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<document id="EDB8E162E34A3E0CC859C32561D0BC3A" ID-CLB-Dataset="29539" ID-DOI="10.11646/zootaxa.4486.3.4" ID-GBIF-Dataset="543c2b3e-ded2-4af3-a7bb-ff803ef3e93a" ID-ISSN="1175-5326" ID-Zenodo-Dep="1437082" ID-ZooBank="19F1EDCF-19AA-47F9-BC1C-BAEAA8206DAE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1538115753658" checkinUser="plazi" docAuthor="Endruweit, Marco" docDate="2018" docId="C43787EDA26CC63BE1C4FE94235D9D9E" docLanguage="en" docName="zootaxa.4486.3.4.pdf" docOrigin="Zootaxa 4486 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rhinogobius phuongae Endruweit 2018, new species" docType="treatment" docVersion="7" lastPageNumber="305" masterDocId="380EFF95A27EC62EE153FFF021669C33" masterDocTitle="Description of four new species of freshwater gobies from the Black River drainage in China and Vietnam (Teleostei: Gobiidae)" masterLastPageNumber="310" masterPageNumber="284" pageNumber="302" updateTime="1698554753243" updateUser="ExternalLinkService">
<mods:mods id="113B79D515A8E1A1A00E9A2A37A80525" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="A60F73C1A2C2E48157D043F7D6CF524F">Description of four new species of freshwater gobies from the Black River drainage in China and Vietnam (Teleostei: Gobiidae)</mods:title>
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<mods:namePart id="4402E63ADE4652588F4D911DB05A4A50">Endruweit, Marco</mods:namePart>
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<mods:title id="6E81DCD01DC55A284825ADCFF541E28B">Zootaxa</mods:title>
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<mods:date id="B97506D9A2BE0FCCDFC17138A68823BE">2018</mods:date>
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<mods:number id="D67B42BE8FB4D532680C7A8F046E7A50">2018-09-28</mods:number>
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<mods:number id="FB557E5819890D301099323527A84342">4486</mods:number>
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<mods:identifier id="294A2C749ABC9A034306DB480F38BCE8" type="CLB-Dataset">29539</mods:identifier>
<mods:identifier id="1A405E4E29DE3F954A1507BAA7D0339F" type="DOI">10.11646/zootaxa.4486.3.4</mods:identifier>
<mods:identifier id="A21D8126F6F573E14B874F37A41D522D" type="GBIF-Dataset">543c2b3e-ded2-4af3-a7bb-ff803ef3e93a</mods:identifier>
<mods:identifier id="C3C8B477E1CF6148C6DB2B373B8085C6" type="ISSN">1175-5326</mods:identifier>
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<treatment id="C43787EDA26CC63BE1C4FE94235D9D9E" ID-DOI="http://doi.org/10.5281/zenodo.6489453" ID-GBIF-Taxon="148403158" ID-Zenodo-Dep="6489453" LSID="urn:lsid:plazi:treatment:C43787EDA26CC63BE1C4FE94235D9D9E" httpUri="http://treatment.plazi.org/id/C43787EDA26CC63BE1C4FE94235D9D9E" lastPageId="21" lastPageNumber="305" pageId="18" pageNumber="302">
<subSubSection id="04846570A26CC63CE1C4FE94204A9D93" pageId="18" pageNumber="302" type="nomenclature">
<paragraph id="4C2136FBA26CC63CE1C4FE9423369D4C" blockId="18.[151,592,356,416]" box="[151,592,356,383]" pageId="18" pageNumber="302">
<heading id="17698197A26CC63CE1C4FE9423369D4C" bold="true" box="[151,592,356,383]" fontSize="11" level="1" pageId="18" pageNumber="302" reason="1">
<emphasis id="7EEAEAE9A26CC63CE1C4FE9423369D4C" bold="true" box="[151,592,356,383]" pageId="18" pageNumber="302">
<taxonomicName id="8B9E4D78A26CC63CE1C4FE9420D79D4D" authority="Endruweit, 2018" authorityName="Endruweit" authorityYear="2018" box="[151,433,356,382]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="302" phylum="Chordata" rank="species" species="phuongae" status="sp. nov.">
<emphasis id="7EEAEAE9A26CC63CE1C4FE9420D79D4D" bold="true" box="[151,433,356,382]" italics="true" pageId="18" pageNumber="302">Rhinogobius phuongae</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="65D95792A26CC63CE093FE9523369D4C" box="[448,592,357,383]" pageId="18" pageNumber="302" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="4C2136FBA26CC63CE1C4FE77204A9D93" blockId="18.[151,592,356,416]" box="[151,300,391,416]" pageId="18" pageNumber="302">
(
<figureCitation id="D4A52A7EA26CC63CE1CCFE7720429D93" box="[159,292,391,416]" captionStart-0="FIGURE 11" captionStart-1="FIGURE 12" captionStart-2="FIGURE 13" captionStart-3="FIGURE 14" captionStartId-0="19.[151,250,1538,1560]" captionStartId-1="19.[151,250,1985,2007]" captionStartId-2="20.[151,250,1862,1884]" captionStartId-3="22.[151,250,623,645]" captionTargetBox-0="[204,1353,196,1491]" captionTargetBox-1="[187,1369,1656,1935]" captionTargetBox-2="[183,1396,228,1824]" captionTargetBox-3="[253,1324,219,574]" captionTargetId-0="figure@19.[159,1402,163,1514]" captionTargetId-1="figure@19.[151,1436,1624,1969]" captionTargetId-2="figure@20.[151,1435,193,1841]" captionTargetId-3="figure@22.[203,1384,193,602]" captionTargetPageId-0="19" captionTargetPageId-1="19" captionTargetPageId-2="20" captionTargetPageId-3="22" captionText-0="FIGURE 11. Rhinogobius phuongae, ZFMK ICH-104102, holotype, 40.0 mm SL, male; Tan Uyen, Lai Chau, Vietnam; Nam Mu River." captionText-1="FIGURE 12. Rhinogobius phuongae, ZFMK ICH-104104, paratype, 38.2 mm SL, female; Tan Uyen, Lai Chau, Vietnam; Nam Mu River." captionText-2="FIGURE 13. Rhinogobius phuongae, cephalic lateralis system, composite drawing based on holotype and paratypes." captionText-3="FIGURE 14. Snout coloration, females, scale bars 2 mm; left: Rhinogobius coccinella, EPC 5334, 39.5 mm SL; right: R. honghensis, EPC 5774, 47.2 mm SL; Wenshan, Yunnan; Nanxihe River." httpUri-0="https://zenodo.org/record/1437104/files/figure.png" httpUri-1="https://zenodo.org/record/1437106/files/figure.png" httpUri-2="https://zenodo.org/record/1437108/files/figure.png" httpUri-3="https://zenodo.org/record/1437110/files/figure.png" pageId="18" pageNumber="302">Figs. 11–14</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="04846570A26CC63CE1C4FE2024699EAE" pageId="18" pageNumber="302" type="materials_examined">
<paragraph id="4C2136FBA26CC63CE1C4FE2022619E3E" blockId="18.[151,1437,463,2037]" pageId="18" pageNumber="302">
<materialsCitation id="FCF63CA6A26CC63CE1C4FE2022619E3E" ID-GBIF-Occurrence="1913818333" collectingDate="2015-07-07" collectionCode="ZFMK" collectorName="M. Endruweit" country="Vietnam" location="Nam Mu River" pageId="18" pageNumber="302" specimenCode="ICH-104102" specimenCount="1" stateProvince="Lai Chau" typeStatus="holotype">
<emphasis id="7EEAEAE9A26CC63CE1C4FE20206D9DDB" bold="true" box="[151,267,464,488]" pageId="18" pageNumber="302">
<typeStatus id="93258859A26CC63CE1C4FE2020609DDB" box="[151,262,464,488]" pageId="18" pageNumber="302" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="2A8FAE3EA26CC63CE042FE2020399DD4" box="[273,351,464,487]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="18" pageNumber="302">ZFMK</collectionCode>
<specimenCode id="1C389E80A26CC63CE035FE2020909DDB" box="[358,502,464,488]" collectionCode="ICH-" pageId="18" pageNumber="302">ICH-104102</specimenCode>
, 40.0 mm SL, male; Nam Can com., Tan Uyen Cty.,
<collectingRegion id="8E5AF819A26CC63CE510FE3F25CC9DDB" box="[1091,1194,463,488]" country="Vietnam" name="Lai Chau" pageId="18" pageNumber="302">Lai Chau</collectingRegion>
Prov.,
<collectingCountry id="3489766BA26CC63CE5AAFE3F243D9DDB" box="[1273,1371,463,488]" name="Vietnam" pageId="18" pageNumber="302">Vietnam</collectingCountry>
;
<location id="49416020A26CC63CE435FE2020619E3E" LSID="urn:lsid:plazi:treatment:C43787EDA26CC63BE1C4FE94235D9D9E:49416020A26CC63CE435FE2020619E3E" country="Vietnam" name="Nam Mu River" pageId="18" pageNumber="302" stateProvince="Lai Chau">Nam Mu River</location>
; coll.
<collectorName id="E16B532DA26CC63CE019FE0520839E3E" box="[330,485,500,525]" pageId="18" pageNumber="302">M. Endruweit</collectorName>
&amp; S.V. Ngo,
<collectingDate id="2864E9D3A26CC63CE32EFE0522659E3E" box="[637,771,500,525]" pageId="18" pageNumber="302" value="2015-07-07">
<date id="3820103BA26CC63CE32EFE0522659E3E" box="[637,771,500,525]" pageId="18" pageNumber="302" value="2015-07-07">7 July 2015</date>
</collectingDate>
.
</materialsCitation>
</paragraph>
<paragraph id="4C2136FBA26CC63CE194FDE823709E66" blockId="18.[151,1437,463,2037]" pageId="18" pageNumber="302">
<materialsCitation id="FCF63CA6A26CC63CE194FDE823759E66" ID-GBIF-Occurrence="1913818332" collectingDate="2015-07-07" collectionCode="ZFMK" collectorName="M. Endruweit" country="Vietnam" location="Nam Mu River" pageId="18" pageNumber="302" specimenCode="KIZ 2015005368-5371, ICH-104101, 4103, 4104, 3, 38.2- 46.7" specimenCount="4" stateProvince="Lai Chau" typeStatus="paratype">
<emphasis id="7EEAEAE9A26CC63CE194FDE820209E03" bold="true" box="[199,326,536,560]" pageId="18" pageNumber="302">
<typeStatus id="93258859A26CC63CE194FDE820279E03" box="[199,321,536,560]" pageId="18" pageNumber="302" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCode id="1C389E80A26CC63CE01FFDE823379E03" box="[332,593,535,560]" collectionCode="KIZ" country="China" httpUri="http://grbio.org/cool/ncht-ribt" name="Kunming Institute of Zoology, Chinese Academy of Sciences" pageId="18" pageNumber="302">KIZ 2015005368-5371</specimenCode>
,
<specimenCount id="5A98FD72A26CC63CE30EFDE8238E9E03" box="[605,744,535,560]" pageId="18" pageNumber="302" type="generic">4 specimens</specimenCount>
,
<quantity id="8B669B1EA26CC63CE3A0FDE722F09E1C" box="[755,918,535,560]" metricMagnitude="-2" metricUnit="m" metricValue="3.225" metricValueMax="3.51" metricValueMin="2.94" pageId="18" pageNumber="302" unit="mm" value="32.25" valueMax="35.1" valueMin="29.4">29.4–35.1 mm</quantity>
SL;
<collectionCode id="2A8FAE3EA26CC63CE298FDE8257F9E1C" box="[971,1049,536,559]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="18" pageNumber="302">ZFMK</collectionCode>
<specimenCode id="1C389E80A26CC63CE573FDE821AE9E66" pageId="18" pageNumber="302">
ICH-104101, 4103, 4104, 3,
<quantity id="8B669B1EA26CC63CE431FDE821AE9E66" metricMagnitude="-2" metricUnit="m" metricValue="4.245" metricValueMax="4.67" metricValueMin="3.82" pageId="18" pageNumber="302" unit="mm" value="42.45" valueMax="46.7" valueMin="38.2">38.2- 46.7</quantity>
</specimenCode>
mm SL; data as for holotype
</materialsCitation>
.
</paragraph>
<paragraph id="4C2136FBA26CC63CE194FDAF24699EAE" blockId="18.[151,1437,463,2037]" pageId="18" pageNumber="302">
<materialsCitation id="FCF63CA6A26CC63CE194FDAF24699EAE" ID-GBIF-Occurrence="1913818328" collectingDate="2015-07-06" collectorName="M. Endruweit" country="Vietnam" location="Nam So River" municipality="Nam Na River" pageId="18" pageNumber="302" specimenCount="1" stateProvince="Lai Chau">
<emphasis id="7EEAEAE9A26CC63CE194FDAF20D49E4B" bold="true" box="[199,434,607,632]" pageId="18" pageNumber="302">Non-typic material.</emphasis>
EPC 5397-5402, 5405-5408, 10,
<quantity id="8B669B1EA26CC63CE26BFDAF22BB9E44" box="[824,989,607,632]" metricMagnitude="-2" metricUnit="m" metricValue="3.815" metricValueMax="4.58" metricValueMin="3.05" pageId="18" pageNumber="302" unit="mm" value="38.15" valueMax="45.8" valueMin="30.5">30.5–45.8 mm</quantity>
SL; Muong So com., Phong Tho Cty.,
<collectingRegion id="8E5AF819A26CC63CE1C4FD7421989EAE" box="[151,254,644,669]" country="Vietnam" name="Lai Chau" pageId="18" pageNumber="302">Lai Chau</collectingRegion>
Prov.,
<collectingCountry id="3489766BA26CC63CE01EFD7420D69EAE" box="[333,432,644,669]" name="Vietnam" pageId="18" pageNumber="302">Vietnam</collectingCountry>
;
<location id="49416020A26CC63CE0E8FD75233B9EAE" LSID="urn:lsid:plazi:treatment:C43787EDA26CC63BE1C4FE94235D9D9E:49416020A26CC63CE0E8FD75233B9EAE" box="[443,605,644,669]" country="Vietnam" municipality="Nam Na River" name="Nam So River" pageId="18" pageNumber="302" stateProvince="Lai Chau">Nam So River</location>
,
<collectingMunicipality id="AC45AC81A26CC63CE33BFD7522699EAE" box="[616,783,644,669]" pageId="18" pageNumber="302">Nam Na River</collectingMunicipality>
; coll.
<collectorName id="E16B532DA26CC63CE201FD7522889EAE" box="[850,1006,644,669]" pageId="18" pageNumber="302">M. Endruweit</collectorName>
&amp; S.V. Ngo,
<collectingDate id="2864E9D3A26CC63CE5D6FD74246A9EAE" box="[1157,1292,644,669]" pageId="18" pageNumber="302" value="2015-07-06">
<date id="3820103BA26CC63CE5D6FD74246A9EAE" box="[1157,1292,644,669]" pageId="18" pageNumber="302" value="2015-07-06">6 July 2015</date>
</collectingDate>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="04846570A26CC63CE194FD57225E9F63" pageId="18" pageNumber="302" type="diagnosis">
<paragraph id="4C2136FBA26CC63CE194FD57225E9F63" blockId="18.[151,1437,463,2037]" pageId="18" pageNumber="302">
<emphasis id="7EEAEAE9A26CC63CE194FD5720279EF3" bold="true" box="[199,321,679,704]" pageId="18" pageNumber="302">Diagnosis.</emphasis>
<taxonomicName id="8B9E4D78A26CC63CE019FD5723299EF3" authorityName="Endruweit" authorityYear="2018" box="[330,591,679,704]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="302" phylum="Chordata" rank="species" species="phuongae">
<emphasis id="7EEAEAE9A26CC63CE019FD5723299EF3" box="[330,591,679,704]" italics="true" pageId="18" pageNumber="302">Rhinogobius phuongae</emphasis>
</taxonomicName>
can be distinguished from its congeners by the following combination of characters: cheek with about 25 indistinct dark spots and branchiostegal membrane with white spots in males; operculum unmarked; flank plain-colored devoid of any notable marks in males, reticulate in females; unpaired fins with conspicuous very broad white distal band; pectoral fin length 20–21% SL; 8–10 [9] predorsal scales; 8–9 [9] branched rays in the second dorsal fin; and 28 vertebrae.
</paragraph>
</subSubSection>
<subSubSection id="04846570A26CC63CE194FCAB24109BBB" pageId="18" pageNumber="302" type="description">
<paragraph id="4C2136FBA26CC63CE194FCAB233A988B" blockId="18.[151,1437,463,2037]" pageId="18" pageNumber="302">
<emphasis id="7EEAEAE9A26CC63CE194FCAB203F9F47" bold="true" box="[199,345,859,884]" pageId="18" pageNumber="302">Description.</emphasis>
Morphometric proportions given in
<tableCitation id="011C0340A26CC63CE25BFCAC22079F47" box="[776,865,860,885]" captionStart="TABLE 3" captionStartId="16.[147,229,537,558]" captionText="TABLE 3. Morphometrics of Rhinogobius ngutinhoceps and R. phuongae." pageId="18" pageNumber="302">Table 3</tableCitation>
. Caudal peduncle short and moderately shallow; depth 2.1–2.5 length in males, 2.0–
<quantity id="8B669B1EA26CC63CE360FC8F23109FAB" box="[563,630,895,920]" metricMagnitude="-2" metricUnit="m" metricValue="6.604" pageId="18" pageNumber="302" unit="in" value="2.6">2.6 in</quantity>
females. Head short, slightly depressed, wider than deep. Eye small, dorsolateral. Interorbital distance moderate. Cheek inflated; edge of operculum represents maximum body width. Snout rounded. Mouth terminal, oblique, about 30° to body axis in males, about 40° in females. Lips thick, fleshy in males, thin in females. Maxilla extending vertical through the anterior rim of orbit. Jaws equal, each with 3–4 alternating rows of small, villiform, recurved teeth; outer row in upper jaw slightly enlarged. Mental frenum trapezoid. Gill opening relatively small, dorsally not reaching upper origin of pectoral fin, ventrally reaching forward to isthmus. Gill rakers 4 + 6. Genital papilla short, not reaching origin of anal fin. VC (10–11) + (17–18) = 28–29 [11 + 17 = 28]; AP 2; DF 3-2110, 3-22110, 3-1111, 3-22111 [3-22110]. Largest known size
<quantity id="8B669B1EA26CC63CE45FFB8C241598A7" box="[1292,1395,1148,1173]" metricMagnitude="-2" metricUnit="m" metricValue="4.67" pageId="18" pageNumber="302" unit="mm" value="46.7">46.7 mm</quantity>
SL (ZFMK ICH-104103,
<typeStatus id="93258859A26CC63CE0C3FB512095988A" box="[400,499,1185,1209]" pageId="18" pageNumber="302" type="paratype">paratype</typeStatus>
, female).
</paragraph>
<paragraph id="4C2136FBA26CC63CE194FB342470997B" blockId="18.[151,1437,463,2037]" pageId="18" pageNumber="302">
Fins. D
<date id="3820103BA26CC63CE04EFB3420E698EF" box="[285,384,1220,1244]" pageId="18" pageNumber="302">1 IV–VII</date>
[VI]; D
<date id="3820103BA26CC63CE089FB34209E98EF" box="[474,504,1220,1244]" pageId="18" pageNumber="302">2 I</date>
,8–9 [9]; A I,7–8 [8]; P 16–17 [17]; C 7 + 7. First dorsal fin spines III or IV longest, non-filamentous, reaching first branched ray of second dorsal fin when adpressed in males, not reaching origin of second dorsal fin in females. Second dorsal fin and anal fin reaching about midway to caudal-fin base in males, not reaching midway in females. Caudal fin spatulate, oblong in males, with rounded rear edge in females.
</paragraph>
<paragraph id="4C2136FBA26CC63CE194FAA423B499CE" blockId="18.[151,1437,463,2037]" pageId="18" pageNumber="302">
Scales. LL 28–31 [29]; TR 9–10 [10]; PD 8–10 [9]; SDP 7–8 [8]; CPS 12. Flank scales arranged in regular rows; anterior margin of predorsal squamation truncate, laterally reaching forward to about pore Θ, medianly with small cycloid scales anterior to first dorsal fin (
<figureCitation id="D4A52A7EA26CC63CE39CFA6C22409987" box="[719,806,1436,1461]" captionStart="FIGURE 13" captionStartId="20.[151,250,1862,1884]" captionTargetBox="[183,1396,228,1824]" captionTargetId="figure@20.[151,1435,193,1841]" captionTargetPageId="20" captionText="FIGURE 13. Rhinogobius phuongae, cephalic lateralis system, composite drawing based on holotype and paratypes." httpUri="https://zenodo.org/record/1437108/files/figure.png" pageId="18" pageNumber="302">Fig. 13</figureCitation>
); anterior margin of abdominal squamation obliquely bridging space between bases of pectoral and pelvic fins. Head, nape, prepectoral, pre- and subpelvic areas naked. Scales along the abdominal midline small, cycloid.
</paragraph>
<paragraph id="4C2136FBA26CC63CE194F9F724109BBB" blockId="18.[151,1437,463,2037]" pageId="18" pageNumber="302">
Cephalic lateralis system. Canals: Preorbital section of anterior oculoscapular canal with terminal pore σ between nostrils, mesially; pore λ on orbital bulge; pore κ located centrally posteriorly to orbit; postorbital section of oculoscapular canal with pore ω located dorsolaterally, slightly posterior to vertical through pore α; pores α and ρ located laterally in oculoscapular groove. Posterior oculoscapular canal with terminal pores Θ and τ. Terminal pores ρ and Θ separated by distance of about 1/5 eye diameter. Preopercular canal with pores γ, δ and ε along edge of preoperculum. Sensory papillae: Row
<emphasis id="7EEAEAE9A26CC63CE328F94D23EF9AE7" box="[635,649,1725,1748]" italics="true" pageId="18" pageNumber="302">a</emphasis>
long with about 6 papillae slanting downward anteriorly, reaching vertical through mid of orbit. Row
<emphasis id="7EEAEAE9A26CC63CE363F92F23589ACB" box="[560,574,1759,1784]" italics="true" pageId="18" pageNumber="302">b</emphasis>
short, with about 10 papillae, slanting downward posteriorly. Row
<emphasis id="7EEAEAE9A26CC63CE41EF911243F9ACB" box="[1357,1369,1761,1784]" italics="true" pageId="18" pageNumber="302">c</emphasis>
long, from beneath anterior nostril to vertical through posterior rim of orbit with about 15 prominent papillae. Row
<emphasis id="7EEAEAE9A26CC63CE4DDF8F424FA9B2E" box="[1422,1436,1796,1821]" italics="true" pageId="18" pageNumber="302">d</emphasis>
long, with about 19 papillae, running along labial groove to about posterior rim of orbit. Rows
<emphasis id="7EEAEAE9A26CC63CE5A7F8D924669B73" box="[1268,1280,1833,1856]" italics="true" pageId="18" pageNumber="302">c</emphasis>
and
<emphasis id="7EEAEAE9A26CC63CE46EF8D7242D9B73" box="[1341,1355,1831,1856]" italics="true" pageId="18" pageNumber="302">d</emphasis>
nearly diverging. Row
<emphasis id="7EEAEAE9A26CC63CE01CF8BD200C9B57" box="[335,362,1869,1892]" italics="true" pageId="18" pageNumber="302">cp</emphasis>
as single papilla located between the terminals of rows
<emphasis id="7EEAEAE9A26CC63CE2A3F8BD229A9B57" box="[1008,1020,1869,1892]" italics="true" pageId="18" pageNumber="302">c</emphasis>
and
<emphasis id="7EEAEAE9A26CC63CE564F8BC25239B56" box="[1079,1093,1868,1893]" italics="true" pageId="18" pageNumber="302">d</emphasis>
. Opercular rows
<emphasis id="7EEAEAE9A26CC63CE45EF8BD24409B57" box="[1293,1318,1869,1892]" italics="true" pageId="18" pageNumber="302">os</emphasis>
,
<emphasis id="7EEAEAE9A26CC63CE465F8BD242D9B57" box="[1334,1355,1869,1892]" italics="true" pageId="18" pageNumber="302">ot</emphasis>
and
<emphasis id="7EEAEAE9A26CC63CE4D5F8BD24FA9B57" box="[1414,1436,1869,1892]" italics="true" pageId="18" pageNumber="302">oi</emphasis>
present;
<emphasis id="7EEAEAE9A26CC63CE1A5F881206A9BBB" box="[246,268,1905,1928]" italics="true" pageId="18" pageNumber="302">ot</emphasis>
and
<emphasis id="7EEAEAE9A26CC63CE010F881203F9BBB" box="[323,345,1905,1928]" italics="true" pageId="18" pageNumber="302">oi</emphasis>
not interconnected. Approximate pathways of canals and papillae rows depicted in
<figureCitation id="D4A52A7EA26CC63CE450F89F24149BBB" box="[1283,1394,1903,1928]" captionStart="FIGURE 13" captionStartId="20.[151,250,1862,1884]" captionTargetBox="[183,1396,228,1824]" captionTargetId="figure@20.[151,1435,193,1841]" captionTargetPageId="20" captionText="FIGURE 13. Rhinogobius phuongae, cephalic lateralis system, composite drawing based on holotype and paratypes." httpUri="https://zenodo.org/record/1437108/files/figure.png" pageId="18" pageNumber="302">Figure 13</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="04846570A26CC63BE194F86425619D73" lastPageId="21" lastPageNumber="305" pageId="18" pageNumber="302" type="materials_examined">
<paragraph id="4C2136FBA26CC63BE194F86425619D73" blockId="18.[151,1437,463,2037]" lastBlockId="21.[151,1437,151,429]" lastPageId="21" lastPageNumber="305" pageId="18" pageNumber="302">
<emphasis id="7EEAEAE9A26CC63CE194F864233F9B9F" box="[199,601,1940,1965]" italics="true" pageId="18" pageNumber="302">Coloration of preserved specimens.</emphasis>
Body light brown, gradually darker dorsally; abdomen beige. Flank with an inconspicuous dark brown midlateral stripe in males, with black reticulate pattern in females. Scales in longitudinal series light brown with gray mark along its distal edge in males, with black pocket and mark along distal edge. Head and nape gray. Cheek light brown with about 25 indistinct gray spots in males, spots smaller than pupil, more conspicuous in females. Operculum gray, unmarked. Branchiostegal membrane gray with white spots in males, spots smaller than pupil; unmarked in females. Pectoral fin with 3–4 black spots along base; spots indistinct in males, upper spot larger and more conspicuous in females. First dorsal fin with conspicuous black blotch between spines I and III, less conspicuous in females larger than
<quantity id="8B669B1EA26BC63BE28AFF4C254C9CE7" box="[985,1066,188,212]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="21" pageNumber="305" unit="mm" value="40.0">40 mm</quantity>
SL. Second dorsal fin with 5–6 transverse rows of black spots. Caudal fin plain brown to black in males, with six transverse rows of black spots in females. Pectoral and pelvic fin gray. Anal fin light brown with broad black median band in males, band thinner in females. All fins except pectoral fin with conspicuous broad white distal band.
</paragraph>
</subSubSection>
<caption id="18E16673A26DC63DE1C4F9F221989A05" httpUri="https://zenodo.org/record/1437104/files/figure.png" pageId="19" pageNumber="303" startId="19.[151,250,1538,1560]" targetBox="[204,1353,196,1491]" targetPageId="19">
<paragraph id="4C2136FBA26DC63DE1C4F9F221989A05" blockId="19.[151,1435,1538,1590]" pageId="19" pageNumber="303">
<emphasis id="7EEAEAE9A26DC63DE1C4F9F220799A24" bold="true" box="[151,287,1538,1560]" pageId="19" pageNumber="303">FIGURE 11.</emphasis>
<taxonomicName id="8B9E4D78A26DC63DE076F9F223779A2B" authorityName="Endruweit" authorityYear="2018" box="[293,529,1538,1560]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="303" phylum="Chordata" rank="species" species="phuongae">
<emphasis id="7EEAEAE9A26DC63DE076F9F223779A2B" box="[293,529,1538,1560]" italics="true" pageId="19" pageNumber="303">Rhinogobius phuongae</emphasis>
</taxonomicName>
, ZFMK ICH-104102, holotype, 40.0 mm SL, male; Tan Uyen, Lai Chau, Vietnam; Nam Mu River.
</paragraph>
</caption>
<caption id="18E16673A26DC63DE1C4F83121989BC5" httpUri="https://zenodo.org/record/1437106/files/figure.png" pageId="19" pageNumber="303" startId="19.[151,250,1985,2007]" targetBox="[187,1369,1656,1935]" targetPageId="19">
<paragraph id="4C2136FBA26DC63DE1C4F83121989BC5" blockId="19.[151,1435,1985,2038]" pageId="19" pageNumber="303">
<emphasis id="7EEAEAE9A26DC63DE1C4F83120799BE5" bold="true" box="[151,287,1985,2007]" pageId="19" pageNumber="303">FIGURE 12.</emphasis>
<taxonomicName id="8B9E4D78A26DC63DE077F83123689BE4" authorityName="Endruweit" authorityYear="2018" box="[292,526,1985,2007]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="303" phylum="Chordata" rank="species" species="phuongae">
<emphasis id="7EEAEAE9A26DC63DE077F83123689BE4" box="[292,526,1985,2007]" italics="true" pageId="19" pageNumber="303">Rhinogobius phuongae</emphasis>
</taxonomicName>
, ZFMK ICH-104104, paratype, 38.2 mm SL, female; Tan Uyen, Lai Chau, Vietnam; Nam Mu River.
</paragraph>
</caption>
<caption id="18E16673A26AC63AE1C4F8B6245C9B6F" box="[151,1338,1862,1884]" httpUri="https://zenodo.org/record/1437108/files/figure.png" pageId="20" pageNumber="304" startId="20.[151,250,1862,1884]" targetBox="[183,1396,228,1824]" targetPageId="20">
<paragraph id="4C2136FBA26AC63AE1C4F8B6245C9B6F" blockId="20.[151,1338,1862,1884]" box="[151,1338,1862,1884]" pageId="20" pageNumber="304">
<emphasis id="7EEAEAE9A26AC63AE1C4F8B620799B6F" bold="true" box="[151,287,1862,1884]" pageId="20" pageNumber="304">FIGURE 13.</emphasis>
<taxonomicName id="8B9E4D78A26AC63AE076F8B623769B6F" authorityName="Endruweit" authorityYear="2018" box="[293,528,1862,1884]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="20" pageNumber="304" phylum="Chordata" rank="species" species="phuongae">
<emphasis id="7EEAEAE9A26AC63AE076F8B623769B6F" box="[293,528,1862,1884]" italics="true" pageId="20" pageNumber="304">Rhinogobius phuongae</emphasis>
</taxonomicName>
, cephalic lateralis system, composite drawing based on holotype and paratypes.
</paragraph>
</caption>
<subSubSection id="04846570A26BC63BE194FEBB250D9D57" box="[199,1131,331,357]" pageId="21" pageNumber="305" type="distribution">
<paragraph id="4C2136FBA26BC63BE194FEBB250D9D57" blockId="21.[151,1437,151,429]" box="[199,1131,331,357]" pageId="21" pageNumber="305">
<emphasis id="7EEAEAE9A26BC63BE194FEBB20069D57" bold="true" box="[199,352,331,356]" pageId="21" pageNumber="305">Distribution.</emphasis>
Nam Na and Nam Mu rivers in
<collectingRegion id="8E5AF819A26BC63BE399FEBC22F99D56" box="[714,927,332,357]" country="Vietnam" name="Lai Chau" pageId="21" pageNumber="305">Lai Chau Province</collectingRegion>
,
<collectingCountry id="3489766BA26BC63BE2F9FEBC256F9D56" box="[938,1033,332,357]" name="Vietnam" pageId="21" pageNumber="305">Vietnam</collectingCountry>
(
<figureCitation id="D4A52A7EA26BC63BE54AFEBC25389D57" box="[1049,1118,332,357]" captionStart="FIGURE 4" captionStartId="5.[151,250,1834,1856]" captionTargetBox="[181,1405,694,1781]" captionTargetId="figure@5.[150,1436,663,1812]" captionTargetPageId="5" captionText="FIGURE 4. Map of the Red River and adjacent drainages. Rivers: 1, Mekong; 2, Ma River, 3, Black River; 4, Red River; 5, Nanpanjiang River. Type localities: Ǫ, Rhinogobius coccinella; ●, R. nanophyllum; □, R. ngutinhoceps; ■, R. phuongae." httpUri="https://zenodo.org/record/1437090/files/figure.png" pageId="21" pageNumber="305">Fig. 4</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="04846570A26BC63BE194FE80235D9D9E" pageId="21" pageNumber="305" type="etymology">
<paragraph id="4C2136FBA26BC63BE194FE80235D9D9E" blockId="21.[151,1437,151,429]" pageId="21" pageNumber="305">
<emphasis id="7EEAEAE9A26BC63BE194FE8020289DBB" bold="true" box="[199,334,368,392]" pageId="21" pageNumber="305">Etymology.</emphasis>
Named after Mrs Thi Dieu Phuong Nguyen of RIA1 for her enthusiastic interest in the Vietnamese fish fauna; a matronym in apposition.
</paragraph>
</subSubSection>
</treatment>
</document>
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