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<mods:title id="1632EFF5BF8774AEBEBA8DDD35995B0E">Phylogeny of the Troglomorphic Scorpion Genus Troglotayosicus (Scorpiones: Troglotayosicidae) with Description of a New Species from Colombia</mods:title>
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<mods:namePart id="9F07A60F74109D71894A30AAD5DE87FE">Moreno-González, Jairo A.</mods:namePart>
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KEY TO IDENTIFICATION OF THE SPECIES OF 
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<emphasis id="63BB4E28003EFF84A5E5704DFC41FBB4" box="[708,915,1034,1060]" italics="true" pageId="9" pageNumber="10">TROGLOTAYOsicus</emphasis>
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1. Lateral eyespot absent (g. 5C, F); pedipalp chela trichobothrium 
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(
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) situated distal to trichobothrium 
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(
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) (gs. 8C, D, 9D) and trichobothrium 
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(
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) situated midway between trichobothrium 
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(
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) and base of xed nger (gs. 8G, H, 9H)............ 
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, 
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– Lateral eyespot present (
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, g. 4B); pedipalp chela trichobothrium 
<emphasis id="63BB4E28003EFF84A38970F1FB61FB5D" box="[1192,1203,1206,1229]" italics="true" pageId="9" pageNumber="10">e</emphasis>
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(
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) situated proximal to trichobothrium 
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(
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) (
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, 13, gs. 8B, 9B) and trichobothrium 
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(
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) situated closer to base of xed nger than to trichobothrium 
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(
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) (
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, 13, gs. 8B, 9D)..........................................2
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2. 
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carinae present, complete (
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, g. 10D–F)......3
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– 
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carinae absent; VSM and VL carinae vestigial, reduced to few granules at anteroventral margin of segment (
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, gs. 23, 24)............................5
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3. Pedipalp patella trichobothrium 
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situated distal to trichobothrium 
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(
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, g. 7C–H); leg III basitarsus, proventral row of macrosetae and leg IV basitarsus, retroventral row of macrosetae present............................................... 
<taxonomicName id="96CFE9B9003EFF84A37A7198FB6EFA66" ID-CoL="992QV" box="[1115,1212,1502,1526]" class="Arachnida" family="Troglotayosicidae" genus="Troglotayosicus" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="ballvei">
<emphasis id="63BB4E28003EFF84A37A7198FB6EFA66" box="[1115,1212,1502,1526]" italics="true" pageId="9" pageNumber="10">T. ballvei</emphasis>
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– Pedipalp patella trichobothrium 
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aligned with trichobothrium 
<emphasis id="63BB4E28003EFF84A44571B8FC57F987" box="[868,901,1535,1559]" italics="true" pageId="9" pageNumber="10">esb</emphasis>
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(
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, g. 7B, D, F); leg III basitarsus, proventral row of macrosetae and leg IV basitarsus, retroventral row of macrosetae absent..................................................................4
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4. Leg III basitarsus, retroventral row of macrosetae present (obsolete, represented by one or two macrosetae), telotarsus ventral macrosetae arranged into pair of distinct ventrosubmedian rows; metasomal segment III with VSM and VL carinae sparsely granular (
<bibRefCitation id="355EEFCB003EFF84A4B872E2FB5DF92D" author="Ochoa, J. A. &amp; R. Botero-Trujillo &amp; L. Prendini" box="[921,1167,1701,1725]" pageId="9" pageNumber="10" pagination="1 - 19" refId="ref11537" refString="Ochoa, J. A., R. Botero-Trujillo, and L. Prendini. 2010. On the troglomorphic scorpion Troglotayosicus humiculum (Scorpiones, Troglotayosicidae), with rst description of the adults. American Museum Novitates 3691: 1 - 19." type="journal article" year="2010">Ochoa et al., 2010: 12</bibRefCitation>
, 13, gs. 9C, 10C)............................................................. 
<taxonomicName id="96CFE9B9003EFF84A3077280FB6EF94E" ID-CoL="5999P" authorityName="Botero-Trujillo and Francke" authorityYear="2009" box="[1062,1212,1734,1758]" class="Arachnida" family="Troglotayosicidae" genus="Troglotayosicus" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="humiculum">
<emphasis id="63BB4E28003EFF84A3077280FB6EF94E" box="[1062,1212,1734,1758]" italics="true" pageId="9" pageNumber="10">T. humiculum</emphasis>
</taxonomicName>
– Leg III basitarsus, retroventral row of macrosetae absent, telotarsus ventral macrosetae irregularly arranged; metasomal segment III ventral surface acarinate, smooth................. 
<taxonomicName id="96CFE9B9003DFF87A36C7544FB6EFE8B" ID-CoL="5999R" authorityName="Lourenco" authorityYear="1981" box="[1101,1212,259,283]" class="Arachnida" family="Troglotayosicidae" genus="Troglotayosicus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="vachoni">
<emphasis id="63BB4E28003DFF87A36C7544FB6EFE8B" box="[1101,1212,259,283]" italics="true" pageId="10" pageNumber="11">T. vachoni</emphasis>
</taxonomicName>
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</keyLead>
</paragraph>
<keyStep id="EA3B8A9F003DFF87A74D7563FB6EFD92" pageId="10" pageNumber="11">
<paragraph id="5170923A003DFF87A74D7563FB6EFE0F" blockId="10.[108,1212,226,746]" pageId="10" pageNumber="11">
<keyLead id="EA3E310F003DFF87A74D7563FB6EFE0F" pageId="10" pageNumber="11">
5. Lateral eyespot similar in size to PDMi ocellus (
<bibRefCitation id="355EEFCB003DFF87A5BE7563FC25FEAC" author="Botero-Trujillo, R. &amp; J. C. Gonzalez-Gomez &amp; J. C. Valenzuela-Rojas &amp; L. F. Garcia" box="[671,1015,292,316]" pageId="10" pageNumber="11" pagination="568 - 582" refId="ref10491" refString="Botero-Trujillo, R., J. C. Gonzalez-Gomez, J. C. Valenzuela-Rojas, and L. F. Garcia. 2017. A new species in the troglomorphic scorpion genus Troglotayosicus from Colombia, representing the northernmost known record for the genus (Scorpiones, Troglotayosicidae). Zootaxa 4244: 568 - 582." type="journal article" year="2017">Botero-Trujillo et al., 2017: 574</bibRefCitation>
, g. 3B); pedipalp patella trichobothrium 
<emphasis id="63BB4E28003DFF87A6B87501FE79FECD" box="[409,427,326,349]" italics="true" pageId="10" pageNumber="11">V</emphasis>
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situated distal to trichobothrium 
<emphasis id="63BB4E28003DFF87A4067502FC9AFECD" box="[807,840,325,349]" italics="true" pageId="10" pageNumber="11">esb</emphasis>
<subScript id="CD4B907F003DFF87A4697517FC83FEF1" attach="left" box="[840,849,336,353]" fontSize="7" pageId="10" pageNumber="11">1</subScript>
(
<bibRefCitation id="355EEFCB003DFF87A47F7502FB6AFECD" author="Botero-Trujillo, R. &amp; J. C. Gonzalez-Gomez &amp; J. C. Valenzuela-Rojas &amp; L. F. Garcia" box="[862,1208,325,349]" pageId="10" pageNumber="11" pagination="568 - 582" refId="ref10491" refString="Botero-Trujillo, R., J. C. Gonzalez-Gomez, J. C. Valenzuela-Rojas, and L. F. Garcia. 2017. A new species in the troglomorphic scorpion genus Troglotayosicus from Colombia, representing the northernmost known record for the genus (Scorpiones, Troglotayosicidae). Zootaxa 4244: 568 - 582." type="journal article" year="2017">Botero-Trujillo et al., 2017: 578</bibRefCitation>
, g. 7A–C); metasomal segment V with VM carina restricted to posterior two-thirds of segment (
<bibRefCitation id="355EEFCB003DFF87A78175C0FE29FE0F" author="Botero-Trujillo, R. &amp; J. C. Gonzalez-Gomez &amp; J. C. Valenzuela-Rojas &amp; L. F. Garcia" box="[160,507,391,415]" pageId="10" pageNumber="11" pagination="568 - 582" refId="ref10491" refString="Botero-Trujillo, R., J. C. Gonzalez-Gomez, J. C. Valenzuela-Rojas, and L. F. Garcia. 2017. A new species in the troglomorphic scorpion genus Troglotayosicus from Colombia, representing the northernmost known record for the genus (Scorpiones, Troglotayosicidae). Zootaxa 4244: 568 - 582." type="journal article" year="2017">Botero-Trujillo et al., 2017: 579</bibRefCitation>
, g. 8E)....................................... 
<taxonomicName id="96CFE9B9003DFF87A36575CFFB6EFE0F" ID-CoL="5999Q" box="[1092,1212,391,415]" class="Arachnida" family="Troglotayosicidae" genus="Troglotayosicus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="meijdeni">
<emphasis id="63BB4E28003DFF87A36575CFFB6EFE0F" box="[1092,1212,391,415]" italics="true" pageId="10" pageNumber="11">T. meijdeni</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="5170923A003DFF87A74D75EEFB6EFD92" blockId="10.[108,1212,226,746]" pageId="10" pageNumber="11">
<keyLead id="EA3E310F003DFF87A74D75EEFB6EFD92" pageId="10" pageNumber="11">
– Lateral eyespot smaller than PDMi ocellus; pedipalp patella trichobothrium 
<emphasis id="63BB4E28003DFF87A32875EEFBC9FE50" box="[1033,1051,425,448]" italics="true" pageId="10" pageNumber="11">V</emphasis>
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aligned with trichobothrium 
<emphasis id="63BB4E28003DFF87A66B758EFEB9FE71" box="[330,363,457,481]" italics="true" pageId="10" pageNumber="11">esb</emphasis>
<subScript id="CD4B907F003DFF87A64A7592FEA6FE76" attach="left" box="[363,372,469,486]" fontSize="7" pageId="10" pageNumber="11">1</subScript>
(
<bibRefCitation id="355EEFCB003DFF87A6A0758EFD19FE71" author="Botero-Trujillo, R. &amp; J. A. Ochoa &amp; O. A. Tovar &amp; J. E. Souza" box="[385,715,457,481]" pageId="10" pageNumber="11" pagination="63 - 76" refId="ref10437" refString="Botero-Trujillo, R., J. A. Ochoa, O. A. Tovar, and J. E. Souza. 2012. A new species in the scorpion genus Troglotayosicus from forest leaf litter in southwestern Colombia (Scorpiones, Troglotayosicidae). Zootaxa 3506: 63 - 76." type="journal article" year="2012">Botero-Trujillo et al., 2012: 71</bibRefCitation>
, gs. 19–21); metasomal segment V with VM carina complete......................................................................6
</keyLead>
</paragraph>
</keyStep>
<keyStep id="EA3B8A9F003DFF87A74D764CFB6EFD7A" pageId="10" pageNumber="11">
<paragraph id="5170923A003DFF87A74D764CFB6EFDF5" blockId="10.[108,1212,226,746]" pageId="10" pageNumber="11">
<keyLead id="EA3E310F003DFF87A74D764CFB6EFDF5" pageId="10" pageNumber="11">
6. Surfaces densely setose (
<bibRefCitation id="355EEFCB003DFF87A681764CFD3AFDB3" author="Botero-Trujillo, R. &amp; J. A. Ochoa &amp; O. A. Tovar &amp; J. E. Souza" box="[416,744,523,547]" pageId="10" pageNumber="11" pagination="63 - 76" refId="ref10437" refString="Botero-Trujillo, R., J. A. Ochoa, O. A. Tovar, and J. E. Souza. 2012. A new species in the scorpion genus Troglotayosicus from forest leaf litter in southwestern Colombia (Scorpiones, Troglotayosicidae). Zootaxa 3506: 63 - 76." type="journal article" year="2012">Botero-Trujillo et al., 2012: 71</bibRefCitation>
, gs. 8–11); pedipalp chela trichobothrium 
<emphasis id="63BB4E28003DFF87A7B8766AFF7EFDD4" box="[153,172,557,580]" italics="true" pageId="10" pageNumber="11">D</emphasis>
<subScript id="CD4B907F003DFF87A78C767FFF64FDD9" attach="left" box="[173,182,568,585]" fontSize="7" pageId="10" pageNumber="11">1</subScript>
(
<emphasis id="63BB4E28003DFF87A7E4766BFF34FDD4" box="[197,230,556,580]" italics="true" pageId="10" pageNumber="11">Db</emphasis>
) situated proximally on manus, slightly distal to trichobothrium 
<emphasis id="63BB4E28003DFF87A484766BFC13FDD4" box="[933,961,556,580]" italics="true" pageId="10" pageNumber="11">Eb</emphasis>
<subScript id="CD4B907F003DFF87A4E0767FFC18FDD9" attach="left" box="[961,970,568,585]" fontSize="7" pageId="10" pageNumber="11">3</subScript>
(
<bibRefCitation id="355EEFCB003DFF87A4F7766BFF25FDF5" author="Botero-Trujillo, R. &amp; J. A. Ochoa &amp; O. A. Tovar &amp; J. E. Souza" pageId="10" pageNumber="11" pagination="63 - 76" refId="ref10437" refString="Botero-Trujillo, R., J. A. Ochoa, O. A. Tovar, and J. E. Souza. 2012. A new species in the scorpion genus Troglotayosicus from forest leaf litter in southwestern Colombia (Scorpiones, Troglotayosicidae). Zootaxa 3506: 63 - 76." type="journal article" year="2012">Botero-Trujillo et al., 2012: 71</bibRefCitation>
, g. 14); leg I basitarsus, retroventral row of macrosetae present............ 
<taxonomicName id="96CFE9B9003DFF87A36A7609FB6EFDF5" ID-CoL="5999N" box="[1099,1212,589,613]" class="Arachnida" family="Troglotayosicidae" genus="Troglotayosicus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="hirsutus">
<emphasis id="63BB4E28003DFF87A36A7609FB6EFDF5" box="[1099,1212,589,613]" italics="true" pageId="10" pageNumber="11">T. hirsutus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="5170923A003DFF87A74D7628FB6EFD7A" blockId="10.[108,1212,226,746]" pageId="10" pageNumber="11">
<keyLead id="EA3E310F003DFF87A74D7628FB6EFD7A" pageId="10" pageNumber="11">
– Surfaces moderately setose (
<bibRefCitation id="355EEFCB003DFF87A6E67629FCC8FD16" author="Sanchez-Vialas, A. &amp; J. Blasco-Arostegui &amp; J. Garcia-Gila &amp; W. R. Lourenco" box="[455,794,622,647]" pageId="10" pageNumber="11" pagination="612 - 618" refId="ref11924" refString="Sanchez-Vialas, A., J. Blasco-Arostegui, J. Garcia-Gila, and W. R. Lourenco. 2020. A new species of Troglotayosicus Lourenco, 1981 (Scorpiones: Troglotayosicidae) from southern Ecuador. Arachnology 18: 612 - 618." type="journal article" year="2020">Sánchez-Vialas et al., 2020: 615</bibRefCitation>
, g. 2A, B); pedipalp chela trichobothrium 
<emphasis id="63BB4E28003DFF87A7F676D7FF38FD37" box="[215,234,656,679]" italics="true" pageId="10" pageNumber="11">D</emphasis>
<subScript id="CD4B907F003DFF87A7CA76DCFF26FD3C" attach="left" box="[235,244,667,684]" fontSize="7" pageId="10" pageNumber="11">1</subScript>
(
<emphasis id="63BB4E28003DFF87A62376C8FEF1FD37" box="[258,291,655,679]" italics="true" pageId="10" pageNumber="11">Db</emphasis>
) situated submedially on manus, approximately midway between trichobothria 
<emphasis id="63BB4E28003DFF87A3B676C8FB61FD37" box="[1175,1203,655,679]" italics="true" pageId="10" pageNumber="11">Eb</emphasis>
<subScript id="CD4B907F003DFF87A39276DCFB6EFD3C" attach="left" box="[1203,1212,667,684]" fontSize="7" pageId="10" pageNumber="11">3</subScript>
and 
<emphasis id="63BB4E28003DFF87A7EB76F6FF0AFD59" box="[202,216,689,713]" italics="true" pageId="10" pageNumber="11">d</emphasis>
<subScript id="CD4B907F003DFF87A7F676FBFF32FD5D" attach="left" box="[215,224,700,717]" fontSize="7" pageId="10" pageNumber="11">8</subScript>
(
<emphasis id="63BB4E28003DFF87A7CF76F7FEDBFD58" box="[238,265,688,712]" italics="true" pageId="10" pageNumber="11">db</emphasis>
) (
<bibRefCitation id="355EEFCB003DFF87A60076F7FDA9FD58" author="Sanchez-Vialas, A. &amp; J. Blasco-Arostegui &amp; J. Garcia-Gila &amp; W. R. Lourenco" box="[289,635,688,713]" pageId="10" pageNumber="11" pagination="612 - 618" refId="ref11924" refString="Sanchez-Vialas, A., J. Blasco-Arostegui, J. Garcia-Gila, and W. R. Lourenco. 2020. A new species of Troglotayosicus Lourenco, 1981 (Scorpiones: Troglotayosicidae) from southern Ecuador. Arachnology 18: 612 - 618." type="journal article" year="2020">Sánchez-Vialas et al., 2020: 616</bibRefCitation>
, g. 5A); leg I basitarsus, retroventral row of macrosetae absent............................................................ 
<taxonomicName id="96CFE9B9003DFF87A3397695FB6EFD7A" ID-CoL="8LZ2N" box="[1048,1212,722,746]" class="Arachnida" family="Troglotayosicidae" genus="Troglotayosicus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="muranunkae">
<emphasis id="63BB4E28003DFF87A3397695FB6EFD7A" box="[1048,1212,722,746]" italics="true" pageId="10" pageNumber="11">T. muranunkae</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>