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<document id="BC77B67F8FFF8F96D50B541E02BFB1AF" ID-CLB-Dataset="169975" ID-DOI="10.11646/zootaxa.5196.2.2" ID-GBIF-Dataset="96e1a27d-a189-4ed0-97e0-c02192b3828d" ID-ISSN="1175-5326" ID-Zenodo-Dep="7224439" ID-ZooBank="8107F44A-C50A-43BB-9D28-0E990E8AA0FC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1666167075819" checkinUser="plazi" docAuthor="Sittichaya, Wisut &amp; Smith, Sarah M." docDate="2022" docId="03B387EC1E25FFB5FF54FAEFFE16D375" docLanguage="en" docName="zootaxa.5196.2.2.pdf" docOrigin="Zootaxa 5196 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Amasa cognatoi Sittichaya &amp; Smith 2022, sp. nov." docType="treatment" docUuid="147EDD3A-7321-4CAB-824B-1967DE6C3FD3" docUuidSource="ZooBank" docVersion="6" lastPageNumber="200" masterDocId="FF8AFF941E27FFB6FFC3FFB7FFC5D708" masterDocTitle="New species and new records of Amasa Lea, 1894 ambrosia beetles from Thailand (Coleoptera: Curculionidae: Scolytinae: Xyleborini)" masterLastPageNumber="210" masterPageNumber="197" pageNumber="199" updateTime="1699408989087" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="3AD07AB0B1B4CCFCD4CDAB7B203CF1EB" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="657F6BCCE6098AB8DF0948A95FE41A16">
<mods:title id="D8623A282BF2BDFC98021561FBDDE9A1">New species and new records of Amasa Lea, 1894 ambrosia beetles from Thailand (Coleoptera: Curculionidae: Scolytinae: Xyleborini)</mods:title>
</mods:titleInfo>
<mods:name id="07C01D7299C91891DA88AFE343232888" type="personal">
<mods:role id="9D2AF72028B9E754820B5B34E883EE6D">
<mods:roleTerm id="A9A4AE44F2AC7A9FD248BD0A85A05A49">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="76517CE9CBDED379D377DE4A76C4FBFE">Sittichaya, Wisut</mods:namePart>
<mods:affiliation id="662EBC47435F13857BA237043DE22245">Agricultural Innovation and Management Division, Faculty of Natural Resources, Prince of Songkla University, Songkhla, 90110, Thailand</mods:affiliation>
</mods:name>
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<mods:role id="1A933D87007E29E2E35772A3508D81BF">
<mods:roleTerm id="B75088CC21BF99DDD94A1FFD98104444">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D602633F16F9975BDF458F71FC9E9AC7">Smith, Sarah M.</mods:namePart>
<mods:nameIdentifier id="DC39560758C41C739FEA95CA40C5DC8C" type="ORCID">0000-0002-5173-3736</mods:nameIdentifier>
<mods:affiliation id="F985D455E941911D8E77AD0036B4FF5A">Department of Entomology, Michigan State University, 288 Farm Lane, 243 Natural Science Bldg., East Lansing, MI 48824, USA smith 462 @ msu. edu; https: // orcid. org / 0000 - 0002 - 5173 - 3736</mods:affiliation>
<mods:nameIdentifier id="A3ACCE8DC74D54C2CACBB52E9F30BC46" type="email">smith462@msu.edu</mods:nameIdentifier>
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<mods:typeOfResource id="5905BC252C37CECC48A165DB9C53523A">text</mods:typeOfResource>
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<mods:title id="7B7438F0574C357A2F64912302DD84B3">Zootaxa</mods:title>
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<mods:part id="F04A82EEC7C167C94888EB08108C8ACE">
<mods:date id="CDD5F1630D9D0D68C080FC04EDC21161">2022</mods:date>
<mods:detail id="5B93F4D85B5A6BA62FB237A6F86E8D5C" type="pubDate">
<mods:number id="6FFEF51E97DD0BF6843173AAAE8A7AA0">2022-10-19</mods:number>
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<mods:detail id="43F23F4DBDDDB7A5DF594D0CDD120588" type="volume">
<mods:number id="F61624D8A0F29415AB8BF47D4E00C94D">5196</mods:number>
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<mods:classification id="6314088B90FCC396EC4CA52C2AFCED92">journal article</mods:classification>
<mods:identifier id="B1C05F0C7F47FA2D7B18ACD98090E74A" type="CLB-Dataset">169975</mods:identifier>
<mods:identifier id="FC48C4DDE7D9E560C6BEB6A05E9B1BC4" type="DOI">10.11646/zootaxa.5196.2.2</mods:identifier>
<mods:identifier id="FF4F97BC2B7A4C3F2CF7BF3031966051" type="GBIF-Dataset">96e1a27d-a189-4ed0-97e0-c02192b3828d</mods:identifier>
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<treatment id="03B387EC1E25FFB5FF54FAEFFE16D375" ID-DOI="http://doi.org/10.5281/zenodo.7224465" ID-GBIF-Taxon="203119835" ID-Zenodo-Dep="7224465" LSID="urn:lsid:zoobank.org:act:147EDD3A-7321-4CAB-824B-1967DE6C3FD3" httpUri="http://treatment.plazi.org/id/03B387EC1E25FFB5FF54FAEFFE16D375" lastPageId="3" lastPageNumber="200" pageId="2" pageNumber="199">
<subSubSection id="C30065711E25FFB4FF54FAEFFD73D27A" box="[151,694,1368,1394]" pageId="2" pageNumber="199" type="nomenclature">
<paragraph id="8BA536FA1E25FFB4FF54FAEFFD73D27A" blockId="2.[151,1194,1368,1466]" box="[151,694,1368,1394]" pageId="2" pageNumber="199">
<heading id="D0ED81961E25FFB4FF54FAEFFD73D27A" bold="true" box="[151,694,1368,1394]" fontSize="11" level="1" pageId="2" pageNumber="199" reason="1">
<emphasis id="B96EEAE81E25FFB4FF54FAEFFD73D27A" bold="true" box="[151,694,1368,1394]" pageId="2" pageNumber="199">
<taxonomicName id="4C1A4D791E25FFB4FF54FAEFFD94D27A" authority="Sittichaya &amp; Smith" authorityName="Sittichaya &amp; Smith" authorityYear="2022" box="[151,593,1368,1394]" class="Insecta" family="Curculionidae" genus="Amasa" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="199" phylum="Arthropoda" rank="species" species="cognatoi" status="sp. nov.">
<emphasis id="B96EEAE81E25FFB4FF54FAEFFE9FD27A" bold="true" box="[151,346,1368,1394]" italics="true" pageId="2" pageNumber="199">Amasa cognatoi</emphasis>
Sittichaya &amp; Smith
</taxonomicName>
<taxonomicNameLabel id="A25D57931E25FFB4FD9BFAEFFD73D27A" box="[600,694,1368,1394]" pageId="2" pageNumber="199" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30065711E25FFB4FF54FACBFB6FD2B2" pageId="2" pageNumber="199" type="description">
<paragraph id="8BA536FA1E25FFB4FF54FACBFF2ED29E" blockId="2.[151,1194,1368,1466]" box="[151,235,1404,1430]" pageId="2" pageNumber="199">
(
<figureCitation id="13212A7F1E25FFB4FF5CFACBFF27D29E" box="[159,226,1404,1430]" captionStart="FIGURE 3" captionStartId="6.[151,250,1972,1996]" captionTargetBox="[214,1374,181,1948]" captionTargetId="figure-23@6.[214,1374,181,1948]" captionTargetPageId="6" captionText="FIGURE 3. Amasa cognatoi sp. nov. holotype A) dorsal view; B) lateral view; C) postero-lateral view; D) frons; E) venter and antennae." figureDoi="http://doi.org/10.5281/zenodo.7224445" httpUri="https://zenodo.org/record/7224445/files/figure.png" pageId="2" pageNumber="199">Fig. 3</figureCitation>
)
</paragraph>
<paragraph id="8BA536FA1E25FFB4FF54FA17FB6FD2B2" blockId="2.[151,1194,1368,1466]" box="[151,1194,1440,1466]" pageId="2" pageNumber="199">
http://zoobank.org/
<uri id="FF8B3AF81E25FFB4FEB0FA17FB6FD2B2" box="[371,1194,1440,1466]" pageId="2" pageNumber="199">
urn:lsid:zoobank.org:act:
<uuid id="FFBC0C2F1E25FFB4FD47FA17FB6FD2B2" box="[644,1194,1440,1466]" pageId="2" pageNumber="199">147EDD3A-7321-4CAB-824B-1967DE6C3FD3</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C30065711E25FFB4FF54FA5FFAE4D12E" pageId="2" pageNumber="199" type="materials_examined">
<paragraph id="8BA536FA1E25FFB4FF54FA5FFAE4D12E" blockId="2.[151,1436,1512,2043]" pageId="2" pageNumber="199">
<emphasis id="B96EEAE81E25FFB4FF54FA5FFE4DD10A" bold="true" box="[151,392,1512,1538]" pageId="2" pageNumber="199">Material examined.</emphasis>
<materialsCitation id="3B723CA71E25FFB4FE54FA5FFAE4D12E" ID-GBIF-Occurrence="3938604308" collectingDate="2015-04-01" collectorName="W. Sittichaya" country="Thailand" latitude="8.994528" location="National Park" longLatPrecision="1" longitude="98.468864" municipality="Sri" pageId="2" pageNumber="199" specimenCount="1" stateProvince="Phangnga" typeStatus="holotype">
<emphasis id="B96EEAE81E25FFB4FE54FA5FFDB7D10A" bold="true" box="[407,626,1512,1538]" pageId="2" pageNumber="199">
<typeStatus id="54A188581E25FFB4FE54FA5FFDC3D10A" box="[407,518,1512,1538]" pageId="2" pageNumber="199" type="holotype">Holotype</typeStatus>
, female:
</emphasis>
<collectingCountry id="F30D766A1E25FFB4FD43FA5FFCD1D10A" box="[640,788,1512,1538]" name="Thailand" pageId="2" pageNumber="199">THAILAND</collectingCountry>
,
<collectingRegion id="49DEF8181E25FFB4FCE4FA5FFBDAD10A" box="[807,1055,1512,1539]" country="Thailand" name="Phangnga" pageId="2" pageNumber="199">Phang Nga Province</collectingRegion>
,
<collectingMunicipality id="6BC1AC801E25FFB4FBF1FA5FFB96D10A" box="[1074,1107,1512,1538]" pageId="2" pageNumber="199">Sri</collectingMunicipality>
<collectingRegion id="49DEF8181E25FFB4FBA2FA5FFB20D10B" box="[1121,1253,1512,1539]" country="Thailand" name="Phangnga" pageId="2" pageNumber="199">Phang Nga</collectingRegion>
<location id="8EC560211E25FFB4FB30FA5FFA5DD10A" LSID="urn:lsid:plazi:treatment:03B387EC1E25FFB5FF54FAEFFE16D375:8EC560211E25FFB4FB30FA5FFA5DD10A" box="[1267,1432,1512,1538]" country="Thailand" latitude="8.994528" longLatPrecision="1" longitude="98.468864" municipality="Sri" name="National Park" pageId="2" pageNumber="199" stateProvince="Phangnga">National Park</location>
,
<geoCoordinate id="EE2E503D1E25FFB4FF54F9BAFEE6D120" box="[151,291,1549,1576]" degrees="8" direction="north" minutes="59" orientation="latitude" pageId="2" pageNumber="199" precision="1" seconds="40.3" value="8.994528">8°59′40.3″N</geoCoordinate>
<geoCoordinate id="EE2E503D1E25FFB4FEE9F9BAFE01D120" box="[298,452,1549,1576]" degrees="98" direction="east" minutes="28" orientation="longitude" pageId="2" pageNumber="199" precision="1" seconds="07.9" value="98.468864">98°28′07.9″E</geoCoordinate>
, tropical rainforest,
<collectingMethod id="525B4EED1E25FFB4FD66F9BBFCBDD12F" box="[677,888,1548,1575]" pageId="2" pageNumber="199">ethanol baited trap</collectingMethod>
,
<date id="FFA4103A1E25FFB4FC47F9BBFC39D12F" box="[900,1020,1548,1575]" pageId="2" pageNumber="199" value="2015-04-01">
<collectingDate id="EFE0E9D21E25FFB4FC47F9BBFC39D12F" box="[900,1020,1548,1575]" pageId="2" pageNumber="199" value="2015-04-01">01.iv.2015</collectingDate>
</date>
,
<collectorName id="26EF532C1E25FFB4FBC4F9BAFB65D12E" box="[1031,1184,1548,1575]" pageId="2" pageNumber="199">W. Sittichaya</collectorName>
, (NHMW).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30065711E25FFB4FF04F987FD37D1D2" pageId="2" pageNumber="199" type="diagnosis">
<paragraph id="8BA536FA1E25FFB4FF04F987FEBCD1BF" blockId="2.[151,1436,1512,2043]" pageId="2" pageNumber="199">
<emphasis id="B96EEAE81E25FFB4FF04F987FE81D142" bold="true" box="[199,324,1584,1610]" pageId="2" pageNumber="199">Diagnosis:</emphasis>
<quantity id="4CE29B1F1E25FFB4FE88F987FE6AD143" box="[331,431,1584,1611]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="2" pageNumber="199" unit="mm" value="2.3">2.30 mm</quantity>
long (pronotal anterior margin to declivital summit 2.0 mm); 1.05× as long as wide. This species is easily distinguished by very short and stout form, pronotum approximately as long as elytra; pronotum round,
<typeStatus id="54A188581E25FFB4FF2BF9CEFEDDD19B" box="[232,280,1657,1683]" pageId="2" pageNumber="199">type</typeStatus>
1, when viewed dorsally, anterior margin serrate, procoxae moderately separated, declivity with long, erect hair-like setae.
</paragraph>
<paragraph id="8BA536FA1E25FFB4FF04F977FD37D1D2" blockId="2.[151,1436,1512,2043]" box="[199,754,1728,1754]" pageId="2" pageNumber="199">
<emphasis id="B96EEAE81E25FFB4FF04F977FE47D1D2" bold="true" box="[199,386,1728,1754]" pageId="2" pageNumber="199">Similar species:</emphasis>
<taxonomicName id="4C1A4D791E25FFB4FE4AF977FD2BD1D2" authority="(Schedl)" baseAuthorityName="Schedl" box="[393,750,1728,1754]" class="Insecta" family="Curculionidae" genus="Amasa" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="199" phylum="Arthropoda" rank="species" species="cylindrotomica">
<emphasis id="B96EEAE81E25FFB4FE4AF977FD40D1D2" box="[393,645,1728,1754]" italics="true" pageId="2" pageNumber="199">Amasa cylindrotomica</emphasis>
(Schedl)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C30065711E25FFB5FF04F953FCB1D4E5" lastPageId="3" lastPageNumber="200" pageId="2" pageNumber="199" type="description">
<paragraph id="8BA536FA1E25FFB4FF04F953FD1DD04E" blockId="2.[151,1436,1512,2043]" pageId="2" pageNumber="199">
<emphasis id="B96EEAE81E25FFB4FF04F953FE07D1F6" bold="true" box="[199,450,1764,1790]" pageId="2" pageNumber="199">Description (female).</emphasis>
<quantity id="4CE29B1F1E25FFB4FE0AF953FDE8D1F7" box="[457,557,1764,1791]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="2" pageNumber="199" unit="mm" value="2.3">2.30 mm</quantity>
long (pronotal anterior margin to declivital summit 2.0 mm); 1.05× as long as wide. Very stout, cylindrical, yellowish brown, pronotal anterior slope, near elytral lateral margin and declivity dark brown, elytra near declivital margin reddish brown.
</paragraph>
<paragraph id="8BA536FA1E25FFB5FF04F8E7FCB1D4E5" blockId="2.[151,1436,1512,2043]" lastBlockId="3.[151,1437,151,1149]" lastPageId="3" lastPageNumber="200" pageId="2" pageNumber="199">
<emphasis id="B96EEAE81E25FFB4FF04F8E7FECBD062" bold="true" box="[199,270,1872,1898]" pageId="2" pageNumber="199">Head.</emphasis>
Epistoma entire, transverse, with row of hair-like setae. Frons weakly convex to upper level of eyes; median line between eyes flat, not impressed, vertically extended to epistoma, surface dull, impunctate, alutaceous, asperate; asperities longitudinal, sparse, inconspicuous, rounder and slightly denser above epistoma, increasing in size and length and decreasing in density dorsally and laterally. Frons sparsely covered with long very fine hairlike setae. Eyes deeply emarginate just above antennal insertion, upper part smaller than lower part. Submentum small triangular, deeply impressed. Antennal scape regularly thick, slightly shorter than club (1:1.2). Pedicel as wide as scape, shorter than funicle. Funicle 4-segmented, segment 1 shorter than pedicel. Club circular and flat without sutures,
<typeStatus id="54A188581E24FFB5FE90FF68FE46D7F1" box="[339,387,223,249]" pageId="3" pageNumber="200">type</typeStatus>
5, its margins present but softly pubescent.
<emphasis id="B96EEAE81E24FFB5FCBAFF68FC3CD7F1" bold="true" box="[889,1017,223,249]" pageId="3" pageNumber="200">Pronotum.</emphasis>
1.0× as long as wide. In dorsal view short and rounded,
<typeStatus id="54A188581E24FFB5FEB6FEB4FE60D615" box="[373,421,259,285]" pageId="3" pageNumber="200">type</typeStatus>
1, sides subparallel in basal 1/2, rounded anteriorly and dorsally; anterior margin with a row of ten serrations. In lateral view basic,
<typeStatus id="54A188581E24FFB5FDB9FE90FD6FD649" box="[634,682,295,321]" pageId="3" pageNumber="200">type</typeStatus>
0, disc flat, summit approximately at midpoint. Anterior slope with very densely spaced, fine transverse asperities, becoming lower towards summit, bearing moderately long, fine, erect hair-like setae. Some longer hair-like setae at anterior and lateral margins. Disc shining, with very fine sparse punctures, glabrous. Lateral margins obliquely costate. Base transverse, posterior angles broadly rounded.
<emphasis id="B96EEAE81E24FFB5FA8AFE24FA59D6A5" bold="true" box="[1353,1436,403,429]" pageId="3" pageNumber="200">Elytra.</emphasis>
1.0× as long as wide, 1.0× as long as pronotum. Scutellum moderately sized, broad, linguiform, flush with elytra, flat, shining. Elytral base broadly convex, edge carinate, humeral angles rounded, parallel-sided in basal 4/5, then sharply angulate to apex. Disc slightly ascending posteriorly, shining, sparsely covered with very fine and very short inconspicuous setae; striae and interstriae laterally diverging from base to declivital summit; striae small, shallowly impressed; interstriae flat, very fine and inconspicuous punctate, double as wide as striae. Declivity truncate, face slightly ascending posteriorly, circumdeclivital margins sharply carinate; face slightly convex, subshiny, covered with erect long setae; three striae present, strial punctures round, deeply impressed, approximately as large as on disc, punctures subcontiguous, each with a very short fine seta; interstriae impunctate, flat, interstriae 1–3 multiseriate granulate, each granule with a long erect hair-like seta, granules multiseriate, confused. Posterolateral margin forming a circumdeclivital carina; carina setose, setae short, erect, hair-like.
<emphasis id="B96EEAE81E24FFB5FBA7FD4CFB66D41D" bold="true" box="[1124,1187,763,789]" pageId="3" pageNumber="200">Legs.</emphasis>
Procoxae moderately separated, prosternal coxal piece flat, inconspicuous. Protibiae slender, triangular, broadest at apex; posterior face inflated, finely granulate; outer margin three and five small socketed denticles at apex, their length as long as basal width. Mesotibiae broad, triangular, broadest at apical 2/3, flattened, outer margins with eight small socketed denticles. Metatibiae broad, flattened, broadest at middle, outer margins evenly rounded, with ten small socketed denticles. Meso-and metatibiae posterior faces with inner part longitudinally inflated and bearing long fine setae, outer part flat, glabrous, unarmed; anterior faces finely granulate.
</paragraph>
</subSubSection>
<subSubSection id="C30065711E24FFB5FF04FC40FE4FD33D" pageId="3" pageNumber="200" type="etymology">
<paragraph id="8BA536FA1E24FFB5FF04FC40FE4FD33D" blockId="3.[151,1437,151,1149]" pageId="3" pageNumber="200">
<emphasis id="B96EEAE81E24FFB5FF04FC40FE8BD319" bold="true" box="[199,334,1015,1041]" pageId="3" pageNumber="200">Etymology.</emphasis>
The species is named in honor of Dr. Anthony I. Cognato for his great contributions to modern
<taxonomicName id="4C1A4D791E24FFB5FF54FBACFEC8D33D" box="[151,269,1051,1077]" class="Insecta" family="Scolytinae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="200" phylum="Arthropoda" rank="family">Scolytinae</taxonomicName>
taxonomy.
</paragraph>
</subSubSection>
<subSubSection id="C30065711E24FFB5FF04FB89FD32D351" box="[199,759,1086,1113]" pageId="3" pageNumber="200" type="distribution">
<paragraph id="8BA536FA1E24FFB5FF04FB89FD32D351" blockId="3.[151,1437,151,1149]" box="[199,759,1086,1113]" pageId="3" pageNumber="200">
<emphasis id="B96EEAE81E24FFB5FF04FB89FEA6D350" bold="true" box="[199,355,1086,1112]" pageId="3" pageNumber="200">Distribution:</emphasis>
<collectingCountry id="F30D766A1E24FFB5FEA9FB88FE3AD351" box="[362,511,1087,1113]" name="Thailand" pageId="3" pageNumber="200">THAILAND</collectingCountry>
:
<collectingRegion id="49DEF8181E24FFB5FDCAFB88FD36D351" box="[521,755,1087,1113]" country="Thailand" name="Phangnga" pageId="3" pageNumber="200">Phang Nga Province</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C30065711E24FFB5FF04FBD4FE16D375" box="[199,467,1123,1149]" pageId="3" pageNumber="200" type="biology_ecology">
<paragraph id="8BA536FA1E24FFB5FF04FBD4FE16D375" blockId="3.[151,1437,151,1149]" box="[199,467,1123,1149]" pageId="3" pageNumber="200">
<emphasis id="B96EEAE81E24FFB5FF04FBD4FE93D375" bold="true" box="[199,342,1123,1149]" pageId="3" pageNumber="200">Host plants.</emphasis>
Unknown.
</paragraph>
</subSubSection>
</treatment>
</document>
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