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<mods:title id="9971A0F303D439553FBD9BEC6A11AD69">An overview of the extant genera and subgenera of the order Scolopendromorpha (Chilopoda): a new identification key and updated diagnoses</mods:title>
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<mods:namePart id="C1B21DC40A16F1A50955F8C86C0EC952">Schileyko, Arkady A.</mods:namePart>
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<mods:namePart id="67A77C0094EDDBFB37C16862036746F0">Vahtera, Varpu</mods:namePart>
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<mods:namePart id="20D5BBCAFAF64F3C44B2FF36E9B23508">Edgecombe, Gregory D.</mods:namePart>
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<treatment id="03DE092DFFFED707FF13FE8F2992DD5E" ID-DOI="http://doi.org/10.5281/zenodo.4455385" ID-GBIF-Taxon="177020703" ID-Zenodo-Dep="4455385" LSID="urn:lsid:plazi:treatment:03DE092DFFFED707FF13FE8F2992DD5E" httpUri="http://treatment.plazi.org/id/03DE092DFFFED707FF13FE8F2992DD5E" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
<subSubSection id="C36DEBB0FFFED706FF13FE8F2CA9DD97" box="[151,492,366,392]" pageId="12" pageNumber="13" type="nomenclature">
<paragraph id="8BC8B83BFFFED706FF13FE8F2CA9DD97" blockId="12.[151,492,366,392]" box="[151,492,366,392]" pageId="12" pageNumber="13">
<emphasis id="B9036429FFFED706FF13FE8F2CA9DD97" bold="true" box="[151,492,366,392]" pageId="12" pageNumber="13">
(!)
<taxonomicName id="4C77C3B8FFFED706FF38FE8F2CA9DD97" ID-CoL="63F3K" ID-ENA="863624" authority="Gervais, 1847" authorityName="Gervais" authorityYear="1847" box="[188,492,366,392]" class="Chilopoda" family="Scolopocryptopidae" genus="Newportia" kingdom="Animalia" order="Scolopendromorpha" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B9036429FFFED706FF38FE8F2C7CDD97" bold="true" box="[188,313,366,392]" italics="true" pageId="12" pageNumber="13">Newportia</emphasis>
Gervais, 1847
</taxonomicName>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C36DEBB0FFFED706FF13FE572C99DDCF" box="[151,476,438,464]" pageId="12" pageNumber="13" type="diagnosis">
<paragraph id="8BC8B83BFFFED706FF13FE572C99DDCF" blockId="12.[151,1437,438,2013]" box="[151,476,438,464]" pageId="12" pageNumber="13">
<emphasis id="B9036429FFFED706FF13FE572C57DDCF" bold="true" box="[151,274,438,464]" pageId="12" pageNumber="13">Diagnosis.</emphasis>
As for subfamily.
</paragraph>
</subSubSection>
<subSubSection id="C36DEBB0FFFED707FF43FE3B2992DD5E" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13" type="discussion">
<paragraph id="8BC8B83BFFFED706FF43FE3B2EFDDDEA" blockId="12.[151,1437,438,2013]" box="[199,952,474,501]" pageId="12" pageNumber="13">
<emphasis id="B9036429FFFED706FF43FE3B2CECDDEB" bold="true" box="[199,425,474,500]" pageId="12" pageNumber="13">Number of species.</emphasis>
50 species, 57 taxa of species rank (new data).
</paragraph>
<subSubSection id="C36DEBB0FFFED707FF43FE1F2992DD5E" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13" type="discussion">
<paragraph id="8BC8B83BFFFED706FF43FE1F2990DE9B" blockId="12.[151,1437,438,2013]" pageId="12" pageNumber="13">
<emphasis id="B9036429FFFED706FF43FE1F2C7EDE07" bold="true" box="[199,315,510,536]" pageId="12" pageNumber="13">Remarks.</emphasis>
<taxonomicName id="4C77C3B8FFFED706FEC0FE1E2CFFDE07" authorityName="Gervais" authorityYear="1847" box="[324,442,511,536]" class="Chilopoda" family="Scolopocryptopidae" genus="Newportia" kingdom="Animalia" order="Scolopendromorpha" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B9036429FFFED706FEC0FE1E2CFFDE07" box="[324,442,511,536]" italics="true" pageId="12" pageNumber="13">Newportia</emphasis>
</taxonomicName>
(in various senses) treated as a genus in
<bibRefCitation id="EFE6C5CAFFFED706FC16FE1F2858DE06" author="Edgecombe, G. D. &amp; Bonato, L." box="[914,1309,510,537]" pageId="12" pageNumber="13" refId="ref47498" refString="Edgecombe, G. D. &amp; Bonato, L. (2011) Chilopoda-taxonomic overview. Order Scolopendromorpha. In: Minelli, A. (Ed.), Treatise on Zoology-Anatomy, Taxonomy, Biology. The Myriapoda. Vol. 1. Brill, Leiden, pp. 392 - 407." type="book" year="2011">Edgecombe &amp; Bonato (2011: 405)</bibRefCitation>
,
<bibRefCitation id="EFE6C5CAFFFED706FAAEFE1F2C72DE23" author="Vahtera, V. &amp; Edgecombe, G. D. &amp; Giribet, G." pageId="12" pageNumber="13" pagination="4 - 20" refId="ref51271" refString="Vahtera, V., Edgecombe, G. D. &amp; Giribet, G. (2012 a) Evolution of blindness in scolopendromorph centipedes (Chilopoda: Scolopendromorpha): insight from an expanded sampling of molecular data. Cladistics, 28, 4 - 20. https: // doi. org / 10.1111 / j. 1096 - 0031.2011.00361. x" type="journal article" year="2012">
Vahtera
<emphasis id="B9036429FFFED706FA0CFE1E2DF1DE23" italics="true" pageId="12" pageNumber="13">et al.</emphasis>
(2012a: 12
</bibRefCitation>
,
<bibRefCitation id="EFE6C5CAFFFED706FEC7FDC32CFCDE22" author="Vahtera, V. &amp; Edgecombe, G. D. &amp; Giribet, G." box="[323,441,546,573]" pageId="12" pageNumber="13" pagination="578 - 602" refId="ref51397" refString="Vahtera, V., Edgecombe, G. D. &amp; Giribet, G. (2013). Phylogenetics of scolopendromorph centipedes: can denser taxon sampling improve an artificial classification? Invertebrate Systematics, 27 (5), 578 - 602. https: // doi. org / 10.1071 / IS 13035" type="journal article" year="2013">2013: 579</bibRefCitation>
), Schileyko (2013: 40, 2014: 159, 2018: 61), Edgecombe
<emphasis id="B9036429FFFED706FBC2FDC2293DDE23" box="[1094,1144,546,572]" italics="true" pageId="12" pageNumber="13">et al</emphasis>
. (2015: 65),
<bibRefCitation id="EFE6C5CAFFFED706FA8FFDC22DA1DE7E" author="Bonato, L. &amp; Chagas-Junior, A. &amp; Edgecombe, G. D. &amp; Lewis, J. G. E. &amp; Minelli, A. &amp; Pereira, L. A. &amp; Shelley, R. M. &amp; Stoev, P. &amp; Zapparoli, M." pageId="12" pageNumber="13" refId="ref45701" refString="Bonato, L., Chagas-Junior, A., Edgecombe, G. D., Lewis, J. G. E., Minelli, A., Pereira, L. A., Shelley, R. M., Stoev, P. &amp; Zapparoli, M. (2016) ChiloBase 2.0 - A World Catalogue of Centipedes (Chilopoda). Available from: http: // chilobase. biologia. unipd. it (accessed 2 August 2016)" type="url" year="2016">
Bonato
<emphasis id="B9036429FFFED706FAE0FDC228D8DE23" box="[1380,1437,546,572]" italics="true" pageId="12" pageNumber="13">et al.</emphasis>
(2016)
</bibRefCitation>
,
<bibRefCitation id="EFE6C5CAFFFED706FF6BFDA72FF4DE7F" author="Martinez-Munoz, C. A. &amp; Tcherva, T." box="[239,689,582,608]" pageId="12" pageNumber="13" pagination="179 - 184" refId="ref49516" refString="Martinez-Munoz, C. A. &amp; Tcherva, T. (2017) Primer registro de Newportia stolli (Pocock, 1896) (Chilopoda: Scolopendromorpha: Scolopocryptopidae) para Cuba y las Antillas. Boletin de la Sociedad Entomologica Aragonesa, 60, 179 - 184." type="journal article" year="2017">Martínez-Muñoz &amp; Tcherva (2017: 179)</bibRefCitation>
and Chagas-Jr (2018: 154). The most recent morphologic accounts on
<taxonomicName id="4C77C3B8FFFED706FF3EFD8A2C75DE9B" authorityName="Gervais" authorityYear="1847" box="[186,304,619,644]" class="Chilopoda" family="Scolopocryptopidae" genus="Newportia" kingdom="Animalia" order="Scolopendromorpha" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B9036429FFFED706FF3EFD8A2C75DE9B" box="[186,304,619,644]" italics="true" pageId="12" pageNumber="13">Newportia</emphasis>
</taxonomicName>
(in the old sense) are those of Schileyko (2013, 2014, 2018) and Chagas-Jr (2018).
</paragraph>
<paragraph id="8BC8B83BFFFED706FF43FD6F2E89D8DB" blockId="12.[151,1437,438,2013]" pageId="12" pageNumber="13">
There is a longstanding question on subgeneric division of this genus. Reviewing
<taxonomicName id="4C77C3B8FFFED706FBE7FD6E299CDEB7" authorityName="Gervais" authorityYear="1847" box="[1123,1241,655,680]" class="Chilopoda" family="Scolopocryptopidae" genus="Newportia" kingdom="Animalia" order="Scolopendromorpha" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B9036429FFFED706FBE7FD6E299CDEB7" box="[1123,1241,655,680]" italics="true" pageId="12" pageNumber="13">Newportia</emphasis>
</taxonomicName>
(in the long-held sense) solely on the basis of morphology,
<bibRefCitation id="EFE6C5CAFFFED706FDEFFD532EA8DED2" author="Schileyko, A. &amp; Minelli, A." box="[619,1005,690,717]" pageId="12" pageNumber="13" pagination="265 - 299" refId="ref50254" refString="Schileyko, A. &amp; Minelli, A. (1999) On the genus Newportia Gervais, 1847 (Chilopoda: Scolopendromorpha: Newportiidae). Arthropoda Selecta, 7 (4), 265 - 299." type="journal article" year="1999">Schileyko and Minelli (1999: 267)</bibRefCitation>
regarded as problematic “any identification of natural subgroups within this large genus … although, based on the structure of the terminal [= ultimate] tarsus, we practically recognize two groups of species …
<taxonomicName id="4C77C3B8FFFED706FCADFD1A2EDADF0B" authorityName="Gervais" authorityYear="1847" box="[809,927,763,788]" class="Chilopoda" family="Scolopocryptopidae" genus="Newportia" kingdom="Animalia" order="Scolopendromorpha" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B9036429FFFED706FCADFD1A2EDADF0B" box="[809,927,763,788]" italics="true" pageId="12" pageNumber="13">Newportia</emphasis>
</taxonomicName>
[(
<taxonomicName id="4C77C3B8FFFED706FC3EFD1A2975DF0B" authorityName="Gervais" authorityYear="1847" box="[954,1072,763,788]" class="Chilopoda" family="Scolopocryptopidae" genus="Newportia" kingdom="Animalia" order="Scolopendromorpha" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B9036429FFFED706FC3EFD1A2975DF0B" box="[954,1072,763,788]" italics="true" pageId="12" pageNumber="13">Newportia</emphasis>
</taxonomicName>
)] and
<taxonomicName id="4C77C3B8FFFED706FBF8FD1B286CDF0B" authorityName="Saussure" authorityYear="1858" box="[1148,1321,762,788]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopendrides" kingdom="Animalia" order="Scolopendromorpha" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B9036429FFFED706FBF8FD1B286CDF0B" box="[1148,1321,762,788]" italics="true" pageId="12" pageNumber="13">Scolopendrides</emphasis>
</taxonomicName>
”. In 2012 Edgecombe
<emphasis id="B9036429FFFED706FEA7FCFE2C11DF27" box="[291,340,798,824]" italics="true" pageId="12" pageNumber="13">et al</emphasis>
. wrote (p. 779): “The molecular data resolve a clade within
<taxonomicName id="4C77C3B8FFFED706FC7DFCFE292ADF27" authorityName="Gervais" authorityYear="1847" box="[1017,1135,799,824]" class="Chilopoda" family="Scolopocryptopidae" genus="Newportia" kingdom="Animalia" order="Scolopendromorpha" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B9036429FFFED706FC7DFCFE292ADF27" box="[1017,1135,799,824]" italics="true" pageId="12" pageNumber="13">Newportia</emphasis>
</taxonomicName>
that was ambiguous based on morphology alone, one composed of
<taxonomicName id="4C77C3B8FFFED706FDDEFCA22E06DF43" authorityName="Chamberlin" authorityYear="1922" box="[602,835,834,860]" class="Chilopoda" family="Scolopocryptopidae" genus="Newportia" kingdom="Animalia" order="Scolopendromorpha" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="divergens">
<emphasis id="B9036429FFFED706FDDEFCA22E06DF43" box="[602,835,834,860]" italics="true" pageId="12" pageNumber="13">Newportia divergens</emphasis>
</taxonomicName>
,
<taxonomicName id="4C77C3B8FFFED706FCD5FCA2294BDF43" authorityName="Pocock" authorityYear="1891" box="[849,1038,835,860]" class="Chilopoda" family="Scolopocryptopidae" genus="Newportia" kingdom="Animalia" order="Scolopendromorpha" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="ernsti">
<emphasis id="B9036429FFFED706FCD5FCA2294BDF43" box="[849,1038,835,860]" italics="true" pageId="12" pageNumber="13">Newportia ernsti</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C77C3B8FFFED706FBCFFCA22845DF43" baseAuthorityName="Pocock" baseAuthorityYear="1896" box="[1099,1280,834,860]" class="Chilopoda" family="Scolopocryptopidae" genus="Newportia" kingdom="Animalia" order="Scolopendromorpha" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="stolli">
<emphasis id="B9036429FFFED706FBCFFCA22845DF43" box="[1099,1280,834,860]" italics="true" pageId="12" pageNumber="13">Newportia stolli</emphasis>
</taxonomicName>
... This group corresponds to
<taxonomicName id="4C77C3B8FFFED706FEC7FC872FE1DF9E" authority="Saussure, 1858" authorityName="Saussure" authorityYear="1858" box="[323,676,870,897]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopendrides" kingdom="Animalia" order="Scolopendromorpha" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B9036429FFFED706FEC7FC872CB5DF9F" box="[323,496,870,896]" italics="true" pageId="12" pageNumber="13">Scolopendrides</emphasis>
Saussure, 1858
</taxonomicName>
, formerly employed as a subgenus of
<taxonomicName id="4C77C3B8FFFED706FBCAFC862981DF9F" authorityName="Gervais" authorityYear="1847" box="[1102,1220,871,896]" class="Chilopoda" family="Scolopocryptopidae" genus="Newportia" kingdom="Animalia" order="Scolopendromorpha" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B9036429FFFED706FBCAFC862981DF9F" box="[1102,1220,871,896]" italics="true" pageId="12" pageNumber="13">Newportia</emphasis>
</taxonomicName>
… A possible apomorphy for this group is the traditional defining character of
<taxonomicName id="4C77C3B8FFFED706FCE0FC6B2954DFBB" authorityName="Saussure" authorityYear="1858" box="[868,1041,906,932]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopendrides" kingdom="Animalia" order="Scolopendromorpha" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B9036429FFFED706FCE0FC6B2954DFBB" box="[868,1041,906,932]" italics="true" pageId="12" pageNumber="13">Scolopendrides</emphasis>
</taxonomicName>
, irregular boundaries between the tarsomeres of tarsus 2 of the ultimate leg”. Those data were supported by
<bibRefCitation id="EFE6C5CAFFFED706FC57FC4F29B6DFD6" author="Vahtera, V. &amp; Edgecombe, G. D. &amp; Giribet, G." box="[979,1267,942,969]" pageId="12" pageNumber="13" pagination="578 - 602" refId="ref51397" refString="Vahtera, V., Edgecombe, G. D. &amp; Giribet, G. (2013). Phylogenetics of scolopendromorph centipedes: can denser taxon sampling improve an artificial classification? Invertebrate Systematics, 27 (5), 578 - 602. https: // doi. org / 10.1071 / IS 13035" type="journal article" year="2013">
Vahtera
<emphasis id="B9036429FFFED706FBB4FC4E292CDFD7" box="[1072,1129,942,968]" italics="true" pageId="12" pageNumber="13">et al.</emphasis>
(2013: 591)
</bibRefCitation>
who, based on combined analysis, stated that “One of the best-supported clades within
<taxonomicName id="4C77C3B8FFFED706FC56FC32290DDFF3" authorityName="Gervais" authorityYear="1847" box="[978,1096,979,1004]" class="Chilopoda" family="Scolopocryptopidae" genus="Newportia" kingdom="Animalia" order="Scolopendromorpha" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B9036429FFFED706FC56FC32290DDFF3" box="[978,1096,979,1004]" italics="true" pageId="12" pageNumber="13">Newportia</emphasis>
</taxonomicName>
is a grouping of species with irregular boundaries between tarsomeres on the ultimate leg” (p. 591) and that “A reclassification of
<taxonomicName id="4C77C3B8FFFED706FA87FC16283CD80F" authorityName="Gervais" authorityYear="1847" box="[1283,1401,1015,1040]" class="Chilopoda" family="Scolopocryptopidae" genus="Newportia" kingdom="Animalia" order="Scolopendromorpha" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B9036429FFFED706FA87FC16283CD80F" box="[1283,1401,1015,1040]" italics="true" pageId="12" pageNumber="13">Newportia</emphasis>
</taxonomicName>
… might also reconsider the utility of
<taxonomicName id="4C77C3B8FFFED706FDA9FBFB2F9FD82B" authorityName="Saussure" authorityYear="1858" box="[557,730,1050,1076]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopendrides" kingdom="Animalia" order="Scolopendromorpha" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B9036429FFFED706FDA9FBFB2F9FD82B" box="[557,730,1050,1076]" italics="true" pageId="12" pageNumber="13">Scolopendrides</emphasis>
</taxonomicName>
because the group is unambiguously monophyletic” (p. 589). The nomenclatural validity of
<taxonomicName id="4C77C3B8FFFED706FE6AFBDF2E43D847" authority="Saussure" authorityName="Saussure" authorityYear="1858" box="[494,774,1086,1112]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopendrides" kingdom="Animalia" order="Scolopendromorpha" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B9036429FFFED706FE6AFBDF2FDED847" box="[494,667,1086,1112]" italics="true" pageId="12" pageNumber="13">Scolopendrides</emphasis>
Saussure
</taxonomicName>
is disputed in a forthcoming work by C. Martínez-Muñoz. Pending publication of that work, we desist from using this name but recognize the utility of taxonomically separating a group of species with the secondary articles of ultimate tarsus 2 definitely separated (
<figureCitation id="134CA4BEFFFED706FB09FB67299BD8BF" box="[1165,1246,1158,1184]" captionStart="FIGURES 1419" captionStartId="15.[151,264,1706,1730]" captionTargetBox="[203,1386,199,1663]" captionTargetId="figure-17@15.[187,1401,181,1682]" captionTargetPageId="15" captionText="FIGURES 1419. Ectonocryptops kraepelini Crabill, 1977; Holotype 14 figures 5, 6 of Shelley &amp; Mercurio (2008): ultimate LBS and left ultimate leg (dorsally) + left ultimate leg (ventrally); Ectonocryptoides sandrops Schileyko, 2006; Holotype Rc 7197 15 LBS 23 + left ultimate leg dorsally; Ectonocryptoides quadrimeropus Shelley &amp; Mercurio, 2005; AMNH, Puebla specimen of Shelley (2009) 16 Apical part of maxilla 2 laterally; Ectonocryptoides sandrops Schileyko, 2006; Holotype Rc 7197 17 Head + forcipular segment ventrally 18 General view dorso-laterally; Newportia monticola Pocock, 1890; Ad. N 7078 19a left side of ultimate LBS + ultimate legs dorso-laterally; Newportia monticola Pocock, 1890 19b right ultimate leg ventrally (schematic); (am)—anterior margin of forcipular coxosternite, (cp)—coxopleural process, (dp)—distomedial uncinate process of tibia, (p2)—pretarsus of maxilla 2, (pf)—prefemur, (ti)—tibia, (t1)—tarsus 1, (t2g)—reduced globose tarsus 2, (t2r)—tarsus 2 with regularly divided secondary articles, (ut)—ultimate tergite, (vsp)—ventral spinous process." figureDoi="http://doi.org/10.5281/zenodo.4402153" httpUri="https://zenodo.org/record/4402153/files/figure.png" pageId="12" pageNumber="13">Fig. 19</figureCitation>
) as the nominate subgenus and a group with irregularly divided ultimate tarsus 2 (
<figureCitation id="134CA4BEFFFED706FCEFFB4B2EFAD8DB" box="[875,959,1194,1220]" captionStart="FIGURES 2025" captionStartId="18.[151,264,1745,1769]" captionTargetBox="[264,1325,199,1702]" captionTargetId="figure-19@18.[243,1344,181,1721]" captionTargetPageId="18" captionText="FIGURES 2025. Newportia stolli (Pocock, 1896); Ad. Rc 7235 (?) 20a ultimate LBS + ultimate legs dorsally; Newportia divergens Chamberlin, 1922 20b right ultimate leg (schematic); Newportia longitarsis virginensis Lewis, 1989; Ad. Rc 7470 21 head and forcipular segment ventrally; Newportia (Tidops) collaris (Kraepelin, 1903); MCZ 32933 22 forcipular segment ventrally 23 anterior half of head ventrally 24 apical part of maxilla 2 laterally 25 right pleuron of mid body LBS laterally; (am)—anterior margin of forcipular coxosternite, (cl)—clypeus, (cst)— forcipular coxosternite, (f)—femur, (l)—labrum, (p2)—pectinate pretarsus of maxilla 2, (pa)—blunt projection of anterior margin of forcipular coxosternite, (pf)—prefemur, (s)—open spiracle with atrium, (t1)—tarsus 1, (t2i)—tarsus 2 with irregularly divided secondary articles, (th)—trichoid sensillae, (ti)—tibia, (tr)—tarsungula, (trp)—forcipular trochantero-prefemur, (ut)—ultimate tergite." figureDoi="http://doi.org/10.5281/zenodo.4402155" httpUri="https://zenodo.org/record/4402155/files/figure.png" pageId="12" pageNumber="13">Fig. 20</figureCitation>
).
</paragraph>
<paragraph id="8BC8B83BFFFED706FF43FB2F2923D92F" blockId="12.[151,1437,438,2013]" pageId="12" pageNumber="13">
<bibRefCitation id="EFE6C5CAFFFED706FF43FB2F2CF6D8F7" author="Vahtera, V. &amp; Edgecombe, G. D. &amp; Giribet, G." box="[199,435,1230,1256]" pageId="12" pageNumber="13" pagination="578 - 602" refId="ref51397" refString="Vahtera, V., Edgecombe, G. D. &amp; Giribet, G. (2013). Phylogenetics of scolopendromorph centipedes: can denser taxon sampling improve an artificial classification? Invertebrate Systematics, 27 (5), 578 - 602. https: // doi. org / 10.1071 / IS 13035" type="journal article" year="2013">
Vahtera
<emphasis id="B9036429FFFED706FEA2FB2E2C25D8F7" box="[294,352,1230,1256]" italics="true" pageId="12" pageNumber="13">et al.</emphasis>
(2013)
</bibRefCitation>
also reclassified the genera
<taxonomicName id="4C77C3B8FFFED706FD7DFB2F2E01D8F7" authorityName="Chamberlin" authorityYear="1915" box="[761,836,1230,1256]" class="Chilopoda" family="Scolopocryptopidae" genus="Tidops" kingdom="Animalia" order="Scolopendromorpha" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B9036429FFFED706FD7DFB2F2E01D8F7" box="[761,836,1230,1256]" italics="true" pageId="12" pageNumber="13">Tidops</emphasis>
</taxonomicName>
,
<taxonomicName id="4C77C3B8FFFED706FCD0FB2E2941D8F7" authorityName="Crabill" authorityYear="1977" box="[852,1028,1231,1256]" class="Chilopoda" family="Scolopocryptopidae" genus="Ectonocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B9036429FFFED706FCD0FB2E2941D8F7" box="[852,1028,1231,1256]" italics="true" pageId="12" pageNumber="13">Ectonocryptops</emphasis>
</taxonomicName>
and
<taxonomicName id="4C77C3B8FFFED706FBBAFB2F2847D8F7" authorityName="Shelley &amp; Mercurio" authorityYear="2005" box="[1086,1282,1230,1256]" class="Chilopoda" family="Scolopocryptopidae" genus="Ectonocryptoides" kingdom="Animalia" order="Scolopendromorpha" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B9036429FFFED706FBBAFB2F2847D8F7" box="[1086,1282,1230,1256]" italics="true" pageId="12" pageNumber="13">Ectonocryptoides</emphasis>
</taxonomicName>
as subgenera of
<taxonomicName id="4C77C3B8FFFED706FF33FB122C68D913" authorityName="Gervais" authorityYear="1847" box="[183,301,1267,1292]" class="Chilopoda" family="Scolopocryptopidae" genus="Newportia" kingdom="Animalia" order="Scolopendromorpha" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B9036429FFFED706FF33FB122C68D913" box="[183,301,1267,1292]" italics="true" pageId="12" pageNumber="13">Newportia</emphasis>
</taxonomicName>
(see below). Taking into consideration the re-validation of the subgenus
<taxonomicName id="4C77C3B8FFFED706FBF5FB1329BAD913" authorityName="Chamberlin" authorityYear="1912" box="[1137,1279,1266,1292]" class="Chilopoda" family="Scolopocryptopidae" genus="Newportia" kingdom="Animalia" order="Scolopendromorpha" pageId="12" pageNumber="13" phylum="Arthropoda" rank="subGenus" subGenus="Newportides">
<emphasis id="B9036429FFFED706FBF5FB1329BAD913" box="[1137,1279,1266,1292]" italics="true" pageId="12" pageNumber="13">Newportides</emphasis>
</taxonomicName>
by Chagas-Jr (2018) (see below), the genus
<taxonomicName id="4C77C3B8FFFED706FE6EFAF62F25D92F" authorityName="Gervais" authorityYear="1847" box="[490,608,1303,1328]" class="Chilopoda" family="Scolopocryptopidae" genus="Newportia" kingdom="Animalia" order="Scolopendromorpha" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B9036429FFFED706FE6EFAF62F25D92F" box="[490,608,1303,1328]" italics="true" pageId="12" pageNumber="13">Newportia</emphasis>
</taxonomicName>
should at the moment include five subgenera.
</paragraph>
<paragraph id="8BC8B83BFFFED706FF43FADB2FBDDBC2" blockId="12.[151,1437,438,2013]" pageId="12" pageNumber="13">
There is also much confusion concerning the exact number of
<taxonomicName id="4C77C3B8FFFED706FCFDFADA2EAAD94B" authorityName="Gervais" authorityYear="1847" box="[889,1007,1339,1364]" class="Chilopoda" family="Scolopocryptopidae" genus="Newportia" kingdom="Animalia" order="Scolopendromorpha" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B9036429FFFED706FCFDFADA2EAAD94B" box="[889,1007,1339,1364]" italics="true" pageId="12" pageNumber="13">Newportia</emphasis>
</taxonomicName>
species. According to
<bibRefCitation id="EFE6C5CAFFFED706FB6DFADB2C31D966" author="Edgecombe, G. D. &amp; Bonato, L." pageId="12" pageNumber="13" refId="ref47498" refString="Edgecombe, G. D. &amp; Bonato, L. (2011) Chilopoda-taxonomic overview. Order Scolopendromorpha. In: Minelli, A. (Ed.), Treatise on Zoology-Anatomy, Taxonomy, Biology. The Myriapoda. Vol. 1. Brill, Leiden, pp. 392 - 407." type="book" year="2011">Edgecombe and Bonato (2011: 405)</bibRefCitation>
the genus includes “More than 50 species”, the most recent key of Schileyko (2013) contains 45 taxa of species rank,
<bibRefCitation id="EFE6C5CAFFFED706FE2BFA632F93D982" author="Vahtera, V. &amp; Edgecombe, G. D. &amp; Giribet, G." box="[431,726,1410,1437]" pageId="12" pageNumber="13" pagination="578 - 602" refId="ref51397" refString="Vahtera, V., Edgecombe, G. D. &amp; Giribet, G. (2013). Phylogenetics of scolopendromorph centipedes: can denser taxon sampling improve an artificial classification? Invertebrate Systematics, 27 (5), 578 - 602. https: // doi. org / 10.1071 / IS 13035" type="journal article" year="2013">
Vahtera
<emphasis id="B9036429FFFED706FD89FA622F0DD983" box="[525,584,1410,1436]" italics="true" pageId="12" pageNumber="13">et al.</emphasis>
(2013: 589)
</bibRefCitation>
mentioned “50+ species”, Edgecombe
<emphasis id="B9036429FFFED706FB1DFA622989D983" box="[1177,1228,1410,1436]" italics="true" pageId="12" pageNumber="13">et al</emphasis>
. (2015: 65) wrote about “some 60 nominal species or subspecies”,
<bibRefCitation id="EFE6C5CAFFFED706FD3EFA462ED9D9DE" author="Bonato, L. &amp; Chagas-Junior, A. &amp; Edgecombe, G. D. &amp; Lewis, J. G. E. &amp; Minelli, A. &amp; Pereira, L. A. &amp; Shelley, R. M. &amp; Stoev, P. &amp; Zapparoli, M." box="[698,924,1446,1473]" pageId="12" pageNumber="13" refId="ref45701" refString="Bonato, L., Chagas-Junior, A., Edgecombe, G. D., Lewis, J. G. E., Minelli, A., Pereira, L. A., Shelley, R. M., Stoev, P. &amp; Zapparoli, M. (2016) ChiloBase 2.0 - A World Catalogue of Centipedes (Chilopoda). Available from: http: // chilobase. biologia. unipd. it (accessed 2 August 2016)" type="url" year="2016">
Bonato
<emphasis id="B9036429FFFED706FC96FA462E0FD9DF" box="[786,842,1446,1472]" italics="true" pageId="12" pageNumber="13">et al.</emphasis>
(2016)
</bibRefCitation>
gave 57 species,
<bibRefCitation id="EFE6C5CAFFFED706FBE5FA472C5ED9FB" author="Martinez-Munoz, C. A. &amp; Tcherva, T." pageId="12" pageNumber="13" pagination="179 - 184" refId="ref49516" refString="Martinez-Munoz, C. A. &amp; Tcherva, T. (2017) Primer registro de Newportia stolli (Pocock, 1896) (Chilopoda: Scolopendromorpha: Scolopocryptopidae) para Cuba y las Antillas. Boletin de la Sociedad Entomologica Aragonesa, 60, 179 - 184." type="journal article" year="2017">Martínez-Muñoz &amp; Tcherva (2017: 179)</bibRefCitation>
gave 66 species and Chagas-Jr (2018: 154) wrote about “more than 70 taxa at species rank”. It should be noted, however, that the first two papers considered
<taxonomicName id="4C77C3B8FFFED706FD7CFA0E2E2BDA17" authorityName="Gervais" authorityYear="1847" box="[760,878,1519,1544]" class="Chilopoda" family="Scolopocryptopidae" genus="Newportia" kingdom="Animalia" order="Scolopendromorpha" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B9036429FFFED706FD7CFA0E2E2BDA17" box="[760,878,1519,1544]" italics="true" pageId="12" pageNumber="13">Newportia</emphasis>
</taxonomicName>
in the “old” sense (i.e. before
<bibRefCitation id="EFE6C5CAFFFED706FB38FA0F28D5DA17" author="Vahtera, V. &amp; Edgecombe, G. D. &amp; Giribet, G." box="[1212,1424,1518,1545]" pageId="12" pageNumber="13" pagination="578 - 602" refId="ref51397" refString="Vahtera, V., Edgecombe, G. D. &amp; Giribet, G. (2013). Phylogenetics of scolopendromorph centipedes: can denser taxon sampling improve an artificial classification? Invertebrate Systematics, 27 (5), 578 - 602. https: // doi. org / 10.1071 / IS 13035" type="journal article" year="2013">
Vahtera
<emphasis id="B9036429FFFED706FA93FA0E2802DA17" box="[1303,1351,1518,1544]" italics="true" pageId="12" pageNumber="13">et al</emphasis>
. 2013
</bibRefCitation>
), whereas in the rest of them this genus also accommodates
<taxonomicName id="4C77C3B8FFFED706FCB8F9F32EC2DA33" authorityName="Chamberlin" authorityYear="1915" box="[828,903,1554,1580]" class="Chilopoda" family="Scolopocryptopidae" genus="Tidops" kingdom="Animalia" order="Scolopendromorpha" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B9036429FFFED706FCB8F9F32EC2DA33" box="[828,903,1554,1580]" italics="true" pageId="12" pageNumber="13">Tidops</emphasis>
</taxonomicName>
,
<taxonomicName id="4C77C3B8FFFED706FC13F9F22902DA33" authorityName="Crabill" authorityYear="1977" box="[919,1095,1555,1580]" class="Chilopoda" family="Scolopocryptopidae" genus="Ectonocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B9036429FFFED706FC13F9F22902DA33" box="[919,1095,1555,1580]" italics="true" pageId="12" pageNumber="13">Ectonocryptops</emphasis>
</taxonomicName>
and
<taxonomicName id="4C77C3B8FFFED706FB06F9F32803DA33" authorityName="Shelley &amp; Mercurio" authorityYear="2005" box="[1154,1350,1554,1580]" class="Chilopoda" family="Scolopocryptopidae" genus="Ectonocryptoides" kingdom="Animalia" order="Scolopendromorpha" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B9036429FFFED706FB06F9F32803DA33" box="[1154,1350,1554,1580]" italics="true" pageId="12" pageNumber="13">Ectonocryptoides</emphasis>
</taxonomicName>
. Moreover, the authors mentioned above (except for Schileyko 2013 and Chagas-Jr 2018), included in
<taxonomicName id="4C77C3B8FFFED706FB5AF9D62811DA4F" authorityName="Gervais" authorityYear="1847" box="[1246,1364,1591,1616]" class="Chilopoda" family="Scolopocryptopidae" genus="Newportia" kingdom="Animalia" order="Scolopendromorpha" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B9036429FFFED706FB5AF9D62811DA4F" box="[1246,1364,1591,1616]" italics="true" pageId="12" pageNumber="13">Newportia</emphasis>
</taxonomicName>
all 17 “new species” that were poorly defined from
<collectingCountry id="F360F8ABFFFED706FD0AF9BB2E44DA6B" box="[654,769,1626,1652]" name="Venezuela" pageId="12" pageNumber="13">Venezuela</collectingCountry>
by González-Sponga (1997, 2000). The corresponding
<typeStatus id="54CC0699FFFED706FAE8F9BA28D9DA6A" box="[1388,1436,1627,1653]" pageId="12" pageNumber="13">type</typeStatus>
material was re-investigated by Chagas-Jr who informed A.S. (e-mail of 2014) that only three of those “species” are valid. We regard two of them (
<taxonomicName id="4C77C3B8FFFED706FDA5F9422FE0DAA3" authorityName="Gonzalez-Sponga" authorityYear="1998" box="[545,677,1698,1724]" class="Chilopoda" family="Scolopocryptopidae" genus="Newportia" kingdom="Animalia" order="Scolopendromorpha" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="avilensis">
<emphasis id="B9036429FFFED706FDA5F9422FE0DAA3" box="[545,677,1698,1724]" italics="true" pageId="12" pageNumber="13">N. avilensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C77C3B8FFFED706FD5AF9422E06DAA3" authorityName="Gonzalez-Sponga" authorityYear="1998" box="[734,835,1699,1724]" class="Chilopoda" family="Scolopocryptopidae" genus="Newportia" kingdom="Animalia" order="Scolopendromorpha" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="prima">
<emphasis id="B9036429FFFED706FD5AF9422E06DAA3" box="[734,835,1699,1724]" italics="true" pageId="12" pageNumber="13">N. prima</emphasis>
</taxonomicName>
, both of González-Sponga, 1997) as members of the species-group that has previously been referred to the former
<taxonomicName id="4C77C3B8FFFED706FCC5F9272EABDAFF" authorityName="Saussure" authorityYear="1858" box="[833,1006,1734,1760]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopendrides" kingdom="Animalia" order="Scolopendromorpha" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B9036429FFFED706FCC5F9272EABDAFF" box="[833,1006,1734,1760]" italics="true" pageId="12" pageNumber="13">Scolopendrides</emphasis>
</taxonomicName>
(=
<taxonomicName id="4C77C3B8FFFED706FB89F92629C6DAFF" authorityName="Gervais" authorityYear="1847" box="[1037,1155,1735,1760]" class="Chilopoda" family="Scolopocryptopidae" genus="Newportia" kingdom="Animalia" order="Scolopendromorpha" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B9036429FFFED706FB89F92629C6DAFF" box="[1037,1155,1735,1760]" italics="true" pageId="12" pageNumber="13">Newportia</emphasis>
</taxonomicName>
(
<taxonomicName id="4C77C3B8FFFED706FB16F926284DDAFF" authorityName="Gervais" authorityYear="1847" box="[1170,1288,1735,1760]" class="Chilopoda" family="Scolopocryptopidae" genus="Newportia" kingdom="Animalia" order="Scolopendromorpha" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B9036429FFFED706FB16F926284DDAFF" box="[1170,1288,1735,1760]" italics="true" pageId="12" pageNumber="13">Newportia</emphasis>
</taxonomicName>
)) and
<taxonomicName id="4C77C3B8FFFED706FAD4F9262F07DB1B" authority="Gonzalez-Sponga, 2000" authorityName="Gonzalez-Sponga" authorityYear="2000" class="Chilopoda" family="Scolopocryptopidae" genus="Newportia" kingdom="Animalia" order="Scolopendromorpha" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="cerrocopeyensis">
<emphasis id="B9036429FFFED706FAD4F9262C6DDB1B" italics="true" pageId="12" pageNumber="13">N. cerrocopeyensis</emphasis>
González-Sponga, 2000
</taxonomicName>
, as a member of
<taxonomicName id="4C77C3B8FFFED706FC82F90A294BDB1A" authority="(Newportia)" authorityName="Gervais" authorityYear="1847" baseAuthorityName="Newportia" box="[774,1038,1770,1797]" class="Chilopoda" family="Scolopocryptopidae" genus="Newportia" kingdom="Animalia" order="Scolopendromorpha" pageId="12" pageNumber="13" phylum="Arthropoda" rank="subGenus" subGenus="Newportia">
<emphasis id="B9036429FFFED706FC82F90A2E39DB1B" box="[774,892,1771,1796]" italics="true" pageId="12" pageNumber="13">Newportia</emphasis>
(
<emphasis id="B9036429FFFED706FC09F90A2946DB1B" box="[909,1027,1771,1796]" italics="true" pageId="12" pageNumber="13">Newportia</emphasis>
)
</taxonomicName>
. In 2018 Chagas-Jr also confirmed validity of
<taxonomicName id="4C77C3B8FFFED706FE91F8EE2FD4DB37" authority="Gonzalez-Sponga, 1997" authorityName="Gonzalez-Sponga" authorityYear="1997" box="[277,657,1806,1832]" class="Chilopoda" family="Scolopocryptopidae" genus="Newportia" kingdom="Animalia" order="Scolopendromorpha" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="pilosa">
<emphasis id="B9036429FFFED706FE91F8EE2C3EDB37" box="[277,379,1806,1832]" italics="true" pageId="12" pageNumber="13">N. pilosa</emphasis>
González-Sponga, 1997
</taxonomicName>
thus only 4 of those doubtful 17 “species” should currently be valid (the corresponding paper of Chagas-Jr is in preparation; personal communication). Chagas-Jr (2018: 167) also questioned the taxonomic validity of
<taxonomicName id="4C77C3B8FFFED706FE7BF8B62E7DDB6F" authority="Chamberlin, 1922" authorityName="Chamberlin" authorityYear="1922" box="[511,824,1878,1904]" class="Chilopoda" family="Scolopocryptopidae" genus="Newportia" kingdom="Animalia" order="Scolopendromorpha" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="sulana">
<emphasis id="B9036429FFFED706FE7BF8B62F2FDB6F" box="[511,618,1878,1904]" italics="true" pageId="12" pageNumber="13">N. sulana</emphasis>
<bibRefCitation id="EFE6C5CAFFFED706FDEBF8B72E7DDB6F" author="Chamberlin, R. V." box="[623,824,1878,1904]" pageId="12" pageNumber="13" pagination="1 - 17" refId="ref46670" refString="Chamberlin, R. V. (1922) Centipeds of Central America. Proceedings of the United States National Museum, 60 (7), 1 - 17." type="journal article" year="1922">Chamberlin, 1922</bibRefCitation>
</taxonomicName>
(which has previously been synonymised with
<taxonomicName id="4C77C3B8FFFED706FAC1F8B62C06DB8A" authority="(Pocock, 1896)" baseAuthorityName="Pocock" baseAuthorityYear="1896" class="Chilopoda" family="Scolopocryptopidae" genus="Newportia" kingdom="Animalia" order="Scolopendromorpha" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="stolli">
<emphasis id="B9036429FFFED706FAC1F8B628D9DB6F" box="[1349,1436,1878,1904]" italics="true" pageId="12" pageNumber="13">N. stolli</emphasis>
(Pocock, 1896)
</taxonomicName>
by
<bibRefCitation id="EFE6C5CAFFFED706FEEAF89B2FEEDB8B" author="Schileyko, A. &amp; Minelli, A." box="[366,683,1914,1940]" pageId="12" pageNumber="13" pagination="265 - 299" refId="ref50254" refString="Schileyko, A. &amp; Minelli, A. (1999) On the genus Newportia Gervais, 1847 (Chilopoda: Scolopendromorpha: Newportiidae). Arthropoda Selecta, 7 (4), 265 - 299." type="journal article" year="1999">Schileyko and Minelli 1999</bibRefCitation>
). Only two species of
<taxonomicName id="4C77C3B8FFFED706FC23F89A2958DB8B" authorityName="Gervais" authorityYear="1847" box="[935,1053,1915,1940]" class="Chilopoda" family="Scolopocryptopidae" genus="Newportia" kingdom="Animalia" order="Scolopendromorpha" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B9036429FFFED706FC23F89A2958DB8B" box="[935,1053,1915,1940]" italics="true" pageId="12" pageNumber="13">Newportia</emphasis>
</taxonomicName>
were described (both by
<bibRefCitation id="EFE6C5CAFFFED706FAB8F89D2C75DBA7" author="De Azara, L. N. &amp; Ferreira, R. L." pageId="12" pageNumber="13" pagination="267 - 278" refId="ref45646" refString="De Azara, L. N. &amp; Ferreira, R. L. (2014) Two new troglobitic Newportia (Newportia) from Brazil (Chilopoda: Scolopendromorpha). Zootaxa, 3881 (3), 267 - 278. https: // doi. org / 10.11646 / zootaxa. 3881.3.5" type="journal article" year="2014">Ázara &amp; Ferreira 2014</bibRefCitation>
) since publication of Schileykos (2013) paper; Schileyko (2018: 64) lowered the taxonomic status of
<taxonomicName id="4C77C3B8FFFED706FF13F8222C99DBC2" authority="Chamberlin, 1915" authorityName="Chamberlin" authorityYear="1915" box="[151,476,1986,2013]" class="Chilopoda" family="Scolopocryptopidae" genus="Newportia" kingdom="Animalia" order="Scolopendromorpha" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="cubana">
<emphasis id="B9036429FFFED706FF13F8222C4FDBC3" box="[151,266,1986,2012]" italics="true" pageId="12" pageNumber="13">N. cubana</emphasis>
Chamberlin, 1915
</taxonomicName>
to
<taxonomicName id="4C77C3B8FFFED706FE7BF8222FB4DBC3" box="[511,753,1986,2012]" class="Chilopoda" family="Scolopocryptopidae" genus="Newportia" kingdom="Animalia" order="Scolopendromorpha" pageId="12" pageNumber="13" phylum="Arthropoda" rank="subSpecies" species="longitarsis" subSpecies="cubana">
<emphasis id="B9036429FFFED706FE7BF8222FB4DBC3" box="[511,753,1986,2012]" italics="true" pageId="12" pageNumber="13">N. longitarsis cubana</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BC8B83BFFFFD707FF43FF762992DD5E" blockId="13.[151,1437,151,322]" pageId="13" pageNumber="14">
Summing up, given the species of all subgenera, the total number of species in the genus
<taxonomicName id="4C77C3B8FFFFD707FB56FF76280DDCAF" authorityName="Gervais" authorityYear="1847" box="[1234,1352,151,176]" class="Chilopoda" family="Scolopocryptopidae" genus="Newportia" kingdom="Animalia" order="Scolopendromorpha" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="B9036429FFFFD707FB56FF76280DDCAF" box="[1234,1352,151,176]" italics="true" pageId="13" pageNumber="14">Newportia</emphasis>
</taxonomicName>
should currently be 50 (or 51, see below). And namely: 25 species of
<taxonomicName id="4C77C3B8FFFFD707FCD4FF5A2913DCCA" authority="(Newportia)" authorityName="Gervais" authorityYear="1847" baseAuthorityName="Newportia" box="[848,1110,187,213]" class="Chilopoda" family="Scolopocryptopidae" genus="Newportia" kingdom="Animalia" order="Scolopendromorpha" pageId="13" pageNumber="14" phylum="Arthropoda" rank="subGenus" subGenus="Newportia">
<emphasis id="B9036429FFFFD707FCD4FF5A2E83DCCB" box="[848,966,187,212]" italics="true" pageId="13" pageNumber="14">Newportia</emphasis>
(
<emphasis id="B9036429FFFFD707FC53FF5A2908DCCB" box="[983,1101,187,212]" italics="true" pageId="13" pageNumber="14">Newportia</emphasis>
)
</taxonomicName>
(plus 6 subspecies of
<taxonomicName id="4C77C3B8FFFFD707FACAFF5A2CE8DCE6" authority="(Newport, 1845)" authorityName="Newport" authorityYear="1845" baseAuthorityName="Newport" baseAuthorityYear="1845" class="Chilopoda" family="Scolopocryptopidae" genus="Newportia" kingdom="Animalia" order="Scolopendromorpha" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="longitarsis">
<emphasis id="B9036429FFFFD707FACAFF5A2DAEDCE7" italics="true" pageId="13" pageNumber="14">N. longitarsis</emphasis>
(
<collectingRegion id="49B376D9FFFFD707FF7EFF3E2C24DCE6" box="[250,353,223,249]" country="United Kingdom" name="Newport" pageId="13" pageNumber="14">Newport</collectingRegion>
, 1845)
</taxonomicName>
and 2 subspecies of
<taxonomicName id="4C77C3B8FFFFD707FD12FF3E2945DCE6" authority="Chamberlin, 1955" authorityName="Chamberlin" authorityYear="1955" box="[662,1024,222,249]" class="Chilopoda" family="Scolopocryptopidae" genus="Newportia" kingdom="Animalia" order="Scolopendromorpha" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="weyrauchi">
<emphasis id="B9036429FFFFD707FD12FF3E2E69DCE6" box="[662,812,223,249]" italics="true" pageId="13" pageNumber="14">N. weyrauchi</emphasis>
Chamberlin, 1955
</taxonomicName>
), 15 of the group that has previously been referred to
<taxonomicName id="4C77C3B8FFFFD707FECAFEE22CBEDD02" authorityName="Saussure" authorityYear="1858" box="[334,507,259,285]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopendrides" kingdom="Animalia" order="Scolopendromorpha" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="B9036429FFFFD707FECAFEE22CBEDD02" box="[334,507,259,285]" italics="true" pageId="13" pageNumber="14">Scolopendrides</emphasis>
</taxonomicName>
(plus 2 subspecies of
<taxonomicName id="4C77C3B8FFFFD707FD75FEE22EB4DD02" authority="Pocock, 1891" authorityName="Pocock" authorityYear="1891" box="[753,1009,259,285]" class="Chilopoda" family="Scolopocryptopidae" genus="Newportia" kingdom="Animalia" order="Scolopendromorpha" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="ernsti">
<emphasis id="B9036429FFFFD707FD75FEE22E17DD03" box="[753,850,259,284]" italics="true" pageId="13" pageNumber="14">N. ernsti</emphasis>
Pocock, 1891
</taxonomicName>
) + 3 of
<taxonomicName id="4C77C3B8FFFFD707FBCDFEE22992DD02" authorityName="Chamberlin" authorityYear="1912" box="[1097,1239,259,285]" class="Chilopoda" family="Scolopocryptopidae" genus="Newportia" kingdom="Animalia" order="Scolopendromorpha" pageId="13" pageNumber="58" phylum="Arthropoda" rank="subGenus" subGenus="Newportides">
<emphasis id="B9036429FFFFD707FBCDFEE22992DD02" box="[1097,1239,259,285]" italics="true" pageId="13" pageNumber="14">Newportides</emphasis>
</taxonomicName>
(or 4 if
<taxonomicName id="4C77C3B8FFFFD707FAB5FEE228D8DD02" authorityName="Chamberlin" authorityYear="1922" box="[1329,1437,259,285]" class="Chilopoda" family="Scolopocryptopidae" genus="Newportia" kingdom="Animalia" order="Scolopendromorpha" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="sulana">
<emphasis id="B9036429FFFFD707FAB5FEE228D8DD02" box="[1329,1437,259,285]" italics="true" pageId="13" pageNumber="14">N. sulana</emphasis>
</taxonomicName>
is a valid species, which we doubt) + 4 of
<taxonomicName id="4C77C3B8FFFFD707FDEBFEC62FFFDD5E" authorityName="Chamberlin" authorityYear="1915" box="[623,698,295,321]" class="Chilopoda" family="Scolopocryptopidae" genus="Tidops" kingdom="Animalia" order="Scolopendromorpha" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="B9036429FFFFD707FDEBFEC62FFFDD5E" box="[623,698,295,321]" italics="true" pageId="13" pageNumber="14">Tidops</emphasis>
</taxonomicName>
+ 1 of
<taxonomicName id="4C77C3B8FFFFD707FC8FFEC62EFEDD5F" authorityName="Crabill" authorityYear="1977" box="[779,955,295,320]" class="Chilopoda" family="Scolopocryptopidae" genus="Ectonocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="B9036429FFFFD707FC8FFEC62EFEDD5F" box="[779,955,295,320]" italics="true" pageId="13" pageNumber="14">Ectonocryptops</emphasis>
</taxonomicName>
+ 2 of
<taxonomicName id="4C77C3B8FFFFD707FB88FEC62995DD5E" authorityName="Shelley &amp; Mercurio" authorityYear="2005" box="[1036,1232,295,321]" class="Chilopoda" family="Scolopocryptopidae" genus="Ectonocryptoides" kingdom="Animalia" order="Scolopendromorpha" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="B9036429FFFFD707FB88FEC62995DD5E" box="[1036,1232,295,321]" italics="true" pageId="13" pageNumber="14">Ectonocryptoides</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</subSubSection>
</treatment>
</document>