35FF97FEB46104C9F7533E" authorityName="Sharp" authorityYear="1880" box="[316,455,613,638]" class="Insecta" family="Dytiscidae" genus="Hyphoporus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="521" phylum="Arthropoda" rank="genus">
<emphasis id="B941EACA0F35FF97FEB46104C9F7533E" box="[316,455,613,638]" italics="true" pageId="22" pageNumber="521">Hyphoporus</emphasis>
</taxonomicName>
contains 19 species distributed in the 
<collectingRegion id="49F1F83A0F35FF97FCF66104CBEA533E" box="[894,986,613,638]" country="Egypt" name="Ash Sharqiyah" pageId="22" pageNumber="521">Oriental</collectingRegion>
and Palaearctic regions (see 
<figureCitation id="130E2A5D0F35FF97FAA06104CD5B533E" box="[1320,1387,613,638]" captionStart="FIGURE 2" captionStartId="12.[151,250,1941,1963]" captionTargetBox="[208,1378,197,1919]" captionTargetId="figure@12.[189,1396,193,1921]" captionTargetPageId="12" captionText="FIGURE 2. Dendrogram showing the new classification of Hygrotini. A number of species without molecular data (no voucher number) have been inserted near morphologically similar species with molecular data. Acronyms refer to biogeographical regions." httpUri="https://zenodo.org/record/884770/files/figure.png" pageId="22" pageNumber="521">Fig. 2</figureCitation>
and 
<tableCitation id="C6B703630F35FF97FF1F61E8C8D953E2" box="[151,233,649,674]" captionStart="TABLE 1" captionStartId="5.[125,200,512,532]" captionText="TABLE 1. Checklist of all 137 species of Hygrotini with original, new and former (Nilsson &amp; Há ek 2017 a) generic and subgeneric names; last three columns indicating new combinations, distribution and methods applied to material studied. See Villastrigo et al. (in press) for details on the specimens used in the molecular phylogeny and accession numbers of the sequences." pageId="22" pageNumber="521">Table 1</tableCitation>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32F65530F35FF97FF4F61CCCBB15512" pageId="22" pageNumber="521" type="diagnosis">
<paragraph id="8B8A36D80F35FF97FF4F61CCCBB15512" blockId="22.[151,1436,613,1322]" pageId="22" pageNumber="521">
<emphasis id="B941EACA0F35FF97FF4F61CCC9715386" bold="true" box="[199,321,685,710]" pageId="22" pageNumber="521">Diagnosis.</emphasis>
Body shape short oval (TL/MW ca. 1.6–1.7), rather globose; small to medium sized species (TL 
<quantity id="4CCD9B3D0F35FF97FF1F61B0C91253AA" box="[151,290,721,746]" metricMagnitude="-3" metricUnit="m" metricValue="4.55" metricValueMax="5.6" metricValueMin="3.5" pageId="22" pageNumber="521" unit="mm" value="4.55" valueMax="5.6" valueMin="3.5">3.5–5.6 mm</quantity>
) (see 
<figureCitation id="130E2A5D0F35FF97FEE461B0C98F53AA" box="[364,447,721,746]" captionStart="FIGURES 13 – 18" captionStartId="16.[151,264,1903,1925]" captionTargetBox="[157,1428,207,1855]" captionTargetId="figure@16.[151,1436,193,1882]" captionTargetPageId="16" captionText="FIGURES 13 – 18. Habitus of: (13) Hygrotus (Leptolambus) impressopunctatus, (14) H. (Leptolambus) orthogrammus, (15) H. (Leptolambus) obscureplagiatus, (16) H. (s. str.) guineensis, (17) H. (s. str.) descarpentriesi (male syntype and labels) and (18) H. (Hyphoporus) solieri (Figs 14 and 18 are reproduced from Fery et al. 2012 with the permission of F. Gusenleitner, Linz, Austria)." httpUri="https://zenodo.org/record/884778/files/figure.png" pageId="22" pageNumber="521">Fig. 18</figureCitation>
for 
<taxonomicName id="4C354D5B0F35FF97FE7C61B3CB3853AA" authorityName="Aube" authorityYear="1838" box="[500,776,721,746]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="521" phylum="Arthropoda" rank="species" species="solieri" subGenus="Hyphoporus">
<emphasis id="B941EACA0F35FF97FE7C61B3CA2053A9" box="[500,528,722,745]" italics="true" pageId="22" pageNumber="521">H.</emphasis>
(
<emphasis id="B941EACA0F35FF97FDAB61B0CA9D53AA" box="[547,685,721,746]" italics="true" pageId="22" pageNumber="521">Hyphoporus</emphasis>
) 
<emphasis id="B941EACA0F35FF97FD4961B0CB3853AA" box="[705,776,721,746]" italics="true" pageId="22" pageNumber="521">solieri</emphasis>
</taxonomicName>
). Head with anterior clypeal margin truncate, straight or slightly emarginated, border not produced forwards; bead continuous, middle part narrowed in most species (see 
<figureCitation id="130E2A5D0F35FF97FF1F6078C8D65272" box="[151,230,793,818]" captionStart="FIGURES 41 – 48" captionStartId="20.[151,264,1912,1934]" captionTargetBox="[151,1436,193,1890]" captionTargetId="figure@20.[151,1436,193,1890]" captionTargetPageId="20" captionText="FIGURES 41 – 48. Head with details of clypeal rim of (41) Hygrotus (Leptolambus) impressopunctatus, (42) H. (Hyphoporus) solieri, (43) H. (s. str.) guineensis, (44) H. (s. str.) inaequalis, (45) idem in lateral view, (46) H. (Coelambus) confluens (the lighter area before the anterior border of the clypeus is the &quot; clypeal stripe &quot;; see Appendix), (47) H. (Leptolambus) masculinus and (48) idem in lateral view." httpUri="https://zenodo.org/record/884784/files/figure.png" pageId="22" pageNumber="521">Fig. 42</figureCitation>
for 
<taxonomicName id="4C354D5B0F35FF97FE9D607BCA135272" authorityName="Aube" authorityYear="1838" box="[277,547,793,818]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="521" phylum="Arthropoda" rank="species" species="solieri" subGenus="Hyphoporus">
<emphasis id="B941EACA0F35FF97FE9D607BC9195271" box="[277,297,794,817]" italics="true" pageId="22" pageNumber="521">H</emphasis>
. (
<emphasis id="B941EACA0F35FF97FEC96078C9FC5272" box="[321,460,793,818]" italics="true" pageId="22" pageNumber="521">Hyphoporus</emphasis>
) 
<emphasis id="B941EACA0F35FF97FE546078CA135272" box="[476,547,793,818]" italics="true" pageId="22" pageNumber="521">solieri</emphasis>
</taxonomicName>
); (eXcept 
<taxonomicName id="4C354D5B0F35FF97FD1D607BCCA55272" authority="(Severin, 1890)" authorityName="Fery" authorityYear="2017" baseAuthorityName="Severin" baseAuthorityYear="1890" box="[661,1173,793,818]" class="Insecta" family="Dytiscidae" genus="Hygrotus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="521" phylum="Arthropoda" rank="species" species="bengalensis" subGenus="Hyphoporus">
<emphasis id="B941EACA0F35FF97FD1D607BCA815271" box="[661,689,794,817]" italics="true" pageId="22" pageNumber="521">H.</emphasis>
(
<emphasis id="B941EACA0F35FF97FD486078CB7B5272" box="[704,843,793,818]" italics="true" pageId="22" pageNumber="521">Hyphoporus</emphasis>
) 
<emphasis id="B941EACA0F35FF97FCD36078CBEF5272" box="[859,991,793,818]" italics="true" pageId="22" pageNumber="521">bengalensis</emphasis>
(Severin, 1890)
</taxonomicName>
with continuous bead). Antennomeres simple, not broadened. Elytra with margin in lateral view moderately ascending to shoulder (similar to 
<figureCitation id="130E2A5D0F35FF97FF336000C924523A" box="[187,276,865,890]" captionStart="FIGURES 49 – 55" captionStartId="21.[151,264,1345,1367]" captionTargetBox="[191,1436,208,1316]" captionTargetId="figure@21.[151,1436,193,1323]" captionTargetPageId="21" captionText="FIGURES 49 – 55. (49 – 53): Epipleuron with oblique epipleural carina and genicular fossa of: (49) Clemnius (s. str.) berneri, (50) Hygrotus (Leptolambus) polonicus polonicus, (51) H. (Coelambus) caspius, (52) H. (Leptolambus) impressopunctatus and (53) Rhithrodytes agnus; epipleural carina (c) and inner margin of epipleuron (m) including angle mentioned in diagnoses of subgenera. (54 – 55): Elytral margin in lateral view of: (54) H. (Coelambus) caspius (strongly ascending to shoulder) and (55) Clemnius (s. str.) berneri (moderately ascending to shoulder) (Figs 52 and 53 are reproduced from Fery 2013 and from Fery 2016 with the permissions of F. Gusenleitner, Linz, Austria, and G. Foster, Ayr, UK, respectively)." httpUri="https://zenodo.org/record/884786/files/figure.png" pageId="22" pageNumber="521">Fig. 55</figureCitation>
); epipleuron comparably broad, broader than mesotibia distally; carina meeting inner margin of epipleuron forming a comparably small angle (&lt;ca. 135°; similar to 
<figureCitation id="130E2A5D0F35FF97FC3760E4CC2552DE" box="[959,1045,901,926]" captionStart="FIGURES 49 – 55" captionStartId="21.[151,264,1345,1367]" captionTargetBox="[191,1436,208,1316]" captionTargetId="figure@21.[151,1436,193,1323]" captionTargetPageId="21" captionText="FIGURES 49 – 55. (49 – 53): Epipleuron with oblique epipleural carina and genicular fossa of: (49) Clemnius (s. str.) berneri, (50) Hygrotus (Leptolambus) polonicus polonicus, (51) H. (Coelambus) caspius, (52) H. (Leptolambus) impressopunctatus and (53) Rhithrodytes agnus; epipleural carina (c) and inner margin of epipleuron (m) including angle mentioned in diagnoses of subgenera. (54 – 55): Elytral margin in lateral view of: (54) H. (Coelambus) caspius (strongly ascending to shoulder) and (55) Clemnius (s. str.) berneri (moderately ascending to shoulder) (Figs 52 and 53 are reproduced from Fery 2013 and from Fery 2016 with the permissions of F. Gusenleitner, Linz, Austria, and G. Foster, Ayr, UK, respectively)." httpUri="https://zenodo.org/record/884786/files/figure.png" pageId="22" pageNumber="521">Fig. 50</figureCitation>
). Elytra with interrupted vittae or dotted (in some species very darkened); venter black or brown. Last abdominal ventrite without deep depression. Aedeagus with median lobe robust, asymmetric (see 
<figureCitation id="130E2A5D0F35FF97FD7D60ACCB7652A6" box="[757,838,973,998]" captionStart="FIGURES 19 – 28" captionStartId="17.[151,264,1527,1549]" captionTargetBox="[158,1432,199,1502]" captionTargetId="figure@17.[151,1436,193,1506]" captionTargetPageId="17" captionText="FIGURES 19 – 28. Median lobe in ventral and lateral view of: (19) Clemnius (Cyclopius) acaroides, (20) Clemnius (s. str.) berneri, (21) Clemnius (s. str.) decoratus, (22) Hygrotus (s. str.) inaequalis, (23) H. (Coelambus) enneagrammus, (24) H. (Coelambus) confluens, (25) H. (Leptolambus) parallellogrammus, (26) H. (Leptolambus) impressopunctatus, (27) H. (s. str.) guineensis and (28) H. (Hyphoporus) tonkinensis (Fig. 28 is a modified reproduction of figs 3 and 4 in Brancucci &amp; Biström 2013)." httpUri="https://zenodo.org/record/884780/files/figure.png" pageId="22" pageNumber="521">Fig. 28</figureCitation>
for 
<taxonomicName id="4C354D5B0F35FF97FCF060AFC8EB554A" authority="(Regimbart, 1899)" authorityName="Fery" authorityYear="2017" baseAuthorityName="Regimbart" baseAuthorityYear="1899" class="Insecta" family="Dytiscidae" genus="Hygrotus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="521" phylum="Arthropoda" rank="species" species="tonkinensis" subGenus="Hyphoporus">
<emphasis id="B941EACA0F35FF97FCF060AFCBEF52A5" box="[888,991,974,997]" italics="true" pageId="22" pageNumber="521">Hygrotus</emphasis>
(
<emphasis id="B941EACA0F35FF97FC7A60ACCC4C52A6" box="[1010,1148,973,998]" italics="true" pageId="22" pageNumber="521">Hyphoporus</emphasis>
) 
<emphasis id="B941EACA0F35FF97FB0660ACCD3C52A6" box="[1166,1292,973,998]" italics="true" pageId="22" pageNumber="521">tonkinensis</emphasis>
(Régimbart, 1899)
</taxonomicName>
; in 
<taxonomicName id="4C354D5B0F35FF97FE846093C987554A" authorityName="Fery" authorityYear="2017" baseAuthorityName="Severin" baseAuthorityYear="1890" box="[268,439,1009,1034]" class="Insecta" family="Dytiscidae" genus="Hygrotus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="521" phylum="Arthropoda" rank="species" species="bengalensis">
<emphasis id="B941EACA0F35FF97FE846093C987554A" box="[268,439,1009,1034]" italics="true" pageId="22" pageNumber="521">H. bengalensis</emphasis>
</taxonomicName>
median lobe more or less symmetric; see 
<bibRefCitation id="EFA44B290F35FF97FC3A6090CC63554A" author="Vazirani" box="[946,1107,1009,1034]" pageId="22" pageNumber="521" refString="Vazirani, T. G. (1969) Contribution to the study of aquatic beetles (Coleoptera). V. Revision of Indian species of Hyphoporus Sharp (Dysticidae). Bulletin du Museum National d'Histoire Naturelle, Series 2, 41, 203 - 225." type="journal article" year="1969">Vazirani 1969</bibRefCitation>
for several further figures); parameres with condylar process of diverse length, forming an obtuse angle with distal part (see 
<figureCitation id="130E2A5D0F35FF97FB736774CD7D556E" box="[1275,1357,1045,1070]" captionStart="FIGURES 29 – 40" captionStartId="18.[151,264,1665,1687]" captionTargetBox="[151,1430,221,1637]" captionTargetId="figure@18.[144,1443,193,1643]" captionTargetPageId="18" captionText="FIGURES 29 – 40. Left paramere of: (29) Clemnius (Cyclopius) acaroides, (30) Clemnius (s. str.) berneri, (31) Clemnius (s. str.) decoratus, (32) Hygrotus (s. str.) inaequalis, (33) H. (Leptolambus) fresnedai, (34) H. (Leptolambus) impressopunctatus, (35) H. (Leptolambus) parallellogrammus, (36) H. (Leptolambus) nubilus, (37) H. (Coelambus) enneagrammus, (38) H. (Coelambus) confluens, (39) H. (s. str.) guineensis and (40) H. (Hyphoporus) tonkinensis (Fig. 40 is a modified reproduction of fig. 5 in Brancucci &amp; Biström 2013)." httpUri="https://zenodo.org/record/884782/files/figure.png" pageId="22" pageNumber="521">Fig. 40</figureCitation>
for 
<taxonomicName id="4C354D5B0F35FF97FA096777C9895512" authorityName="Fery" authorityYear="2017" baseAuthorityName="Regimbart" baseAuthorityYear="1899" class="Insecta" family="Dytiscidae" genus="Hygrotus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="521" phylum="Arthropoda" rank="species" species="tonkinensis" subGenus="Hyphoporus">
<emphasis id="B941EACA0F35FF97FA096777CDAC556D" box="[1409,1436,1046,1069]" italics="true" pageId="22" pageNumber="521">H.</emphasis>
(
<emphasis id="B941EACA0F35FF97FF286758C91B5512" box="[160,299,1081,1106]" italics="true" pageId="22" pageNumber="521">Hyphoporus</emphasis>
) 
<emphasis id="B941EACA0F35FF97FEB36758C9895512" box="[315,441,1081,1106]" italics="true" pageId="22" pageNumber="521">tonkinensis</emphasis>
</taxonomicName>
). Male metatarsal claws of equal length.
</paragraph>
</subSubSection>
<subSubSection id="C32F65530F35FF97FF4F673CCB46546A" pageId="22" pageNumber="521" type="distribution">
<paragraph id="8B8A36D80F35FF97FF4F673CCAC355DA" blockId="22.[151,1436,613,1322]" pageId="22" pageNumber="521">
<emphasis id="B941EACA0F35FF97FF4F673CC9505536" bold="true" box="[199,352,1117,1142]" pageId="22" pageNumber="521">Distribution.</emphasis>
Palaearctic and 
<collectingRegion id="49F1F83A0F35FF97FD95673CCA4B5536" box="[541,635,1117,1142]" country="Egypt" name="Ash Sharqiyah" pageId="22" pageNumber="521">Oriental</collectingRegion>
, from 
<collectingCountry id="F32276480F35FF97FD4F673FCAC45535" box="[711,756,1118,1141]" name="Iran" pageId="22" pageNumber="521">Iran</collectingCountry>
to 
<collectingCountry id="F32276480F35FF97FC93673CCB645536" box="[795,852,1117,1142]" name="India" pageId="22" pageNumber="521">India</collectingCountry>
and south-east Asia; one species– 
<taxonomicName id="4C354D5B0F35FF97FB53673FC8EF55DA" authorityName="Aube" authorityYear="1838" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="521" phylum="Arthropoda" rank="species" species="solieri" subGenus="Hyphoporus">
<emphasis id="B941EACA0F35FF97FB53673FCCC75535" box="[1243,1271,1118,1141]" italics="true" pageId="22" pageNumber="521">H.</emphasis>
(
<emphasis id="B941EACA0F35FF97FA80673CCDA35536" box="[1288,1427,1117,1142]" italics="true" pageId="22" pageNumber="521">Hyphoporus</emphasis>
) 
<emphasis id="B941EACA0F35FF97FF1F67E0C8EF55DA" box="[151,223,1153,1178]" italics="true" pageId="22" pageNumber="521">solieri</emphasis>
</taxonomicName>
–from 
<collectingCountry id="F32276480F35FF97FEA267E3C96755D9" box="[298,343,1154,1177]" name="Iran" pageId="22" pageNumber="521">Iran</collectingCountry>
to the Arabian Peninsula and 
<collectingCountry id="F32276480F35FF97FD2067E3CADF55DA" box="[680,751,1154,1178]" name="Egypt" pageId="22" pageNumber="521">Egypt</collectingCountry>
.
</paragraph>
<paragraph id="8B8A36D80F35FF97FF4F67C4CB46546A" blockId="22.[151,1436,613,1322]" pageId="22" pageNumber="521">
<emphasis id="B941EACA0F35FF97FF4F67C4C98255FE" bold="true" box="[199,434,1189,1214]" pageId="22" pageNumber="521">Main habitat types.</emphasis>
There is no information on the habitat of most of the species of the subgenus, although they are likely to be mostly associated with freshwater environments. However, it shall be mentioned that 
<bibRefCitation id="EFA44B290F35FF97FAD167A8C93C5446" author="Hajek" pageId="22" pageNumber="521" refString="Hajek, J. (2006) The westernmost record of Neptosternus circumductus, and a review of Dytiscidae (Coleoptera) of Baluchistan (Iran, Pakistan). Acta Entomologica Musei Nationalis Pragae, 46, 43 - 56." type="journal article" year="2006">Hájek (2006: 48)</bibRefCitation>
illustrated a rest-pool in a wadi (Pir Sohrab, Iran) where he has collected 
<taxonomicName id="4C354D5B0F35FF97FBC1678FC8E8546A" authority="(Sharp, 1882)" authorityName="Fery" authorityYear="2017" baseAuthorityName="Sharp" baseAuthorityYear="1882" class="Insecta" family="Dytiscidae" genus="Hygrotus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="521" phylum="Arthropoda" rank="species" species="aper" subGenus="Hyphoporus">
<emphasis id="B941EACA0F35FF97FBC1678FCC6D5445" box="[1097,1117,1262,1285]" italics="true" pageId="22" pageNumber="521">H</emphasis>
. (
<emphasis id="B941EACA0F35FF97FBFD678CCD305446" box="[1141,1280,1261,1286]" italics="true" pageId="22" pageNumber="521">Hyphoporus</emphasis>
) 
<emphasis id="B941EACA0F35FF97FA98678FCD745445" box="[1296,1348,1262,1285]" italics="true" pageId="22" pageNumber="521">aper</emphasis>
(
<bibRefCitation id="EFA44B290F35FF97FADA678CC8E0546A" author="Sharp" pageId="22" pageNumber="521" refString="Sharp, D. (1882) On aquatic carnivorous Coleoptera or Dytiscidae. The Scientific Transactions of the Royal Dublin Society, 2, 179 - 1003, pls. 7 - 18." type="journal article" year="1882">Sharp, 1882</bibRefCitation>
)
</taxonomicName>
together with 
<taxonomicName id="4C354D5B0F35FF97FEF36673CB41546A" authority="Regimbart, 1899" authorityName="Regimbart" authorityYear="1899" box="[379,881,1297,1322]" class="Insecta" family="Dytiscidae" genus="Neptosternus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="521" phylum="Arthropoda" rank="species" species="circumductus">
<emphasis id="B941EACA0F35FF97FEF36673CA9A546A" box="[379,682,1297,1322]" italics="true" pageId="22" pageNumber="521">Neptosternus circumductus</emphasis>
Régimbart, 1899
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>