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<document id="CCE5F378E0D620D387788864FAD33571" ID-CLB-Dataset="29447" ID-DOI="10.11646/zootaxa.4472.2.8" ID-GBIF-Dataset="afe65a87-5307-4488-8450-89036d9e018e" ID-ISSN="1175-5326" ID-Zenodo-Dep="1440241" ID-ZooBank="B17E81FB-C118-494D-A4A9-A57F291EA24F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1538372220167" checkinUser="plazi" docAuthor="Bartlett, Charles R., Passos, Eliana Maria Dos, Silva, Flaviana Gonçalves Da, Diniz, Leandro Eugenio Cardamone &amp; Dollet, Michel" docDate="2018" docId="039C7735AC6FFFC047C8F9CBFC98BBA7" docLanguage="en" docName="zootaxa.4472.2.8.pdf" docOrigin="Zootaxa 4472 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Oecleus Stal 1862" docType="treatment" docVersion="7" lastPageNumber="360" masterDocId="FFA50F4DAC6EFFC2475FFFFFFFFDB90F" masterDocTitle="A new species of Oecleus Stål (Hemiptera: Fulgoroidea: Cixiidae) from coconut in Brazil" masterLastPageNumber="364" masterPageNumber="358" pageNumber="359" updateTime="1698559022165" updateUser="ExternalLinkService">
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<mods:title id="DAB7623131509A2C701FEB83851AA348">A new species of Oecleus Stål (Hemiptera: Fulgoroidea: Cixiidae) from coconut in Brazil</mods:title>
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<mods:namePart id="7675B1BDA9A73CD106052744CBD3FCAB">Bartlett, Charles R.</mods:namePart>
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<mods:namePart id="D07ADD39411BEDAE54ABA22A9497FBBC">Passos, Eliana Maria Dos</mods:namePart>
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<mods:namePart id="2A1FA789233632EA8CA4CCAE725080D3">Silva, Flaviana Gonçalves Da</mods:namePart>
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<mods:namePart id="0EA873D98768B6785E7F6F95C895F5DF">Diniz, Leandro Eugenio Cardamone</mods:namePart>
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<mods:namePart id="E16E8980A445F1D301920BEA787EC33B">Dollet, Michel</mods:namePart>
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<treatment id="039C7735AC6FFFC047C8F9CBFC98BBA7" ID-DOI="http://doi.org/10.5281/zenodo.5967730" ID-GBIF-Taxon="148421371" ID-Zenodo-Dep="5967730" LSID="urn:lsid:plazi:treatment:039C7735AC6FFFC047C8F9CBFC98BBA7" httpUri="http://treatment.plazi.org/id/039C7735AC6FFFC047C8F9CBFC98BBA7" lastPageId="2" lastPageNumber="360" pageId="1" pageNumber="359">
<subSubSection id="C32F95A8AC6FFFC347C8F9CBFDEFBF41" box="[151,530,1587,1614]" pageId="1" pageNumber="359" type="nomenclature">
<paragraph id="8B8AC623AC6FFFC347C8F9CBFDEFBF41" blockId="1.[151,530,1587,1614]" box="[151,530,1587,1614]" pageId="1" pageNumber="359">
<heading id="D0C2714FAC6FFFC347C8F9CBFDEFBF41" bold="true" box="[151,530,1587,1614]" fontSize="11" level="1" pageId="1" pageNumber="359" reason="1">
<treatmentCitationGroup id="AB25E10DAC6FFFC347C8F9CBFDEFBF41" box="[151,530,1587,1614]" pageId="1" pageNumber="359">
<emphasis id="B9411A31AC6FFFC347C8F9CBFDEFBF41" bold="true" box="[151,530,1587,1614]" pageId="1" pageNumber="359">
Genus
<taxonomicName id="4C35BDA0AC6FFFC347B1F9CCFDF0BF41" ID-CoL="675D" ID-ENA="390100" authority="Stal, 1862: 306" authorityName="Stal" authorityPageNumber="306" authorityYear="1862" box="[238,525,1587,1614]" class="Insecta" family="Cixiidae" genus="Oecleus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="359" phylum="Arthropoda" rank="genus">
<emphasis id="B9411A31AC6FFFC347B1F9CCFEB3BF42" bold="true" box="[238,334,1587,1613]" italics="true" pageId="1" pageNumber="359">Oecleus</emphasis>
<treatmentCitation id="0A94E032AC6FFFC3460AF9CBFDF0BF41" author="" box="[341,525,1588,1614]" page="306" pageId="1" pageNumber="359">
<bibRefCitation id="EFA4BBD2AC6FFFC3460AF9CBFE2FBF41" box="[341,466,1588,1614]" pageId="1" pageNumber="359" refId="ref4248" refString="Stal, C. (1862) Novae vel minus cognitae Homopterorum formae et species. Berliner Entomologische Zeitschrift, 6, 303 - 315. https: // doi. org / 10.1002 / mmnd. 47918620303" type="journal article">Stål, 1862</bibRefCitation>
: 306
</treatmentCitation>
</taxonomicName>
.
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</paragraph>
</subSubSection>
<subSubSection id="C32F95A8AC6FFFC047C8F983FC80B888" lastPageId="2" lastPageNumber="360" pageId="1" pageNumber="359" type="materials_examined">
<paragraph id="8B8AC623AC6FFFC347C8F983FD7CBF9E" blockId="1.[151,641,1659,1681]" box="[151,641,1659,1681]" pageId="1" pageNumber="359">
Type species
<taxonomicName id="4C35BDA0AC6FFFC34640F984FD80BF9E" authority="Stal, 1862: 307" authorityName="Stal" authorityPageNumber="307" authorityYear="1862" box="[287,637,1659,1681]" class="Insecta" family="Cixiidae" genus="Oecleus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="359" phylum="Arthropoda" rank="species" species="seminiger">
<emphasis id="B9411A31AC6FFFC34640F984FE26BF9E" box="[287,475,1659,1681]" italics="true" pageId="1" pageNumber="359">Oecleus seminiger</emphasis>
<bibRefCitation id="EFA4BBD2AC6FFFC346BEF984FDB7BF9E" box="[481,586,1659,1681]" pageId="1" pageNumber="359" refId="ref4248" refString="Stal, C. (1862) Novae vel minus cognitae Homopterorum formae et species. Berliner Entomologische Zeitschrift, 6, 303 - 315. https: // doi. org / 10.1002 / mmnd. 47918620303" type="journal article">Stål, 1862</bibRefCitation>
: 307
</taxonomicName>
.
</paragraph>
<paragraph id="8B8AC623AC6FFFC047C8F942FC80B888" blockId="1.[151,1437,1725,2038]" lastBlockId="2.[151,1436,151,680]" lastPageId="2" lastPageNumber="360" pageId="1" pageNumber="359">
<emphasis id="B9411A31AC6FFFC347C8F942FE6BBFD9" bold="true" box="[151,406,1725,1750]" pageId="1" pageNumber="359">Diagnostic summary</emphasis>
(modified from
<bibRefCitation id="EFA4BBD2AC6FFFC34539F941FCFABFD9" box="[614,775,1726,1750]" pageId="1" pageNumber="359" refId="ref3806" refString="Kramer, J. P. (1977) Taxonomic study of the planthopper genus Oecleus in the United States (Homoptera: Fulgoroidea: Cixiidae). Transactions of the American Entomological Society, 103, 379 - 449." type="journal article">Kramer 1977</bibRefCitation>
). Small to midsize (
<quantity id="4CCD6BC6AC6FFFC3435BF942FB64BFD9" box="[1028,1177,1725,1750]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" metricValueMax="8.5" metricValueMin="3.3" pageId="1" pageNumber="359" unit="mm" value="5.9" valueMax="8.5" valueMin="3.3">3.3–8.5 mm</quantity>
); head narrower than pronotum (in dorsal view) with eyes large. Vertex narrow, trough-like and parallel-sided, lateral and anterior margin carinate. Slightly raised; proximally narrowed and distally produced beyond eyes for a variable distance. In lateral view, apex of head acutely or obtusely angled, eyes emarginate, ocellus present under each eye and near midline above frontoclypeal suture. In frontal view, frons elongated and narrowed toward vertex, carina on midline of frons present (sometimes obsolete). Clypeus triangular or subtriangular. Antennae originating from large socket, scape reduced and collar-like, pedicel rounded with sensoria, flagellum beadlike basally and filamentous distally. Pronotum collar-like with irregular ridges, narrowest on midline, indented on posterior margin, carinate on posterior and lateral margins. Mesonotum longer at midline than combined lengths of vertex and pronotum, mesonotum flattened, usually bearing five longitudinal carinae, sometimes carinae flanking central carina absent or reduced to pigmented lines. Hind tibiae without lateral spines. Forewings clear, rarely with color pattern, veins usually dotted with dark pustules, often bearing setae. Male terminalia with pygofer longest on ventral margin, anal tube elongated, paired slender symmetrical gonostyli (‘styles’ of
<bibRefCitation id="EFA4BBD2AC6CFFC04423FEFAFBE9B813" box="[892,1044,260,284]" pageId="2" pageNumber="360" refId="ref3806" refString="Kramer, J. P. (1977) Taxonomic study of the planthopper genus Oecleus in the United States (Homoptera: Fulgoroidea: Cixiidae). Transactions of the American Entomological Society, 103, 379 - 449." type="journal article">Kramer 1977</bibRefCitation>
) flanking median lobe of pygofer; gonostyli and basal portion of asymmetrical aedeagus articulated by vertical connective. Aedeagus consisting of a ridged basal phallotheca (periandrium, “shaft” of
<bibRefCitation id="EFA4BBD2AC6CFFC045E0FEB2FCA9B86B" box="[703,852,332,356]" pageId="2" pageNumber="360" refId="ref3806" refString="Kramer, J. P. (1977) Taxonomic study of the planthopper genus Oecleus in the United States (Homoptera: Fulgoroidea: Cixiidae). Transactions of the American Entomological Society, 103, 379 - 449." type="journal article">Kramer 1977</bibRefCitation>
) and distal, movable retrorse flagellum (endosoma); phallotheca with 1 to 3 processes, flagellum with 0 to 2 processes.
</paragraph>
</subSubSection>
<subSubSection id="C32F95A8AC6CFFC04798FE6BFC98BBA7" pageId="2" pageNumber="360" type="discussion">
<paragraph id="8B8AC623AC6CFFC04798FE6BFC98BBA7" blockId="2.[151,1436,151,680]" pageId="2" pageNumber="360">
<emphasis id="B9411A31AC6CFFC04798FE6BFEC6B8A2" bold="true" box="[199,315,404,429]" pageId="2" pageNumber="360">Remarks.</emphasis>
<taxonomicName id="4C35BDA0AC6CFFC0461DFE6BFE61B8A3" box="[322,412,404,428]" class="Insecta" family="Cixiidae" genus="Oecleus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="360" phylum="Arthropoda" rank="genus">
<emphasis id="B9411A31AC6CFFC0461DFE6BFE61B8A3" box="[322,412,404,428]" italics="true" pageId="2" pageNumber="360">Oecleus</emphasis>
</taxonomicName>
is distinctive and can be separated from all other
<taxonomicName id="4C35BDA0AC6CFFC04494FE6BFBD7B8A2" box="[971,1066,404,429]" class="Insecta" family="Cixiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="360" phylum="Arthropoda" rank="family">Cixiidae</taxonomicName>
in our fauna by characters of the head in dorsal view. The vertex, is greatly narrowed and trough-like, mainly parallel sided but narrower proximally than apically and is separated from the front by a short transverse carina. These features separate
<taxonomicName id="4C35BDA0AC6CFFC043BFFE23FAC7B8FB" box="[1248,1338,476,500]" class="Insecta" family="Cixiidae" genus="Oecleus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="360" phylum="Arthropoda" rank="genus">
<emphasis id="B9411A31AC6CFFC043BFFE23FAC7B8FB" box="[1248,1338,476,500]" italics="true" pageId="2" pageNumber="360">Oecleus</emphasis>
</taxonomicName>
from all genera in
<taxonomicName id="4C35BDA0AC6CFFC0464BFE00FE8EBB17" authorityName="Muir" authorityYear="1922" box="[276,371,511,536]" class="Insecta" family="Cixiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="360" phylum="Arthropoda" rank="tribe" tribe="Oecleini">Oecleini</taxonomicName>
except possibly
<taxonomicName id="4C35BDA0AC6CFFC0451AFDFFFD2ABB17" authorityName="Fowler" authorityYear="1904" box="[581,727,512,536]" class="Insecta" family="Cixiidae" genus="Rhamphixius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="360" phylum="Arthropoda" rank="genus">
<emphasis id="B9411A31AC6CFFC0451AFDFFFD2ABB17" box="[581,727,512,536]" italics="true" pageId="2" pageNumber="360">Rhamphixius</emphasis>
</taxonomicName>
, which (e.g.) has a greatly projected vertex, and possibly
<taxonomicName id="4C35BDA0AC6CFFC047C8FDDBFEF7BB33" authorityName="Myers" authorityYear="1928" box="[151,266,548,572]" class="Insecta" family="Cixiidae" genus="Antillixius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="360" phylum="Arthropoda" rank="genus">
<emphasis id="B9411A31AC6CFFC047C8FDDBFEF7BB33" box="[151,266,548,572]" italics="true" pageId="2" pageNumber="360">Antillixius</emphasis>
</taxonomicName>
.
<emphasis id="B9411A31AC6CFFC04644FDDBFE68BB33" box="[283,405,548,572]" italics="true" pageId="2" pageNumber="360">
<taxonomicName id="4C35BDA0AC6CFFC04644FDDBFE6CBB33" authorityName="Myers" authorityYear="1928" box="[283,401,548,572]" class="Insecta" family="Cixiidae" genus="Antillixius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="360" phylum="Arthropoda" rank="genus">Antillixius</taxonomicName>
,
</emphasis>
known only from
<collectingCountry id="F32286B3AC6CFFC0452CFDDBFD4CBB32" box="[627,689,548,573]" name="Cuba" pageId="2" pageNumber="360">Cuba</collectingCountry>
, which has a sharply pointed head (in lateral view), vertex very deeply trough-like, with the lips of the trough contiguous all the way between the eyes, 3 carinae on the mesonotum, veins of wings without granulations (
<bibRefCitation id="EFA4BBD2AC6CFFC04595FD93FCAABB8B" box="[714,855,620,644]" pageId="2" pageNumber="360" refId="ref4083" refString="Myers J. G. (1928) Notes on Cuban fulgoroid Homoptera. Studies on Cuban insects, 1928, 13 - 28 + 2 unpaginated figure pages." type="journal article">Myers 1928</bibRefCitation>
).
<taxonomicName id="4C35BDA0AC6CFFC04434FD93FC02BB8B" authorityName="Van Duzee" authorityYear="1923" box="[875,1023,620,644]" class="Insecta" family="Cixiidae" genus="Nymphocixia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="360" phylum="Arthropoda" rank="genus">
<emphasis id="B9411A31AC6CFFC04434FD93FC02BB8B" box="[875,1023,620,644]" italics="true" pageId="2" pageNumber="360">Nymphocixia</emphasis>
</taxonomicName>
and
<taxonomicName id="4C35BDA0AC6CFFC0436AFD93FB15BB8B" authorityName="Emeljanov" authorityYear="2007" box="[1077,1256,620,644]" class="Insecta" family="Cixiidae" genus="Nymphomyndus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="360" phylum="Arthropoda" rank="genus">
<emphasis id="B9411A31AC6CFFC0436AFD93FB15BB8B" box="[1077,1256,620,644]" italics="true" pageId="2" pageNumber="360">Nymphomyndus</emphasis>
</taxonomicName>
have the vertex narrow, but the head appears broadly rounded from lateral view.
</paragraph>
</subSubSection>
</treatment>
</document>
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