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<document id="AB2B1AA6DB139DB47165333909E5FB80" ID-CLB-Dataset="26506" ID-DOI="10.11646/zootaxa.4615.2.5" ID-GBIF-Dataset="f2ea6903-8250-4cef-b245-7d0358639a32" ID-ISSN="1175-5326" ID-Zenodo-Dep="3244718" ID-ZooBank="41436B41-169B-4228-AAEC-29C26BCFD77A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1560402730588" checkinUser="plazi" docAuthor="Larrazábal-Filho, Alexandre L., Neres, Patrícia F., Silva, Maria Cristina Da &amp; Esteves, André M." docDate="2019" docId="03AFF861FFEEFF97329CA951FC2C539B" docLanguage="en" docName="zootaxa.4615.2.5.pdf" docOrigin="Zootaxa 4615 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Spinonema absente Larrazábal-Filho &amp; Neres &amp; Silva &amp; Esteves 2019, gen. et sp. n." docType="treatment" docVersion="10" lastPageNumber="339" masterDocId="FF968019FFE3FF85320BAE01FFF25074" masterDocTitle="A new genus with three new species of free-living marine nematodes of the subfamily Desmodorinae (Nematoda: Desmodoridae), from the continental shelf off northeastern Brazil" masterLastPageNumber="342" masterPageNumber="321" pageNumber="334" updateTime="1698734701748" updateUser="ExternalLinkService">
<mods:mods id="BAB91A25D4E81A27C58151FA5F2A450D" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="DB7FC7B5BC38BC09EE47447DAAE07F4F">
<mods:title id="D8BD120108DA391493CBFA6CE40A6C28">A new genus with three new species of free-living marine nematodes of the subfamily Desmodorinae (Nematoda: Desmodoridae), from the continental shelf off northeastern Brazil</mods:title>
</mods:titleInfo>
<mods:name id="47A3ED0CB68186DE760A3C2BFE07D176" type="personal">
<mods:role id="9087E8DA1BEA5AB6A616752CEF3BFCD7">
<mods:roleTerm id="A2958ED5701694E40583C94B4EDD4EBD">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5964EA0F2C1AFD39E02FB973E6145FE3">Larrazábal-Filho, Alexandre L.</mods:namePart>
</mods:name>
<mods:name id="5F1540C90D36C3202B96220E39AEE481" type="personal">
<mods:role id="40F1AFDC783487A8565E6C01A30BC8B5">
<mods:roleTerm id="7FA30AE9643E543B99BB5F320827221B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9F7A1E6CC0B7C4E36DE423E9A3CA7A7B">Neres, Patrícia F.</mods:namePart>
</mods:name>
<mods:name id="A070331F7EBED96819B5A67B983DA804" type="personal">
<mods:role id="D0BD7C45046054A2552DBF0A3826DF8E">
<mods:roleTerm id="A046ECC1D4AE4DC09D7B355E0A61EDBF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F630C921A53B9B7528AFF71A8EE28C23">Silva, Maria Cristina Da</mods:namePart>
</mods:name>
<mods:name id="8D5E2EF7EEA90D2B5E569CC4B417978A" type="personal">
<mods:role id="07AFB4254354FAD05987A86DBD45853A">
<mods:roleTerm id="BB2F33045C478FC74035BEF913D4D4CF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B7A7D27DD9F75A142AC47B461E8D64D0">Esteves, André M.</mods:namePart>
</mods:name>
<mods:typeOfResource id="262F5B7C2B8512D7EE3460B9C3388058">text</mods:typeOfResource>
<mods:relatedItem id="EFB1F761EC7D499E5A67E0E4A8283459" type="host">
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<mods:title id="EAA0CE75CFBF48DD4B1695220FCF167F">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="D2C303C49ADBE541E262D664C8CCFBFB">
<mods:date id="310319975D06F865438815C1E06D1920">2019</mods:date>
<mods:detail id="C361C85554B96A6B6831D98DCA8DC93D" type="pubDate">
<mods:number id="FD2033089BA44112989EC059DADAC892">2019-06-13</mods:number>
</mods:detail>
<mods:detail id="0872776C6A253C1E52D572D59BCE9BDE" type="volume">
<mods:number id="CB5EB4FF2E12B920D73387EA3F6F5D2D">4615</mods:number>
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<mods:detail id="0A176B07D406864CAD3C170F02AC1E96" type="issue">
<mods:number id="1C21679F8FACD05242A47311DD679C19">2</mods:number>
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<mods:classification id="A8362D71B88FDCC66E0B6A6CBA12212C">journal article</mods:classification>
<mods:identifier id="3666A8D38F1833F6FCF53CBE239541CA" type="CLB-Dataset">26506</mods:identifier>
<mods:identifier id="A18341930A81D4A8BDB7E2CEC1D51A90" type="DOI">10.11646/zootaxa.4615.2.5</mods:identifier>
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<mods:identifier id="4737E0E4C06E4A53779C60086CB1DD35" type="ISSN">1175-5326</mods:identifier>
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</mods:mods>
<treatment id="03AFF861FFEEFF97329CA951FC2C539B" ID-GBIF-Taxon="157533059" LSID="urn:lsid:plazi:treatment:03AFF861FFEEFF97329CA951FC2C539B" httpUri="http://treatment.plazi.org/id/03AFF861FFEEFF97329CA951FC2C539B" lastPageId="18" lastPageNumber="339" pageId="13" pageNumber="334">
<subSubSection id="C31C1AFCFFEEFF88329CA951FE5657FA" pageId="13" pageNumber="334" type="nomenclature">
<paragraph id="8BB94977FFEEFF88329CA951FDD7571E" blockId="13.[151,549,1872,1934]" box="[151,549,1872,1899]" pageId="13" pageNumber="334">
<heading id="D0F1FE1BFFEEFF88329CA951FDD7571E" bold="true" box="[151,549,1872,1899]" fontSize="11" level="1" pageId="13" pageNumber="334" reason="1">
<emphasis id="B9729565FFEEFF88329CA951FDD7571E" bold="true" box="[151,549,1872,1899]" pageId="13" pageNumber="334">
<taxonomicName id="4C0632F4FFEEFF88329CA951FE70571F" authority="Larrazábal-Filho &amp; Neres &amp; Silva &amp; Esteves, 2019" authorityName="Larrazábal-Filho &amp; Neres &amp; Silva &amp; Esteves" authorityYear="2019" box="[151,386,1872,1899]" class="Adenophorea" family="Desmodoridae" genus="Spinonema" kingdom="Animalia" order="Desmodorida" pageId="13" pageNumber="334" phylum="Nematoda" rank="species" species="absente" status="gen. et sp. n.">
<emphasis id="B9729565FFEEFF88329CA951FE70571F" bold="true" box="[151,386,1872,1899]" italics="true" pageId="13" pageNumber="334">Spinonema absente</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A241281EFFEEFF883381A951FDD7571E" box="[394,549,1872,1898]" pageId="13" pageNumber="334" rank="species">gen. et sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB94977FFEEFF88329CA975FE5657FA" blockId="13.[151,549,1872,1934]" box="[151,420,1908,1934]" pageId="13" pageNumber="334">
(
<tableCitation id="C6847CCCFFEEFF883294A975FF0657FA" box="[159,244,1908,1934]" captionStart="TABLE 3" captionStartId="14.[151,239,442,466]" captionTargetPageId="14" captionText="TABLE 3. Measurements (µm) of Spinonema absente gen. et sp. n. in the Potiguar Basin, Brazil. “-“ = not applicable. The mean and standard deviation are followed by the range for paratype measurements." pageId="13" pageNumber="334">Table 3</tableCitation>
,
<figureCitation id="133D55F2FFEEFF88330AA975FE6557FA" box="[257,407,1908,1934]" captionStart-0="FIGURE 9" captionStart-1="FIGURE 10" captionStart-2="FIGURE 11" captionStartId-0="15.[151,250,1961,1986]" captionStartId-1="16.[151,250,1909,1934]" captionStartId-2="17.[151,250,1962,1987]" captionTargetBox-0="[221,1379,172,1917]" captionTargetBox-1="[166,1248,176,1882]" captionTargetBox-2="[219,1359,183,1869]" captionTargetId-0="figure@15.[206,1382,166,1941]" captionTargetId-1="figure@16.[151,1260,166,1890]" captionTargetId-2="figure@17.[202,1386,166,1942]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionTargetPageId-2="17" captionText-0="FIGURE 9. Spinonema absente gen. et sp. n. Holotype MNRJ 400. A: overview; B, C and F: anterior region; D: tail region; E: spicules; H: lateral alae (ANT anterior part and POS posterior part); G: Inversion of direction of annuli." captionText-1="FIGURE 10. Spinonema absente gen. et sp. n. A and C, Holotype male MNRJ 400; A: Cephalic region; C: Tail region. B, D, E and F, Paratype female MNRJ 401; B: Tail region; D: Lateral alae (ANT anterior part and POS posterior part); E: Vulva opening; F: Anterior region." captionText-2="FIGURE 11. Spinonema absente gen. et sp. n. Paratype female MNRJ 401. A: overview; C: Ovary and vulva opening; B and D: Anterior region. Paratype juvenile in stage 4. E: Anterior region; F: Tail region; G: Ovary of juvenile." httpUri-0="https://zenodo.org/record/3244734/files/figure.png" httpUri-1="https://zenodo.org/record/3244736/files/figure.png" httpUri-2="https://zenodo.org/record/3244738/files/figure.png" pageId="13" pageNumber="334">Figures 9–11</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C31C1AFCFFEEFF8B329CA9B9FC5350A3" lastPageId="14" lastPageNumber="335" pageId="13" pageNumber="334" type="materials_examined">
<paragraph id="8BB94977FFEEFF88329CA9B9FB2C5782" blockId="13.[151,1436,1975,2039]" pageId="13" pageNumber="334">
<emphasis id="B9729565FFEEFF88329CA9B9FEBB57A6" bold="true" box="[151,329,1975,2002]" pageId="13" pageNumber="334">Type material:</emphasis>
<materialsCitation id="3B6E432AFFEEFF88335AA9B9FB2C5782" ID-GBIF-Occurrence="2252532241" collectingDate="2009-06" collectionCode="MNRJ" collectorName="Basin &amp; Faciology &amp; Gear &amp; Van Veen" elevation="35" latitude="-4.75" longLatPrecision="21" longitude="-36.75" pageId="13" pageNumber="334" specimenCode="MNRJ 400" specimenCount="1" typeStatus="holotype">
<emphasis id="B9729565FFEEFF88335AA9B9FE3757A6" bold="true" box="[337,453,1976,2002]" pageId="13" pageNumber="334">
<typeStatus id="54BDF7D5FFEEFF88335AA9B9FE3257A6" box="[337,448,1976,2002]" pageId="13" pageNumber="334" type="holotype">Holotype</typeStatus>
,
</emphasis>
Male adult
<specimenCode id="DBA0E10CFFEEFF88305AA9B8FD3C57A6" box="[593,718,1976,2003]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="13" pageNumber="334">MNRJ 400</specimenCode>
(
<geoCoordinate id="EE322FB0FFEEFF8830D4A9B6FC9557A6" box="[735,871,1975,2002]" degrees="04" direction="south" minutes="45" orientation="latitude" pageId="13" pageNumber="334" precision="15" seconds="00" value="-4.75">04°45’00”S</geoCoordinate>
,
<geoCoordinate id="EE322FB0FFEEFF88317FA9B6FBE057A6" box="[884,1042,1975,2002]" degrees="036" direction="west" minutes="45" orientation="longitude" pageId="13" pageNumber="334" precision="15" seconds="00" value="-36.75">036°45’00”W</geoCoordinate>
) collected in
<collectingDate id="EFFC965FFFEEFF8836BBA9B8FAD157A6" box="[1200,1315,1976,2003]" pageId="13" pageNumber="334" value="2009-06">
<date id="FFB86FB7FFEEFF8836BBA9B8FAD157A6" box="[1200,1315,1976,2003]" pageId="13" pageNumber="334" value="2009-06">
June
<quantity id="4CFEE492FFEEFF8836E0A9B9FAD157A6" box="[1259,1315,1976,2002]" metricMagnitude="1" metricUnit="m" metricValue="5.10286" pageId="13" pageNumber="334" unit="in" value="2009.0">2009</quantity>
</date>
</collectingDate>
in the Po- tiguar
<collectorName id="26F32CA1FFEEFF8832EBA9DDFED15782" box="[224,291,2012,2038]" pageId="13" pageNumber="334">Basin</collectorName>
, between 30 and
<elevation id="002BAE44FFEEFF8833EFA9DAFDEF5783" box="[484,541,2011,2039]" metricMagnitude="1" metricUnit="m" metricValue="3.5" pageId="13" pageNumber="334" unit="m" value="35.0">
<quantity id="4CFEE492FFEEFF8833EFA9DAFDEF5783" box="[484,541,2011,2039]" metricMagnitude="1" metricUnit="m" metricValue="3.5" pageId="13" pageNumber="334" unit="m" value="35.0">35 m</quantity>
</elevation>
deep.
<collectorName id="26F32CA1FFEEFF88306CA9DDFD295782" box="[615,731,2012,2038]" pageId="13" pageNumber="334">Faciology</collectorName>
: coarse bioclastic sand.
<collectorName id="26F32CA1FFEEFF8831E3A9DDFBD15782" box="[1000,1059,2012,2038]" pageId="13" pageNumber="334">Gear</collectorName>
:
<collectorName id="26F32CA1FFEEFF883627A9DCFB6A5782" box="[1068,1176,2012,2039]" pageId="13" pageNumber="334">Van Veen</collectorName>
/corer.
</materialsCitation>
</paragraph>
<paragraph id="8BB94977FFEDFF8B32CCAE98FC5350A3" blockId="14.[151,1437,152,395]" pageId="14" pageNumber="335">
<materialsCitation id="3B6E432AFFEDFF8B32CCAE98FC6E50A3" ID-GBIF-Occurrence="2252532237" collectionCode="MNRJ" collectorName="Potiguar Basin &amp; Faciology &amp; Gear &amp; Van Veen" elevation="35" latitude="-5.0223336" longLatPrecision="1" longitude="-36.18722" pageId="14" pageNumber="335" specimenCode="MNRJ 401" specimenCount="1" typeStatus="paratype">
<emphasis id="B9729565FFEDFF8B32CCAE98FE6150C7" bold="true" box="[199,403,152,179]" pageId="14" pageNumber="335">
<typeStatus id="54BDF7D5FFEDFF8B32CCAE98FEC650C7" box="[199,308,153,179]" pageId="14" pageNumber="335" type="paratype">Paratype</typeStatus>
female:
</emphasis>
Adult,
<specimenCode id="DBA0E10CFFEDFF8B33EFAE98FDAD50C7" box="[484,607,153,179]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="14" pageNumber="335">MNRJ 401</specimenCode>
(
<geoCoordinate id="EE322FB0FFEDFF8B3065AE99FCFE50C7" box="[622,780,152,179]" degrees="05" direction="south" minutes="01" orientation="latitude" pageId="14" pageNumber="335" precision="1" seconds="20.4" value="-5.0223336">05°01’20.4’’S</geoCoordinate>
<geoCoordinate id="EE322FB0FFEDFF8B3119AE99FC3B50C7" box="[786,969,152,179]" degrees="036" direction="west" minutes="11" orientation="longitude" pageId="14" pageNumber="335" precision="1" seconds="14.0" value="-36.18722">036°11’14.0’’W</geoCoordinate>
) from the
<collectorName id="26F32CA1FFEDFF8B3637AE99FB1650C6" box="[1084,1252,152,178]" pageId="14" pageNumber="335">Potiguar Basin</collectorName>
, between 30 and
<elevation id="002BAE44FFEDFF8B329CAEBDFF2250A3" box="[151,208,188,215]" metricMagnitude="1" metricUnit="m" metricValue="3.5" pageId="14" pageNumber="335" unit="m" value="35.0">
<quantity id="4CFEE492FFEDFF8B329CAEBDFF2250A3" box="[151,208,188,215]" metricMagnitude="1" metricUnit="m" metricValue="3.5" pageId="14" pageNumber="335" unit="m" value="35.0">35 m</quantity>
</elevation>
deep.
<collectorName id="26F32CA1FFEDFF8B3311AEBDFE7C50A2" box="[282,398,188,214]" pageId="14" pageNumber="335">Faciology</collectorName>
: coarse lithoclastic sand.
<collectorName id="26F32CA1FFEDFF8B30A1AEBCFD1750A3" box="[682,741,189,215]" pageId="14" pageNumber="335">Gear</collectorName>
:
<collectorName id="26F32CA1FFEDFF8B30E4AEBCFCA950A3" box="[751,859,189,215]" pageId="14" pageNumber="335">Van Veen</collectorName>
/corer
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31C1AFCFFEDFF8B32CCAEE1FDA25113" pageId="14" pageNumber="335" type="description">
<paragraph id="8BB94977FFEDFF8B32CCAEE1FDA25113" blockId="14.[151,1437,152,395]" pageId="14" pageNumber="335">
<emphasis id="B9729565FFEDFF8B32CCAEE1FEFC508F" bold="true" box="[199,270,224,251]" pageId="14" pageNumber="335">Other</emphasis>
<materialsCitation id="3B6E432AFFEDFF8B331CAEE0FEBD5137" ID-GBIF-Occurrence="2252532245" collectionCode="LMZOO-UFPE" pageId="14" pageNumber="335" specimenCount="14" specimenCount-female="4" specimenCount-juvenile="5" specimenCount-male="5" typeStatus="paratype">
<emphasis id="B9729565FFEDFF8B331CAEE0FE65508F" bold="true" box="[279,407,225,251]" pageId="14" pageNumber="335">
<typeStatus id="54BDF7D5FFEDFF8B331CAEE0FE60508F" box="[279,402,225,251]" pageId="14" pageNumber="335" type="paratype">paratypes</typeStatus>
:
</emphasis>
<specimenCount id="9D0082FEFFEDFF8B33ABAEE0FDFC508E" box="[416,526,224,251]" pageId="14" pageNumber="335" type="female">4 females</specimenCount>
(210-213
<collectionCode id="ED17D1B2FFEDFF8B308CAEE0FCCF508F" box="[647,829,225,251]" pageId="14" pageNumber="335">LMZOO-UFPE</collectionCode>
),
<specimenCount id="9D0082FEFFEDFF8B3159AEE1FC58508E" box="[850,938,224,251]" pageId="14" pageNumber="335" type="male">5 males</specimenCount>
(214-218
<collectionCode id="ED17D1B2FFEDFF8B362FAEE0FB25508F" box="[1060,1239,225,251]" pageId="14" pageNumber="335">LMZOO-UFPE</collectionCode>
) and
<specimenCount id="9D0082FEFFEDFF8B3712AEE1FA6B508E" box="[1305,1433,224,251]" pageId="14" pageNumber="335" type="juvenile">5 juveniles</specimenCount>
. Four in first stage of development (J1) and one in fourth stage of development (J4) (219
<collectionCode id="ED17D1B2FFEDFF8B3664AF04FAD0516B" box="[1135,1314,261,287]" pageId="14" pageNumber="335">LMZOO-UFPE</collectionCode>
), collected on same date as
</materialsCitation>
<typeStatus id="54BDF7D5FFEDFF8B3352AF29FE4D5136" box="[345,447,296,322]" pageId="14" pageNumber="335" type="holotype">holotype</typeStatus>
. Juvenile stages were identified based on the developmental stage of the reproductive system, based on
<bibRefCitation id="EF973486FFEDFF8B3356AF4DFDBE5113" author="Bird, A. &amp; Bird J." box="[349,588,332,359]" pageId="14" pageNumber="335" refId="ref8230" refString="Bird, A. &amp; Bird J. (1991) The Structure of Nematodes. 2 nd Edition. Academic Press, San Diego, California, 317 pp." type="book" year="1991">Bird and Bird (1991)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31C1AFCFFEDFF8B32CCAF70FC4851FE" box="[199,954,368,395]" pageId="14" pageNumber="335" type="etymology">
<paragraph id="8BB94977FFEDFF8B32CCAF70FC4851FE" blockId="14.[151,1437,152,395]" box="[199,954,368,395]" pageId="14" pageNumber="335">
<emphasis id="B9729565FFEDFF8B32CCAF70FEBC51FF" bold="true" box="[199,334,369,395]" pageId="14" pageNumber="335">Etymology.</emphasis>
This species lacks a dorsal spine. L.
<emphasis id="B9729565FFEDFF8B30E6AF71FCBE51FE" box="[749,844,368,394]" italics="true" pageId="14" pageNumber="335">absentia</emphasis>
, absence.
</paragraph>
</subSubSection>
<caption id="DF7919FFFFEDFF8B329CAFBBFBCD5182" pageId="14" pageNumber="335" targetBox="[161,1408,514,1539]" targetIsTable="true" targetPageId="14">
<paragraph id="8BB94977FFEDFF8B329CAFBBFBCD5182" blockId="14.[151,1437,442,503]" pageId="14" pageNumber="335">
<emphasis id="B9729565FFEDFF8B329CAFBBFEF651A6" bold="true" box="[151,260,442,466]" pageId="14" pageNumber="335">TABLE 3</emphasis>
. Measurements (µm) of
<taxonomicName id="4C0632F4FFEDFF8B301AAFBAFD1251A6" authority="Larrazábal-Filho &amp; Neres &amp; Silva &amp; Esteves, 2019" authorityName="Larrazábal-Filho &amp; Neres &amp; Silva &amp; Esteves" authorityYear="2019" box="[529,736,442,466]" class="Adenophorea" family="Desmodoridae" genus="Spinonema" kingdom="Animalia" order="Desmodorida" pageId="14" pageNumber="335" phylum="Nematoda" rank="species" species="absente" status="gen. et sp. n.">
<emphasis id="B9729565FFEDFF8B301AAFBAFD1251A6" box="[529,736,442,466]" italics="true" pageId="14" pageNumber="335">Spinonema absente</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A241281EFFEDFF8B30E3AFBBFC8551A6" box="[744,887,442,466]" pageId="14" pageNumber="335" rank="species">
<emphasis id="B9729565FFEDFF8B30E3AFBBFC8551A6" bold="true" box="[744,887,442,466]" pageId="14" pageNumber="335">gen. et sp. n.</emphasis>
</taxonomicNameLabel>
in the Potiguar Basin, Brazil. “-“ = not applicable. The mean and standard deviation are followed by the range for paratype measurements.
</paragraph>
</caption>
<paragraph id="8BB94977FFEDFF8B32AAAC02FAC75676" pageId="14" pageNumber="335">
<table id="F906BBD7FFED007A32AAAC03FA725677" box="[161,1408,514,1539]" colsContinueFrom="13.[161,1410,612,1764]" gridcols="5" gridrows="25" pageId="14" pageNumber="335">
<tr id="35364B35FFED007A32AAAC03FA72526F" box="[161,1408,514,539]" gridrow="0" pageId="14" pageNumber="335">
<th id="76E72249FFED007A32AAAC03FDF9526F" box="[161,523,514,539]" gridcol="0" gridrow="0" pageId="14" pageNumber="335">Sex</th>
<th id="76E72249FFED007A3021AC03FD7A526F" box="[554,648,514,539]" gridcol="1" gridrow="0" pageId="14" pageNumber="335">Male</th>
<th id="76E72249FFED007A30A2AC03FC6A526F" box="[681,920,514,539]" gridcol="2" gridrow="0" pageId="14" pageNumber="335">Male</th>
<th id="76E72249FFED007A31BFAC03FB7D526F" box="[948,1167,514,539]" gridcol="3" gridrow="0" pageId="14" pageNumber="335">Females</th>
<th id="76E72249FFED007A36B9AC03FA72526F" box="[1202,1408,514,539]" gridcol="4" gridrow="0" pageId="14" pageNumber="335">Juvenile</th>
</tr>
<tr id="35364B35FFED007A32AAAC29FA725235" box="[161,1408,552,577]" gridrow="1" pageId="14" pageNumber="335" rowspan-0="1">
<td id="76E72249FFED007A3021AC29FD7A5235" box="[554,648,552,577]" gridcol="1" gridrow="1" pageId="14" pageNumber="335">Holotype</td>
<td id="76E72249FFED007A30A2AC29FC6A5235" box="[681,920,552,577]" gridcol="2" gridrow="1" pageId="14" pageNumber="335">Paratypes</td>
<td id="76E72249FFED007A31BFAC29FB7D5235" box="[948,1167,552,577]" gridcol="3" gridrow="1" pageId="14" pageNumber="335">Paratypes</td>
<td id="76E72249FFED007A36B9AC29FA725235" box="[1202,1408,552,577]" gridcol="4" gridrow="1" pageId="14" pageNumber="335">Paratypes</td>
</tr>
<tr id="35364B35FFED007A32AAAC4CFA725212" box="[161,1408,589,614]" gridrow="2" pageId="14" pageNumber="335">
<th id="76E72249FFED007A32AAAC4CFDF95212" box="[161,523,589,614]" gridcol="0" gridrow="2" pageId="14" pageNumber="335">Number of specimens</th>
<td id="76E72249FFED007A3021AC4CFD7A5212" box="[554,648,589,614]" gridcol="1" gridrow="2" pageId="14" pageNumber="335">1</td>
<td id="76E72249FFED007A30A2AC4CFC6A5212" box="[681,920,589,614]" gridcol="2" gridrow="2" pageId="14" pageNumber="335">5</td>
<td id="76E72249FFED007A31BFAC4CFB7D5212" box="[948,1167,589,614]" gridcol="3" gridrow="2" pageId="14" pageNumber="335">5</td>
<td id="76E72249FFED007A36B9AC4CFA725212" box="[1202,1408,589,614]" gridcol="4" gridrow="2" pageId="14" pageNumber="335">5</td>
</tr>
<tr id="35364B35FFED007A32AAAC72FA7252F8" box="[161,1408,627,652]" gridrow="3" pageId="14" pageNumber="335">
<th id="76E72249FFED007A32AAAC72FDF952F8" box="[161,523,627,652]" gridcol="0" gridrow="3" pageId="14" pageNumber="335">Body length</th>
<td id="76E72249FFED007A3021AC72FD7A52F8" box="[554,648,627,652]" gridcol="1" gridrow="3" pageId="14" pageNumber="335">1020</td>
<td id="76E72249FFED007A30A2AC72FC6A52F8" box="[681,920,627,652]" gridcol="2" gridrow="3" pageId="14" pageNumber="335">1014±102(806–1163.5)</td>
<td id="76E72249FFED007A31BFAC72FB7D52F8" box="[948,1167,627,652]" gridcol="3" gridrow="3" pageId="14" pageNumber="335">1069±132(825–1196)</td>
<td id="76E72249FFED007A36B9AC72FA7252F8" box="[1202,1408,627,652]" gridcol="4" gridrow="3" pageId="14" pageNumber="335">939±110(871–1135)</td>
</tr>
<tr id="35364B35FFED007A32AAAC98FA7252C6" box="[161,1408,665,690]" gridrow="4" pageId="14" pageNumber="335">
<th id="76E72249FFED007A32AAAC98FDF952C6" box="[161,523,665,690]" gridcol="0" gridrow="4" pageId="14" pageNumber="335">Pharynx length</th>
<td id="76E72249FFED007A3021AC98FD7A52C6" box="[554,648,665,690]" gridcol="1" gridrow="4" pageId="14" pageNumber="335">127.5</td>
<td id="76E72249FFED007A30A2AC98FC6A52C6" box="[681,920,665,690]" gridcol="2" gridrow="4" pageId="14" pageNumber="335">127±9(103.5–141)</td>
<td id="76E72249FFED007A31BFAC98FB7D52C6" box="[948,1167,665,690]" gridcol="3" gridrow="4" pageId="14" pageNumber="335">127±8(129–141)</td>
<td id="76E72249FFED007A36B9AC98FA7252C6" box="[1202,1408,665,690]" gridcol="4" gridrow="4" pageId="14" pageNumber="335">119±13(102–130)</td>
</tr>
<tr id="35364B35FFED007A32AAACBEFA7252AC" box="[161,1408,703,728]" gridrow="5" pageId="14" pageNumber="335">
<th id="76E72249FFED007A32AAACBEFDF952AC" box="[161,523,703,728]" gridcol="0" gridrow="5" pageId="14" pageNumber="335">Maximum body diameter</th>
<td id="76E72249FFED007A3021ACBEFD7A52AC" box="[554,648,703,728]" gridcol="1" gridrow="5" pageId="14" pageNumber="335">34.5</td>
<td id="76E72249FFED007A30A2ACBEFC6A52AC" box="[681,920,703,728]" gridcol="2" gridrow="5" pageId="14" pageNumber="335">35±3(27–40.5)</td>
<td id="76E72249FFED007A31BFACBEFB7D52AC" box="[948,1167,703,728]" gridcol="3" gridrow="5" pageId="14" pageNumber="335">36±3(31–40.5)</td>
<td id="76E72249FFED007A36B9ACBEFA7252AC" box="[1202,1408,703,728]" gridcol="4" gridrow="5" pageId="14" pageNumber="335">39±5(32–45)</td>
</tr>
<tr id="35364B35FFED007A32AAACE4FA72528A" box="[161,1408,741,766]" gridrow="6" pageId="14" pageNumber="335">
<th id="76E72249FFED007A32AAACE4FDF9528A" box="[161,523,741,766]" gridcol="0" gridrow="6" pageId="14" pageNumber="335">Anal body diameter</th>
<td id="76E72249FFED007A3021ACE4FD7A528A" box="[554,648,741,766]" gridcol="1" gridrow="6" pageId="14" pageNumber="335">34</td>
<td id="76E72249FFED007A30A2ACE4FC6A528A" box="[681,920,741,766]" gridcol="2" gridrow="6" pageId="14" pageNumber="335">32±5(21–40)</td>
<td id="76E72249FFED007A31BFACE4FB7D528A" box="[948,1167,741,766]" gridcol="3" gridrow="6" pageId="14" pageNumber="335">23±2(21–31.5)</td>
<td id="76E72249FFED007A36B9ACE4FA72528A" box="[1202,1408,741,766]" gridcol="4" gridrow="6" pageId="14" pageNumber="335">27±3(23–30)</td>
</tr>
<tr id="35364B35FFED007A32AAAD0BFA725357" box="[161,1408,778,803]" gridrow="7" pageId="14" pageNumber="335">
<th id="76E72249FFED007A32AAAD0BFDF95357" box="[161,523,778,803]" gridcol="0" gridrow="7" pageId="14" pageNumber="335">Tail length</th>
<td id="76E72249FFED007A3021AD0BFD7A5357" box="[554,648,778,803]" gridcol="1" gridrow="7" pageId="14" pageNumber="335">81</td>
<td id="76E72249FFED007A30A2AD0BFC6A5357" box="[681,920,778,803]" gridcol="2" gridrow="7" pageId="14" pageNumber="335">74±4(70.5–79.5)</td>
<td id="76E72249FFED007A31BFAD0BFB7D5357" box="[948,1167,778,803]" gridcol="3" gridrow="7" pageId="14" pageNumber="335">77±5(76–87)</td>
<td id="76E72249FFED007A36B9AD0BFA725357" box="[1202,1408,778,803]" gridcol="4" gridrow="7" pageId="14" pageNumber="335">81±10(68–91)</td>
</tr>
<tr id="35364B35FFED007A32AAAD31FA72533D" box="[161,1408,816,841]" gridrow="8" pageId="14" pageNumber="335">
<th id="76E72249FFED007A32AAAD31FDF9533D" box="[161,523,816,841]" gridcol="0" gridrow="8" pageId="14" pageNumber="335">Length of unstriated tail tip</th>
<td id="76E72249FFED007A3021AD31FD7A533D" box="[554,648,816,841]" gridcol="1" gridrow="8" pageId="14" pageNumber="335">36</td>
<td id="76E72249FFED007A30A2AD31FC6A533D" box="[681,920,816,841]" gridcol="2" gridrow="8" pageId="14" pageNumber="335">35±5(29–39)</td>
<td id="76E72249FFED007A31BFAD31FB7D533D" box="[948,1167,816,841]" gridcol="3" gridrow="8" pageId="14" pageNumber="335">39±2(35–41)</td>
<td id="76E72249FFED007A36B9AD31FA72533D" box="[1202,1408,816,841]" gridcol="4" gridrow="8" pageId="14" pageNumber="335">35±2(33–38)</td>
</tr>
<tr id="35364B35FFED007A32AAAD57FA72531B" box="[161,1408,854,879]" gridrow="9" pageId="14" pageNumber="335">
<th id="76E72249FFED007A32AAAD57FDF9531B" box="[161,523,854,879]" gridcol="0" gridrow="9" pageId="14" pageNumber="335">Head diameter</th>
<td id="76E72249FFED007A3021AD57FD7A531B" box="[554,648,854,879]" gridcol="1" gridrow="9" pageId="14" pageNumber="335">21.5</td>
<td id="76E72249FFED007A30A2AD57FC6A531B" box="[681,920,854,879]" gridcol="2" gridrow="9" pageId="14" pageNumber="335">24±3(19.5–30)</td>
<td id="76E72249FFED007A31BFAD57FB7D531B" box="[948,1167,854,879]" gridcol="3" gridrow="9" pageId="14" pageNumber="335">24±3(19–27)</td>
<td id="76E72249FFED007A36B9AD57FA72531B" box="[1202,1408,854,879]" gridcol="4" gridrow="9" pageId="14" pageNumber="335">19±2(17–21)</td>
</tr>
<tr id="35364B35FFED007A32AAAD7DFA7253E1" box="[161,1408,892,917]" gridrow="10" pageId="14" pageNumber="335">
<th id="76E72249FFED007A32AAAD7DFDF953E1" box="[161,523,892,917]" gridcol="0" gridrow="10" pageId="14" pageNumber="335">Cephalic capsule length</th>
<td id="76E72249FFED007A3021AD7DFD7A53E1" box="[554,648,892,917]" gridcol="1" gridrow="10" pageId="14" pageNumber="335">21.5</td>
<td id="76E72249FFED007A30A2AD7DFC6A53E1" box="[681,920,892,917]" gridcol="2" gridrow="10" pageId="14" pageNumber="335">23±2(19.5–27)</td>
<td id="76E72249FFED007A31BFAD7DFB7D53E1" box="[948,1167,892,917]" gridcol="3" gridrow="10" pageId="14" pageNumber="335">24±3(19–27)</td>
<td id="76E72249FFED007A36B9AD7DFA7253E1" box="[1202,1408,892,917]" gridcol="4" gridrow="10" pageId="14" pageNumber="335">16±2(12–18)</td>
</tr>
<tr id="35364B35FFED007A32AAADA3FA7253CF" box="[161,1408,930,955]" gridrow="11" pageId="14" pageNumber="335">
<th id="76E72249FFED007A32AAADA3FDF953CF" box="[161,523,930,955]" gridcol="0" gridrow="11" pageId="14" pageNumber="335">Spicule length along arc</th>
<td id="76E72249FFED007A3021ADA3FD7A53CF" box="[554,648,930,955]" gridcol="1" gridrow="11" pageId="14" pageNumber="335">51</td>
<td id="76E72249FFED007A30A2ADA3FC6A53CF" box="[681,920,930,955]" gridcol="2" gridrow="11" pageId="14" pageNumber="335">43±5(33–52.5)</td>
<td id="76E72249FFED007A31BFADA3FB7D53CF" box="[948,1167,930,955]" gridcol="3" gridrow="11" pageId="14" pageNumber="335">-</td>
<td id="76E72249FFED007A36B9ADA3FA7253CF" box="[1202,1408,930,955]" gridcol="4" gridrow="11" pageId="14" pageNumber="335">-</td>
</tr>
<tr id="35364B35FFED007A32AAADC6FA725394" box="[161,1408,967,992]" gridrow="12" pageId="14" pageNumber="335">
<th id="76E72249FFED007A32AAADC6FDF95394" box="[161,523,967,992]" gridcol="0" gridrow="12" pageId="14" pageNumber="335">Gubernaculum length</th>
<td id="76E72249FFED007A3021ADC6FD7A5394" box="[554,648,967,992]" gridcol="1" gridrow="12" pageId="14" pageNumber="335">24</td>
<td id="76E72249FFED007A30A2ADC6FC6A5394" box="[681,920,967,992]" gridcol="2" gridrow="12" pageId="14" pageNumber="335">20±3(15–22.5)</td>
<td id="76E72249FFED007A31BFADC6FB7D5394" box="[948,1167,967,992]" gridcol="3" gridrow="12" pageId="14" pageNumber="335">-</td>
<td id="76E72249FFED007A36B9ADC6FA725394" box="[1202,1408,967,992]" gridcol="4" gridrow="12" pageId="14" pageNumber="335">-</td>
</tr>
<tr id="35364B35FFED007A32AAADECFA725472" box="[161,1408,1005,1030]" gridrow="13" pageId="14" pageNumber="335">
<th id="76E72249FFED007A32AAADECFDF95472" box="[161,523,1005,1030]" gridcol="0" gridrow="13" pageId="14" pageNumber="335">Velum length</th>
<td id="76E72249FFED007A3021ADECFD7A5472" box="[554,648,1005,1030]" gridcol="1" gridrow="13" pageId="14" pageNumber="335">32</td>
<td id="76E72249FFED007A30A2ADECFC6A5472" box="[681,920,1005,1030]" gridcol="2" gridrow="13" pageId="14" pageNumber="335">31±3(25–37.5)</td>
<td id="76E72249FFED007A31BFADECFB7D5472" box="[948,1167,1005,1030]" gridcol="3" gridrow="13" pageId="14" pageNumber="335">-</td>
<td id="76E72249FFED007A36B9ADECFA725472" box="[1202,1408,1005,1030]" gridcol="4" gridrow="13" pageId="14" pageNumber="335">-</td>
</tr>
<tr id="35364B35FFED007A32AAAA12FA725458" box="[161,1408,1043,1068]" gridrow="14" pageId="14" pageNumber="335">
<th id="76E72249FFED007A32AAAA12FDF95458" box="[161,523,1043,1068]" gridcol="0" gridrow="14" pageId="14" pageNumber="335">Lateral alae length</th>
<td id="76E72249FFED007A3021AA12FD7A5458" box="[554,648,1043,1068]" gridcol="1" gridrow="14" pageId="14" pageNumber="335">67.5</td>
<td id="76E72249FFED007A30A2AA12FC6A5458" box="[681,920,1043,1068]" gridcol="2" gridrow="14" pageId="14" pageNumber="335">63±5(55–72)</td>
<td id="76E72249FFED007A31BFAA12FB7D5458" box="[948,1167,1043,1068]" gridcol="3" gridrow="14" pageId="14" pageNumber="335">68±6(52–62)</td>
<td id="76E72249FFED007A36B9AA12FA725458" box="[1202,1408,1043,1068]" gridcol="4" gridrow="14" pageId="14" pageNumber="335">-</td>
</tr>
<tr id="35364B35FFED007A32AAAA38FA725426" box="[161,1408,1081,1106]" gridrow="15" pageId="14" pageNumber="335">
<th id="76E72249FFED007A32AAAA38FDF95426" box="[161,523,1081,1106]" gridcol="0" gridrow="15" pageId="14" pageNumber="335">Length of cephalic setae</th>
<td id="76E72249FFED007A3021AA38FD7A5426" box="[554,648,1081,1106]" gridcol="1" gridrow="15" pageId="14" pageNumber="335">7</td>
<td id="76E72249FFED007A30A2AA38FC6A5426" box="[681,920,1081,1106]" gridcol="2" gridrow="15" pageId="14" pageNumber="335">7±2(5.5–9)</td>
<td id="76E72249FFED007A31BFAA38FB7D5426" box="[948,1167,1081,1106]" gridcol="3" gridrow="15" pageId="14" pageNumber="335">5±1(5–6)</td>
<td id="76E72249FFED007A36B9AA38FA725426" box="[1202,1408,1081,1106]" gridcol="4" gridrow="15" pageId="14" pageNumber="335">5±1(3.4–5.5)</td>
</tr>
<tr id="35364B35FFED007A32AAAA5EFA72540C" box="[161,1408,1119,1144]" gridrow="16" pageId="14" pageNumber="335">
<th id="76E72249FFED007A32AAAA5EFDF9540C" box="[161,523,1119,1144]" gridcol="0" gridrow="16" pageId="14" pageNumber="335">Percentage of bulb diameter (cbd)*</th>
<td id="76E72249FFED007A3021AA5EFD7A540C" box="[554,648,1119,1144]" gridcol="1" gridrow="16" pageId="14" pageNumber="335">65</td>
<td id="76E72249FFED007A30A2AA5EFC6A540C" box="[681,920,1119,1144]" gridcol="2" gridrow="16" pageId="14" pageNumber="335">60±5(57–67)</td>
<td id="76E72249FFED007A31BFAA5EFB7D540C" box="[948,1167,1119,1144]" gridcol="3" gridrow="16" pageId="14" pageNumber="335">60±4(55–66)</td>
<td id="76E72249FFED007A36B9AA5EFA72540C" box="[1202,1408,1119,1144]" gridcol="4" gridrow="16" pageId="14" pageNumber="335">65±6(55–70)</td>
</tr>
<tr id="35364B35FFED007A32AAAA85FA7254E9" box="[161,1408,1156,1181]" gridrow="17" pageId="14" pageNumber="335">
<th id="76E72249FFED007A32AAAA85FDF954E9" box="[161,523,1156,1181]" gridcol="0" gridrow="17" pageId="14" pageNumber="335">
Diameter of
<emphasis id="B9729565FFEDFF8B332AAA84FE2454E9" box="[289,470,1157,1181]" italics="true" pageId="14" pageNumber="335">fovea amphidialis</emphasis>
in %
</th>
<td id="76E72249FFED007A3021AA85FD7A54E9" box="[554,648,1156,1181]" gridcol="1" gridrow="17" pageId="14" pageNumber="335">58</td>
<td id="76E72249FFED007A30A2AA85FC6A54E9" box="[681,920,1156,1181]" gridcol="2" gridrow="17" pageId="14" pageNumber="335">56±6(50–62)</td>
<td id="76E72249FFED007A31BFAA85FB7D54E9" box="[948,1167,1156,1181]" gridcol="3" gridrow="17" pageId="14" pageNumber="335">52±7(44–60)</td>
<td id="76E72249FFED007A36B9AA85FA7254E9" box="[1202,1408,1156,1181]" gridcol="4" gridrow="17" pageId="14" pageNumber="335">49±5(40–53)</td>
</tr>
<tr id="35364B35FFED007A32AAAAA9FA725496" box="[161,1408,1192,1250]" gridrow="18" pageId="14" pageNumber="335">
<th id="76E72249FFED007A32AAAAA9FDF95496" box="[161,523,1192,1250]" gridcol="0" gridrow="18" pageId="14" pageNumber="335">of cephalic capsule width Striation of the cuticle</th>
<td id="76E72249FFED007A3021AAA9FD7A5496" box="[554,648,1192,1250]" gridcol="1" gridrow="18" pageId="14" pageNumber="335">5.5</td>
<td id="76E72249FFED007A30A2AAA9FC6A5496" box="[681,920,1192,1250]" gridcol="2" gridrow="18" pageId="14" pageNumber="335">5±1(4.5–6)</td>
<td id="76E72249FFED007A31BFAAA9FB7D5496" box="[948,1167,1192,1250]" gridcol="3" gridrow="18" pageId="14" pageNumber="335">6±0(5.5)</td>
<td id="76E72249FFED007A36B9AAA9FA725496" box="[1202,1408,1192,1250]" gridcol="4" gridrow="18" pageId="14" pageNumber="335">5±0(4.5–5)</td>
</tr>
<tr id="35364B35FFED007A32AAAAEEFA72557C" box="[161,1408,1263,1288]" gridrow="19" pageId="14" pageNumber="335">
<th id="76E72249FFED007A32AAAAEEFDF9557C" box="[161,523,1263,1288]" gridcol="0" gridrow="19" pageId="14" pageNumber="335">Distance from anterior end to the</th>
<td id="76E72249FFED007A3021AAEEFD7A557C" box="[554,648,1263,1288]" gridcol="1" gridrow="19" pageId="14" pageNumber="335">-</td>
<td id="76E72249FFED007A30A2AAEEFC6A557C" box="[681,920,1263,1288]" gridcol="2" gridrow="19" pageId="14" pageNumber="335">-</td>
<td id="76E72249FFED007A31BFAAEEFB7D557C" box="[948,1167,1263,1288]" gridcol="3" gridrow="19" pageId="14" pageNumber="335">709±113(600–832)</td>
<td id="76E72249FFED007A36B9AAEEFA72557C" box="[1202,1408,1263,1288]" gridcol="4" gridrow="19" pageId="14" pageNumber="335">710±63(620–779)</td>
</tr>
<tr id="35364B35FFED007A32AAAB12FA725539" box="[161,1408,1299,1357]" gridrow="20" pageId="14" pageNumber="335">
<th id="76E72249FFED007A32AAAB12FDF95539" box="[161,523,1299,1357]" gridcol="0" gridrow="20" pageId="14" pageNumber="335">vulva V%</th>
<td id="76E72249FFED007A3021AB12FD7A5539" box="[554,648,1299,1357]" gridcol="1" gridrow="20" pageId="14" pageNumber="335">-</td>
<td id="76E72249FFED007A30A2AB12FC6A5539" box="[681,920,1299,1357]" gridcol="2" gridrow="20" pageId="14" pageNumber="335">-</td>
<td id="76E72249FFED007A31BFAB12FB7D5539" box="[948,1167,1299,1357]" gridcol="3" gridrow="20" pageId="14" pageNumber="335">43±2(40–56)</td>
<td id="76E72249FFED007A36B9AB12FA725539" box="[1202,1408,1299,1357]" gridcol="4" gridrow="20" pageId="14" pageNumber="335">66±4(60–71)</td>
</tr>
<tr id="35364B35FFED007A32AAAB58FA725506" box="[161,1408,1369,1394]" gridrow="21" pageId="14" pageNumber="335">
<th id="76E72249FFED007A32AAAB58FDF95506" box="[161,523,1369,1394]" gridcol="0" gridrow="21" pageId="14" pageNumber="335">Ring with the inversion of direction</th>
<td id="76E72249FFED007A3021AB58FD7A5506" box="[554,648,1369,1394]" gridcol="1" gridrow="21" pageId="14" pageNumber="335">80</td>
<td id="76E72249FFED007A30A2AB58FC6A5506" box="[681,920,1369,1394]" gridcol="2" gridrow="21" pageId="14" pageNumber="335">85±4(80–89)</td>
<td id="76E72249FFED007A31BFAB58FB7D5506" box="[948,1167,1369,1394]" gridcol="3" gridrow="21" pageId="14" pageNumber="335">77±2(75–89)</td>
<td id="76E72249FFED007A36B9AB58FA725506" box="[1202,1408,1369,1394]" gridcol="4" gridrow="21" pageId="14" pageNumber="335">-</td>
</tr>
<tr id="35364B35FFED007A32AAAB7CFA7255C3" box="[161,1408,1405,1463]" gridrow="22" pageId="14" pageNumber="335">
<th id="76E72249FFED007A32AAAB7CFDF955C3" box="[161,523,1405,1463]" gridcol="0" gridrow="22" pageId="14" pageNumber="335">of annulation a</th>
<td id="76E72249FFED007A3021AB7CFD7A55C3" box="[554,648,1405,1463]" gridcol="1" gridrow="22" pageId="14" pageNumber="335">29.5</td>
<td id="76E72249FFED007A30A2AB7CFC6A55C3" box="[681,920,1405,1463]" gridcol="2" gridrow="22" pageId="14" pageNumber="335">29±3(25–32)</td>
<td id="76E72249FFED007A31BFAB7CFB7D55C3" box="[948,1167,1405,1463]" gridcol="3" gridrow="22" pageId="14" pageNumber="335">30±4(24–31)</td>
<td id="76E72249FFED007A36B9AB7CFA7255C3" box="[1202,1408,1405,1463]" gridcol="4" gridrow="22" pageId="14" pageNumber="335">26±2(25–30)</td>
</tr>
<tr id="35364B35FFED007A32AAABC5FA7255A9" box="[161,1408,1476,1501]" gridrow="23" pageId="14" pageNumber="335">
<th id="76E72249FFED007A32AAABC5FDF955A9" box="[161,523,1476,1501]" gridcol="0" gridrow="23" pageId="14" pageNumber="335">b</th>
<td id="76E72249FFED007A3021ABC5FD7A55A9" box="[554,648,1476,1501]" gridcol="1" gridrow="23" pageId="14" pageNumber="335">8</td>
<td id="76E72249FFED007A30A2ABC5FC6A55A9" box="[681,920,1476,1501]" gridcol="2" gridrow="23" pageId="14" pageNumber="335">8±0(7.5–9)</td>
<td id="76E72249FFED007A31BFABC5FB7D55A9" box="[948,1167,1476,1501]" gridcol="3" gridrow="23" pageId="14" pageNumber="335">8±1(6.5–10)</td>
<td id="76E72249FFED007A36B9ABC5FA7255A9" box="[1202,1408,1476,1501]" gridcol="4" gridrow="23" pageId="14" pageNumber="335">8±1(7.5–9)</td>
</tr>
<tr id="35364B35FFED007A32AAABEBFA725677" box="[161,1408,1514,1539]" gridrow="24" pageId="14" pageNumber="335">
<th id="76E72249FFED007A32AAABEBFDF95677" box="[161,523,1514,1539]" gridcol="0" gridrow="24" pageId="14" pageNumber="335">c</th>
<td id="76E72249FFED007A3021ABEBFD7A5677" box="[554,648,1514,1539]" gridcol="1" gridrow="24" pageId="14" pageNumber="335">12.5</td>
<td id="76E72249FFED007A30A2ABEBFC6A5677" box="[681,920,1514,1539]" gridcol="2" gridrow="24" pageId="14" pageNumber="335">14±1(11.5–16)</td>
<td id="76E72249FFED007A31BFABEBFB7D5677" box="[948,1167,1514,1539]" gridcol="3" gridrow="24" pageId="14" pageNumber="335">3±0(10–16)</td>
<td id="76E72249FFED007A36B9ABEBFA725677" box="[1202,1408,1514,1539]" gridcol="4" gridrow="24" pageId="14" pageNumber="335">12±1(11–14)</td>
</tr>
</table>
</paragraph>
<paragraph id="8BB94977FFEDFF8B32AAA80EFE24565C" blockId="14.[161,470,1551,1576]" box="[161,470,1551,1576]" pageId="14" pageNumber="335">
<tableNote id="76E048F9FFEDFF8B32AAA80EFE24565C" box="[161,470,1551,1576]" pageId="14" pageNumber="335" targetBox="[161,1408,514,1539]" targetPageId="14">*corresponding body diameter</tableNote>
</paragraph>
<subSubSection id="C31C1AFCFFEDFF9732CCA859FB6252F3" lastPageId="18" lastPageNumber="339" pageId="14" pageNumber="335" type="description">
<paragraph id="8BB94977FFEDFF9732CCA859FEDD5113" blockId="14.[151,1437,1624,1975]" lastBlockId="18.[151,1437,153,1007]" lastPageId="18" lastPageNumber="339" pageId="14" pageNumber="335">
<emphasis id="B9729565FFEDFF8B32CCA859FD765607" bold="true" box="[199,644,1624,1651]" pageId="14" pageNumber="335">
Description.
<typeStatus id="54BDF7D5FFEDFF8B3354A858FE3E5607" box="[351,460,1625,1651]" pageId="14" pageNumber="335" type="holotype">Holotype</typeStatus>
(
<figureCitation id="133D55F2FFEDFF8B33D1A859FD855607" box="[474,631,1624,1651]" captionStart-0="FIGURE 9" captionStart-1="FIGURE 10" captionStartId-0="15.[151,250,1961,1986]" captionStartId-1="16.[151,250,1909,1934]" captionTargetBox-0="[221,1379,172,1917]" captionTargetBox-1="[166,1248,176,1882]" captionTargetId-0="figure@15.[206,1382,166,1941]" captionTargetId-1="figure@16.[151,1260,166,1890]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionText-0="FIGURE 9. Spinonema absente gen. et sp. n. Holotype MNRJ 400. A: overview; B, C and F: anterior region; D: tail region; E: spicules; H: lateral alae (ANT anterior part and POS posterior part); G: Inversion of direction of annuli." captionText-1="FIGURE 10. Spinonema absente gen. et sp. n. A and C, Holotype male MNRJ 400; A: Cephalic region; C: Tail region. B, D, E and F, Paratype female MNRJ 401; B: Tail region; D: Lateral alae (ANT anterior part and POS posterior part); E: Vulva opening; F: Anterior region." httpUri-0="https://zenodo.org/record/3244734/files/figure.png" httpUri-1="https://zenodo.org/record/3244736/files/figure.png" pageId="14" pageNumber="335">Figures 9–10</figureCitation>
).
</emphasis>
Body cylindrical, yellowish brown. Cuticle annulated, with irregularly distributed vacuoles that start to appear from 6
<superScript id="7C73E43FFFEDFF8B3097A87AFD5B56FD" attach="left" box="[668,681,1659,1673]" fontSize="6" pageId="14" pageNumber="335">th</superScript>
(5-12
<superScript id="7C73E43FFFEDFF8B30E2A87AFD0456FD" attach="left" box="[745,758,1659,1673]" fontSize="6" pageId="14" pageNumber="335">th</superScript>
for
<typeStatus id="54BDF7D5FFEDFF8B312AA87CFC7C56E3" box="[801,910,1661,1687]" pageId="14" pageNumber="335" type="paratype">paratypes</typeStatus>
) cuticular annule. The vacuoles are in the inner layer of the cuticle and not in its exterior. Annulations absent on tail end with vacuoles. Cuticle with lateral alae, thicker than rest of body, reaching only 19 (17-25) rings of total body length. Lateral alae begin
<quantity id="4CFEE492FFEDFF8B36DDA8C5FADC56AB" box="[1238,1326,1732,1759]" metricMagnitude="-1" metricUnit="m" metricValue="4.68" pageId="14" pageNumber="335" unit="mm" value="468.0">468 µm</quantity>
(
<quantity id="4CFEE492FFEDFF8B3736A8C4FF4F5777" metricMagnitude="-1" metricUnit="m" metricValue="4.255" metricValueMax="4.7299999999999995" metricValueMin="3.78" pageId="14" pageNumber="335" unit="mm" value="425.5" valueMax="473.0" valueMin="378.0">378-473 µm</quantity>
for
<typeStatus id="54BDF7D5FFEDFF8B32E7A8E8FEAB5777" box="[236,345,1769,1795]" pageId="14" pageNumber="335" type="paratype">paratypes</typeStatus>
) from head. Annules in anterior region are
<quantity id="4CFEE492FFEDFF8B3134A8E9FC625777" box="[831,912,1768,1795]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="14" pageNumber="335" unit="mm" value="4.5">4.5 µm</quantity>
wide, at midline
<quantity id="4CFEE492FFEDFF8B365FA8E8FB635777" box="[1108,1169,1769,1795]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="14" pageNumber="335" unit="mm" value="4.0">4 µm</quantity>
, and in posterior region
<quantity id="4CFEE492FFEDFF8B329CA90DFF1E5753" box="[151,236,1804,1831]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="14" pageNumber="335" unit="mm" value="3.5">3.5 µm</quantity>
. Sparse somatic setae irregularly distributed along body. Somatic setae indistinct or barely visible in pharyngeal region. Inversion of direction of cuticular annuli occurring after 80
<superScript id="7C73E43FFFEDFF8B31D4A92EFC1E5749" attach="left" box="[991,1004,1839,1853]" fontSize="6" pageId="14" pageNumber="335">th</superScript>
(80
<superScript id="7C73E43FFFEDFF8B3612A92EFBDF573F" attach="right" box="[1049,1069,1839,1867]" fontSize="6" pageId="14" pageNumber="335">th-</superScript>
89
<superScript id="7C73E43FFFEDFF8B3640A92EFBAA5749" attach="left" box="[1099,1112,1839,1853]" fontSize="6" pageId="14" pageNumber="335">th</superScript>
for
<typeStatus id="54BDF7D5FFEDFF8B3688A930FB02573F" box="[1155,1264,1841,1867]" pageId="14" pageNumber="335" type="paratype">paratypes</typeStatus>
) cuticular ring. Cephalic capsule long and well developed, with thick cuticle without annulations. Anterior sensilla arrangement: six inner and six outer labial sensilla, both papilliform. Four cephalic setiform sensilla located at level with upper portion of
<emphasis id="B9729565FFEDFF8B3304A99CFE2557C2" box="[271,471,1948,1975]" italics="true" pageId="14" pageNumber="335">fovea amphidialis</emphasis>
.
<emphasis id="B9729565FFEDFF8B33EDA99CFD4A57C2" box="[486,696,1948,1974]" italics="true" pageId="14" pageNumber="335">Fovea amphidialis</emphasis>
cryptospiral, occupying 58% (50-62%) of diameter of head. Oral cavity with one dorsal tooth and one (or two, difficult to see) ventral tooth. Cylindrical pharynx with muscular and pyriform basal bulb. Ventral gland and secretory-excretory pore not observed. Cardia inserted in intestine. Reproductive system monorchic, with single anterior testis situated to left of intestine, outstretched. Globular sperm cells. Spicules arched, cuticularized and capitulum rounded.
<emphasis id="B9729565FFF1FF973118AF04FCA5516B" box="[787,855,261,287]" italics="true" pageId="18" pageNumber="339">Velum</emphasis>
absent; gubernaculum laminar, without apophysis (
<figureCitation id="133D55F2FFF1FF973294AF28FEE75137" box="[159,277,297,323]" captionStart="FIGURE 9" captionStartId="15.[151,250,1961,1986]" captionTargetBox="[221,1379,172,1917]" captionTargetId="figure@15.[206,1382,166,1941]" captionTargetPageId="15" captionText="FIGURE 9. Spinonema absente gen. et sp. n. Holotype MNRJ 400. A: overview; B, C and F: anterior region; D: tail region; E: spicules; H: lateral alae (ANT anterior part and POS posterior part); G: Inversion of direction of annuli." httpUri="https://zenodo.org/record/3244734/files/figure.png" pageId="18" pageNumber="339">Figure 9C</figureCitation>
). Three caudal glands. Tail conical with numerous vacuoles. Three precloacal setae (
<figureCitation id="133D55F2FFF1FF9736C1AF28FABE5137" box="[1226,1356,297,323]" captionStart="FIGURE 10" captionStartId="16.[151,250,1909,1934]" captionTargetBox="[166,1248,176,1882]" captionTargetId="figure@16.[151,1260,166,1890]" captionTargetPageId="16" captionText="FIGURE 10. Spinonema absente gen. et sp. n. A and C, Holotype male MNRJ 400; A: Cephalic region; C: Tail region. B, D, E and F, Paratype female MNRJ 401; B: Tail region; D: Lateral alae (ANT anterior part and POS posterior part); E: Vulva opening; F: Anterior region." httpUri="https://zenodo.org/record/3244736/files/figure.png" pageId="18" pageNumber="339">Figure 10F</figureCitation>
). Spinneret conical.
</paragraph>
<caption id="DF7919FFFFECFF8A329CA9A8FB205792" httpUri="https://zenodo.org/record/3244734/files/figure.png" pageId="15" pageNumber="336" startId="15.[151,250,1961,1986]" targetBox="[221,1379,172,1917]" targetPageId="15">
<paragraph id="8BB94977FFECFF8A329CA9A8FB205792" blockId="15.[151,1436,1961,2022]" pageId="15" pageNumber="336">
<emphasis id="B9729565FFECFF8A329CA9A8FEE657B6" bold="true" box="[151,276,1961,1986]" pageId="15" pageNumber="336">FIGURE 9.</emphasis>
<taxonomicName id="4C0632F4FFECFF8A3310A9ABFE1257B5" authority="Larrazábal-Filho &amp; Neres &amp; Silva &amp; Esteves, 2019" authorityName="Larrazábal-Filho &amp; Neres &amp; Silva &amp; Esteves" authorityYear="2019" box="[283,480,1961,1985]" class="Adenophorea" family="Desmodoridae" genus="Spinonema" kingdom="Animalia" order="Desmodorida" pageId="15" pageNumber="336" phylum="Nematoda" rank="species" species="absente" status="gen. et sp. n.">
<emphasis id="B9729565FFECFF8A3310A9ABFE1257B5" box="[283,480,1961,1985]" italics="true" pageId="15" pageNumber="336">Spinonema absente</emphasis>
</taxonomicName>
<emphasis id="B9729565FFECFF8A33ECA9ABFD9D57B6" bold="true" box="[487,623,1962,1986]" pageId="15" pageNumber="336">
<taxonomicNameLabel id="A241281EFFECFF8A33ECA9ABFD9D57B6" box="[487,623,1962,1986]" pageId="15" pageNumber="336" rank="species">gen. et sp. n.</taxonomicNameLabel>
</emphasis>
Holotype MNRJ 400.
<emphasis id="B9729565FFECFF8A3151A9A8FC9E57B6" bold="true" box="[858,876,1961,1986]" pageId="15" pageNumber="336">A</emphasis>
: overview;
<emphasis id="B9729565FFECFF8A31ECA9ABFC0A57B6" bold="true" box="[999,1016,1962,1986]" pageId="15" pageNumber="336">B</emphasis>
,
<emphasis id="B9729565FFECFF8A360EA9A8FBE557B6" bold="true" box="[1029,1047,1961,1986]" pageId="15" pageNumber="336">C</emphasis>
and
<emphasis id="B9729565FFECFF8A3641A9ABFBAB57B6" bold="true" box="[1098,1113,1962,1986]" pageId="15" pageNumber="336">F</emphasis>
: anterior region;
<emphasis id="B9729565FFECFF8A3706A9ABFAED57B6" bold="true" box="[1293,1311,1962,1986]" pageId="15" pageNumber="336">D</emphasis>
: tail region;
<emphasis id="B9729565FFECFF8A329CA9CFFF5A5792" bold="true" box="[151,168,1998,2022]" pageId="15" pageNumber="336">E</emphasis>
: spicules;
<emphasis id="B9729565FFECFF8A331FA9CFFEDA5792" bold="true" box="[276,296,1998,2022]" pageId="15" pageNumber="336">H</emphasis>
: lateral alae (ANT anterior part and POS posterior part);
<emphasis id="B9729565FFECFF8A3166A9CCFC735792" bold="true" box="[877,897,1997,2022]" pageId="15" pageNumber="336">G</emphasis>
: Inversion of direction of annuli.
</paragraph>
</caption>
<caption id="DF7919FFFFF3FF95329CA974FE4457A2" httpUri="https://zenodo.org/record/3244736/files/figure.png" pageId="16" pageNumber="337" startId="16.[151,250,1909,1934]" targetBox="[166,1248,176,1882]" targetPageId="16">
<paragraph id="8BB94977FFF3FF95329CA974FE4457A2" blockId="16.[151,1436,1909,2006]" pageId="16" pageNumber="337">
<emphasis id="B9729565FFF3FF95329CA974FED057FA" bold="true" box="[151,290,1909,1934]" pageId="16" pageNumber="337">FIGURE 10.</emphasis>
<taxonomicName id="4C0632F4FFF3FF953321A977FE1D57FA" authority="Larrazábal-Filho &amp; Neres &amp; Silva &amp; Esteves, 2019" authorityName="Larrazábal-Filho &amp; Neres &amp; Silva &amp; Esteves" authorityYear="2019" box="[298,495,1910,1934]" class="Adenophorea" family="Desmodoridae" genus="Spinonema" kingdom="Animalia" order="Desmodorida" pageId="16" pageNumber="337" phylum="Nematoda" rank="species" species="absente" status="gen. et sp. n.">
<emphasis id="B9729565FFF3FF953321A977FE1D57FA" box="[298,495,1910,1934]" italics="true" pageId="16" pageNumber="337">Spinonema absente</emphasis>
</taxonomicName>
<emphasis id="B9729565FFF3FF9533F3A977FD6E57FA" bold="true" box="[504,668,1909,1934]" pageId="16" pageNumber="337">
<taxonomicNameLabel id="A241281EFFF3FF9533F3A977FD7157FA" box="[504,643,1910,1934]" pageId="16" pageNumber="337" rank="species">gen. et sp. n.</taxonomicNameLabel>
A
</emphasis>
and
<emphasis id="B9729565FFF3FF9530DAA974FD1157FA" bold="true" box="[721,739,1909,1934]" pageId="16" pageNumber="337">C</emphasis>
, Holotype male MNRJ 400;
<emphasis id="B9729565FFF3FF953619A974FBD657FA" bold="true" box="[1042,1060,1909,1934]" pageId="16" pageNumber="337">A</emphasis>
: Cephalic region;
<emphasis id="B9729565FFF3FF9536EDA974FB0A57FA" bold="true" box="[1254,1272,1909,1934]" pageId="16" pageNumber="337">C</emphasis>
: Tail region.
<emphasis id="B9729565FFF3FF95378EA977FA6457FA" bold="true" box="[1413,1430,1910,1934]" pageId="16" pageNumber="337">B</emphasis>
,
<emphasis id="B9729565FFF3FF95329CA99BFF5B57C6" bold="true" box="[151,169,1946,1970]" pageId="16" pageNumber="337">D</emphasis>
,
<emphasis id="B9729565FFF3FF9532BCA99BFF3A57C6" bold="true" box="[183,200,1946,1970]" pageId="16" pageNumber="337">E</emphasis>
and
<emphasis id="B9729565FFF3FF9532F1A99BFEFB57C6" bold="true" box="[250,265,1946,1970]" pageId="16" pageNumber="337">F</emphasis>
, Paratype female MNRJ 401;
<emphasis id="B9729565FFF3FF95304AA99BFDA057C6" bold="true" box="[577,594,1946,1970]" pageId="16" pageNumber="337">B</emphasis>
: Tail region;
<emphasis id="B9729565FFF3FF9530D7A99BFD1C57C6" bold="true" box="[732,750,1946,1970]" pageId="16" pageNumber="337">D</emphasis>
: Lateral alae (ANT anterior part and POS posterior part);
<emphasis id="B9729565FFF3FF95374AA99BFAA057C6" bold="true" box="[1345,1362,1946,1970]" pageId="16" pageNumber="337">E</emphasis>
: Vulva opening;
<emphasis id="B9729565FFF3FF9532FDA9BFFEF757A2" bold="true" box="[246,261,1982,2006]" pageId="16" pageNumber="337">F</emphasis>
: Anterior region.
</paragraph>
</caption>
<caption id="DF7919FFFFF2FF94329CA9ABFB4C5793" httpUri="https://zenodo.org/record/3244738/files/figure.png" pageId="17" pageNumber="338" startId="17.[151,250,1962,1987]" targetBox="[219,1359,183,1869]" targetPageId="17">
<paragraph id="8BB94977FFF2FF94329CA9ABFB4C5793" blockId="17.[151,1437,1962,2023]" pageId="17" pageNumber="338">
<emphasis id="B9729565FFF2FF94329CA9ABFEED57B7" bold="true" box="[151,287,1962,1987]" pageId="17" pageNumber="338">FIGURE 11.</emphasis>
<taxonomicName id="4C0632F4FFF2FF94332EA9AAFE1B57B7" authority="Larrazábal-Filho &amp; Neres &amp; Silva &amp; Esteves, 2019" authorityName="Larrazábal-Filho &amp; Neres &amp; Silva &amp; Esteves" authorityYear="2019" box="[293,489,1963,1987]" class="Adenophorea" family="Desmodoridae" genus="Spinonema" kingdom="Animalia" order="Desmodorida" pageId="17" pageNumber="338" phylum="Nematoda" rank="species" species="absente" status="gen. et sp. n.">
<emphasis id="B9729565FFF2FF94332EA9AAFE1B57B7" box="[293,489,1963,1987]" italics="true" pageId="17" pageNumber="338">Spinonema absente</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A241281EFFF2FF9433FBA9AAFD8557B7" box="[496,631,1963,1987]" pageId="17" pageNumber="338" rank="species">
<emphasis id="B9729565FFF2FF9433FBA9AAFD8557B7" bold="true" box="[496,631,1963,1987]" pageId="17" pageNumber="338">gen. et sp. n.</emphasis>
</taxonomicNameLabel>
Paratype female MNRJ 401.
<emphasis id="B9729565FFF2FF9431AEA9ABFC4557B7" bold="true" box="[933,951,1962,1987]" pageId="17" pageNumber="338">A</emphasis>
: overview;
<emphasis id="B9729565FFF2FF94363AA9ABFBB157B7" bold="true" box="[1073,1091,1962,1987]" pageId="17" pageNumber="338">C</emphasis>
: Ovary and vulva opening;
<emphasis id="B9729565FFF2FF94376BA9AAFA8357B7" bold="true" box="[1376,1393,1963,1987]" pageId="17" pageNumber="338">B</emphasis>
and
<emphasis id="B9729565FFF2FF94329CA9CEFF5B5793" bold="true" box="[151,169,1999,2023]" pageId="17" pageNumber="338">D</emphasis>
: Anterior region. Paratype juvenile in stage 4.
<emphasis id="B9729565FFF2FF94308FA9CEFD675793" bold="true" box="[644,661,1999,2023]" pageId="17" pageNumber="338">E</emphasis>
: Anterior region;
<emphasis id="B9729565FFF2FF943140A9CEFCA85793" bold="true" box="[843,858,1999,2023]" pageId="17" pageNumber="338">F</emphasis>
: Tail region;
<emphasis id="B9729565FFF2FF9431E8A9CFFC055793" bold="true" box="[995,1015,1998,2023]" pageId="17" pageNumber="338">G</emphasis>
: Ovary of juvenile.
</paragraph>
</caption>
<paragraph id="8BB94977FFF1FF9732CCAF70FCE8526F" blockId="18.[151,1437,153,1007]" pageId="18" pageNumber="339">
<emphasis id="B9729565FFF1FF9732CCAF70FE1D51FF" bold="true" box="[199,495,368,395]" pageId="18" pageNumber="339">
Females (
<figureCitation id="133D55F2FFF1FF973333AF71FE1051FF" box="[312,482,368,395]" captionStart-0="FIGURE 10" captionStart-1="FIGURE 11" captionStartId-0="16.[151,250,1909,1934]" captionStartId-1="17.[151,250,1962,1987]" captionTargetBox-0="[166,1248,176,1882]" captionTargetBox-1="[219,1359,183,1869]" captionTargetId-0="figure@16.[151,1260,166,1890]" captionTargetId-1="figure@17.[202,1386,166,1942]" captionTargetPageId-0="16" captionTargetPageId-1="17" captionText-0="FIGURE 10. Spinonema absente gen. et sp. n. A and C, Holotype male MNRJ 400; A: Cephalic region; C: Tail region. B, D, E and F, Paratype female MNRJ 401; B: Tail region; D: Lateral alae (ANT anterior part and POS posterior part); E: Vulva opening; F: Anterior region." captionText-1="FIGURE 11. Spinonema absente gen. et sp. n. Paratype female MNRJ 401. A: overview; C: Ovary and vulva opening; B and D: Anterior region. Paratype juvenile in stage 4. E: Anterior region; F: Tail region; G: Ovary of juvenile." httpUri-0="https://zenodo.org/record/3244736/files/figure.png" httpUri-1="https://zenodo.org/record/3244738/files/figure.png" pageId="18" pageNumber="339">Figures 10–11</figureCitation>
).
</emphasis>
Similar to male in body size and with minor differences: inversion occurs after 79
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(75
<superScript id="7C73E43FFFF1FF9732B6AF95FF2351DB" attach="right" box="[189,209,404,431]" fontSize="6" pageId="18" pageNumber="339">th-</superScript>
89
<superScript id="7C73E43FFFF1FF9732E4AF95FF0E51D6" attach="right" box="[239,252,404,418]" fontSize="6" pageId="18" pageNumber="339">th</superScript>
) cuticular ring. Lateral alae begin
<quantity id="4CFEE492FFF1FF97309BAF94FD1851DB" box="[656,746,405,431]" metricMagnitude="-1" metricUnit="m" metricValue="4.42" pageId="18" pageNumber="339" unit="mm" value="442.0">442 µm</quantity>
(
<quantity id="4CFEE492FFF1FF9730F0AF95FC7951DB" box="[763,907,404,431]" metricMagnitude="-1" metricUnit="m" metricValue="4.2299999999999995" metricValueMax="4.59" metricValueMin="3.87" pageId="18" pageNumber="339" unit="mm" value="423.0" valueMax="459.0" valueMin="387.0">387-459 µm</quantity>
) from head. Reproductive system with paired ovaries, opposite and reflexed antidromous; both genital branches to right of intestine. Vulva as simple transverse slit-like aperture. Vagina vera and vagina uterina weakly cuticularized. No egg found. Vulva located at 57% (40- 56%) of total body length from anterior end. Tail conical.
</paragraph>
<paragraph id="8BB94977FFF1FF9732CCAC25FB6252F3" blockId="18.[151,1437,153,1007]" pageId="18" pageNumber="339">
<emphasis id="B9729565FFF1FF9732CCAC25FE34524B" bold="true" box="[199,454,548,575]" pageId="18" pageNumber="339">
Juveniles (
<figureCitation id="133D55F2FFF1FF97334FAC25FE4A524B" box="[324,440,548,575]" captionStart="FIGURE 11" captionStartId="17.[151,250,1962,1987]" captionTargetBox="[219,1359,183,1869]" captionTargetId="figure@17.[202,1386,166,1942]" captionTargetPageId="17" captionText="FIGURE 11. Spinonema absente gen. et sp. n. Paratype female MNRJ 401. A: overview; C: Ovary and vulva opening; B and D: Anterior region. Paratype juvenile in stage 4. E: Anterior region; F: Tail region; G: Ovary of juvenile." httpUri="https://zenodo.org/record/3244738/files/figure.png" pageId="18" pageNumber="339">Figure 11</figureCitation>
).
</emphasis>
Similar to adults, except lacking some characteristics including supplements and lateral alae; reproductive system immature.
<emphasis id="B9729565FFF1FF973030AC4BFCFF5217" box="[571,781,585,611]" italics="true" pageId="18" pageNumber="339">Fovea amphidialis</emphasis>
is very similar to the adults. Juveniles in stage 1 lack so- matic setae arranged in rows. Five juveniles were found, four in stage 1 and one in stage 4.
</paragraph>
</subSubSection>
<subSubSection id="C31C1AFCFFF1FF9732CCAC91FC2C539B" pageId="18" pageNumber="339" type="diagnosis">
<paragraph id="8BB94977FFF1FF9732CCAC91FC4F5287" blockId="18.[151,1437,153,1007]" pageId="18" pageNumber="339">
<emphasis id="B9729565FFF1FF9732CCAC91FEB052DF" bold="true" box="[199,322,656,683]" pageId="18" pageNumber="339">Diagnosis.</emphasis>
<taxonomicName id="4C0632F4FFF1FF973341AC90FDD752DF" authority="Larrazábal-Filho &amp; Neres &amp; Silva &amp; Esteves, 2019" authorityName="Larrazábal-Filho &amp; Neres &amp; Silva &amp; Esteves" authorityYear="2019" box="[330,549,657,683]" class="Adenophorea" family="Desmodoridae" genus="Spinonema" kingdom="Animalia" order="Desmodorida" pageId="18" pageNumber="339" phylum="Nematoda" rank="species" species="absente" status="gen. et sp. n.">
<emphasis id="B9729565FFF1FF973341AC90FDD752DF" box="[330,549,657,683]" italics="true" pageId="18" pageNumber="339">Spinonema absente</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A241281EFFF1FF973025AC90FD3552DF" box="[558,711,657,683]" pageId="18" pageNumber="339" rank="species">
<emphasis id="B9729565FFF1FF973025AC90FD3552DF" bold="true" box="[558,711,657,683]" pageId="18" pageNumber="339">gen. et sp. n.</emphasis>
</taxonomicNameLabel>
is characterized by having a cuticle with irregularly distributed vacuoles that start to appear from the 6
<superScript id="7C73E43FFFF1FF973056ACB5FD9852B6" attach="left" box="[605,618,692,706]" fontSize="6" pageId="18" pageNumber="339">th</superScript>
cuticular ring, a cryptospiral
<emphasis id="B9729565FFF1FF9731C9ACB4FB7E52BB" box="[962,1164,693,719]" italics="true" pageId="18" pageNumber="339">fovea amphidialis</emphasis>
, absence of pharyngeal dorsal spine, presence of lateral alae, and three precloacal setae in male.
</paragraph>
<paragraph id="8BB94977FFF1FF9732CCACFDFC2C539B" blockId="18.[151,1437,153,1007]" pageId="18" pageNumber="339">
<emphasis id="B9729565FFF1FF9732CCACFDFE825363" bold="true" box="[199,368,764,791]" pageId="18" pageNumber="339">Relationships.</emphasis>
<taxonomicName id="4C0632F4FFF1FF97337CACFCFDA35363" authority="Larrazábal-Filho &amp; Neres &amp; Silva &amp; Esteves, 2019" authorityName="Larrazábal-Filho &amp; Neres &amp; Silva &amp; Esteves" authorityYear="2019" box="[375,593,765,791]" class="Adenophorea" family="Desmodoridae" genus="Spinonema" kingdom="Animalia" order="Desmodorida" pageId="18" pageNumber="339" phylum="Nematoda" rank="species" species="absente" status="gen. et sp. n.">
<emphasis id="B9729565FFF1FF97337CACFCFDA35363" box="[375,593,765,791]" italics="true" pageId="18" pageNumber="339">Spinonema absente</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A241281EFFF1FF973053ACFCFD1E5363" box="[600,748,765,791]" pageId="18" pageNumber="339" rank="species">
<emphasis id="B9729565FFF1FF973053ACFCFD1E5363" bold="true" box="[600,748,765,791]" pageId="18" pageNumber="339">gen. et sp. n.</emphasis>
</taxonomicNameLabel>
,
<taxonomicName id="4C0632F4FFF1FF9730F1ACFCFC965363" authority="Larrazábal-Filho &amp; Neres &amp; Silva &amp; Esteves, 2019" authorityName="Larrazábal-Filho &amp; Neres &amp; Silva &amp; Esteves" authorityYear="2019" box="[762,868,765,791]" class="Adenophorea" family="Desmodoridae" genus="Spinonema" kingdom="Animalia" order="Desmodorida" pageId="18" pageNumber="339" phylum="Nematoda" rank="species" species="spirale" status="gen. et sp. n.">
<emphasis id="B9729565FFF1FF9730F1ACFCFC965363" box="[762,868,765,791]" italics="true" pageId="18" pageNumber="339">S. spirale</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A241281EFFF1FF973160ACFCFBF25363" box="[875,1024,765,791]" pageId="18" pageNumber="339" rank="species">
<emphasis id="B9729565FFF1FF973160ACFCFBF25363" bold="true" box="[875,1024,765,791]" pageId="18" pageNumber="339">gen. et sp. n.</emphasis>
</taxonomicNameLabel>
and
<emphasis id="B9729565FFF1FF97363CACFCFB255363" box="[1079,1239,765,791]" italics="true" pageId="18" pageNumber="339">
S,
<taxonomicName id="4C0632F4FFF1FF973658ACFCFB255363" authority="Larrazábal-Filho &amp; Neres &amp; Silva &amp; Esteves, 2019" authorityName="Larrazábal-Filho &amp; Neres &amp; Silva &amp; Esteves" authorityYear="2019" box="[1107,1239,765,791]" class="Adenophorea" family="Desmodoridae" genus="Spinonema" kingdom="Animalia" order="Desmodorida" pageId="18" pageNumber="339" phylum="Nematoda" rank="species" species="cuticulatum" status="gen. et sp. n.">cuticulatum</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A241281EFFF1FF9736D5ACFCFA815363" box="[1246,1395,765,791]" pageId="18" pageNumber="339" rank="species">
<emphasis id="B9729565FFF1FF9736D5ACFCFA815363" bold="true" box="[1246,1395,765,791]" pageId="18" pageNumber="339">gen. et sp. n.</emphasis>
</taxonomicNameLabel>
are similar in the presence of vacuoles, the inversion ring and the lateral alae, although varying in length (these three features are on the cuticle).
<taxonomicName id="4C0632F4FFF1FF9733DDAD44FD43532B" authority="Larrazábal-Filho &amp; Neres &amp; Silva &amp; Esteves, 2019" authorityName="Larrazábal-Filho &amp; Neres &amp; Silva &amp; Esteves" authorityYear="2019" box="[470,689,837,863]" class="Adenophorea" family="Desmodoridae" genus="Spinonema" kingdom="Animalia" order="Desmodorida" pageId="18" pageNumber="339" phylum="Nematoda" rank="species" species="absente" status="gen. et sp. n.">
<emphasis id="B9729565FFF1FF9733DDAD44FD43532B" box="[470,689,837,863]" italics="true" pageId="18" pageNumber="339">Spinonema absente</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A241281EFFF1FF9730B2AD44FCA0532B" box="[697,850,837,863]" pageId="18" pageNumber="339" rank="species">
<emphasis id="B9729565FFF1FF9730B2AD44FCA0532B" bold="true" box="[697,850,837,863]" pageId="18" pageNumber="339">gen. et sp. n.</emphasis>
</taxonomicNameLabel>
differs from the others in the absence of the dorsal spine, shape of the
<emphasis id="B9729565FFF1FF97337BAD68FDCB53F7" box="[368,569,873,899]" italics="true" pageId="18" pageNumber="339">fovea amphidialis</emphasis>
(
<taxonomicName id="4C0632F4FFF1FF973041AD68FD1853F7" authority="Larrazábal-Filho &amp; Neres &amp; Silva &amp; Esteves, 2019" authorityName="Larrazábal-Filho &amp; Neres &amp; Silva &amp; Esteves" authorityYear="2019" box="[586,746,873,899]" class="Adenophorea" family="Desmodoridae" genus="Spinonema" kingdom="Animalia" order="Desmodorida" pageId="18" pageNumber="339" phylum="Nematoda" rank="species" species="cuticulatum" status="gen. et sp. n.">
<emphasis id="B9729565FFF1FF973041AD68FD1853F7" box="[586,746,873,899]" italics="true" pageId="18" pageNumber="339">S. cuticulatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A241281EFFF1FF9730F9AD68FC7553F7" box="[754,903,873,899]" pageId="18" pageNumber="339" rank="species">
<emphasis id="B9729565FFF1FF9730F9AD68FC7553F7" bold="true" box="[754,903,873,899]" pageId="18" pageNumber="339">gen. et sp. n.</emphasis>
</taxonomicNameLabel>
spiral and
<taxonomicName id="4C0632F4FFF1FF97360FAD68FB9D53F7" authority="Larrazábal-Filho &amp; Neres &amp; Silva &amp; Esteves, 2019" authorityName="Larrazábal-Filho &amp; Neres &amp; Silva &amp; Esteves" authorityYear="2019" box="[1028,1135,873,899]" class="Adenophorea" family="Desmodoridae" genus="Spinonema" kingdom="Animalia" order="Desmodorida" pageId="18" pageNumber="339" phylum="Nematoda" rank="species" species="spirale" status="gen. et sp. n.">
<emphasis id="B9729565FFF1FF97360FAD68FB9D53F7" box="[1028,1135,873,899]" italics="true" pageId="18" pageNumber="339">S. spirale</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A241281EFFF1FF97367DAD68FAFE53F7" box="[1142,1292,873,899]" pageId="18" pageNumber="339" rank="species">
<emphasis id="B9729565FFF1FF97367DAD68FAFE53F7" bold="true" box="[1142,1292,873,899]" pageId="18" pageNumber="339">gen. et sp. n.</emphasis>
</taxonomicNameLabel>
multispiral), rounded capitulum (
<taxonomicName id="4C0632F4FFF1FF973376AD8CFDEC53D3" authority="Larrazábal-Filho &amp; Neres &amp; Silva &amp; Esteves, 2019" authorityName="Larrazábal-Filho &amp; Neres &amp; Silva &amp; Esteves" authorityYear="2019" box="[381,542,909,935]" class="Adenophorea" family="Desmodoridae" genus="Spinonema" kingdom="Animalia" order="Desmodorida" pageId="18" pageNumber="339" phylum="Nematoda" rank="species" species="cuticulatum" status="gen. et sp. n.">
<emphasis id="B9729565FFF1FF973376AD8CFDEC53D3" box="[381,542,909,935]" italics="true" pageId="18" pageNumber="339">S. cuticulatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A241281EFFF1FF97302CAD8CFD3253D3" box="[551,704,909,935]" pageId="18" pageNumber="339" rank="species">
<emphasis id="B9729565FFF1FF97302CAD8CFD3253D3" bold="true" box="[551,704,909,935]" pageId="18" pageNumber="339">gen. et sp. n.</emphasis>
</taxonomicNameLabel>
hook-shaped), gubernaculum without apophysis (
<taxonomicName id="4C0632F4FFF1FF9736F3AD8CFA9153D3" authority="Larrazábal-Filho &amp; Neres &amp; Silva &amp; Esteves, 2019" authorityName="Larrazábal-Filho &amp; Neres &amp; Silva &amp; Esteves" authorityYear="2019" box="[1272,1379,909,935]" class="Adenophorea" family="Desmodoridae" genus="Spinonema" kingdom="Animalia" order="Desmodorida" pageId="18" pageNumber="339" phylum="Nematoda" rank="species" species="spirale" status="gen. et sp. n.">
<emphasis id="B9729565FFF1FF9736F3AD8CFA9153D3" box="[1272,1379,909,935]" italics="true" pageId="18" pageNumber="339">S. spirale</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A241281EFFF1FF973760AD8CFF0553BF" pageId="18" pageNumber="339" rank="species">
<emphasis id="B9729565FFF1FF973760AD8CFF0553BF" bold="true" pageId="18" pageNumber="339">gen. et sp. n.</emphasis>
</taxonomicNameLabel>
gubernaculum intricate with lateral
<emphasis id="B9729565FFF1FF97309CADB3FD1053BF" box="[663,738,946,971]" italics="true" pageId="18" pageNumber="339">crurae</emphasis>
and dorsal apophysis) and presence of three precloacal setae (absent in
<taxonomicName id="4C0632F4FFF1FF973307ADD4FE84539B" authority="Larrazábal-Filho &amp; Neres &amp; Silva &amp; Esteves, 2019" authorityName="Larrazábal-Filho &amp; Neres &amp; Silva &amp; Esteves" authorityYear="2019" box="[268,374,981,1007]" class="Adenophorea" family="Desmodoridae" genus="Spinonema" kingdom="Animalia" order="Desmodorida" pageId="18" pageNumber="339" phylum="Nematoda" rank="species" species="spirale" status="gen. et sp. n.">
<emphasis id="B9729565FFF1FF973307ADD4FE84539B" box="[268,374,981,1007]" italics="true" pageId="18" pageNumber="339">S. spirale</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A241281EFFF1FF973376ADD4FDE3539B" box="[381,529,981,1007]" pageId="18" pageNumber="339" rank="species">
<emphasis id="B9729565FFF1FF973376ADD4FDE3539B" bold="true" box="[381,529,981,1007]" pageId="18" pageNumber="339">gen. et sp. n.</emphasis>
</taxonomicNameLabel>
and one in
<taxonomicName id="4C0632F4FFF1FF973098ADD4FCC1539B" authority="Larrazábal-Filho &amp; Neres &amp; Silva &amp; Esteves, 2019" authorityName="Larrazábal-Filho &amp; Neres &amp; Silva &amp; Esteves" authorityYear="2019" box="[659,819,981,1007]" class="Adenophorea" family="Desmodoridae" genus="Spinonema" kingdom="Animalia" order="Desmodorida" pageId="18" pageNumber="339" phylum="Nematoda" rank="species" species="cuticulatum" status="gen. et sp. n.">
<emphasis id="B9729565FFF1FF973098ADD4FCC1539B" box="[659,819,981,1007]" italics="true" pageId="18" pageNumber="339">S. cuticulatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A241281EFFF1FF973131ADD4FC3C539B" box="[826,974,981,1007]" pageId="18" pageNumber="339" rank="species">
<emphasis id="B9729565FFF1FF973131ADD4FC3C539B" bold="true" box="[826,974,981,1007]" pageId="18" pageNumber="339">gen. et sp. n.</emphasis>
</taxonomicNameLabel>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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