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<document id="25BB812E6255205B4FA3A5E9AA5F45F0" ID-CLB-Dataset="25322" ID-DOI="10.11646/zootaxa.4646.3.2" ID-GBIF-Dataset="b5150fe8-e62c-4754-8daa-641db7533221" ID-ISSN="1175-5326" ID-Zenodo-Dep="3351022" ID-ZooBank="9179E6F0-AAD8-4FC6-86C6-B489F9462A0F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1564032218894" checkinUser="plazi" docAuthor="Chatzaki, Maria &amp; Keer, Johan Van" docDate="2019" docId="03EA8818F133B61CA98EA403FDB0F746" docLanguage="en" docName="zootaxao.4646.3.2.pdf" docOrigin="Zootaxa 4646 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Echemus kaltsasi Chatzaki &amp; Keer 2019, sp.n." docType="treatment" docVersion="9" lastPageNumber="440" masterDocId="FFD3F060F136B61AA919A13EFFE2F55D" masterDocTitle="Ground spiders (Araneae: Gnaphosidae, Liocranidae, Prodidomidae) from the Greek islands Rodos, Symi and Karpathos, with the description of new species" masterLastPageNumber="460" masterPageNumber="434" pageNumber="439" updateTime="1698747319675" updateUser="ExternalLinkService">
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<mods:title id="EA955DC7168D37F4FD3DB0E8A58C61C3">Ground spiders (Araneae: Gnaphosidae, Liocranidae, Prodidomidae) from the Greek islands Rodos, Symi and Karpathos, with the description of new species</mods:title>
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<mods:namePart id="4472B4EA5E75DB2E347FFAF889E52674">Chatzaki, Maria</mods:namePart>
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<mods:namePart id="A819B3D5C248072F07B253BA1398A4B5">Keer, Johan Van</mods:namePart>
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<treatment id="03EA8818F133B61CA98EA403FDB0F746" ID-DOI="http://doi.org/10.5281/zenodo.5628761" ID-GBIF-Taxon="159628364" ID-Zenodo-Dep="5628761" LSID="urn:lsid:plazi:treatment:03EA8818F133B61CA98EA403FDB0F746" httpUri="http://treatment.plazi.org/id/03EA8818F133B61CA98EA403FDB0F746" lastPageId="6" lastPageNumber="440" pageId="5" pageNumber="439">
<subSubSection id="C3596A85F133B61FA98EA403FDC4F00A" box="[151,550,1341,1368]" pageId="5" pageNumber="439" type="nomenclature">
<paragraph id="8BFC390EF133B61FA98EA403FDC4F00A" blockId="5.[151,550,1341,1403]" box="[151,550,1341,1368]" pageId="5" pageNumber="439">
<heading id="D0B48E62F133B61FA98EA403FDC4F00A" bold="true" box="[151,550,1341,1368]" fontSize="11" level="1" pageId="5" pageNumber="439" reason="1">
<emphasis id="B937E51CF133B61FA98EA403FDC4F00A" bold="true" box="[151,550,1341,1368]" pageId="5" pageNumber="439">
<taxonomicName id="4C43428DF133B61FA98EA403FE06F00A" ID-CoL="95FMP" authority="Chatzaki" authorityName="Chatzaki &amp; Keer" authorityYear="2019" box="[151,484,1341,1368]" class="Arachnida" family="Gnaphosidae" genus="Echemus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="439" phylum="Arthropoda" rank="species" species="kaltsasi" status="sp. nov.">
<emphasis id="B937E51CF133B61FA98EA403FE88F005" bold="true" box="[151,362,1341,1368]" italics="true" pageId="5" pageNumber="439">Echemus kaltsasi</emphasis>
Chatzaki
</taxonomicName>
<taxonomicNameLabel id="A2045867F133B61FA8F2A403FDC4F00A" box="[491,550,1341,1367]" pageId="5" pageNumber="439" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C3596A85F133B61FA98EA45FFE89F026" box="[151,363,1377,1403]" pageId="5" pageNumber="439" type="description">
<paragraph id="8BFC390EF133B61FA98EA45FFE89F026" blockId="5.[151,550,1341,1403]" box="[151,363,1377,1403]" pageId="5" pageNumber="439">
<figureCitation id="1378258BF133B61FA98EA45FFEF5F026" box="[151,279,1377,1403]" captionStart="FIGURES 1316" captionStartId="7.[151,264,1994,2019]" captionTargetBox="[151,1418,812,1975]" captionTargetId="figure@7.[151,1418,812,1975]" captionTargetPageId="7" captionText="FIGURES 1316. Echemus species: E. kaltsasi sp.n. (1314), E. levyi (1516). Left column epigynes, right column vulvae." httpUri="https://zenodo.org/record/3351030/files/figure.png" pageId="5" pageNumber="439">Figs 1314</figureCitation>
,
<figureCitation id="1378258BF133B61FA83BA45FFE89F026" box="[290,363,1377,1403]" captionStart="FIGURES 1720" captionStartId="8.[151,264,1603,1628]" captionTargetBox="[164,1417,197,1575]" captionTargetId="figure@8.[151,1436,181,1583]" captionTargetPageId="8" captionText="FIGURES 1720. Echemus species: E. kaltsasi sp.n. (1718), E. levyi (1920). 17, 19 Epigyne. 18, 20 Vulva. AM: anterior margin, at: atrium, g: glandular head." httpUri="https://zenodo.org/record/3351032/files/figure.png" pageId="5" pageNumber="439">1718</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3596A85F133B61CA98EA496FDB0F746" lastPageId="6" lastPageNumber="440" pageId="5" pageNumber="439" type="materials_examined">
<paragraph id="8BFC390EF133B61FA98EA496FE3EF30E" blockId="5.[151,1437,1448,2015]" pageId="5" pageNumber="439">
<emphasis id="B937E51CF133B61FA98EA496FEF0F09E" bold="true" box="[151,274,1448,1475]" pageId="5" pageNumber="439">Diagnosis.</emphasis>
Females of this species are very similar to
<taxonomicName id="4C43428DF133B61FAA13A494FB9DF099" authority="(Westring, 1861)" baseAuthorityName="Westring" baseAuthorityYear="1861" box="[778,1151,1449,1476]" class="Arachnida" family="Gnaphosidae" genus="Echemus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="439" phylum="Arthropoda" rank="species" species="angustifrons">
<emphasis id="B937E51CF133B61FAA13A494FC54F09E" box="[778,950,1449,1475]" italics="true" pageId="5" pageNumber="439">E. angustifrons</emphasis>
(Westring, 1861)
</taxonomicName>
but differ from it by the mushroom-like epigynal frame formed by the widely open lateral margins which connect to the anterior margin (
<figureCitation id="1378258BF133B61FA986A4CFFEC0F356" box="[159,290,1521,1547]" captionStart="FIGURES 1316" captionStartId="7.[151,264,1994,2019]" captionTargetBox="[151,1418,812,1975]" captionTargetId="figure@7.[151,1418,812,1975]" captionTargetPageId="7" captionText="FIGURES 1316. Echemus species: E. kaltsasi sp.n. (1314), E. levyi (1516). Left column epigynes, right column vulvae." httpUri="https://zenodo.org/record/3351030/files/figure.png" pageId="5" pageNumber="439">Figs 1314</figureCitation>
), in
<taxonomicName id="4C43428DF133B61FA84DA4CCFE1DF356" baseAuthorityName="Westring" baseAuthorityYear="1861" box="[340,511,1521,1547]" class="Arachnida" family="Gnaphosidae" genus="Echemus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="439" phylum="Arthropoda" rank="species" species="angustifrons">
<emphasis id="B937E51CF133B61FA84DA4CCFE1DF356" box="[340,511,1521,1547]" italics="true" pageId="5" pageNumber="439">E. angustifrons</emphasis>
</taxonomicName>
continuing anteriorly as parallel S-shaped lines and not connecting to the anterior margin (
<figureCitation id="1378258BF133B61FA9EDA72BFE8BF372" box="[244,361,1557,1583]" captionStart="FIGURES 912" captionStartId="6.[151,264,1793,1818]" captionTargetBox="[151,1435,561,1774]" captionTargetId="figure@6.[151,1435,561,1774]" captionText="FIGURES 912. Echemus species: E. angustifrons (912)." figureDoi="http://doi.org/10.5281/zenodo.3470819" httpUri="https://zenodo.org/record/3470819/files/figure.png" pageId="5" pageNumber="439">Figs 912</figureCitation>
). It differs from
<taxonomicName id="4C43428DF133B61FAB39A728FCD2F372" authority="Kovblyuk, 2009" authorityName="Kovblyuk &amp; Seyyar" authorityYear="2009" box="[544,816,1557,1583]" class="Arachnida" family="Gnaphosidae" genus="Echemus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="439" phylum="Arthropoda" rank="species" species="levyi">
<emphasis id="B937E51CF133B61FAB39A728FD91F372" box="[544,627,1557,1583]" italics="true" pageId="5" pageNumber="439">E. levyi</emphasis>
<bibRefCitation id="EFD244FFF133B61FAB63A72BFCD2F372" author="Kovblyuk, M. M. &amp; Seyyar, O. &amp; Demir, H. &amp; Topcu, A." box="[634,816,1557,1583]" pageId="5" pageNumber="439" pagination="169 - 187" refId="ref12910" refString="Kovblyuk, M. M., Seyyar, O., Demir, H. &amp; Topcu, A. (2009) New taxonomic and faunistic data on the gnaphosid spiders of Turkey (Aranei: Gnaphosidae). Arthropoda Selecta, 18, 169 - 187." type="journal article" year="2009">Kovblyuk, 2009</bibRefCitation>
</taxonomicName>
by the larger atrium of the epigynal plate and the larger glandular heads of the vulva.
</paragraph>
<paragraph id="8BFC390EF133B61FA9DEA763FED2F3C6" blockId="5.[151,1437,1448,2015]" pageId="5" pageNumber="439">
<materialsCitation id="3B2B3353F133B61FA9DEA763FED2F3C6" ID-GBIF-Occurrence="2423618956" collectingDate="2006-05-16" collectingDateMax="2006-07-12" collectingDateMin="2006-05-16" collectionCode="NHMC" collectorName="M. Chatzaki &amp; D. Kaltsas." pageId="5" pageNumber="439" specimenCount="1" typeStatus="holotype">
<emphasis id="B937E51CF133B61FA9DEA763FEDFF32A" bold="true" box="[199,317,1629,1655]" pageId="5" pageNumber="439">
<typeStatus id="54F887ACF133B61FA9DEA763FEDAF32A" box="[199,312,1629,1655]" pageId="5" pageNumber="439" type="holotype">Holotype</typeStatus>
:
</emphasis>
female, Symi, Panormitis, pitfalls in phrygana,
<collectingDate id="EFB9E626F133B61FAA50A763FBCBF32A" box="[841,1065,1629,1655]" pageId="5" pageNumber="439" value="2006-05-16" valueMax="2006-07-12" valueMin="2006-05-16">
<date id="FFFD1FCEF133B61FAA50A763FBCBF32A" box="[841,1065,1629,1655]" pageId="5" pageNumber="439" value="2006-05-16" valueMax="2006-07-12" valueMin="2006-05-16">16.V to 12.VII.2006</date>
</collectingDate>
(
<collectionCode id="ED52A1CBF133B61FAD2FA763FB72F32A" box="[1078,1168,1629,1655]" country="Myanmar" httpUri="http://grbio.org/cool/4819-0jtd" name="Natural History Museum, Rangoon" pageId="5" pageNumber="439">NHMC</collectionCode>
: 8473), leg.
<collectorName id="26B65CD8F133B61FAC0DA763FA7FF32A" box="[1300,1437,1629,1655]" pageId="5" pageNumber="439">M. Chatzaki</collectorName>
&amp;
<collectorName id="26B65CD8F133B61FA9ADA7BFFED2F3C6" box="[180,304,1665,1691]" pageId="5" pageNumber="439">D. Kaltsas.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="8BFC390EF133B61FA9DEA79AFD51F25A" blockId="5.[151,1437,1448,2015]" pageId="5" pageNumber="439">
<emphasis id="B937E51CF133B61FA9DEA79AFDAFF3E2" bold="true" box="[199,589,1700,1727]" pageId="5" pageNumber="439">Comparative material examined.</emphasis>
<taxonomicName id="4C43428DF133B61FAB48A798FCA4F3E2" authority=": Evros" authorityName="Evros" box="[593,838,1701,1727]" class="Arachnida" family="Gnaphosidae" genus="Echemus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="439" phylum="Arthropoda" rank="species" species="angustifrons">
<emphasis id="B937E51CF133B61FAB48A798FD1BF3E2" box="[593,761,1701,1727]" italics="true" pageId="5" pageNumber="439">E. angustifrons</emphasis>
: Evros
</taxonomicName>
district, Dadia forest,
<specimenCount id="9D45F287F133B61FAD21A79BFBBDF3E2" box="[1080,1119,1701,1727]" pageId="5" pageNumber="439" type="female">1 ♀</specimenCount>
, pitfalls
<date id="FFFD1FCEF133B61FADA4A79AFA7FF3E2" box="[1213,1437,1700,1727]" pageId="5" pageNumber="439" value="2009-07-05" valueMax="2009-08-05" valueMin="2009-07-05">5.VII to 5.VIII.2009</date>
(
<collectionCode id="ED52A1CBF133B61FA986A7F7FF1BF3BE" box="[159,249,1737,1763]" country="Myanmar" httpUri="http://grbio.org/cool/4819-0jtd" name="Natural History Museum, Rangoon" pageId="5" pageNumber="439">NHMC</collectionCode>
: 12116), leg. M. Chatzaki (
<figureCitation id="1378258BF133B61FAB31A7F7FD8CF3BE" box="[552,622,1737,1763]" captionStart="FIGURES 912" captionStartId="6.[151,264,1793,1818]" captionTargetBox="[151,1435,561,1774]" captionTargetId="figure@6.[151,1435,561,1774]" captionText="FIGURES 912. Echemus species: E. angustifrons (912)." figureDoi="http://doi.org/10.5281/zenodo.3470819" httpUri="https://zenodo.org/record/3470819/files/figure.png" pageId="5" pageNumber="439">Figs 9</figureCitation>
─10);
<materialsCitation id="3B2B3353F133B61FABAEA7F7FD51F25A" ID-GBIF-Occurrence="2423618957" collectingDate="2015-06-22" collectingDateMax="2015-07-21" collectingDateMin="2015-06-22" collectionCode="NHMC" collectorName="Loutros N &amp; K. Zografou." pageId="5" pageNumber="439" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<collectorName id="26B65CD8F133B61FABAEA7F7FCCBF3BE" box="[695,809,1737,1763]" pageId="5" pageNumber="439">Loutros N</collectorName>
to Dadia, 17 Km after Loutros,
<specimenCount id="9D45F287F133B61FAD89A7F7FB51F3BE" box="[1168,1203,1737,1763]" pageId="5" pageNumber="439" type="male">1♂</specimenCount>
<specimenCount id="9D45F287F133B61FADA3A7F7FB3EF3BE" box="[1210,1244,1737,1763]" pageId="5" pageNumber="439" type="female">1♀</specimenCount>
, pitfalls
<collectingDate id="EFB9E626F133B61FAC26A7F7FEC2F25A" pageId="5" pageNumber="439" value="2015-06-22" valueMax="2015-07-21" valueMin="2015-06-22">
<date id="FFFD1FCEF133B61FAC26A7F7FEC2F25A" pageId="5" pageNumber="439" value="2015-06-22" valueMax="2015-07-21" valueMin="2015-06-22">22.VI to 21.VII.2015</date>
</collectingDate>
(
<collectionCode id="ED52A1CBF133B61FA836A7D3FE6BF25A" box="[303,393,1773,1799]" country="Myanmar" httpUri="http://grbio.org/cool/4819-0jtd" name="Natural History Museum, Rangoon" pageId="5" pageNumber="439">NHMC</collectionCode>
: 17189), leg.
<collectorName id="26B65CD8F133B61FAB38A7D3FD51F25A" box="[545,691,1773,1799]" pageId="5" pageNumber="439">K. Zografou.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="8BFC390EF133B61FA9DEA62FFD68F212" blockId="5.[151,1437,1448,2015]" pageId="5" pageNumber="439">
<emphasis id="B937E51CF133B61FA9DEA62FFEACF276" bold="true" box="[199,334,1809,1835]" pageId="5" pageNumber="439">Etymology.</emphasis>
The name is given in honor of the biologist and colleague Dr. Dimitris Kaltsas, who contributed to the collection of the material presented here.
</paragraph>
<paragraph id="8BFC390EF133B61CA9DEA666FB1CF442" blockId="5.[151,1437,1448,2015]" lastBlockId="6.[151,1437,153,539]" lastPageId="6" lastPageNumber="440" pageId="5" pageNumber="439">
<emphasis id="B937E51CF133B61FA9DEA666FDCAF22E" bold="true" box="[199,552,1880,1907]" pageId="5" pageNumber="439">
Description. Female
<typeStatus id="54F887ACF133B61FA8A3A667FDC1F22E" box="[442,547,1881,1907]" pageId="5" pageNumber="439" type="holotype">holotype</typeStatus>
.
</emphasis>
Yellow spiders of medium to small size. Body and legs uniformly colored. Measurements: TL 4.965; CL 2.075; CW 1.555; AL 2.627. Eye sizes and interdistances:
<collectionCode id="ED52A1CBF133B61FAD23A643FB9AF2CA" box="[1082,1144,1917,1943]" pageId="5" pageNumber="439">AME</collectionCode>
0.117, PME 0.092x0.119, PLE 0.107, ALE 0.123, AME-ALE almost touching, AME-AME, PME-PME and PME-PLE
<emphasis id="B937E51CF133B61FADD2A69CFB07F2E6" box="[1227,1253,1954,1979]" italics="true" pageId="5" pageNumber="439">ca</emphasis>
equidistant and separated by less than 1/2 their diameter.All eyes round and pearly white with black surroundings, especially
<collectionCode id="ED52A1CBF133B61FAC4EA6FBFA7BF282" box="[1367,1433,1989,2015]" pageId="5" pageNumber="439">AME</collectionCode>
. PER procurved, AER straight to slightly recurved. Prosoma with widening thoracic region, fovea at posterior half of its length. Labium triangular, longer than wide. Chelicerae with 1 and 3 denticles on
<collectingRegion id="4987F7ECF130B61CAD62A183FB45F58A" box="[1147,1191,189,215]" country="Peru" name="Lima" pageId="6" pageNumber="440">RM</collectingRegion>
and
<collectionCode id="ED52A1CBF130B61CADC6A183FAE5F58A" box="[1247,1287,189,215]" country="USA" name="Pratt Museum" pageId="6" pageNumber="440">PM</collectionCode>
respectively. Leg formula is IV&gt;I&gt;II&gt;III. Legs spination: Leg I: Fe 2d 1pl; leg II: Fe 2d 1pl, Mt
<date id="FFFD1FCEF130B61CAD34A1DFFBA9F5A6" box="[1069,1099,225,251]" pageId="6" pageNumber="440">1v</date>
; leg III: Fe 4-5d, Pa 1rl, Ti/Mt spinose; leg IV: Fe 3d, Pa 1rl, Ti/Mt spinose. Preening brush at Mt III and IV, Ta densely scopulated.
</paragraph>
<paragraph id="8BFC390EF130B61CA9DEA017FEF3F4D6" blockId="6.[151,1437,153,539]" pageId="6" pageNumber="440">
Epigyne (
<figureCitation id="1378258BF130B61CA82CA017FE6FF41E" box="[309,397,297,323]" captionStart="FIGURES 1316" captionStartId="7.[151,264,1994,2019]" captionTargetBox="[151,1418,812,1975]" captionTargetId="figure@7.[151,1418,812,1975]" captionTargetPageId="7" captionText="FIGURES 1316. Echemus species: E. kaltsasi sp.n. (1314), E. levyi (1516). Left column epigynes, right column vulvae." httpUri="https://zenodo.org/record/3351030/files/figure.png" pageId="6" pageNumber="440">Figs 13</figureCitation>
,
<figureCitation id="1378258BF130B61CA883A017FE5BF41E" box="[410,441,297,323]" captionStart="FIGURES 1720" captionStartId="8.[151,264,1603,1628]" captionTargetBox="[164,1417,197,1575]" captionTargetId="figure@8.[151,1436,181,1583]" captionTargetPageId="8" captionText="FIGURES 1720. Echemus species: E. kaltsasi sp.n. (1718), E. levyi (1920). 17, 19 Epigyne. 18, 20 Vulva. AM: anterior margin, at: atrium, g: glandular head." httpUri="https://zenodo.org/record/3351032/files/figure.png" pageId="6" pageNumber="440">17</figureCitation>
). Mushroom like atrium with large cup and short base formed by lateral margins which open widely to meet anterior margin and bend medially to form the base. Sclerotized median lines also present in the atrium.
</paragraph>
<paragraph id="8BFC390EF130B61CA9DEA0ABFCC6F4A5" blockId="6.[151,1437,153,539]" pageId="6" pageNumber="440">
Vulva (
<figureCitation id="1378258BF130B61CA803A0ABFE90F4F2" box="[282,370,405,431]" captionStart="FIGURES 1316" captionStartId="7.[151,264,1994,2019]" captionTargetBox="[151,1418,812,1975]" captionTargetId="figure@7.[151,1418,812,1975]" captionTargetPageId="7" captionText="FIGURES 1316. Echemus species: E. kaltsasi sp.n. (1314), E. levyi (1516). Left column epigynes, right column vulvae." httpUri="https://zenodo.org/record/3351030/files/figure.png" pageId="6" pageNumber="440">Figs 14</figureCitation>
,
<figureCitation id="1378258BF130B61CA866A0ABFE7CF4F2" box="[383,414,405,431]" captionStart="FIGURES 1720" captionStartId="8.[151,264,1603,1628]" captionTargetBox="[164,1417,197,1575]" captionTargetId="figure@8.[151,1436,181,1583]" captionTargetPageId="8" captionText="FIGURES 1720. Echemus species: E. kaltsasi sp.n. (1718), E. levyi (1920). 17, 19 Epigyne. 18, 20 Vulva. AM: anterior margin, at: atrium, g: glandular head." httpUri="https://zenodo.org/record/3351032/files/figure.png" pageId="6" pageNumber="440">18</figureCitation>
). Coiled copulatory ducts with long transverse glandular heads on anterior ends. Enlarged spermathecal sacs connected to copulatory ducts and attached to anterior margin. Spermathecae of rather irregular shape (not easily distinguished in the available specimen).
</paragraph>
<paragraph id="8BFC390EF130B61CA9DEA33EFDB0F746" blockId="6.[151,1437,153,539]" box="[199,594,512,539]" pageId="6" pageNumber="440">
<emphasis id="B937E51CF130B61CA9DEA33EFEBBF746" bold="true" box="[199,345,512,539]" pageId="6" pageNumber="440">Distribution</emphasis>
:
<collectingCountry id="F354799EF130B61CA871A33FFE58F746" box="[360,442,513,539]" name="Greece" pageId="6" pageNumber="440">Greece</collectingCountry>
, Symi island.
</paragraph>
</subSubSection>
</treatment>
</document>