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<document id="30B5E784A72BC544C4187EEFBBCD5CEC" ID-DOI="10.5281/zenodo.280626" ID-GBIF-Dataset="bdb9a932-2643-48f1-8545-cfeb75a683cd" ID-ISSN="1175-5326" ID-Zenodo-Dep="280626" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460462984094" checkinUser="plazi" docAuthor="Huang, Hao &amp; Chen, Chang-Chin" docDate="2012" docId="03FBD04AFFCC6013FF0546E5B7C1F92B" docLanguage="en" docName="zt03255p036.pdf" docOrigin="Zootaxa 3255" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Prismognathus nigricolor Boucher 1996" docType="treatment" docVersion="13" lastPageNumber="9" masterDocId="FFC2A832FFC4601BFF924672B73FFFDC" masterDocTitle="A review of the genera Prismognathus Motschulsky and Cladophyllus Houlbert (Coleoptera: Scarabaeoidea: Lucanidae) from China, with the description of two new species" masterLastPageNumber="36" masterPageNumber="1" pageNumber="9" updateTime="1698304529053" updateUser="plazi">
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<mods:title id="18B985763B8C8F73CBF0C73C2970C7FE">A review of the genera Prismognathus Motschulsky and Cladophyllus Houlbert (Coleoptera: Scarabaeoidea: Lucanidae) from China, with the description of two new species</mods:title>
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<paragraph id="8BED615CFFCC6013FF0546E5B5B8FF6D" blockId="8.[151,749,151,210]" box="[151,647,151,177]" pageId="8" pageNumber="9">
<heading id="D0A5D630FFCC6013FF0546E5B5B8FF6D" bold="true" box="[151,647,151,177]" fontSize="11" level="1" pageId="8" pageNumber="9" reason="1">
<taxonomicName id="4C521ADFFFCC6013FF0546E5B5B8FF6D" ID-CoL="4MHQY" authority="Boucher, 1996" authorityName="Boucher" authorityYear="1996" box="[151,647,151,177]" class="Insecta" family="Lucanidae" genus="Prismognathus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="nigricolor">
<emphasis id="B926BD4EFFCC6013FF0546E5B5B8FF6D" bold="true" box="[151,647,151,177]" pageId="8" pageNumber="9">
<emphasis id="B926BD4EFFCC6013FF0546E5B6F1FF6D" bold="true" box="[151,462,151,177]" italics="true" pageId="8" pageNumber="9">Prismognathus nigricolor</emphasis>
<bibRefCitation id="EFC31CADFFCC6013FE4446E5B5B8FF6D" author="Boucher" box="[470,647,151,177]" pageId="8" pageNumber="9" refString="Boucher, S. (1996) Deux especes inedites sino-birmanes des genres Prismognathus Motschulsky et Cladophyllus Houlbert, et mise en synonymie de C. flavus Didier (Coleoptera, Lucanidae). Coleopteres, 2 (7), 69 - 78." type="journal article" year="1996">Boucher, 1996</bibRefCitation>
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(
<figureCitation id="13697DD9FFCC6013FF0D46C8B619FF0E" box="[159,294,186,210]" captionStart="FIGURES 21 – 34" captionStartId="22.[151,257,819,840]" captionTargetBox="[350,1238,194,798]" captionTargetId="figure@22.[340,1247,175,807]" captionTargetPageId="22" captionText="FIGURES 21 – 34. Male mandible. 21 – 30 — dorsal view; 21 — Prismognathus dauricus (Liaoning); 22 — P. davidis davidis (Beijing); 23 — P. davidis davidis (Zhejiang); 24 — P. klapperichi (Fujian); 25 — P. s i n i a e v i (Gongshan, Yunnan); 26 — P. siniaevi (Jinping, Yunnan); 27 — P. prossi (Sichuan); 28 — P. nigricolor (holotype); 29 — P. s h a n i (paratype, Sichuan); 30 — P. nigricolor (Gongshan); 31 – 34 — ventral view; 31 — P. davidis davidis (Zhejiang); 32 — P. klapperichi (Chongqing); 33 — P. klapperichi (Fujian); 34 — P. siniaevi (Gongshan, Yunnan). Arrow mark directed to dorsal subbasal tooth or gap on ventral teeth of the mandible." httpUri="https://zenodo.org/record/280628/files/figure.png" pageId="8" pageNumber="9">Figs. 28, 30</figureCitation>
, 
<figureCitation id="13697DD9FFCC6013FEA046C8B661FF0E" box="[306,350,186,210]" captionStart="FIGURES 111 – 131" captionStartId="28.[151,257,1303,1324]" captionTargetBox="[219,1365,191,1298]" captionTargetId="figure@28.[219,1366,191,1307]" captionTargetPageId="28" captionText="FIGURES 111 – 131. Female canthus (not under the same scale). 111 — P. s h a n i, paratype, Labahe, Sichuan; 112 — P. nigricolor, paratype; 113 — P. arcuatus, Siping, Sichuan; 114 – 116 — P. p ro s s i; 114 — Siping, Sichuan; 115 — Fanjingshan, Guizhou; 116 — Maoershan, Guangxi; 117 – 119 — P. davidis davidis; 117 — Siping, Sichuan; 118 — Tianmushan, Zhejiang; 119 — Dabashan, Chongqing; 120 – 123 — P. triapicalis; 120 — Sichuan; 121 — Dabashan, Chongqing; 122 — Fengyangshan, Zhejiang; 123 — Dabashan, Chongqing; 124 — P. haojiani, paratype, Fanjingshan; 125 — P. dauricus, Liaoning; 126 — P. kanghianus kanghianus, Hekou, Yunnan; 127 — P. alessandrae, Zhongdian, Yunnan; 128 — P. sinen sis, holotype, Kuatun, Fujian; 129 — P. sukk ito rum, Tengchong, Yunnan; 130 — P. formosanus, Taiwan; 131 — P. siniaevi, Gongshan, Yunnan." httpUri="https://zenodo.org/record/280634/files/figure.png" pageId="8" pageNumber="9">112</figureCitation>
, 
<figureCitation id="13697DD9FFCC6013FEFB46C8B6F4FF0E" box="[361,459,186,210]" captionStart="FIGURES 134 – 149" captionStartId="29.[151,257,1583,1604]" captionTargetBox="[151,1427,211,1495]" captionTargetId="figure@29.[151,1436,191,1503]" captionTargetPageId="29" captionText="FIGURES 134 – 149. Aedeagus in ventral view. 134 – 136 — P. klapperichi; 134 – 135 — Wuyishan, Fujian; 136 — Baishanzu, Zhejiang; 137 — P. dauricus, Liaoning; 138 – 141 — P. davidis davidis; 138 — Qinling, Shaanxi; 139 — Siping, Sichuan; 140 — Tianmushan, Zhejiang; 141 — Dabieshan, Anhui; 142 — P. davidis cheni, Taiwan; 143 — P. formosanus, Taiwan; 144 – 145 — P. nigricolor; 144 — holotype, after Boucher 1996, without scale; 145 — Gongshan, Yunnan, same specimen as in Fig. 226; 146 – 147 — P. s h a n i; 146 — holotype, Erlangshan, Sichuan; 147 — paratypes, Siping, Sichuan; 148 – 149 — P. alessandrae; 148 — Zhongdian, Yunnan; 149 — Weixi, Yunnan. Scale bars 1 mm for all." httpUri="https://zenodo.org/record/280635/files/figure.png" pageId="8" pageNumber="9">144–145</figureCitation>
, 
<figureCitation id="13697DD9FFCC6013FE4B46C8B53AFF0E" box="[473,517,186,210]" captionStart="FIGURES 168 – 199" captionStartId="31.[151,258,1264,1285]" captionTargetBox="[151,1436,191,1213]" captionTargetId="figure@31.[151,1436,191,1213]" captionTargetPageId="31" captionText="FIGURES 168 – 199. Flagellum of the median lobe. 168 – 171 — P. davidis davidis; 168 — Qinling, Shaanxi; 169 — Erlangshan, Sichuan; 170 — Tianmushan, Zhejiang; 171 — Dabieshan, Anhui; 172 — P. davidis cheni, Taiwan; 173 — P. nigricolor, Gongshan, Yunnan, same specimen as in Fig. 226; 174 – 178 — P. klapperichi; 174 — Simianshan, Chongqing; 175 — Wuyishan, Fujian; 176 — Maoershan, Guangxi; 177 — Nanling, Guangdong; 178 — Baishanzu, Zhejiang; 179 — P. dauricus, Liaoning; 180 — P. s h a n i, paratype, Siping; 181 — P. alessandrae, Zhongdian, Yunnan; 182 – 184 — P. p ro s s i; 182 — Siping, Sichuan; 183 — Fanjingshan, Guizhou; 184 — Maoershan, Guangxi; 185 — P. siniaevi, Gongshan; 186 – 187 — P. arcuatus, Sichuan; 188 – 189 — P. haojiani; 188 — holotype, Fanjingshan; 189 — paratype, Leigongshan, Guizhou; 190 — P. kanghianus kanghianus, Hekou, Yunnan; 191 — P. sukkitorum, Tengchong, Yunnan; 192 – 194 — P. triapicalis triapicalis, Sichuan; 195 – 196 — P. triapicalis subspecies; 195 — Dabashan, Chongqing; 196 — Fengyangshan, Zhejiang; 197 — P. formosanus, Taiwan; 198 — Tetrarthrius sukkiti, Gongshan, Yunnan; 199 — Cyclorasis fukinukii, Motuo, Tibet. Figs. 168 – 198 under the same scale. Scale bars 1 mm for all." httpUri="https://zenodo.org/record/280637/files/figure.png" pageId="8" pageNumber="9">173</figureCitation>
, 
<figureCitation id="13697DD9FFCC6013FD8346C8B501FF0E" box="[529,574,186,210]" captionStart="FIGURES 200 – 211" captionStartId="32.[151,257,1891,1912]" captionTargetBox="[193,1398,196,1847]" captionTargetId="figure@32.[193,1398,191,1848]" captionTargetPageId="32" captionText="FIGURES 200 – 211. Female genitalia and the central protrusion of the 9 th tergites (at left bottom or right bottom). 200 — P. davidis davidis, Beijing; 201 — P. davidis davidis, Tianmushan; 202 — P. davidis cheni, Taiwan; 203 — P. davidis davidis, Nanling; 204 — P. nigricolor, Gongshan, Yunnan; 205 – 206 — P. klapperichi; 205 — Wuyishan, Fujian; 206 — Damingshan, Guangxi; 207 — P. dauricus, Liaoning; 208 — P. s h a n i, paratype, Labahe; 209 — P. alessandrae, Zhongdian, Yunnan; 210 – 211 — P. prossi; 210 — Siping, Sichuan; 211 — Fanjingshan, Guizhou. Scale bars 1 mm for all." httpUri="https://zenodo.org/record/280638/files/figure.png" pageId="8" pageNumber="9">204</figureCitation>
, 
<figureCitation id="13697DD9FFCC6013FDDB46C8B594FF0E" box="[585,683,186,210]" captionStart-0="FIGURES 224 – 231" captionStart-1="FIGURES 232 – 233" captionStartId-0="34.[151,258,1829,1850]" captionStartId-1="34.[151,258,1946,1967]" captionTargetBox-0="[157,1411,201,1780]" captionTargetId-0="figure@34.[151,1436,191,1807]" captionTargetPageId-0="34" captionText-0="FIGURES 224 – 231. Habitus of P. nigricolor (under the same scale). 224 — male, Yaojiaping, between Pianma and Lushui; 225 — male, Gongshan; 226 — male, Gongshan; 227 — male, holotype; 228 — male, holotype in ventral view; 229 — female, paratype in ventral view; 230 — female, Gongshan; 231 — female, paratype. Scale bar = 1 cm." captionText-1="FIGURES 232 – 233. Female head and pronotum of P. nigricolor (under the same scale). 232 — Gongshan; 233 — paratype. Scale bar = 1 cm." httpUri-0="https://zenodo.org/record/280640/files/figure.png" pageId="8" pageNumber="9">224–233</figureCitation>
, 
<figureCitation id="13697DD9FFCC6013FD2B46C8B5DBFF0E" box="[697,740,186,210]" captionStart="FIGURE 235" captionStartId="35.[151,244,2001,2022]" captionTargetBox="[264,1307,1289,1992]" captionTargetId="figure@35.[264,1323,1289,1992]" captionTargetPageId="35" captionText="FIGURE 235. Distribution of the other Chinese Prismognathus." httpUri="https://zenodo.org/record/280642/files/figure.png" pageId="8" pageNumber="9">235</figureCitation>
)
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<taxonomicName id="4C521ADFFFCC6013FF054770B561FEC4" ID-CoL="4MHQY" authority="Boucher, 1996: 70" authorityName="Boucher" authorityPageNumber="70" authorityYear="1996" box="[151,606,258,280]" class="Insecta" family="Lucanidae" genus="Prismognathus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="nigricolor">
<emphasis id="B926BD4EFFCC6013FF054770B6A2FEC4" box="[151,413,258,280]" italics="true" pageId="8" pageNumber="9">Prismognathus nigricolor</emphasis>
<treatmentCitation id="0AF3474DFFCC6013FE314770B561FEC4" author="Boucher" box="[419,606,258,280]" page="70" pageId="8" pageNumber="9" year="1996">
<bibRefCitation id="EFC31CADFFCC6013FE314770B504FEC4" author="Boucher" box="[419,571,258,280]" pageId="8" pageNumber="9" refString="Boucher, S. (1996) Deux especes inedites sino-birmanes des genres Prismognathus Motschulsky et Cladophyllus Houlbert, et mise en synonymie de C. flavus Didier (Coleoptera, Lucanidae). Coleopteres, 2 (7), 69 - 78." type="journal article" year="1996">Boucher, 1996</bibRefCitation>
: 70
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[
<typeStatus id="54E9DFFEFFCC6013FDF84771B5A7FEC4" box="[618,664,259,280]" pageId="8" pageNumber="9">type</typeStatus>
locality: Chinese Gaoligong, China-Myanmar frontier], 
<figureCitation id="13697DD9FFCC6013FB444770B22EFEC4" box="[1238,1297,258,280]" captionStart="FIGURES 1 – 20" captionStartId="21.[151,257,1392,1413]" captionTargetBox="[156,1436,191,1371]" captionTargetId="figure@21.[151,1436,191,1371]" captionTargetPageId="21" captionText="FIGURES 1 – 20. Cladophyllus, Tetrarthrius and Cyclorasis species. 1 – 4 — Cladophyllus oberthuri, male (Weixi, Yunnan); 5 – 8 — Cladophyllus bousqueti, male (Pianma, Yunnan); 9 – 12 — Cladophyllus nobuhikoi, male (Jinping, Yunnan); 13 – 14 — Cladophyllus oberthuri, female (Weixi, Yunnan); 15 – 16 — Cladophyllus bousqueti, female (Pianma, Yunnan); 17 – 18 — Tetrarthrius sukkiti, female (Motuo, Tibet); 19 – 20 — Cyclorasis nosei, female (Motuo, Tibet). 1, 5, 9, 13, 15, 17, 19 — head and pronotum in dorsal view; 2, 6, 10 — 9 th abdominal segment; 3, 7, 11 — aedeagus; 4, 8, 12 — flagellum; 14, 16, 18, 20 — female genitalia. Scale bars 5 mm for habitus, 1 mm for genitalia." httpUri="https://zenodo.org/record/280627/files/figure.png" pageId="8" pageNumber="9">Fig. 1</figureCitation>
, head; 
<figureCitation id="13697DD9FFCC6013FAC94770B2A8FEC4" box="[1371,1431,258,280]" captionStart="FIGURES 1 – 20" captionStartId="21.[151,257,1392,1413]" captionTargetBox="[156,1436,191,1371]" captionTargetId="figure@21.[151,1436,191,1371]" captionTargetPageId="21" captionText="FIGURES 1 – 20. Cladophyllus, Tetrarthrius and Cyclorasis species. 1 – 4 — Cladophyllus oberthuri, male (Weixi, Yunnan); 5 – 8 — Cladophyllus bousqueti, male (Pianma, Yunnan); 9 – 12 — Cladophyllus nobuhikoi, male (Jinping, Yunnan); 13 – 14 — Cladophyllus oberthuri, female (Weixi, Yunnan); 15 – 16 — Cladophyllus bousqueti, female (Pianma, Yunnan); 17 – 18 — Tetrarthrius sukkiti, female (Motuo, Tibet); 19 – 20 — Cyclorasis nosei, female (Motuo, Tibet). 1, 5, 9, 13, 15, 17, 19 — head and pronotum in dorsal view; 2, 6, 10 — 9 th abdominal segment; 3, 7, 11 — aedeagus; 4, 8, 12 — flagellum; 14, 16, 18, 20 — female genitalia. Scale bars 5 mm for habitus, 1 mm for genitalia." httpUri="https://zenodo.org/record/280627/files/figure.png" pageId="8" pageNumber="9">Fig. 5</figureCitation>
, antenna; 
<figureCitation id="13697DD9FFCC6013FEB14753B653FEEB" box="[291,364,289,311]" captionStart="FIGURES 1 – 20" captionStartId="21.[151,257,1392,1413]" captionTargetBox="[156,1436,191,1371]" captionTargetId="figure@21.[151,1436,191,1371]" captionTargetPageId="21" captionText="FIGURES 1 – 20. Cladophyllus, Tetrarthrius and Cyclorasis species. 1 – 4 — Cladophyllus oberthuri, male (Weixi, Yunnan); 5 – 8 — Cladophyllus bousqueti, male (Pianma, Yunnan); 9 – 12 — Cladophyllus nobuhikoi, male (Jinping, Yunnan); 13 – 14 — Cladophyllus oberthuri, female (Weixi, Yunnan); 15 – 16 — Cladophyllus bousqueti, female (Pianma, Yunnan); 17 – 18 — Tetrarthrius sukkiti, female (Motuo, Tibet); 19 – 20 — Cyclorasis nosei, female (Motuo, Tibet). 1, 5, 9, 13, 15, 17, 19 — head and pronotum in dorsal view; 2, 6, 10 — 9 th abdominal segment; 3, 7, 11 — aedeagus; 4, 8, 12 — flagellum; 14, 16, 18, 20 — female genitalia. Scale bars 5 mm for habitus, 1 mm for genitalia." httpUri="https://zenodo.org/record/280627/files/figure.png" pageId="8" pageNumber="9">Fig. 10</figureCitation>
, mesotibia of male; 
<figureCitation id="13697DD9FFCC6013FDA84753B5BBFEEB" box="[570,644,289,311]" captionStart="FIGURES 1 – 20" captionStartId="21.[151,257,1392,1413]" captionTargetBox="[156,1436,191,1371]" captionTargetId="figure@21.[151,1436,191,1371]" captionTargetPageId="21" captionText="FIGURES 1 – 20. Cladophyllus, Tetrarthrius and Cyclorasis species. 1 – 4 — Cladophyllus oberthuri, male (Weixi, Yunnan); 5 – 8 — Cladophyllus bousqueti, male (Pianma, Yunnan); 9 – 12 — Cladophyllus nobuhikoi, male (Jinping, Yunnan); 13 – 14 — Cladophyllus oberthuri, female (Weixi, Yunnan); 15 – 16 — Cladophyllus bousqueti, female (Pianma, Yunnan); 17 – 18 — Tetrarthrius sukkiti, female (Motuo, Tibet); 19 – 20 — Cyclorasis nosei, female (Motuo, Tibet). 1, 5, 9, 13, 15, 17, 19 — head and pronotum in dorsal view; 2, 6, 10 — 9 th abdominal segment; 3, 7, 11 — aedeagus; 4, 8, 12 — flagellum; 14, 16, 18, 20 — female genitalia. Scale bars 5 mm for habitus, 1 mm for genitalia." httpUri="https://zenodo.org/record/280627/files/figure.png" pageId="8" pageNumber="9">Fig. 13</figureCitation>
, male genitalia.
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<emphasis id="B926BD4EFFCC6013FF054711B520FEA0" bold="true" box="[151,543,355,380]" pageId="8" pageNumber="9">
Material examined. 
<collectingCountry id="F34521CCFFCC6013FE1A4711B520FEA0" box="[392,543,355,380]" name="Myanmar" pageId="8" pageNumber="9">MYANMAR</collectingCountry>
</emphasis>
: 1 3 (
<collectionCode id="ED43F999FFCC6013FDF44716B5FFFEA0" LSID="urn:lsid:biocol.org:col:34988" box="[614,704,356,380]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="8" pageNumber="9">MNHN</collectionCode>
, figs. 28, 227–228), 
<typeStatus id="54E9DFFEFFCC6013FC3B4716B331FEA0" box="[937,1038,356,380]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
, labeled as “Chaine GAOLIGONG front. Sino-Birm. 
<quantity id="4CAACCB9FFCC6013FE1E47F5B6E1FE43" box="[396,478,391,415]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="9" unit="m" value="2000.0">2000m</quantity>
. vol. 
<date id="FFEC479CFFCC6013FDD447F5B5AEFE43" box="[582,657,391,415]" pageId="8" pageNumber="9" value="1995-07">VII.95</date>
J.H.BOUSQUET/ collection S. BOUCHER/ 
<typeStatus id="54E9DFFEFFCC6013FA9247F5B2ACFE43" box="[1280,1427,391,415]" pageId="8" pageNumber="9" type="holotype">HOLOTYPE</typeStatus>
/ 
<taxonomicName id="4C521ADFFFCC6013FF0547DEB6C6FE18" box="[151,505,428,452]" class="Insecta" family="Lucanidae" genus="Prismognathus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="nigricolor">PRISMOGNATHUS nigricolor</taxonomicName>
<typeStatus id="54E9DFFEFFCC6013FD9147DEB5A8FE18" box="[515,663,428,452]" pageId="8" pageNumber="9" type="holotype">HOLOTYPE</typeStatus>
male S. Boucher det. 96”; 1 Ƥ (
<collectionCode id="ED43F999FFCC6013FB8347DEB354FE18" LSID="urn:lsid:biocol.org:col:34988" box="[1041,1131,428,452]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="8" pageNumber="9">MNHN</collectionCode>
, figs. 112, 229, 231, 233), 
<typeStatus id="54E9DFFEFFCC6013FF0547A2B7C5FE34" box="[151,250,464,488]" pageId="8" pageNumber="9" type="paratype">paratype</typeStatus>
, labeled as “Chaine GAOLIGONG front. Sino-Birm. 
<quantity id="4CAACCB9FFCC6013FCCE47BDB491FE3B" box="[860,942,463,487]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="9" unit="m" value="2000.0">2000m</quantity>
. vol. 
<date id="FFEC479CFFCC6013FC7947BDB309FE3B" box="[1003,1078,463,487]" pageId="8" pageNumber="9" value="1995-07">VII.95</date>
J.H.BOUSQUET/ collection S. BOUCHER/ 
<typeStatus id="54E9DFFEFFCC6013FEA24786B684FDD0" box="[304,443,500,524]" pageId="8" pageNumber="9" type="paratype">PARATYPE</typeStatus>
/ 
<taxonomicName id="4C521ADFFFCC6013FE5D4786B40BFDD0" box="[463,820,500,524]" class="Insecta" family="Lucanidae" genus="Prismognathus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="nigricolor">PRISMOGNATHUS nigricolor</taxonomicName>
<typeStatus id="54E9DFFEFFCC6013FCD24786B4F4FDD0" box="[832,971,500,524]" pageId="8" pageNumber="9" type="paratype">PARATYPE</typeStatus>
female S. Boucher det. 96/ MUSEUM 
<collectionCode id="ED43F999FFCC6013FF054465B7DEFDF3" box="[151,225,535,559]" pageId="8" pageNumber="9">PARIS</collectionCode>
”. 
<emphasis id="B926BD4EFFCC6013FE904465B6E4FDEC" bold="true" box="[258,475,535,560]" pageId="8" pageNumber="9">
<collectingCountry id="F34521CCFFCC6013FE904465B65CFDEC" box="[258,355,535,560]" name="China" pageId="8" pageNumber="9">CHINA</collectingCountry>
: Yunnan:
</emphasis>
1 3 (
<collectionCode id="ED43F999FFCC6013FDB44465B549FDF3" box="[550,630,535,559]" httpUri="http://grbio.org/cool/3dng-x47a" name="Carthage College" pageId="8" pageNumber="9">CCCC</collectionCode>
), Nujiang, Lushui County, Yaojiaping (on route between Lushui and Pianma), 
<date id="FFEC479CFFCC6013FE9D444EB6A4FD88" box="[271,411,572,596]" pageId="8" pageNumber="9" value="2005-07-25">25.VII.2005</date>
, Ben-Yong Mao &amp; Ji-Shan Xu leg.; 3 3, 1 Ƥ (
<collectionCode id="ED43F999FFCC6013FBD2444EB3B1FD88" box="[1088,1166,572,596]" pageId="8" pageNumber="9">CLYB</collectionCode>
), Nujiang, Gongshan, 
<date id="FFEC479CFFCC6013FF05442DB66BFDAB" box="[151,340,607,631]" pageId="8" pageNumber="9" value="2011-08-18" valueMax="2011-08-19" valueMin="2011-08-18">18–19.VIII.2011</date>
, Ying-Bing Li leg.
</paragraph>
<paragraph id="8BED615CFFCC6013FF5544F1B308FC4B" blockId="8.[151,1436,355,1783]" pageId="8" pageNumber="9">
<emphasis id="B926BD4EFFCC6013FF5544F1B656FD40" bold="true" box="[199,361,643,668]" pageId="8" pageNumber="9">Identification</emphasis>
. Only one male and one female are known in the 
<typeStatus id="54E9DFFEFFCC6013FC3D44F7B4E0FD41" box="[943,991,645,669]" pageId="8" pageNumber="9">type</typeStatus>
series collected by Mr. Bousquet. The 
<typeStatus id="54E9DFFEFFCC6013FF0544DAB7F8FD1C" box="[151,199,680,704]" pageId="8" pageNumber="9">type</typeStatus>
locality, as described by Boucher (personal communication), is actually in 
<collectingCountry id="F34521CCFFCC6013FBA044DAB3A0FD1C" box="[1074,1183,680,704]" name="Myanmar" pageId="8" pageNumber="9">Myanmar</collectingCountry>
and a few kilometers away from the Chinese town of Pianma. The specimens from northern 
<collectingCountry id="F34521CCFFCC6013FC3E44BFB326FD39" box="[940,1049,717,741]" name="Myanmar" pageId="8" pageNumber="9">Myanmar</collectingCountry>
figured in last publications (
<bibRefCitation id="EFC31CADFFCC6013FAC544BEB7E4FCDB" author="Nagai" pageId="8" pageNumber="9" refString="Nagai, S. (2000 a) Twelve new species, three new subspecies, two new status and with the check list of the family Lucanidae of northern Myanmar (Coleoptera: Lamellicornia), pp. 74 - 108. In: Nose Y. (ed.), The Insects of Myanmar, collected by Nose's Expedition. Notes on Eurasian Insects, 3. Y. Aoki, Osaka, 115 pp, 31 plates." type="book chapter" year="2000" yearSuffix="a">Nagai 2000a</bibRefCitation>
, 
<bibRefCitation id="EFC31CADFFCC6013FF78449DB652FCDB" author="Fujita" box="[234,365,751,775]" pageId="8" pageNumber="9" refString="Fujita, H. (2010) The Lucanid Beetles of the World. Mushi-Sha, Tokyo, 472 pp, 248 plates." type="book" year="2010">Fujita 2010</bibRefCitation>
) can not belong to this species, because of the markedly larger antennal club; they may belong to an undescribed species. This species was recently rediscovered by some Chinese collectors from Pianma area, only a few kilometers away from the 
<typeStatus id="54E9DFFEFFCC6013FDD6454AB54BFC8C" box="[580,628,824,848]" pageId="8" pageNumber="9">type</typeStatus>
locality, and Gongshan area, with the small, the medium-sized and the large males and the females collected together by light traps. The male 
<typeStatus id="54E9DFFEFFCC6013FC2F452EB320FCA8" box="[957,1055,860,884]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
is proved to be a small form, and the large form of this species is somewhat similar to 
<taxonomicName id="4C521ADFFFCC6013FD7745F3B46DFC44" box="[741,850,896,920]" class="Insecta" family="Lucanidae" genus="Prismognathus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="davidis">
<emphasis id="B926BD4EFFCC6013FD7745F3B46DFC44" box="[741,850,896,920]" italics="true" pageId="8" pageNumber="9">P. davidis</emphasis>
</taxonomicName>
in external features.
</paragraph>
<paragraph id="8BED615CFFCC6013FF5545D6B266FB90" blockId="8.[151,1436,355,1783]" pageId="8" pageNumber="9">
The recently collected small male specimen (
<figureCitation id="13697DD9FFCC6013FD5D45D6B40EFC60" box="[719,817,932,956]" captionStart="FIGURES 224 – 231" captionStartId="34.[151,258,1829,1850]" captionTargetBox="[157,1411,201,1780]" captionTargetId="figure@34.[151,1436,191,1807]" captionTargetPageId="34" captionText="FIGURES 224 – 231. Habitus of P. nigricolor (under the same scale). 224 — male, Yaojiaping, between Pianma and Lushui; 225 — male, Gongshan; 226 — male, Gongshan; 227 — male, holotype; 228 — male, holotype in ventral view; 229 — female, paratype in ventral view; 230 — female, Gongshan; 231 — female, paratype. Scale bar = 1 cm." httpUri="https://zenodo.org/record/280640/files/figure.png" pageId="8" pageNumber="9">Fig. 226</figureCitation>
) from Gongshan is nearly identical with the 
<typeStatus id="54E9DFFEFFCC6013FAA845D6B2A3FC60" box="[1338,1436,932,956]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
(
<figureCitation id="13697DD9FFCC6013FF0D45B5B7C1FC03" box="[159,254,967,991]" captionStart="FIGURES 224 – 231" captionStartId="34.[151,258,1829,1850]" captionTargetBox="[157,1411,201,1780]" captionTargetId="figure@34.[151,1436,191,1807]" captionTargetPageId="34" captionText="FIGURES 224 – 231. Habitus of P. nigricolor (under the same scale). 224 — male, Yaojiaping, between Pianma and Lushui; 225 — male, Gongshan; 226 — male, Gongshan; 227 — male, holotype; 228 — male, holotype in ventral view; 229 — female, paratype in ventral view; 230 — female, Gongshan; 231 — female, paratype. Scale bar = 1 cm." httpUri="https://zenodo.org/record/280640/files/figure.png" pageId="8" pageNumber="9">Fig. 227</figureCitation>
) of 
<taxonomicName id="4C521ADFFFCC6013FEB945BBB687FC3C" box="[299,440,968,992]" class="Insecta" family="Lucanidae" genus="Prismognathus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="nigricolor">
<emphasis id="B926BD4EFFCC6013FEB945BBB687FC3C" box="[299,440,968,992]" italics="true" pageId="8" pageNumber="9">P. nigricolor</emphasis>
</taxonomicName>
in all the external features; its male genitalia (
<figureCitation id="13697DD9FFCC6013FC2E45B5B324FC03" box="[956,1051,967,991]" captionStart="FIGURES 134 – 149" captionStartId="29.[151,257,1583,1604]" captionTargetBox="[151,1427,211,1495]" captionTargetId="figure@29.[151,1436,191,1503]" captionTargetPageId="29" captionText="FIGURES 134 – 149. Aedeagus in ventral view. 134 – 136 — P. klapperichi; 134 – 135 — Wuyishan, Fujian; 136 — Baishanzu, Zhejiang; 137 — P. dauricus, Liaoning; 138 – 141 — P. davidis davidis; 138 — Qinling, Shaanxi; 139 — Siping, Sichuan; 140 — Tianmushan, Zhejiang; 141 — Dabieshan, Anhui; 142 — P. davidis cheni, Taiwan; 143 — P. formosanus, Taiwan; 144 – 145 — P. nigricolor; 144 — holotype, after Boucher 1996, without scale; 145 — Gongshan, Yunnan, same specimen as in Fig. 226; 146 – 147 — P. s h a n i; 146 — holotype, Erlangshan, Sichuan; 147 — paratypes, Siping, Sichuan; 148 – 149 — P. alessandrae; 148 — Zhongdian, Yunnan; 149 — Weixi, Yunnan. Scale bars 1 mm for all." httpUri="https://zenodo.org/record/280635/files/figure.png" pageId="8" pageNumber="9">Fig. 145</figureCitation>
) are generally identical with those of the 
<typeStatus id="54E9DFFEFFCC6013FF4D459EB67EFBD8" box="[223,321,1004,1028]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
(
<figureCitation id="13697DD9FFCC6013FEC2459EB68FFBD8" box="[336,432,1004,1028]" captionStart="FIGURES 134 – 149" captionStartId="29.[151,257,1583,1604]" captionTargetBox="[151,1427,211,1495]" captionTargetId="figure@29.[151,1436,191,1503]" captionTargetPageId="29" captionText="FIGURES 134 – 149. Aedeagus in ventral view. 134 – 136 — P. klapperichi; 134 – 135 — Wuyishan, Fujian; 136 — Baishanzu, Zhejiang; 137 — P. dauricus, Liaoning; 138 – 141 — P. davidis davidis; 138 — Qinling, Shaanxi; 139 — Siping, Sichuan; 140 — Tianmushan, Zhejiang; 141 — Dabieshan, Anhui; 142 — P. davidis cheni, Taiwan; 143 — P. formosanus, Taiwan; 144 – 145 — P. nigricolor; 144 — holotype, after Boucher 1996, without scale; 145 — Gongshan, Yunnan, same specimen as in Fig. 226; 146 – 147 — P. s h a n i; 146 — holotype, Erlangshan, Sichuan; 147 — paratypes, Siping, Sichuan; 148 – 149 — P. alessandrae; 148 — Zhongdian, Yunnan; 149 — Weixi, Yunnan. Scale bars 1 mm for all." httpUri="https://zenodo.org/record/280635/files/figure.png" pageId="8" pageNumber="9">Fig. 144</figureCitation>
) except for a wider flagellum apex (
<figureCitation id="13697DD9FFCC6013FCD8459EB493FBD8" box="[842,940,1004,1028]" captionStart="FIGURES 168 – 199" captionStartId="31.[151,258,1264,1285]" captionTargetBox="[151,1436,191,1213]" captionTargetId="figure@31.[151,1436,191,1213]" captionTargetPageId="31" captionText="FIGURES 168 – 199. Flagellum of the median lobe. 168 – 171 — P. davidis davidis; 168 — Qinling, Shaanxi; 169 — Erlangshan, Sichuan; 170 — Tianmushan, Zhejiang; 171 — Dabieshan, Anhui; 172 — P. davidis cheni, Taiwan; 173 — P. nigricolor, Gongshan, Yunnan, same specimen as in Fig. 226; 174 – 178 — P. klapperichi; 174 — Simianshan, Chongqing; 175 — Wuyishan, Fujian; 176 — Maoershan, Guangxi; 177 — Nanling, Guangdong; 178 — Baishanzu, Zhejiang; 179 — P. dauricus, Liaoning; 180 — P. s h a n i, paratype, Siping; 181 — P. alessandrae, Zhongdian, Yunnan; 182 – 184 — P. p ro s s i; 182 — Siping, Sichuan; 183 — Fanjingshan, Guizhou; 184 — Maoershan, Guangxi; 185 — P. siniaevi, Gongshan; 186 – 187 — P. arcuatus, Sichuan; 188 – 189 — P. haojiani; 188 — holotype, Fanjingshan; 189 — paratype, Leigongshan, Guizhou; 190 — P. kanghianus kanghianus, Hekou, Yunnan; 191 — P. sukkitorum, Tengchong, Yunnan; 192 – 194 — P. triapicalis triapicalis, Sichuan; 195 – 196 — P. triapicalis subspecies; 195 — Dabashan, Chongqing; 196 — Fengyangshan, Zhejiang; 197 — P. formosanus, Taiwan; 198 — Tetrarthrius sukkiti, Gongshan, Yunnan; 199 — Cyclorasis fukinukii, Motuo, Tibet. Figs. 168 – 198 under the same scale. Scale bars 1 mm for all." httpUri="https://zenodo.org/record/280637/files/figure.png" pageId="8" pageNumber="9">Fig. 173</figureCitation>
); it is obvious that the flagellum apex of the 
<typeStatus id="54E9DFFEFFCC6013FF05427DB7C6FBFB" box="[151,249,1039,1063]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
was damaged as it is very short and not enlarged at all. A medium-sized male (
<figureCitation id="13697DD9FFCC6013FBE6427DB3EAFBFB" box="[1140,1237,1039,1063]" captionStart="FIGURES 224 – 231" captionStartId="34.[151,258,1829,1850]" captionTargetBox="[157,1411,201,1780]" captionTargetId="figure@34.[151,1436,191,1807]" captionTargetPageId="34" captionText="FIGURES 224 – 231. Habitus of P. nigricolor (under the same scale). 224 — male, Yaojiaping, between Pianma and Lushui; 225 — male, Gongshan; 226 — male, Gongshan; 227 — male, holotype; 228 — male, holotype in ventral view; 229 — female, paratype in ventral view; 230 — female, Gongshan; 231 — female, paratype. Scale bar = 1 cm." httpUri="https://zenodo.org/record/280640/files/figure.png" pageId="8" pageNumber="9">Fig. 225</figureCitation>
) and a large male (
<figureCitation id="13697DD9FFCC6013FF0D4246B7C0FB90" box="[159,255,1076,1100]" captionStart="FIGURES 224 – 231" captionStartId="34.[151,258,1829,1850]" captionTargetBox="[157,1411,201,1780]" captionTargetId="figure@34.[151,1436,191,1807]" captionTargetPageId="34" captionText="FIGURES 224 – 231. Habitus of P. nigricolor (under the same scale). 224 — male, Yaojiaping, between Pianma and Lushui; 225 — male, Gongshan; 226 — male, Gongshan; 227 — male, holotype; 228 — male, holotype in ventral view; 229 — female, paratype in ventral view; 230 — female, Gongshan; 231 — female, paratype. Scale bar = 1 cm." httpUri="https://zenodo.org/record/280640/files/figure.png" pageId="8" pageNumber="9">Fig. 224</figureCitation>
) are also dissected for a comparison and they show no differences from the small male in genitalia.
</paragraph>
<paragraph id="8BED615CFFCC6013FF554225B7D0FB6B" blockId="8.[151,1436,355,1783]" pageId="8" pageNumber="9">
The recently collected female specimen (
<figureCitation id="13697DD9FFCC6013FD004225B5C1FBB3" box="[658,766,1111,1135]" captionStart="FIGURES 224 – 231" captionStartId="34.[151,258,1829,1850]" captionTargetBox="[157,1411,201,1780]" captionTargetId="figure@34.[151,1436,191,1807]" captionTargetPageId="34" captionText="FIGURES 224 – 231. Habitus of P. nigricolor (under the same scale). 224 — male, Yaojiaping, between Pianma and Lushui; 225 — male, Gongshan; 226 — male, Gongshan; 227 — male, holotype; 228 — male, holotype in ventral view; 229 — female, paratype in ventral view; 230 — female, Gongshan; 231 — female, paratype. Scale bar = 1 cm." httpUri="https://zenodo.org/record/280640/files/figure.png" pageId="8" pageNumber="9">Figs. 230</figureCitation>
, 
<figureCitation id="13697DD9FFCC6013FC9B4225B40BFBB3" box="[777,820,1111,1135]" captionStart="FIGURES 232 – 233" captionStartId="34.[151,258,1946,1967]" captionText="FIGURES 232 – 233. Female head and pronotum of P. nigricolor (under the same scale). 232 — Gongshan; 233 — paratype. Scale bar = 1 cm." pageId="8" pageNumber="9">232</figureCitation>
) is only slightly different from the 
<typeStatus id="54E9DFFEFFCC6013FB2F422AB222FBAC" box="[1213,1309,1112,1136]" pageId="8" pageNumber="9" type="paratype">paratype</typeStatus>
(
<figureCitation id="13697DD9FFCC6013FABE4225B2A8FBB3" box="[1324,1431,1111,1135]" captionStart="FIGURES 224 – 231" captionStartId="34.[151,258,1829,1850]" captionTargetBox="[157,1411,201,1780]" captionTargetId="figure@34.[151,1436,191,1807]" captionTargetPageId="34" captionText="FIGURES 224 – 231. Habitus of P. nigricolor (under the same scale). 224 — male, Yaojiaping, between Pianma and Lushui; 225 — male, Gongshan; 226 — male, Gongshan; 227 — male, holotype; 228 — male, holotype in ventral view; 229 — female, paratype in ventral view; 230 — female, Gongshan; 231 — female, paratype. Scale bar = 1 cm." httpUri="https://zenodo.org/record/280640/files/figure.png" pageId="8" pageNumber="9">Figs. 231</figureCitation>
, 
<figureCitation id="13697DD9FFCC6013FF05420EB7FCFB48" box="[151,195,1148,1172]" captionStart="FIGURES 232 – 233" captionStartId="34.[151,258,1946,1967]" captionText="FIGURES 232 – 233. Female head and pronotum of P. nigricolor (under the same scale). 232 — Gongshan; 233 — paratype. Scale bar = 1 cm." pageId="8" pageNumber="9">233</figureCitation>
) in the length-width ratio of the pronotum; however such variation is commonly found in the 
<taxonomicName id="4C521ADFFFCC6013FB60420EB2A3FB48" box="[1266,1436,1148,1172]" class="Insecta" family="Lucanidae" genus="Prismognathus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B926BD4EFFCC6013FB60420EB2A3FB48" box="[1266,1436,1148,1172]" italics="true" pageId="8" pageNumber="9">Prismognathus</emphasis>
</taxonomicName>
species.
</paragraph>
</subSubSection>
<subSubSection id="C34832D7FFCC6013FF5542B1B266F973" pageId="8" pageNumber="9" type="diagnosis">
<paragraph id="8BED615CFFCC6013FF5542B1B343FAF8" blockId="8.[151,1436,355,1783]" pageId="8" pageNumber="9">
<emphasis id="B926BD4EFFCC6013FF5542B1B605FB00" bold="true" box="[199,314,1219,1244]" pageId="8" pageNumber="9">Diagnosis</emphasis>
. The large and medium-sized males of 
<taxonomicName id="4C521ADFFFCC6013FC9742B7B4AAFB00" box="[773,917,1220,1244]" class="Insecta" family="Lucanidae" genus="Prismognathus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="nigricolor">
<emphasis id="B926BD4EFFCC6013FC9742B7B4AAFB00" box="[773,917,1220,1244]" italics="true" pageId="8" pageNumber="9">P. nigricolor</emphasis>
</taxonomicName>
are similar to those of 
<taxonomicName id="4C521ADFFFCC6013FB3742B7B22BFB00" box="[1189,1300,1220,1244]" class="Insecta" family="Lucanidae" genus="Prismognathus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="davidis">
<emphasis id="B926BD4EFFCC6013FB3742B7B22BFB00" box="[1189,1300,1220,1244]" italics="true" pageId="8" pageNumber="9">P. davidis</emphasis>
</taxonomicName>
, but can be distinguished by the following combination of characters: 1) outer margin of the mandible less sinuate; 2) the gap between the ventral subbasal teeth and the median teeth of the mandible markedly larger.
</paragraph>
<paragraph id="8BED615CFFCC6013FF55435DB4D5FA68" blockId="8.[151,1436,355,1783]" pageId="8" pageNumber="9">
The small males of 
<taxonomicName id="4C521ADFFFCC6013FE374343B50DFA94" box="[421,562,1328,1352]" class="Insecta" family="Lucanidae" genus="Prismognathus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="nigricolor">
<emphasis id="B926BD4EFFCC6013FE374343B50DFA94" box="[421,562,1328,1352]" italics="true" pageId="8" pageNumber="9">P. nigricolor</emphasis>
</taxonomicName>
are similar to 
<taxonomicName id="4C521ADFFFCC6013FD444343B410FA94" box="[726,815,1328,1352]" class="Insecta" family="Lucanidae" genus="Prismognathus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="shani">
<emphasis id="B926BD4EFFCC6013FD444343B410FA94" box="[726,815,1328,1352]" italics="true" pageId="8" pageNumber="9">P. shani</emphasis>
</taxonomicName>
(described below) and the small males of 
<taxonomicName id="4C521ADFFFCC6013FA9E4343B246FA94" box="[1292,1401,1328,1352]" class="Insecta" family="Lucanidae" genus="Prismognathus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="davidis">
<emphasis id="B926BD4EFFCC6013FA9E4343B246FA94" box="[1292,1401,1328,1352]" italics="true" pageId="8" pageNumber="9">P. davidis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C521ADFFFCC6013FA154343B628FAB0" class="Insecta" family="Lucanidae" genus="Prismognathus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="klapperichi">
<emphasis id="B926BD4EFFCC6013FA154343B628FAB0" italics="true" pageId="8" pageNumber="9">P. klapperichi</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C521ADFFFCC6013FEDE4327B6F6FAB0" box="[332,457,1364,1388]" class="Insecta" family="Lucanidae" genus="Prismognathus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="dauricus">
<emphasis id="B926BD4EFFCC6013FEDE4327B6F6FAB0" box="[332,457,1364,1388]" italics="true" pageId="8" pageNumber="9">P. dauricus</emphasis>
</taxonomicName>
, but can be distinguished from all of them by a markedly larger gap between the ventral subbasal teeth and the median teeth of the mandible. The male flagellum apex of 
<taxonomicName id="4C521ADFFFCC6013FBBE430BB386FA4C" box="[1068,1209,1400,1424]" class="Insecta" family="Lucanidae" genus="Prismognathus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="nigricolor">
<emphasis id="B926BD4EFFCC6013FBBE430BB386FA4C" box="[1068,1209,1400,1424]" italics="true" pageId="8" pageNumber="9">P. nigricolor</emphasis>
</taxonomicName>
is nearly as wide as that of 
<emphasis id="B926BD4EFFCC6013FF7443EFB670FA68" box="[230,335,1436,1460]" italics="true" pageId="8" pageNumber="9">P. d a v i d i s</emphasis>
, wider than that of 
<emphasis id="B926BD4EFFCC6013FDBC43EFB414FA68" box="[558,811,1436,1460]" italics="true" pageId="8" pageNumber="9">P. s h a n i, P. k l a p p e r i c h i</emphasis>
and 
<taxonomicName id="4C521ADFFFCC6013FCF743EFB4DCFA68" box="[869,995,1436,1460]" class="Insecta" family="Lucanidae" genus="Prismognathus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="dauricus">
<emphasis id="B926BD4EFFCC6013FCF743EFB4DCFA68" box="[869,995,1436,1460]" italics="true" pageId="8" pageNumber="9">P. dauricus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BED615CFFCC6013FF5543CDB4B0F998" blockId="8.[151,1436,355,1783]" pageId="8" pageNumber="9">
The females of 
<taxonomicName id="4C521ADFFFCC6013FEEA43B3B539FA04" box="[376,518,1472,1496]" class="Insecta" family="Lucanidae" genus="Prismognathus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="nigricolor">
<emphasis id="B926BD4EFFCC6013FEEA43B3B539FA04" box="[376,518,1472,1496]" italics="true" pageId="8" pageNumber="9">P. nigricolor</emphasis>
</taxonomicName>
are similar to those of 
<taxonomicName id="4C521ADFFFCC6013FC9E43B3B4A3FA04" box="[780,924,1472,1496]" class="Insecta" family="Lucanidae" genus="Gonometopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="triapicalis">
<emphasis id="B926BD4EFFCC6013FC9E43B3B4A3FA04" box="[780,924,1472,1496]" italics="true" pageId="8" pageNumber="9">P. triapicalis</emphasis>
</taxonomicName>
by having a narrow and sharp tip of canthus, but can be distinguished by the black appearance of the body, the shorter intermandibular projection and the larger hemisternite in female genitalia. The females of 
<taxonomicName id="4C521ADFFFCC6013FD20407BB401F9FC" box="[690,830,1544,1568]" class="Insecta" family="Lucanidae" genus="Prismognathus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="nigricolor">
<emphasis id="B926BD4EFFCC6013FD20407BB401F9FC" box="[690,830,1544,1568]" italics="true" pageId="8" pageNumber="9">P. nigricolor</emphasis>
</taxonomicName>
can be distinguished from those of 
<emphasis id="B926BD4EFFCC6013FB59407BB2A3F9FC" box="[1227,1436,1544,1568]" italics="true" pageId="8" pageNumber="9">
<taxonomicName id="4C521ADFFFCC6013FB59407BB205F9FC" box="[1227,1338,1544,1568]" class="Insecta" family="Lucanidae" genus="Prismognathus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="davidis">P. davidis</taxonomicName>
, 
<taxonomicName id="4C521ADFFFCC6013FAD6407BB2A3F9FC" box="[1348,1436,1544,1568]" class="Insecta" family="Lucanidae" genus="Prismognathus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="shani">P. shani</taxonomicName>
</emphasis>
and 
<taxonomicName id="4C521ADFFFCC6013FF54405FB656F998" box="[198,361,1580,1604]" class="Insecta" family="Lucanidae" genus="Prismognathus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="alessandrae">
<emphasis id="B926BD4EFFCC6013FF54405FB656F998" box="[198,361,1580,1604]" italics="true" pageId="8" pageNumber="9">P. alessandrae</emphasis>
</taxonomicName>
by having a narrower and sharper tip of canthus.
</paragraph>
<paragraph id="8BED615CFFCC6013FF55403DB266F973" blockId="8.[151,1436,355,1783]" pageId="8" pageNumber="9">
<emphasis id="B926BD4EFFCC6013FF55403DB6A8F9B4" bold="true" box="[199,407,1615,1640]" pageId="8" pageNumber="9">Taxonomic notes.</emphasis>
The male genitalic differences between 
<taxonomicName id="4C521ADFFFCC6013FCF64023B4CEF9B4" box="[868,1009,1616,1640]" class="Insecta" family="Lucanidae" genus="Prismognathus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="nigricolor">
<emphasis id="B926BD4EFFCC6013FCF64023B4CEF9B4" box="[868,1009,1616,1640]" italics="true" pageId="8" pageNumber="9">P. nigricolor</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C521ADFFFCC6013FBB94023B3A7F9B4" box="[1067,1176,1616,1640]" class="Insecta" family="Lucanidae" genus="Prismognathus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="davidis">
<emphasis id="B926BD4EFFCC6013FBB94023B3A7F9B4" box="[1067,1176,1616,1640]" italics="true" pageId="8" pageNumber="9">P. davidis</emphasis>
</taxonomicName>
are hardly recognized, but the female genitalic differences between the two species are well marked in size of the hemisternite and length of the spermathecal and spermathecal duct. Therefore it is reasonable to treat 
<taxonomicName id="4C521ADFFFCC6013FC6940EBB3B6F96C" box="[1019,1161,1688,1712]" class="Insecta" family="Lucanidae" genus="Prismognathus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="nigricolor">
<emphasis id="B926BD4EFFCC6013FC6940EBB3B6F96C" box="[1019,1161,1688,1712]" italics="true" pageId="8" pageNumber="9">P. nigricolor</emphasis>
</taxonomicName>
as a valid species.
</paragraph>
</subSubSection>
<subSubSection id="C34832D7FFCC6013FF5540C9B7C1F92B" pageId="8" pageNumber="9" type="distribution">
<paragraph id="8BED615CFFCC6013FF5540C9B7C1F92B" blockId="8.[151,1436,355,1783]" pageId="8" pageNumber="9">
<emphasis id="B926BD4EFFCC6013FF5540C9B65FF908" bold="true" box="[199,352,1723,1748]" pageId="8" pageNumber="9">Distribution.</emphasis>
Western Yunnan (Pianma, Gongshan); northeastern 
<collectingCountry id="F34521CCFFCC6013FC5340CFB311F909" box="[961,1070,1725,1749]" name="Myanmar" pageId="8" pageNumber="9">Myanmar</collectingCountry>
(area close to Pianma, western Yunnan).
</paragraph>
</subSubSection>
</treatment>
</document>