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<document id="8FA2FB92FFA4D063A81C041ABB8F2BBD" ID-CLB-Dataset="22877" ID-DOI="10.5852/ejt.2020.629" ID-GBIF-Dataset="3722aa65-b935-4294-ac20-0d1179da9015" ID-Zenodo-Dep="3762280" ID-ZooBank="ECEBD539-6E3E-45BE-A0CB-264DF3270CC0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1587578640316" checkinUser="valdenar" docAuthor="Herbert, David G." docDate="2020" docId="03C78785463CFFF34239419B35A5FB67" docLanguage="en" docName="ejt-629_herbert.pdf.imf" docOrigin="European Journal of Taxonomy 629" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Phialodonta rivalalea Herbert 2020, gen. et sp. nov." docType="treatment" docUuid="4F0EED11-370A-4828-9A7A-9C320AA7BCFB" docUuidSource="ZooBank" docVersion="7" lastPageNumber="50" masterDocId="FFFEFFFD460CFFC2400243213138FFD9" masterDocTitle="Revision of the aperturally dentate Charopidae (Gastropoda: Stylommatophora) of southern Africa - genus Afrodonta s. lat., with description of five new genera, twelve new species and one new subspecies" masterLastPageNumber="55" masterPageNumber="1" pageNumber="49" updateTime="1698835565728" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="4128A522A5D06B015B6DE2EDCE8E3FF2" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B7BFE62FE71445E46C88C6CB0D59A3E8">
<mods:title id="F395DBF1BC73FA5AFA100086D1DE2356">Revision of the aperturally dentate Charopidae (Gastropoda: Stylommatophora) of southern Africa - genus Afrodonta s. lat., with description of five new genera, twelve new species and one new subspecies</mods:title>
</mods:titleInfo>
<mods:name id="AF064FC58EAE9A23B16B6247EAE0F4B2" type="personal">
<mods:role id="DE28374A837106B536A1ADC06095B5A1">
<mods:roleTerm id="99AEB3F9FBFE8A34995560C67695DD6A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EFF2BD152CB57650E549A6359F0D49B1">Herbert, David G.</mods:namePart>
<mods:nameIdentifier id="D84066F978312F63722C3D5DCA980FFE" type="LSID">urn:lsid:zoobank.org:author:0C09EE45-6198-482E-857A-EF690C2A016F</mods:nameIdentifier>
<mods:affiliation id="2DCE4B1C23D7B47393B57E64DF37CCD2">Department of Natural Sciences, National Museum Wales, Cathays Park, Cardiff, CF 10 3 NP, UK. Formerly at KwaZulu-Natal Museum, P. Bag 9070, Pietermaritzburg 3201, South Africa</mods:affiliation>
<mods:nameIdentifier id="28DAEC5F0F0D66F5B02D3E5228383F4C" type="email">phasianella@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="45F0136F51778F80910539B140191E95">text</mods:typeOfResource>
<mods:relatedItem id="EC4E075707EADEC0F608D9014CAABCAB" type="host">
<mods:titleInfo id="6DD2368A2306F460114F966264FF383A">
<mods:title id="2C125D04BD90C7B4970B08379FF5B58B">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="C76574415C07A85C79B898A2995D54CE">
<mods:date id="3E07C2317A146F3B40683E1C330E623E">2020</mods:date>
<mods:detail id="CAAA2C6CF5C2625522211D8CF6698AE0" type="pubDate">
<mods:number id="23118D3E3BA6408EB689DA9CC229B6E3">2020-04-17</mods:number>
</mods:detail>
<mods:detail id="708F285D9268E45D19617F335F6788DF" type="volume">
<mods:number id="FD828557874EEEDAD3784F5130456777">629</mods:number>
</mods:detail>
<mods:extent id="1C27E61348B5F6C0B92292B369A7651F" unit="page">
<mods:start id="3C5C47EBAEA9C95A336059FA30E05009">1</mods:start>
<mods:end id="7F335E38CCFA6562163F8945853FB27D">55</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="58663D672D887A4A3C1BFE6D14EA9214">journal article</mods:classification>
<mods:identifier id="F7077DDC3C9F608A9AC87D56FB6D4AA9" type="CLB-Dataset">22877</mods:identifier>
<mods:identifier id="4014763C0497B1C66CDC290FEFE4F7F5" type="DOI">10.5852/ejt.2020.629</mods:identifier>
<mods:identifier id="1EB5CCFA1121E211568EB4A297B77F16" type="GBIF-Dataset">3722aa65-b935-4294-ac20-0d1179da9015</mods:identifier>
<mods:identifier id="CF47091E35E97D8422E722D88A396F92" type="Zenodo-Dep">3762280</mods:identifier>
<mods:identifier id="9398FABDFF78F7F2D344E05510B9A189" type="ZooBank">ECEBD539-6E3E-45BE-A0CB-264DF3270CC0</mods:identifier>
</mods:mods>
<treatment id="03C78785463CFFF34239419B35A5FB67" ID-DOI="http://doi.org/10.5281/zenodo.3804768" ID-GBIF-Taxon="163732358" ID-Zenodo-Dep="3804768" LSID="urn:lsid:zoobank.org:act:4F0EED11-370A-4828-9A7A-9C320AA7BCFB" httpUri="http://treatment.plazi.org/id/03C78785463CFFF34239419B35A5FB67" lastPageId="49" lastPageNumber="50" pageId="48" pageNumber="49">
<subSubSection id="C3746518463CFFF24239419B35F4FD20" pageId="48" pageNumber="49" type="nomenclature">
<paragraph id="8BD13693463CFFF24239419B32C0FD0C" blockId="48.[359,1228,698,797]" box="[571,1016,698,725]" pageId="48" pageNumber="49">
<heading id="D09981FF463CFFF24239419B32C0FD0C" box="[571,1016,698,725]" centered="true" fontSize="11" level="2" pageId="48" pageNumber="49" reason="2">
<taxonomicName id="4C6E4D10463CFFF24239419B3279FD0C" ID-CoL="7SP56" authority="Herbert, 2020" authorityName="Herbert" authorityYear="2020" box="[571,833,698,725]" class="Gastropoda" family="Charopidae" genus="Phialodonta" kingdom="Animalia" order="Stylommatophora" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="rivalalea" status="gen. et sp. nov.">
<emphasis id="B91AEA81463CFFF24239419B3279FD0C" bold="true" box="[571,833,698,725]" italics="true" pageId="48" pageNumber="49">Phialodonta rivalalea</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22957FA463CFFF2434A419A32C0FD0C" box="[840,1016,699,725]" pageId="48" pageNumber="49" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="8BD13693463CFFF2416541FE35F4FD20" blockId="48.[359,1228,698,797]" box="[359,1228,735,761]" pageId="48" pageNumber="49">
<uri id="FFFF3A91463CFFF2416541FE35F4FD20" box="[359,1228,735,761]" pageId="48" pageNumber="49">
urn:lsid:zoobank.org:act:
<uuid id="FFC80C46463CFFF2428741FE35F4FD20" box="[645,1228,735,761]" pageId="48" pageNumber="49">4F0EED11-370A-4828-9A7A-9C320AA7BCFB</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C3746518463CFFF2429140223299FCC4" box="[659,929,771,797]" pageId="48" pageNumber="49" type="description">
<paragraph id="8BD13693463CFFF2429140223299FCC4" blockId="48.[359,1228,698,797]" box="[659,929,771,797]" pageId="48" pageNumber="49">
<figureCitation id="13552A16463CFFF24291402233D4FCC4" box="[659,748,771,797]" captionStart="Fig" captionStartId="36.[189,232,1860,1886]" captionTargetBox="[302,1285,1199,1820]" captionTargetId="figure@36.[302,1285,1199,1820]" captionTargetPageId="36" captionText="Fig. 14. Distribution of Iterodonta gen. nov. and Phialodonta gen. nov. species. Iterodonta ammonita gen. et sp. nov. (brown triangles), Phialodonta agulhasae gen. et sp. nov. (orange square), P. atromontana gen. et sp. nov. (turquoise square), P. aviana gen. et sp. nov. (royal blue squares), P. introtuberculata (Connolly, 1933) gen. et comb. nov. (red squares), P. perfida (Burnup, 1912) gen. et comb. nov. (green square), P. rivalalea gen. et sp. nov. (pink squares). Contour at 1000 m." figureDoi="http://doi.org/10.5281/zenodo.3762308" httpUri="https://zenodo.org/record/3762308/files/figure.png" pageId="48" pageNumber="49">Figs 14</figureCitation>
,
<figureCitation id="13552A16463CFFF242FA4022322CFCC4" box="[760,788,771,797]" captionStart="Fig" captionStartId="42.[189,232,1895,1921]" captionTargetBox="[190,1395,677,1852]" captionTargetId="figure@42.[189,1399,667,1855]" captionTargetPageId="42" captionText="Fig. 16. Shells of Phialodonta gen. nov. species. A–D. P. introtuberculata (Connolly, 1933) gen. et comb. nov., Karkloof, KwaZulu-Natal, diameter 1.87 mm (NMSA W2438). E–H. P. perfida (Burnup, 1912) gen. et comb. nov., paratype, diameter 1.84 mm (NMSA 2602/T617). I–L. P. rivalalea gen. et sp. nov., holotype, diameter 1.70 mm (NMSA W3114 /T4251)." figureDoi="http://doi.org/10.5281/zenodo.3762312" httpUri="https://zenodo.org/record/3762312/files/figure.png" pageId="48" pageNumber="49">16</figureCitation>
I–L, 19K–L
</paragraph>
</subSubSection>
<subSubSection id="C3746518463CFFF240BF406430F7FBBE" pageId="48" pageNumber="49" type="diagnosis">
<paragraph id="8BD13693463CFFF240BF4064300EFC86" blockId="48.[189,310,837,863]" box="[189,310,837,863]" pageId="48" pageNumber="49">
<heading id="D09981FF463CFFF240BF4064300EFC86" bold="true" box="[189,310,837,863]" fontSize="11" level="3" pageId="48" pageNumber="49" reason="3">
<emphasis id="B91AEA81463CFFF240BF4064300EFC86" bold="true" box="[189,310,837,863]" pageId="48" pageNumber="49">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BD13693463CFFF240BF405530F7FBBE" blockId="48.[189,1399,884,1127]" pageId="48" pageNumber="49">
Shell small, spire slightly to distinctly raised; protoconch lacking axial sculpture; teleoconch sculpture of close-set, compound axial riblets, intervals with 4–5 finer intermediaries and extremely fine, close-set spiral threads; aperture with two narrow, in-running parietal lamellae and a narrow, in-running basocolumellar ridge; palatal region with three denticles recessed
<superScript id="7C1B9BDB463CFFF2438840C032ABFC36" attach="right" box="[906,915,993,1007]" fontSize="6" pageId="48" pageNumber="49">1</superScript>
⁄
<subScript id="17EA34D6463CFFF2439540C83298FC2E" attach="none" box="[919,928,1001,1015]" fontSize="6" pageId="48" pageNumber="49">5</subScript>
–
<superScript id="7C1B9BDB463CFFF243AC40C0328FFC36" attach="none" box="[942,951,993,1007]" fontSize="6" pageId="48" pageNumber="49">1</superScript>
⁄
<subScript id="17EA34D6463CFFF243BE40C832FDFC2E" attach="left" box="[956,965,1001,1015]" fontSize="6" pageId="48" pageNumber="49">4</subScript>
whorl behind outer lip; an additional narrow, thread-like ridge immediately below suture evident by transparency; umbilicus very wide, its margin relatively sharply rounded. Shell pale corneous-brown to pale honey-brown when fresh; diameter up to
<quantity id="4C969B76463CFFF2416C476C30F3FBBE" box="[366,459,1101,1127]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="48" pageNumber="49" unit="mm" value="1.7">1.7 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C3746518463CFFF240BF47AE3507FB25" pageId="48" pageNumber="49" type="etymology">
<paragraph id="8BD13693463CFFF240BF47AE307CFB70" blockId="48.[189,324,1167,1193]" box="[189,324,1167,1193]" pageId="48" pageNumber="49">
<heading id="D09981FF463CFFF240BF47AE307CFB70" bold="true" box="[189,324,1167,1193]" fontSize="11" level="3" pageId="48" pageNumber="49" reason="3">
<emphasis id="B91AEA81463CFFF240BF47AE307CFB70" bold="true" box="[189,324,1167,1193]" pageId="48" pageNumber="49">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BD13693463CFFF240BF479E3507FB25" blockId="48.[189,1399,1214,1277]" pageId="48" pageNumber="49">
From the Latin
<emphasis id="B91AEA81463CFFF24182479E3083FB01" box="[384,443,1215,1240]" italics="true" pageId="48" pageNumber="49">rivus</emphasis>
: a stream, and the Greek
<emphasis id="B91AEA81463CFFF242FC479F3201FB01" box="[766,825,1214,1240]" italics="true" pageId="48" pageNumber="49">laleo</emphasis>
(
<emphasis id="B91AEA81463CFFF2434F479F32A8FB01" box="[845,912,1214,1240]" italics="true" pageId="48" pageNumber="49">λαλεΟ</emphasis>
): to talk, to utter; with reference to the Tsitsikamma region. Tsitsikamma (Outeniqua Khoi): ‘the water that speaks’.
</paragraph>
</subSubSection>
<subSubSection id="C3746518463CFFF240BF46053023F832" pageId="48" pageNumber="49" type="materials_examined">
<paragraph id="8BD13693463CFFF240BF46053092FAE7" blockId="48.[189,426,1316,1342]" box="[189,426,1316,1342]" pageId="48" pageNumber="49">
<heading id="D09981FF463CFFF240BF46053092FAE7" bold="true" box="[189,426,1316,1342]" fontSize="11" level="3" pageId="48" pageNumber="49" reason="3">
<emphasis id="B91AEA81463CFFF240BF46053092FAE7" bold="true" box="[189,426,1316,1342]" pageId="48" pageNumber="49">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BD13693463CFFF240E846753064FAB7" blockId="48.[189,1399,1364,1499]" box="[234,348,1364,1390]" pageId="48" pageNumber="49">
<emphasis id="B91AEA81463CFFF240E846753064FAB7" bold="true" box="[234,348,1364,1390]" pageId="48" pageNumber="49">
<typeStatus id="54D58831463CFFF240E846753064FAB7" box="[234,348,1364,1390]" pageId="48" pageNumber="49" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BD13693463CFFF240BF4659355AFA02" blockId="48.[189,1399,1364,1499]" pageId="48" pageNumber="49">
<materialsCitation id="3B063CCE463CFFF240BF4659355AFA02" ID-GBIF-Occurrence="2605305683" collectedFrom="indigenous forest, in leaf-litter" collectingDate="2004-09-20" collectionCode="NMSA" collectorName="A. Moussalli &amp; D. Stuart-Fox" country="South Africa" elevation="10" latitude="-33.9723" location="Tsitsikamma Nat. Park" longLatPrecision="5" longitude="23.56306" municipality="Nature's Valley" pageId="48" pageNumber="49" specimenCode="W3114/T4251" specimenCount="1" stateProvince="W. Cape" typeStatus="holotype">
<collectingCountry id="F3797603463CFFF240BF465930AAFA4B" box="[189,402,1400,1426]" name="South Africa" pageId="48" pageNumber="49">SOUTH AFRICA</collectingCountry>
•
<collectingRegion id="49AAF871463CFFF241B146593325FA4B" box="[435,541,1400,1426]" pageId="48" pageNumber="49">W. Cape</collectingRegion>
,
<collectingMunicipality id="6BB5ACE9463CFFF2422F465933DFFA4B" box="[557,743,1400,1426]" pageId="48" pageNumber="49">Nature’s Valley</collectingMunicipality>
,
<location id="8EB16048463CFFF242F446593535FA4B" LSID="urn:lsid:plazi:treatment:03C78785463CFFF34239419B35A5FB67:8EB16048463CFFF242F446593535FA4B" box="[758,1037,1400,1426]" country="South Africa" latitude="-33.9723" longLatPrecision="5" longitude="23.56306" municipality="Nature's Valley" name="Tsitsikamma Nat. Park" pageId="48" pageNumber="49" stateProvince="W. Cape">Tsitsikamma Nat. Park</location>
, at camping ground entrance;
<geoCoordinate id="EE5A5054463CFFF240BF46BD307BFA6F" box="[189,323,1436,1462]" degrees="33.9723" direction="south" orientation="latitude" pageId="48" pageNumber="49" precision="5" value="-33.9723">33.9723° S</geoCoordinate>
,
<geoCoordinate id="EE5A5054463CFFF2415046BD30D2FA6F" box="[338,490,1436,1463]" degrees="23.56306" direction="east" orientation="longitude" pageId="48" pageNumber="49" precision="1" value="23.56306">23.56306° E</geoCoordinate>
;
<elevation id="0043D1A0463CFFF241FB46BD3340FA6F" box="[505,632,1436,1462]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="48" pageNumber="49" unit="m" value="10.0">
<quantity id="4C969B76463CFFF241FB46BD330FFA6F" box="[505,567,1436,1462]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="48" pageNumber="49" unit="m" value="10.0">10 m</quantity>
a.s.l.
</elevation>
;
<date id="FFD01053463CFFF2428A46BD3211FA6F" box="[648,809,1436,1462]" pageId="48" pageNumber="49" value="2004-09-20">
<collectingDate id="EF94E9BB463CFFF2428A46BD3211FA6F" box="[648,809,1436,1462]" pageId="48" pageNumber="49" value="2004-09-20">20 Sep. 2004</collectingDate>
</date>
;
<collectorName id="269B5345463CFFF2433446BD32E8FA6F" box="[822,976,1436,1462]" pageId="48" pageNumber="49">A. Moussalli</collectorName>
and
<collectorName id="269B5345463CFFF2440E46BD3597FA6F" box="[1036,1199,1436,1462]" pageId="48" pageNumber="49">D. Stuart-Fox</collectorName>
leg.;
<collectedFrom id="3872756E463CFFF244F446BD30A6FA03" pageId="48" pageNumber="49">indigenous forest, in leaf-litter</collectedFrom>
; diameter
<quantity id="4C969B76463CFFF2421546E133BBFA03" box="[535,643,1472,1498]" metricMagnitude="-3" metricUnit="m" metricValue="1.68" pageId="48" pageNumber="49" unit="mm" value="1.68">1.68 mm</quantity>
, height
<quantity id="4C969B76463CFFF242DD46E13273FA03" box="[735,843,1472,1498]" metricMagnitude="-4" metricUnit="m" metricValue="7.9" pageId="48" pageNumber="49" unit="mm" value="0.79">0.79 mm</quantity>
;
<collectionCode id="ED7FAE56463CFFF2435546E13292FA03" box="[855,938,1472,1498]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="48" pageNumber="49" type="Museum">NMSA</collectionCode>
<specimenCode id="DBC89EE8463CFFF243B246E13566FA02" box="[944,1118,1472,1499]" pageId="48" pageNumber="49">W3114/T4251</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BD13693463CFFF240E84523305FF9C5" blockId="48.[189,1399,1538,1853]" box="[234,359,1538,1564]" pageId="48" pageNumber="49">
<emphasis id="B91AEA81463CFFF240E84523305FF9C5" bold="true" box="[234,359,1538,1564]" pageId="48" pageNumber="49">
<typeStatus id="54D58831463CFFF240E84523305FF9C5" box="[234,359,1538,1564]" pageId="48" pageNumber="49" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BD13693463CFFF240BF450733F7F8E4" blockId="48.[189,1399,1538,1853]" pageId="48" pageNumber="49">
<materialsCitation id="3B063CCE463CFFF240BF45073289F950" ID-GBIF-Occurrence="2605305721" collectedFrom="indigenous forest, in leaf-litter" collectingDate="2004-09-16" collectionCode="NMSA" collectorName="A. Moussalli &amp; D. Stuart- Fox" country="South Africa" elevation="200" latitude="-33.9703" location="Tsitsikamma Nat. Park" longLatPrecision="5" longitude="23.53936" municipality="Nature's Valley" pageId="48" pageNumber="49" specimenCode="W3125/T4253" specimenCount="7" stateProvince="W. Cape" typeStatus="paratype">
<collectingCountry id="F3797603463CFFF240BF450730A8F999" box="[189,400,1574,1600]" name="South Africa" pageId="48" pageNumber="49">SOUTH AFRICA</collectingCountry>
–
<emphasis id="B91AEA81463CFFF241B345073325F999" bold="true" box="[433,541,1574,1600]" pageId="48" pageNumber="49">
<collectingRegion id="49AAF871463CFFF241B345073325F999" box="[433,541,1574,1600]" pageId="48" pageNumber="49">W. Cape</collectingRegion>
</emphasis>
•
<specimenCount id="9D68FD1A463CFFF24238450633E8F999" box="[570,720,1574,1601]" pageId="48" pageNumber="49" type="generic">7 specimens</specimenCount>
;
<collectingMunicipality id="6BB5ACE9463CFFF242DF450732ADF999" box="[733,917,1574,1600]" pageId="48" pageNumber="49">Nature’s Valley</collectingMunicipality>
,
<location id="8EB16048463CFFF243A04507358CF999" LSID="urn:lsid:plazi:treatment:03C78785463CFFF34239419B35A5FB67:8EB16048463CFFF243A04507358CF999" box="[930,1204,1574,1601]" country="South Africa" latitude="-33.9703" longLatPrecision="5" longitude="23.53936" municipality="Nature's Valley" name="Tsitsikamma Nat. Park" pageId="48" pageNumber="49" stateProvince="W. Cape">Tsitsikamma Nat. Park</location>
, scenic drive on the way to park office;
<geoCoordinate id="EE5A5054463CFFF241CB456B3374F9BD" box="[457,588,1610,1636]" degrees="33.9703" direction="south" orientation="latitude" pageId="48" pageNumber="49" precision="5" value="-33.9703">33.9703° S</geoCoordinate>
,
<geoCoordinate id="EE5A5054463CFFF2425B456B33D6F9BC" box="[601,750,1610,1637]" degrees="23.53936" direction="east" orientation="longitude" pageId="48" pageNumber="49" precision="1" value="23.53936">23.53936° E</geoCoordinate>
;
<elevation id="0043D1A0463CFFF242F8456B32BAF9BD" box="[762,898,1610,1637]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="48" pageNumber="49" unit="m" value="200.0">
<quantity id="4C969B76463CFFF242F8456B327BF9BC" box="[762,835,1610,1637]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="48" pageNumber="49" unit="m" value="200.0">200 m</quantity>
a.s.l.
</elevation>
;
<date id="FFD01053463CFFF2438D456B3513F9BD" box="[911,1067,1610,1636]" pageId="48" pageNumber="49" value="2004-09-16">
<collectingDate id="EF94E9BB463CFFF2438D456B3513F9BD" box="[911,1067,1610,1636]" pageId="48" pageNumber="49" value="2004-09-16">16 Sep. 2004</collectingDate>
</date>
;
<collectorName id="269B5345463CFFF24437456B35F4F9BD" box="[1077,1228,1610,1636]" pageId="48" pageNumber="49">A. Moussalli</collectorName>
and
<collectorName id="269B5345463CFFF24501456A31D3F950" pageId="48" pageNumber="49">D. Stuart- Fox</collectorName>
leg.;
<collectedFrom id="3872756E463CFFF24132454F33A5F951" box="[304,669,1646,1672]" pageId="48" pageNumber="49">indigenous forest, in leaf-litter</collectedFrom>
;
<collectionCode id="ED7FAE56463CFFF242A8454F33C5F951" box="[682,765,1646,1672]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="48" pageNumber="49" type="Museum">NMSA</collectionCode>
<specimenCode id="DBC89EE8463CFFF24307454F3289F950" box="[773,945,1646,1673]" pageId="48" pageNumber="49">W3125/T4253</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B063CCE463CFFF243CE454F33CAF975" ID-GBIF-Occurrence="2605305678" collectingDate="2004-09-20" collectionCode="NMSA" collectorName="A. Moussalli &amp; D. Stuart-Fox" country="South Africa" elevation="10" latitude="-33.9723" location="E. Cape" longLatPrecision="5" longitude="23.56306" municipality="Nature's Valley" pageId="48" pageNumber="49" specimenCode="P1012/T4252." specimenCount="1" stateProvince="E. Cape" typeStatus="paratype">
<specimenCount id="9D68FD1A463CFFF243CE454F356EF951" box="[972,1110,1646,1672]" pageId="48" pageNumber="49" type="generic">1 specimen</specimenCount>
, same collection data as for holotype;
<collectionCode id="ED7FAE56463CFFF2416145B3308EF975" box="[355,438,1682,1708]" collectionName="South Africa, Kwa-Zulu Natal, Pietermaritzburg, Natal Museum" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="48" pageNumber="49" type="Museum">NMSA</collectionCode>
<specimenCode id="DBC89EE8463CFFF241C345B33350F974" box="[449,616,1682,1709]" pageId="48" pageNumber="49">P1012/T4252.</specimenCode>
–
<emphasis id="B91AEA81463CFFF2428E45B333CAF975" bold="true" box="[652,754,1682,1708]" pageId="48" pageNumber="49">
<collectingRegion id="49AAF871463CFFF2428E45B333CAF975" box="[652,754,1682,1708]" pageId="48" pageNumber="49">E. Cape</collectingRegion>
</emphasis>
</materialsCitation>
•
<materialsCitation id="3B063CCE463CFFF2431045B3326BF92D" ID-GBIF-Occurrence="2605305715" collectingDate="2003-09-21" collectionCode="NMSA" collectorName="D. G. Herbert" country="South Africa" elevation="140" latitude="-33.95377" location="Bloukrans River Valley" longLatPrecision="1" longitude="23.65423" pageId="48" pageNumber="49" specimenCode="W1171/T4249" specimenCount="5" typeStatus="paratype">
<specimenCount id="9D68FD1A463CFFF2431045B33291F975" box="[786,937,1682,1709]" pageId="48" pageNumber="49" type="generic">5 specimens</specimenCount>
;
<location id="8EB16048463CFFF243BA45B335EAF975" LSID="urn:lsid:plazi:treatment:03C78785463CFFF34239419B35A5FB67:8EB16048463CFFF243BA45B335EAF975" box="[952,1234,1682,1708]" country="South Africa" latitude="-33.95377" longLatPrecision="1" longitude="23.65423" name="Bloukrans River Valley" pageId="48" pageNumber="49">Bloukrans River Valley</location>
, eastern side;
<geoCoordinate id="EE5A5054463CFFF240BF4597306AF909" box="[189,338,1718,1745]" degrees="33.95377" direction="south" orientation="latitude" pageId="48" pageNumber="49" precision="1" value="-33.95377">33.95377° S</geoCoordinate>
,
<geoCoordinate id="EE5A5054463CFFF24162459730C0F908" box="[352,504,1718,1745]" degrees="23.65423" direction="east" orientation="longitude" pageId="48" pageNumber="49" precision="1" value="23.65423">23.65423° E</geoCoordinate>
;
<elevation id="0043D1A0463CFFF24204459733ABF909" box="[518,659,1718,1745]" metricMagnitude="2" metricUnit="m" metricValue="1.4" pageId="48" pageNumber="49" unit="m" value="140.0">
<quantity id="4C969B76463CFFF2420445973369F908" box="[518,593,1718,1745]" metricMagnitude="2" metricUnit="m" metricValue="1.4" pageId="48" pageNumber="49" unit="m" value="140.0">140 m</quantity>
a.s.l.
</elevation>
;
<date id="FFD01053463CFFF242A04597327AF909" box="[674,834,1718,1744]" pageId="48" pageNumber="49" value="2003-09-21">
<collectingDate id="EF94E9BB463CFFF242A04597327AF909" box="[674,834,1718,1744]" pageId="48" pageNumber="49" value="2003-09-21">21 Sep. 2003</collectingDate>
</date>
;
<collectorName id="269B5345463CFFF24352459732D4F909" box="[848,1004,1718,1744]" pageId="48" pageNumber="49">D.G. Herbert</collectorName>
leg.; in leaf-litter at base of road cutting through indigenous forest;
<collectionCode id="ED7FAE56463CFFF2424D45FB339AF92D" box="[591,674,1754,1780]" collectionName="South Africa, Kwa-Zulu Natal, Pietermaritzburg, Natal Museum" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="48" pageNumber="49" type="Museum">NMSA</collectionCode>
<specimenCode id="DBC89EE8463CFFF242AB45FB326BF92D" box="[681,851,1754,1780]" pageId="48" pageNumber="49">W1171/T4249</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B063CCE463CFFF2436845FB33F3F8E4" ID-GBIF-Occurrence="2605305730" collectingDate="1997-04-28" collectionCode="NMSA" collectorName="D. G. Herbert" country="South Africa" county="Tsitsikamma Forest" elevation="260" latitude="-33.9682" location="Tsitsikamma Forest" longLatPrecision="5" longitude="23.92634" municipality="Paul Sauer Bridge" pageId="48" pageNumber="49" specimenCode="V4734/T4250" specimenCount="14" typeStatus="paratype">
<specimenCount id="9D68FD1A463CFFF2436845FB3534F92D" box="[874,1036,1754,1780]" pageId="48" pageNumber="49" type="generic">14 specimens</specimenCount>
;
<location id="8EB16048463CFFF2441445FB35C6F92C" LSID="urn:lsid:plazi:treatment:03C78785463CFFF34239419B35A5FB67:8EB16048463CFFF2441445FB35C6F92C" box="[1046,1278,1754,1781]" country="South Africa" county="Tsitsikamma Forest" latitude="-33.9682" longLatPrecision="5" longitude="23.92634" municipality="Paul Sauer Bridge" name="Tsitsikamma Forest" pageId="48" pageNumber="49">Tsitsikamma Forest</location>
,
<locationDeviation id="9A632FB1463CFFF2450B45FA330CF8C1" pageId="48" pageNumber="49">near Paul Sauer Bridge over Storms River</locationDeviation>
;
<geoCoordinate id="EE5A5054463CFFF2423C45DF33F9F8C1" box="[574,705,1790,1816]" degrees="33.9682" direction="south" orientation="latitude" pageId="48" pageNumber="49" precision="5" value="-33.9682">33.9682° S</geoCoordinate>
,
<geoCoordinate id="EE5A5054463CFFF242CF45DF325AF8C0" box="[717,866,1790,1817]" degrees="23.92634" direction="east" orientation="longitude" pageId="48" pageNumber="49" precision="1" value="23.92634">23.92634° E</geoCoordinate>
;
<elevation id="0043D1A0463CFFF2436F45DF32CCF8C1" box="[877,1012,1790,1817]" metricMagnitude="2" metricUnit="m" metricValue="2.6" pageId="48" pageNumber="49" unit="m" value="260.0">
<quantity id="4C969B76463CFFF2436F45DF328EF8C0" box="[877,950,1790,1817]" metricMagnitude="2" metricUnit="m" metricValue="2.6" pageId="48" pageNumber="49" unit="m" value="260.0">260 m</quantity>
a.s.l.
</elevation>
;
<date id="FFD01053463CFFF2440245DF35A2F8C1" box="[1024,1178,1790,1817]" pageId="48" pageNumber="49" value="1997-04-28">
<collectingDate id="EF94E9BB463CFFF2440245DF35A2F8C1" box="[1024,1178,1790,1817]" pageId="48" pageNumber="49" value="1997-04-28">28 Apr. 1997</collectingDate>
</date>
;
<collectorName id="269B5345463CFFF244A645DF3405F8C1" box="[1188,1341,1790,1816]" pageId="48" pageNumber="49">D.G. Herbert</collectorName>
leg.; sorted from leaf-litter;
<collectionCode id="ED7FAE56463CFFF241CB44033324F8E5" box="[457,540,1826,1852]" collectionName="South Africa, Kwa-Zulu Natal, Pietermaritzburg, Natal Museum" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="48" pageNumber="49" type="Museum">NMSA</collectionCode>
<specimenCode id="DBC89EE8463CFFF24221440333F3F8E4" box="[547,715,1826,1853]" pageId="48" pageNumber="49">V4734/T4250</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BD13693463CFFF240E84445309EF8A7" blockId="48.[189,1398,1892,2027]" box="[234,422,1892,1918]" pageId="48" pageNumber="49">
<emphasis id="B91AEA81463CFFF240E84445309EF8A7" bold="true" box="[234,422,1892,1918]" pageId="48" pageNumber="49">Other material</emphasis>
</paragraph>
<paragraph id="8BD13693463CFFF240BF44A83023F832" blockId="48.[189,1398,1892,2027]" pageId="48" pageNumber="49">
<materialsCitation id="3B063CCE463CFFF240BF44A8302FF832" ID-GBIF-Occurrence="2605305659" collectedFrom="sorted from leaf-litter" collectingDate="2002-10-01" collectionCode="NMSA" collectorName="J. P. Marais" country="South Africa" elevation="260" latitude="-33.9682" location="Tsitsikamma Forest" longLatPrecision="6" longitude="23.8982" municipality="Tsitsikamma Forest" pageId="48" pageNumber="49" specimenCode="W1768" specimenCount="1" stateProvince="E. Cape">
<collectingCountry id="F3797603463CFFF240BF44A830A8F87A" box="[189,400,1929,1955]" name="South Africa" pageId="48" pageNumber="49">SOUTH AFRICA</collectingCountry>
•
<specimenCount id="9D68FD1A463CFFF241AE44A8330EF87A" box="[428,566,1929,1955]" pageId="48" pageNumber="49" type="generic">1 specimen</specimenCount>
;
<collectingRegion id="49AAF871463CFFF2424144A8339CF87A" box="[579,676,1929,1955]" pageId="48" pageNumber="49">E. Cape</collectingRegion>
,
<location id="8EB16048463CFFF242B344A832A3F87A" LSID="urn:lsid:plazi:treatment:03C78785463CFFF34239419B35A5FB67:8EB16048463CFFF242B344A832A3F87A" box="[689,923,1929,1955]" country="South Africa" latitude="-33.9682" longLatPrecision="6" longitude="23.8982" municipality="Tsitsikamma Forest" name="Tsitsikamma Forest" pageId="48" pageNumber="49" stateProvince="E. Cape">Tsitsikamma Forest</location>
,
<locationDeviation id="9A632FB1463CFFF243AB44A8344AF87A" box="[937,1394,1929,1955]" pageId="48" pageNumber="49">‘big tree’ west of Storms River bridge</locationDeviation>
;
<geoCoordinate id="EE5A5054463CFFF240BF448C307EF81E" box="[189,326,1965,1991]" degrees="33.9682" direction="south" orientation="latitude" pageId="48" pageNumber="49" precision="5" value="-33.9682">33.9682° S</geoCoordinate>
,
<geoCoordinate id="EE5A5054463CFFF24155448C30DAF81E" box="[343,482,1965,1991]" degrees="23.8982" direction="east" orientation="longitude" pageId="48" pageNumber="49" precision="5" value="23.8982">23.8982° E</geoCoordinate>
;
<elevation id="0043D1A0463CFFF241F1448C33BDF81E" box="[499,645,1965,1991]" metricMagnitude="2" metricUnit="m" metricValue="2.6" pageId="48" pageNumber="49" unit="m" value="260.0">
<quantity id="4C969B76463CFFF241F1448C3379F81E" box="[499,577,1965,1991]" metricMagnitude="2" metricUnit="m" metricValue="2.6" pageId="48" pageNumber="49" unit="m" value="260.0">260 m</quantity>
a.s.l.
</elevation>
;
<date id="FFD01053463CFFF24295448C3214F81E" box="[663,812,1965,1991]" pageId="48" pageNumber="49" value="2002-10-01">
<collectingDate id="EF94E9BB463CFFF24295448C3214F81E" box="[663,812,1965,1991]" pageId="48" pageNumber="49" value="2002-10-01">1 Oct. 2002</collectingDate>
</date>
;
<collectorName id="269B5345463CFFF2433F448C32F8F81E" box="[829,960,1965,1991]" pageId="48" pageNumber="49">J.P. Marais</collectorName>
leg.;
<collectedFrom id="3872756E463CFFF24409448C342CF81E" box="[1035,1300,1965,1991]" pageId="48" pageNumber="49">sorted from leaf-litter</collectedFrom>
;
<collectionCode id="ED7FAE56463CFFF24521448C344EF81E" box="[1315,1398,1965,1991]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="48" pageNumber="49" type="Museum">NMSA</collectionCode>
<specimenCode id="DBC89EE8463CFFF240BF44F0302FF832" box="[189,279,2001,2027]" collectionCode="NMSA" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="48" pageNumber="49" type="Museum">W1768</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3746518463DFFF340BF422A32BAFC92" pageId="49" pageNumber="50" type="description">
<paragraph id="8BD13693463DFFF340BF422A3076FEFC" blockId="49.[189,334,267,293]" box="[189,334,267,293]" pageId="49" pageNumber="50">
<heading id="D09981FF463DFFF340BF422A3076FEFC" bold="true" box="[189,334,267,293]" fontSize="11" level="3" pageId="49" pageNumber="50" reason="3">
<emphasis id="B91AEA81463DFFF340BF422A3076FEFC" bold="true" box="[189,334,267,293]" pageId="49" pageNumber="50">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BD13693463DFFF340BF421B32BAFC92" blockId="49.[189,1399,314,844]" pageId="49" pageNumber="50">
Shell small, diameter up to
<quantity id="4C969B76463DFFF34205421B335EFE8C" box="[519,614,314,341]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="49" pageNumber="50" unit="mm" value="1.7">1.7 mm</quantity>
, H/D ratio ±0.47; spire slightly to distinctly raised; whorls tightly coiled; last adult whorl descendant; suture narrowly indented, somewhat sunken; periphery evenly convex. Protoconch comprising apical cap plus approx. 1.0 whorl; diameter ±410 μm; microscopically shagreened, lacking axial sculpture. Teleoconch of up to 3.5 whorls; sculptured by distinct, close-set, compound axial riblets, with 4–5 finer, intermediary axial threads; intervals between riblets 1–2 times width of riblets at whorl periphery; spiral sculpture of extremely fine, close-set threads more or less throughout. Umbilicus very wide, its margin relatively sharply rounded.Aperture broadly lunate; parietal region with two narrow, in-running lamellae, the upper one weaker and slightly more recessed; basocolumellar region with a similar recessed, narrow, in-running, lamella-like ridge (sometimes scarcely evident in apertural view); palatal region with three denticles recessed
<superScript id="7C1B9BDB463DFFF34419415E351CFD54" attach="right" box="[1051,1060,639,653]" fontSize="6" pageId="49" pageNumber="50">1</superScript>
⁄5–
<superScript id="7C1B9BDB463DFFF3443D415E3570FD54" attach="none" box="[1087,1096,639,653]" fontSize="6" pageId="49" pageNumber="50">1</superScript>
⁄4 whorl behind outer lip, denticles evident by transparency, but hardly visible in apertural view, one just above mid-whorl, one just below mid-whorl, the third more basal, middle denticle situated slightly closer to aperture and often more elongate (particularly in juveniles); a fourth palatal structure in the form of a narrow, thread-like ridge lying immediately below the suture is evident by transparency in fresh juveniles and subadults. Shell pale corneous-brown to pale honey-brown when fresh.
</paragraph>
</subSubSection>
<subSubSection id="C3746518463DFFF340BF4052344FFBDC" pageId="49" pageNumber="50" type="distribution">
<paragraph id="8BD13693463DFFF340BF4052330CFC54" blockId="49.[189,564,883,910]" box="[189,564,883,910]" pageId="49" pageNumber="50">
<heading id="D09981FF463DFFF340BF4052330CFC54" bold="true" box="[189,564,883,910]" fontSize="11" level="3" pageId="49" pageNumber="50" reason="3">
<emphasis id="B91AEA81463DFFF340BF4052330CFC54" bold="true" box="[189,564,883,910]" pageId="49" pageNumber="50">Distribution and conservation</emphasis>
</heading>
</paragraph>
<paragraph id="8BD13693463DFFF340BF4082344FFBDC" blockId="49.[189,1399,931,1029]" pageId="49" pageNumber="50">
A narrow-range endemic (
<figureCitation id="13552A16463DFFF341F140823373FC64" box="[499,587,931,957]" captionStart="Fig" captionStartId="36.[189,232,1860,1886]" captionTargetBox="[302,1285,1199,1820]" captionTargetId="figure@36.[302,1285,1199,1820]" captionTargetPageId="36" captionText="Fig. 14. Distribution of Iterodonta gen. nov. and Phialodonta gen. nov. species. Iterodonta ammonita gen. et sp. nov. (brown triangles), Phialodonta agulhasae gen. et sp. nov. (orange square), P. atromontana gen. et sp. nov. (turquoise square), P. aviana gen. et sp. nov. (royal blue squares), P. introtuberculata (Connolly, 1933) gen. et comb. nov. (red squares), P. perfida (Burnup, 1912) gen. et comb. nov. (green square), P. rivalalea gen. et sp. nov. (pink squares). Contour at 1000 m." figureDoi="http://doi.org/10.5281/zenodo.3762308" httpUri="https://zenodo.org/record/3762308/files/figure.png" pageId="49" pageNumber="50">Fig. 14</figureCitation>
), known only from the coastal hinterland in the Tsitsikamma region, southern
<collectingRegion id="49AAF871463DFFF3412A40E6305FFC38" box="[296,359,967,993]" country="South Africa" name="Western Cape" pageId="49" pageNumber="50">Cape</collectingRegion>
, from the coast to
<quantity id="4C969B76463DFFF3423540E633B8FC38" box="[567,640,967,993]" metricMagnitude="2" metricUnit="m" metricValue="2.6" pageId="49" pageNumber="50" unit="m" value="260.0">260 m</quantity>
a.s.l.; in leaf-litter of southern afrotemperate forest. The forests in this region fall within the Garden Route National Park and are thus afforded a high degree of protection.
</paragraph>
</subSubSection>
<subSubSection id="C3746518463DFFF340BF470C35A5FB67" pageId="49" pageNumber="50" type="discussion">
<paragraph id="8BD13693463DFFF340BF470C3017FB9E" blockId="49.[189,303,1069,1095]" box="[189,303,1069,1095]" pageId="49" pageNumber="50">
<heading id="D09981FF463DFFF340BF470C3017FB9E" bold="true" box="[189,303,1069,1095]" fontSize="11" level="3" pageId="49" pageNumber="50" reason="3">
<emphasis id="B91AEA81463DFFF340BF470C3017FB9E" bold="true" box="[189,303,1069,1095]" pageId="49" pageNumber="50">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BD13693463DFFF340BF477C35A5FB67" blockId="49.[189,1399,1116,1215]" pageId="49" pageNumber="50">
In terms of the overall facies of the shell,
<taxonomicName id="4C6E4D10463DFFF342AD477D328AFBAF" authority="Herbert, 2020" authorityName="Herbert" authorityYear="2020" box="[687,946,1116,1142]" class="Gastropoda" family="Charopidae" genus="Phialodonta" kingdom="Animalia" order="Stylommatophora" pageId="49" pageNumber="50" phylum="Mollusca" rank="species" species="rivalalea" status="gen. et sp. nov.">
<emphasis id="B91AEA81463DFFF342AD477D328AFBAF" box="[687,946,1116,1142]" italics="true" pageId="49" pageNumber="50">Phialodonta rivalalea</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22957FA463DFFF343B8477C3556FBAE" box="[954,1134,1117,1143]" pageId="49" pageNumber="50" rank="species">gen. et sp. nov.</taxonomicNameLabel>
is closest to
<taxonomicName id="4C6E4D10463DFFF3450B477C344FFBAF" authority="Herbert, 2020" authorityName="Herbert" authorityYear="2020" box="[1289,1399,1117,1142]" class="Gastropoda" family="Charopidae" genus="Phialodonta" kingdom="Animalia" order="Stylommatophora" pageId="49" pageNumber="50" phylum="Mollusca" rank="species" species="aviana" status="gen. et sp. nov.">
<emphasis id="B91AEA81463DFFF3450B477C344FFBAF" box="[1289,1399,1117,1142]" italics="true" pageId="49" pageNumber="50">P. aviana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22957FA463DFFF340BF47A03055FB42" box="[189,365,1153,1179]" pageId="49" pageNumber="50" rank="species">gen. et sp. nov.</taxonomicNameLabel>
It differs from the latter in having parietal lamellae that are visible through the aperture, and in possessing a narrow, in-running baso-columellar ridge and a wider umbilicus.
</paragraph>
</subSubSection>
</treatment>
</document>
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