281 lines
36 KiB
XML
281 lines
36 KiB
XML
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<document id="5B034706655434F5ED58CEBD8650F2B8" ID-DOI="10.11646/zootaxa.5566.1.11" ID-ISSN="1175-5326" ID-Zenodo-Dep="14612158" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1736289691392" checkinUser="plazi" docAuthor="Shavrin, Alexey V." docDate="2025" docId="4518D35DFFFCFFDB7DB6FA74FABFCEDD" docLanguage="en" docName="zootaxa.5566.1.11.pdf" docOrigin="Zootaxa 5566 (1)" docSource="https://doi.org/10.11646/zootaxa.5566.1.11" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Trichodromeus ottoi Shavrin, 2025, sp. nov." docType="treatment" docVersion="2" lastPageNumber="199" masterDocId="B921AB25FFFCFFD97D21FFE8FFA2C96A" masterDocTitle="A new species of Trichodromeus Luze, 1903 from Afghanistan, and a new combination (Coleoptera: Staphylinidae: Anthophagini)" masterLastPageNumber="200" masterPageNumber="197" pageNumber="197" updateTime="1736290056554" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="FE4BAA35763D075A25BE35CB70E48AC5">
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<mods:title id="42E1A63AADEF3DC86DAC64AD75F1D94D">A new species of Trichodromeus Luze, 1903 from Afghanistan, and a new combination (Coleoptera: Staphylinidae: Anthophagini)</mods:title>
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<mods:namePart id="08F3B470950B223CCA5E88D800FA298C">Shavrin, Alexey V.</mods:namePart>
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<mods:typeOfResource id="7337A2B5F750B92D2E48771D3ACD9E20">text</mods:typeOfResource>
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<mods:title id="97792B19C86C3C30A8A56318CC3BF141">Zootaxa</mods:title>
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<mods:part id="3C7F8B9C25A032D049F9824F8BF73DD8">
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<mods:date id="EC0B4974790EF11C7B2D5768CDCE4854">2025</mods:date>
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<mods:number id="F54D1871548C71EB95A1679374F7D0BD">2025-01-07</mods:number>
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<mods:number id="42F710FB06876E2FC9EDFA1591F487C3">5566</mods:number>
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<mods:number id="3D1DBAD5423A01E87362BB5D3A38D7AB">1</mods:number>
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<mods:url id="4E6E50AAEC7C4853B434E7A81DF8DD6C">https://doi.org/10.11646/zootaxa.5566.1.11</mods:url>
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<mods:classification id="D26CD7E2EC16C6BEC5FC238EB4FED7A5">journal article</mods:classification>
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<mods:identifier id="9F45F3849A379F1652B420D1E7C26E8F" type="DOI">10.11646/zootaxa.5566.1.11</mods:identifier>
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<mods:identifier id="C6CD79733B2D78B3D7049A571C25121F" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="BECA1244DA269FA639CE24080818CCCD" type="Zenodo-Dep">14612158</mods:identifier>
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<treatment id="4518D35DFFFCFFDB7DB6FA74FABFCEDD" ID-DOI="http://doi.org/10.5281/zenodo.14612160" ID-Zenodo-Dep="14612160" LSID="urn:lsid:plazi:treatment:4518D35DFFFCFFDB7DB6FA74FABFCEDD" httpUri="http://treatment.plazi.org/id/4518D35DFFFCFFDB7DB6FA74FABFCEDD" lastPageId="2" lastPageNumber="199" pageId="0" pageNumber="197">
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<subSubSection id="85AB31C0FFFCFFD97DB6FA74FE50CCDD" box="[151,498,1436,1463]" pageId="0" pageNumber="197" type="nomenclature">
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<paragraph id="CD0E624BFFFCFFD97DB6FA74FE50CCDD" blockId="0.[151,1437,1436,1967]" box="[151,498,1436,1463]" pageId="0" pageNumber="197">
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<heading id="9646D527FFFCFFD97DB6FA74FE50CCDD" bold="true" box="[151,498,1436,1463]" fontSize="11" level="1" pageId="0" pageNumber="197" reason="1">
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<emphasis id="FFC5BE59FFFCFFD97DB6FA74FE50CCDD" bold="true" box="[151,498,1436,1463]" pageId="0" pageNumber="197">
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<taxonomicName id="0AB119C8FFFCFFD97DB6FA74FE2FCCDC" authorityName="Shavrin" authorityYear="2025" box="[151,397,1436,1463]" class="Insecta" family="Staphylinidae" genus="Trichodromeus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="197" phylum="Arthropoda" rank="species" species="ottoi" status="sp. nov.">
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<emphasis id="FFC5BE59FFFCFFD97DB6FA74FE2FCCDC" bold="true" box="[151,397,1436,1463]" italics="true" pageId="0" pageNumber="197">Trichodromeus ottoi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="E4F60322FFFCFFD97CB5FA75FE50CCDD" box="[404,498,1437,1463]" pageId="0" pageNumber="197" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="85AB31C0FFFCFFD97DB6FA2AFEA4CCB1" box="[151,262,1474,1499]" pageId="0" pageNumber="197" type="description">
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<paragraph id="CD0E624BFFFCFFD97DB6FA2AFEA4CCB1" blockId="0.[151,1437,1436,1967]" box="[151,262,1474,1499]" pageId="0" pageNumber="197">
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(
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<figureCitation id="558A7ECEFFFCFFD97DBEFA2AFF5ECCB1" box="[159,252,1474,1499]" captionStart="FIGURES 1–3" captionStartId="1.[151,264,1140,1164]" captionTargetBox="[182,1416,201,1082]" captionTargetId="figure-15@1.[151,1436,181,1117]" captionTargetPageId="1" captionText="FIGURES 1–3. Trichodromeus ottoi: 1–habitus (paratype), 2–aedeagus, parameral view, 3–aedeagus, lateral view. Scale bars: 1.0 mm (Fig. 1), 0.1 mm (Figs 2–3)." figureDoi="http://doi.org/10.5281/zenodo.14612162" httpUri="https://zenodo.org/record/14612162/files/figure.png" pageId="0" pageNumber="197">Figs 1–3</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="85AB31C0FFFCFFD97DB6FA0FFE0BCEC5" pageId="0" pageNumber="197" type="materials_examined">
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<paragraph id="CD0E624BFFFCFFD97DB6FA0FFBCDCF01" blockId="0.[151,1437,1436,1967]" pageId="0" pageNumber="197">
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<emphasis id="FFC5BE59FFFCFFD97DB6FA0FFE0ECC94" bold="true" box="[151,428,1510,1535]" inLineHeading="true" pageId="0" pageNumber="197" reason="1">Type material examined.</emphasis>
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<typeStatus id="120ADCE9FFFCFFD97C91FA0EFDB1CC94" box="[432,531,1510,1534]" pageId="0" pageNumber="197" type="holotype">Holotype</typeStatus>
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<specimenCount id="DBB7A9C2FFFCFFD97F39FA0FFD8DCC94" box="[536,559,1511,1534]" count="1" pageId="0" pageNumber="197" type="male">♂</specimenCount>
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: ‘
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<specimenCount id="DBB7A9C2FFFCFFD97F1CFA0FFDF4CC95" box="[573,598,1511,1535]" count="1" pageId="0" pageNumber="197" type="male">♂</specimenCount>
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’ <printed>, ‘J. Klapperich | Anjuman-Pass |
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<quantity id="0A49CFAEFFFCFFD9793EFA0FFBCFCC94" box="[1055,1133,1511,1535]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="0" pageNumber="197" unit="m" value="4000.0">4000 m</quantity>
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Anjuman.’ <printed>, ‘Geb., Badakschan | 11.8.[19]52, NO. Afghan.’ <printed>, ‘
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<collectionCode id="ABA0FA8EFFFCFFD97FF4F9E3FC8CCF49" box="[725,814,1547,1571]" pageId="0" pageNumber="197">TYPUS</collectionCode>
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<taxonomicName id="0AB119C8FFFCFFD97E66F9E2FC75CF48" box="[839,983,1546,1570]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="197" phylum="Arthropoda" rank="genus">Geodromicus</taxonomicName>
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[handwritten in black] | badagschanensis [handwritten in black] | O. Scheerpeltz’ <red, printed>, ‘
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<taxonomicName id="0AB119C8FFFCFFD97FCAF9C6FC22CF2C" box="[747,896,1582,1606]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="197" phylum="Arthropoda" rank="genus">Geodromicus</taxonomicName>
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| badagschanensis | n. sp.’ <handwritten in black>, ‘
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<typeStatus id="120ADCE9FFFCFFD97DBAF9BBFE8FCF01" box="[155,301,1619,1643]" pageId="0" pageNumber="197" type="holotype">HOLOTYPE</typeStatus>
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<taxonomicName id="0AB119C8FFFCFFD97C1EF9BAFE42CF00" authorityName="Luze" authorityYear="1903" box="[319,480,1618,1642]" class="Insecta" family="Staphylinidae" genus="Trichodromeus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="197" phylum="Arthropoda" rank="genus">
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<emphasis id="FFC5BE59FFFCFFD97C1EF9BAFE42CF00" box="[319,480,1618,1642]" italics="true" pageId="0" pageNumber="197">Trichodromeus</emphasis>
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</taxonomicName>
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<taxonomicName id="0AB119C8FFFCFFD97CD2F9BBFD86CF00" authorityName="Shavrin" authorityYear="2025" box="[499,548,1619,1642]" class="Insecta" family="Staphylinidae" genus="Trichodromeus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="197" phylum="Arthropoda" rank="species" species="ottoi" status="sp. nov.">
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<emphasis id="FFC5BE59FFFCFFD97CD2F9BBFD86CF00" box="[499,548,1619,1642]" italics="true" pageId="0" pageNumber="197">ottoi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="E4F60322FFFCFFD97F0BF9BBFDFFCF00" box="[554,605,1619,1642]" pageId="0" pageNumber="197" rank="species">sp.n.</taxonomicNameLabel>
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| Shavrin A.
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<collectionCode id="ABA0FA8EFFFCFFD97FC1F9BBFD57CF01" box="[736,757,1619,1643]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="0" pageNumber="197" type="Museum">V</collectionCode>
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. des. 2024’ <red, printed> (
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<collectionCode id="ABA0FA8EFFFCFFD97938F9BBFBC1CF01" box="[1049,1123,1619,1643]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="0" pageNumber="197">NMW</collectionCode>
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).
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</paragraph>
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<paragraph id="CD0E624BFFFCFFD97DE6F99FFE0BCEC5" blockId="0.[151,1437,1436,1967]" pageId="0" pageNumber="197">
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<typeStatus id="120ADCE9FFFCFFD97DE6F99FFE91CFE5" box="[199,307,1655,1679]" pageId="0" pageNumber="197" type="paratype">Paratypes</typeStatus>
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:
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<specimenCount id="DBB7A9C2FFFCFFD97C1CF99FFE2ACFE4" box="[317,392,1655,1679]" count="10" pageId="0" pageNumber="197" type="female">10 ♀♀</specimenCount>
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: ‘
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<specimenCount id="DBB7A9C2FFFCFFD97CBDF99FFE10CFE5" box="[412,434,1655,1679]" count="1" pageId="0" pageNumber="197" type="female">♀</specimenCount>
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’ <printed>, ‘J. Klapperich | Anjuman-Pass |
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<quantity id="0A49CFAEFFFCFFD97EACF99FFC7CCFE4" box="[909,990,1655,1679]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="0" pageNumber="197" unit="m" value="4000.0">4000 m</quantity>
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Anjuman.’ <printed>, ‘Geb., Badakschan | 11.8.[19]52, NO. Afghan.’ <printed>, ‘
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<collectionCode id="ABA0FA8EFFFCFFD97F1FF973FD35CFD9" box="[574,663,1691,1715]" pageId="0" pageNumber="197">TYPUS</collectionCode>
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<taxonomicName id="0AB119C8FFFCFFD97F8BF972FC98CFD8" box="[682,826,1690,1714]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="197" phylum="Arthropoda" rank="genus">Geodromicus</taxonomicName>
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[handwritten in black] | badagschanensis [handwritten in black] | O. Scheerpeltz’ <red printed label> (
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<specimenCount id="DBB7A9C2FFFCFFD97F51F957FD3ACFBC" box="[624,664,1727,1751]" count="1" pageId="0" pageNumber="197" type="female">1 ♀</specimenCount>
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: cSh;
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<specimenCount id="DBB7A9C2FFFCFFD97FF8F957FCB4CFBC" box="[729,790,1727,1751]" count="9" pageId="0" pageNumber="197" type="female">9 ♀♀</specimenCount>
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:
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<collectionCode id="ABA0FA8EFFFCFFD97E03F957FCC9CFBD" box="[802,875,1727,1751]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="0" pageNumber="197">NMW</collectionCode>
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);
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<specimenCount id="DBB7A9C2FFFCFFD97E5DF956FC67CFBC" box="[892,965,1726,1750]" count="15" pageId="0" pageNumber="197" type="male">15 ♂♂</specimenCount>
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(
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<specimenCount id="DBB7A9C2FFFCFFD97EF3F957FBD3CFBC" box="[978,1137,1726,1751]" count="2" pageId="0" pageNumber="197" type="generic">two specimens</specimenCount>
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dissected): same four labels as in the
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<typeStatus id="120ADCE9FFFCFFD97DD7F90AFEF1CF90" box="[246,339,1762,1786]" pageId="0" pageNumber="197" type="holotype">holotype</typeStatus>
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(
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<specimenCount id="DBB7A9C2FFFCFFD97C45F90BFE2FCF90" box="[356,397,1763,1787]" count="1" pageId="0" pageNumber="197" type="male">1 ♂</specimenCount>
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: cSh;
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<specimenCount id="DBB7A9C2FFFCFFD97CEFF90BFDB9CF90" box="[462,539,1763,1787]" count="14" pageId="0" pageNumber="197" type="male">14 ♂♂</specimenCount>
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:
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<collectionCode id="ABA0FA8EFFFCFFD97F04F90BFDCCCF91" box="[549,622,1763,1787]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="0" pageNumber="197">NMW</collectionCode>
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);
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<specimenCount id="DBB7A9C2FFFCFFD97FA1F90AFD1ECF90" box="[640,700,1762,1786]" count="5" pageId="0" pageNumber="197" type="male">5 ♂♂</specimenCount>
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(
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<specimenCount id="DBB7A9C2FFFCFFD97FEBF90BFCCBCF90" box="[714,873,1762,1787]" count="2" pageId="0" pageNumber="197" type="generic">two specimens</specimenCount>
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dissected): ‘
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<specimenCount id="DBB7A9C2FFFCFFD97ECCF90BFBA4CF91" box="[1005,1030,1763,1787]" count="1" pageId="0" pageNumber="197" type="male">♂</specimenCount>
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’ <printed>, ‘J. Klapperich | Anjuman-Pass |
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<quantity id="0A49CFAEFFFCFFD97DFCF8EFFE92CE74" box="[221,304,1799,1823]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="0" pageNumber="197" unit="m" value="4000.0">4000 m</quantity>
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Anjuman.’ <printed>, ‘Geb., Badakschan | 11.8.[19]52, NO. Afghan.’ <printed>, ‘ex coll. | Scheerpeltz’ <blue, printed>, ‘
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<collectionCode id="ABA0FA8EFFFCFFD97C6AF8C3FE06CE29" box="[331,420,1835,1859]" pageId="0" pageNumber="197">TYPUS</collectionCode>
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<taxonomicName id="0AB119C8FFFCFFD97C94F8C2FDE7CE28" box="[437,581,1834,1858]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="197" phylum="Arthropoda" rank="genus">Geodromicus</taxonomicName>
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[handwritten in black] | badagschanensis [handwritten in black] | O. Scheerpeltz’ <red, printed>, ‘
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<taxonomicName id="0AB119C8FFFCFFD97C67F8A6FE79CE0C" box="[326,475,1870,1894]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="197" phylum="Arthropoda" rank="genus">Geodromicus</taxonomicName>
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| badagschanensis | n. sp.’ <handwritten in black> (
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<collectionCode id="ABA0FA8EFFFCFFD97932F8A7FBFFCE0D" box="[1043,1117,1871,1895]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="0" pageNumber="197">NMW</collectionCode>
|
|||
|
|
);
|
|||
|
|
<specimenCount id="DBB7A9C2FFFCFFD97950F8A7FB10CE0C" box="[1137,1202,1871,1895]" count="4" pageId="0" pageNumber="197" type="female">4 ♀♀</specimenCount>
|
|||
|
|
: ‘
|
|||
|
|
<specimenCount id="DBB7A9C2FFFCFFD979E6F8A7FB7FCE0D" box="[1223,1245,1871,1895]" count="1" pageId="0" pageNumber="197" type="female">♀</specimenCount>
|
|||
|
|
’ <printed>, other labels as the previous (
|
|||
|
|
<collectionCode id="ABA0FA8EFFFCFFD97CABF89BFE76CEE1" box="[394,468,1907,1931]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="0" pageNumber="197">NMW</collectionCode>
|
|||
|
|
). All
|
|||
|
|
<typeStatus id="120ADCE9FFFCFFD97F2FF89BFDD6CEE1" box="[526,628,1907,1931]" pageId="0" pageNumber="197" type="paratype">paratypes</typeStatus>
|
|||
|
|
with additional red printed label: ‘
|
|||
|
|
<typeStatus id="120ADCE9FFFCFFD97EC2F89BFBCFCEE1" box="[995,1133,1907,1931]" pageId="0" pageNumber="197" type="paratype">PARATYPE</typeStatus>
|
|||
|
|
|
|
|||
|
|
<taxonomicName id="0AB119C8FFFCFFD9795EF89AFA82CEE0" authorityName="Luze" authorityYear="1903" box="[1151,1312,1906,1930]" class="Insecta" family="Staphylinidae" genus="Trichodromeus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="197" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="FFC5BE59FFFCFFD9795EF89AFA82CEE0" box="[1151,1312,1906,1930]" italics="true" pageId="0" pageNumber="197">Trichodromeus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
|
|
|||
|
|
<taxonomicName id="0AB119C8FFFCFFD97813F89BFAC1CEE0" authorityName="Shavrin" authorityYear="2025" box="[1330,1379,1907,1930]" class="Insecta" family="Staphylinidae" genus="Trichodromeus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="197" phylum="Arthropoda" rank="species" species="ottoi" status="sp. nov.">
|
|||
|
|
<emphasis id="FFC5BE59FFFCFFD97813F89BFAC1CEE0" box="[1330,1379,1907,1930]" italics="true" pageId="0" pageNumber="197">ottoi</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="E4F60322FFFCFFD9784BF89BFA3ECEE0" box="[1386,1436,1907,1930]" pageId="0" pageNumber="197" rank="species">sp.n.</taxonomicNameLabel>
|
|||
|
|
| Shavrin A.
|
|||
|
|
<collectionCode id="ABA0FA8EFFFCFFD97C35F87FFE8BCEC5" box="[276,297,1943,1967]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="0" pageNumber="197" type="Museum">V</collectionCode>
|
|||
|
|
. des. 2024’.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<caption id="99CE32C3FFFDFFD87DB6FB9CFDA5CDDB" ID-DOI="http://doi.org/10.5281/zenodo.14612162" ID-Zenodo-Dep="14612162" httpUri="https://zenodo.org/record/14612162/files/figure.png" pageId="1" pageNumber="198" startId="1.[151,264,1140,1164]" targetBox="[182,1416,201,1082]" targetPageId="1" targetType="figure">
|
|||
|
|
<paragraph id="CD0E624BFFFDFFD87DB6FB9CFDA5CDDB" blockId="1.[151,1437,1140,1201]" pageId="1" pageNumber="198">
|
|||
|
|
<emphasis id="FFC5BE59FFFDFFD87DB6FB9CFE99CDE6" bold="true" box="[151,315,1140,1164]" pageId="1" pageNumber="198">FIGURES 1–3.</emphasis>
|
|||
|
|
<taxonomicName id="0AB119C8FFFDFFD87C63FB9DFDADCDE6" authorityName="Shavrin" authorityYear="2025" box="[322,527,1141,1165]" class="Insecta" family="Staphylinidae" genus="Trichodromeus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="198" phylum="Arthropoda" rank="species" species="ottoi">
|
|||
|
|
<emphasis id="FFC5BE59FFFDFFD87C63FB9DFDADCDE6" box="[322,527,1141,1165]" italics="true" pageId="1" pageNumber="198">Trichodromeus ottoi</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
: 1–habitus (paratype), 2–aedeagus, parameral view, 3–aedeagus, lateral view. Scale bars: 1.0 mm (Fig. 1), 0.1 mm (Figs 2–3).
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<subSubSection id="85AB31C0FFFDFFDB7DE6FAECFEE7CE69" lastPageId="2" lastPageNumber="199" pageId="1" pageNumber="198" type="description">
|
|||
|
|
<paragraph id="CD0E624BFFFDFFD87DE6FAECFCB9CF72" blockId="1.[151,1437,1284,2029]" pageId="1" pageNumber="198">
|
|||
|
|
<emphasis id="FFC5BE59FFFDFFD87DE6FAECFEF0CC76" bold="true" box="[199,338,1284,1308]" inLineHeading="true" pageId="1" pageNumber="198" reason="1">Description.</emphasis>
|
|||
|
|
Measurements (n=37): maximum width of head including eyes: 0.69–0.76; length of head (from base of labrum to posterior constriction along head midline): 0.49–0.56; ocular length (longitudinal): 0.18–0.22; length of temple: 0.12–0.15; length of antenna (
|
|||
|
|
<typeStatus id="120ADCE9FFFDFFD87CF9FAA5FD9ECC0F" box="[472,572,1357,1381]" pageId="1" pageNumber="198" type="holotype">holotype</typeStatus>
|
|||
|
|
): 2.11; length of pronotum: 0.69–0.75; maximum width of pronotum: 0.85–0.96; minimum width of pronotum: 0.71–0.74; sutural length of elytra (length of elytra from apex of scutellum to posterior margin of sutural angle): 1.29–1.38; maximum width of elytra: 1.34–1.41; length of metatibia (
|
|||
|
|
<typeStatus id="120ADCE9FFFDFFD879AAFA7DFB4DCCC7" box="[1163,1263,1429,1453]" pageId="1" pageNumber="198" type="holotype">holotype</typeStatus>
|
|||
|
|
): 1.00; length of metatrsus (
|
|||
|
|
<typeStatus id="120ADCE9FFFDFFD87C2DFA51FED2CCBB" box="[268,368,1465,1489]" pageId="1" pageNumber="198" type="holotype">holotype</typeStatus>
|
|||
|
|
): 0.39 (metatarsomeres 1–4: 0.27; metatarsomere 5: 0.12); maximum width of abdomen: 1.34–1.38; length of aedeagus (from base of median lobe to apex of parameres): 0.68–0.82; total length of body (from anterior margin of clypeus to apex of abdomen): 4.00–5.34 (
|
|||
|
|
<typeStatus id="120ADCE9FFFDFFD87F4EF9E9FD70CF73" box="[623,722,1537,1561]" pageId="1" pageNumber="198" type="holotype">holotype</typeStatus>
|
|||
|
|
: 4.15).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="CD0E624BFFFDFFD87DE6F9CDFE9BCEEA" blockId="1.[151,1437,1284,2029]" pageId="1" pageNumber="198">
|
|||
|
|
Habitus as in
|
|||
|
|
<figureCitation id="558A7ECEFFFDFFD87C45F9CDFE04CF57" box="[356,422,1573,1597]" captionStart="FIGURES 1–3" captionStartId="1.[151,264,1140,1164]" captionTargetBox="[182,1416,201,1082]" captionTargetId="figure-15@1.[151,1436,181,1117]" captionTargetPageId="1" captionText="FIGURES 1–3. Trichodromeus ottoi: 1–habitus (paratype), 2–aedeagus, parameral view, 3–aedeagus, lateral view. Scale bars: 1.0 mm (Fig. 1), 0.1 mm (Figs 2–3)." figureDoi="http://doi.org/10.5281/zenodo.14612162" httpUri="https://zenodo.org/record/14612162/files/figure.png" pageId="1" pageNumber="198">Fig. 1</figureCitation>
|
|||
|
|
. Body brown to reddish-brown, with yellow-brown to brown elytra, usually with distinctly darkened mediobasal and median portions; mouthparts, antenna and legs yellowish to yellow-brown (antenna sometimes darker); apical maxillary palpomere and tarsi yellow. Head with distinct and irregular microsculpture: dense, fine and transverse on clypeus, isodiametric in portions between anteriomedian depression and eyes, with dense and coarse transverse meshes in middle and dense isodiametric microreticulation on infraorbital portions; neck with dense and coarse isodiametric microsculpture; pronotum with dense isodiametric sculpture, finer and sometimes indistinct in mediobasal portion; scutellum with dense transverse microreticulation; elytra without meshes; abdominal tergites with dense transverse microsculpture, sometimes finer on abdominal tergite VII. Pubescence of forebody moderately dense, somewhat longer in frontal part of head; pubescence of abdomen dense and fine, longer and and sometimes denser in middle of abdominal tergites IV–VI.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="CD0E624BFFFDFFDB7DE6F865FAA8CB7D" blockId="1.[151,1437,1284,2029]" lastBlockId="2.[151,1437,151,1976]" lastPageId="2" lastPageNumber="199" pageId="1" pageNumber="198">
|
|||
|
|
Head 1.3–1.4 times as broad as long, with strongly elevated supra-antennal prominences and infraorbital portions; anteriomedian depression wide and deep, strongly narrowed basad; interocellar depression subrectangular, deep, narrow or moderately wide; anteocellar foveae indistinct or distinct, deep and long, strongly convergent latero-anteriad toward level of middle or anterior third of eyes (some
|
|||
|
|
<typeStatus id="120ADCE9FFFEFFDB7FA3FF70FD4AC9DA" box="[642,744,152,176]" pageId="2" pageNumber="199" type="paratype">paratypes</typeStatus>
|
|||
|
|
without distinct anteocellar foveae); temples convex, moderately long, 1.4–1.5 times as long as longitudinal length of eyes if seen dorsally. Ocelli small, but well visible, located at level or slightly below level of posterior margins of eyes; distance between ocelli 1.3–1.6 times as long as distance between ocellus and posterior margin of eye. Eyes moderately large, convex. Punctation irregular and fine, usually denser, larger and deeper on infraorbital portions, but some
|
|||
|
|
<typeStatus id="120ADCE9FFFEFFDB7F5BFEC0FD42C82A" box="[634,736,296,320]" pageId="2" pageNumber="199" type="paratype">paratypes</typeStatus>
|
|||
|
|
with finer and sparser punctation in these parts, median portion in front of interocellar depression without or with several fine punctures; neck without punctures.Apical maxillary palpomere slightly shorter than preceding segment, from widest basal portion strongly narrowed toward acute apex. Antenna reaching basal part of elytra when reclined, with slightly elongate antennomeres 3–10; basal antennomere wide, about two and a half times as long as broad, antennomere 2 distinctly shorter and narrower than basal antennomere, 3 distinctly longer and slightly broader (in apical part) than 3, 4–5 distinctly shorter than 3, 6–10 slightly longer and indistinctly broader than 5, apical antennomere 1.3–1.4 times as long as 10, from about middle gradually narrowed toward acute apex.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="CD0E624BFFFEFFDB7DE6FDCCFD05CA5D" blockId="2.[151,1437,151,1976]" pageId="2" pageNumber="199">
|
|||
|
|
Pronotum convex, 1.2 times as broad as long, 1.2 times as broad as head, from widest anterior part strongly narrowed posteriad toward rounded hind angles; frontal part slightly protruded apicad, with widely rounded apical margin, about as long as straight or slightly concaved posterior margin; laterobasal margins somewhat subparallel or indistinctly concave in some
|
|||
|
|
<typeStatus id="120ADCE9FFFEFFDB7DF7FD78FEE2CBC2" box="[214,320,656,680]" pageId="2" pageNumber="199" type="paratype">paratypes</typeStatus>
|
|||
|
|
; middle potion with indistinct or distinct longitudinal depression (some
|
|||
|
|
<typeStatus id="120ADCE9FFFEFFDB791BFD78FB02CBC2" box="[1082,1184,656,680]" pageId="2" pageNumber="199" type="paratype">paratypes</typeStatus>
|
|||
|
|
without it); mediobasal portion without depression, but sometimes indistinctly impressed in some specimens; lateral margins narrowly bordered; laterobasal portions widely depressed. Punctation dense, moderately large and deep, finer and sparser in middle and mediobasal portions; interstices between punctures in middle about as long as diameters of two-three nearest punctures. Scutellum without or with several fine punctures.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="CD0E624BFFFEFFDB7DE6FCABFDB4CAAD" blockId="2.[151,1437,151,1976]" pageId="2" pageNumber="199">Elytra somewhat flattened, slightly broader than long and slightly broadened posteriad, 1.8 times as long as pronotum; lateral margins narrowly impressed and sometimes indistinctly reflexed; hind margins widely rounded. Punctation dense, slightly larger and deeper than that on pronotum, finer, denser and coarser around scutellum, finer and sparser along suture. Hind wings fully developed.</paragraph>
|
|||
|
|
<paragraph id="CD0E624BFFFEFFDB7DE6FC3CFEA0CD7A" blockId="2.[151,1437,151,1976]" pageId="2" pageNumber="199">Metatarsus more than twice shorter than metatibia; apical metatarsomere more than twice shorter than preceding four segments.</paragraph>
|
|||
|
|
<paragraph id="CD0E624BFFFEFFDB7DE6FBF3FEF9CD11" blockId="2.[151,1437,151,1976]" pageId="2" pageNumber="199">Abdomen about as broad as elytra; middle portions of abdominal tergites IV with two transverse and moderately large tomentose spots; middle portion of abdominal tergite V with two indistinct or distinct oval tomentose spots, smaller than these in tergite IV.</paragraph>
|
|||
|
|
<paragraph id="CD0E624BFFFEFFDB7DE6FB6FFEDECC12" blockId="2.[151,1437,151,1976]" pageId="2" pageNumber="199">
|
|||
|
|
Male. Profemora and protarsomeres 1–4 wide. Posterior margin of abdominal tergite VIII somewhat rounded. Posterior margin of abdominal sternite VIII widely concave. Aedeagus with broad basal part, gradually narrowed toward wide median lobe; median lobe narrowed in preapical part, gradually narrowed toward subacute apex; parameres short and very narrow, not reaching preapical part of median lobe, without apical setae; dorsal lobes strongly sclerotized, curved in preapical portion; internal sac moderately short and wide, with two fields of short sclerotized spines and long flagellum between them, spirally folded in basal part; preapical part of flagellum widely broadened (
|
|||
|
|
<figureCitation id="558A7ECEFFFEFFDB7949FAD3FB09CC3E" box="[1128,1195,1339,1364]" captionStart="FIGURES 1–3" captionStartId="1.[151,264,1140,1164]" captionTargetBox="[182,1416,201,1082]" captionTargetId="figure-15@1.[151,1436,181,1117]" captionTargetPageId="1" captionText="FIGURES 1–3. Trichodromeus ottoi: 1–habitus (paratype), 2–aedeagus, parameral view, 3–aedeagus, lateral view. Scale bars: 1.0 mm (Fig. 1), 0.1 mm (Figs 2–3)." figureDoi="http://doi.org/10.5281/zenodo.14612162" httpUri="https://zenodo.org/record/14612162/files/figure.png" pageId="2" pageNumber="199">Fig. 2</figureCitation>
|
|||
|
|
). Lateral aspect of the aedeagus as in
|
|||
|
|
<figureCitation id="558A7ECEFFFEFFDB7C19FAB7FEDBCC12" box="[312,377,1375,1400]" captionStart="FIGURES 1–3" captionStartId="1.[151,264,1140,1164]" captionTargetBox="[182,1416,201,1082]" captionTargetId="figure-15@1.[151,1436,181,1117]" captionTargetPageId="1" captionText="FIGURES 1–3. Trichodromeus ottoi: 1–habitus (paratype), 2–aedeagus, parameral view, 3–aedeagus, lateral view. Scale bars: 1.0 mm (Fig. 1), 0.1 mm (Figs 2–3)." figureDoi="http://doi.org/10.5281/zenodo.14612162" httpUri="https://zenodo.org/record/14612162/files/figure.png" pageId="2" pageNumber="199">Fig. 3</figureCitation>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="CD0E624BFFFEFFDB7DE6FA6BFCE4CCD5" blockId="2.[151,1437,151,1976]" pageId="2" pageNumber="199">Female. Profemora and protarsomeres 1–4 narrow. Posterior margins of abdominal tergite VIII somewhat rounded. Posterior margin of abdominal sternite VIII truncate or rounded.</paragraph>
|
|||
|
|
<paragraph id="CD0E624BFFFEFFDB7DE6FA23FEE7CE69" blockId="2.[151,1437,151,1976]" pageId="2" pageNumber="199">
|
|||
|
|
<emphasis id="FFC5BE59FFFEFFDB7DE6FA23FE0ACC8E" bold="true" box="[199,424,1483,1508]" inLineHeading="true" pageId="2" pageNumber="199" reason="1">Comparative notes.</emphasis>
|
|||
|
|
Based on the general shapes of the body, antennomeres and narrow shortened parameres,
|
|||
|
|
<taxonomicName id="0AB119C8FFFEFFDB78A8FA24FF6ACF6D" authorityName="Shavrin" authorityYear="2025" class="Insecta" family="Staphylinidae" genus="Trichodromeus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="199" phylum="Arthropoda" rank="species" species="ottoi" status="sp. nov.">
|
|||
|
|
<emphasis id="FFC5BE59FFFEFFDB78A8FA24FF6ACF6D" italics="true" pageId="2" pageNumber="199">T. ottoi</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="FFC5BE59FFFEFFDB7DF0FA18FE8BCF62" bold="true" box="[209,297,1520,1544]" pageId="2" pageNumber="199">
|
|||
|
|
<taxonomicNameLabel id="E4F60322FFFEFFDB7DF0FA18FE8BCF62" box="[209,297,1520,1544]" pageId="2" pageNumber="199" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
is similar to
|
|||
|
|
<taxonomicName id="0AB119C8FFFEFFDB7C9CFA18FDF7CF6D" authorityName="Shavrin" authorityYear="2025" baseAuthorityName="Tronquet" baseAuthorityYear="1981" box="[445,597,1519,1543]" class="Insecta" family="Staphylinidae" genus="Trichodromeus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="199" phylum="Arthropoda" rank="species" species="salangensis">
|
|||
|
|
<emphasis id="FFC5BE59FFFEFFDB7C9CFA18FDF7CF6D" box="[445,597,1519,1543]" italics="true" pageId="2" pageNumber="199">T. salangensis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, also described from Hindu Kush Mts. (see below). The new species can be distinguished from the latter species by the slightly narrower pronotum, broader elytra and median lobe, broader and shorter apical part of the aedeagus, and the shape of strongly sclerotized dorsal lobes in apical portion of the median lobe. Besides that, based on the general shape and coloration of the body, and the general shape of the median lobe,
|
|||
|
|
<taxonomicName id="0AB119C8FFFEFFDB780AF9B4FAD4CF19" authorityName="Shavrin" authorityYear="2025" box="[1323,1398,1628,1651]" class="Insecta" family="Staphylinidae" genus="Trichodromeus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="199" phylum="Arthropoda" rank="species" species="ottoi" status="sp. nov.">
|
|||
|
|
<emphasis id="FFC5BE59FFFEFFDB780AF9B4FAD4CF19" box="[1323,1398,1628,1651]" italics="true" pageId="2" pageNumber="199">T. ottoi</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="FFC5BE59FFFEFFDB785DF9B4FF64CFF2" bold="true" pageId="2" pageNumber="199">
|
|||
|
|
<taxonomicNameLabel id="E4F60322FFFEFFDB785DF9B4FF64CFF2" pageId="2" pageNumber="199" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
is somewhat similar to the Middle Asian
|
|||
|
|
<taxonomicName id="0AB119C8FFFEFFDB7FA6F968FBA3CFFD" authority="(A. Bordoni, 1985)" baseAuthorityName="A. Bordoni" baseAuthorityYear="1985" box="[647,1025,1663,1688]" class="Insecta" family="Staphylinidae" genus="Trichodromeus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="199" phylum="Arthropoda" rank="species" species="ketmeniensis">
|
|||
|
|
<emphasis id="FFC5BE59FFFEFFDB7FA6F968FC88CFFD" box="[647,810,1663,1687]" italics="true" pageId="2" pageNumber="199">T. ketmeniensis</emphasis>
|
|||
|
|
(A. Bordoni, 1985)
|
|||
|
|
</taxonomicName>
|
|||
|
|
, known from
|
|||
|
|
<collectingCountry id="B5A622DBFFFEFFDB79B6F997FAB4CFFD" box="[1175,1302,1663,1687]" name="Kazakhstan" pageId="2" pageNumber="199">Kazakhstan</collectingCountry>
|
|||
|
|
,
|
|||
|
|
<collectingCountry id="B5A622DBFFFEFFDB7803F968FA3ECFF2" box="[1314,1436,1664,1688]" name="Kyrgyzstan" pageId="2" pageNumber="199">Kyrgyzstan</collectingCountry>
|
|||
|
|
and
|
|||
|
|
<collectingCountry id="B5A622DBFFFEFFDB7DE2F94BFEF4CFD1" box="[195,342,1699,1723]" name="Turkmenistan" pageId="2" pageNumber="199">Turkmenistan</collectingCountry>
|
|||
|
|
(
|
|||
|
|
<bibRefCitation id="A9201FBAFFFEFFDB7C42F94BFE54CFD6" author="Shavrin, A. V." box="[355,502,1699,1724]" pageId="2" pageNumber="199" pagination="1 - 62" refId="ref2446" refString="Shavrin, A. V. (2024) Review of Hygrogeus Mulsant & Rey, 1880 and Trichodromeus Luze, 1903, with descriptions of three new genera from the eastern Palaearctic Region (Coleoptera: Staphylinidae: Anthophagini: Omaliinae). Zootaxa, 5531 (1), 1 - 62. https: // doi. org / 10.11646 / zootaxa. 5531.1.1" type="journal article" year="2024">Shavrin 2024</bibRefCitation>
|
|||
|
|
), but differs from it by the smaller body, significantly shorter antennomeres 3–10, longer temples, slightly narrower and longer elytra, significantly shorter and narrower parameres, and details of the morphology of the aedeagus.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="85AB31C0FFFEFFDB7DE6F8E7FDDBCE21" pageId="2" pageNumber="199" type="distribution">
|
|||
|
|
<paragraph id="CD0E624BFFFEFFDB7DE6F8E7FDDBCE21" blockId="2.[151,1437,151,1976]" pageId="2" pageNumber="199">
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<emphasis id="FFC5BE59FFFEFFDB7DE6F8E7FEF8CE4D" bold="true" box="[199,346,1807,1831]" inLineHeading="true" pageId="2" pageNumber="199" reason="1">Distribution.</emphasis>
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<taxonomicName id="0AB119C8FFFEFFDB7C42F8E7FD9DCE4D" authorityName="Shavrin" authorityYear="2025" box="[355,575,1807,1831]" class="Insecta" family="Staphylinidae" genus="Trichodromeus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="199" phylum="Arthropoda" rank="species" species="ottoi" status="sp. nov.">
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<emphasis id="FFC5BE59FFFEFFDB7C42F8E7FD9DCE4D" box="[355,575,1807,1831]" italics="true" pageId="2" pageNumber="199">Trichodromeus ottoi</emphasis>
|
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</taxonomicName>
|
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<emphasis id="FFC5BE59FFFEFFDB7F68F8F8FD00CE42" bold="true" box="[585,674,1808,1832]" pageId="2" pageNumber="199">
|
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|
|
<taxonomicNameLabel id="E4F60322FFFEFFDB7F68F8F8FD00CE42" box="[585,674,1808,1832]" pageId="2" pageNumber="199" rank="species">sp. nov.</taxonomicNameLabel>
|
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|
|
</emphasis>
|
|||
|
|
is known only from the
|
|||
|
|
<typeStatus id="120ADCE9FFFEFFDB7E9DF8F8FC48CE42" box="[956,1002,1808,1832]" pageId="2" pageNumber="199">type</typeStatus>
|
|||
|
|
locality in Anjuman (=Anjoman) Pass, Hindu Kush Mts., north-eastern
|
|||
|
|
<collectingCountry id="B5A622DBFFFEFFDB7CCEF8DBFDD7CE21" box="[495,629,1843,1867]" name="Afghanistan" pageId="2" pageNumber="199">Afghanistan</collectingCountry>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="85AB31C0FFFEFFDB7DE6F8B0FABFCEDD" pageId="2" pageNumber="199" type="etymology">
|
|||
|
|
<paragraph id="CD0E624BFFFEFFDB7DE6F8B0FD49CEF9" blockId="2.[151,1437,151,1976]" pageId="2" pageNumber="199">
|
|||
|
|
<emphasis id="FFC5BE59FFFEFFDB7DE6F8B0FEEACE1A" bold="true" box="[199,328,1880,1904]" inLineHeading="true" pageId="2" pageNumber="199" reason="1">Etymology.</emphasis>
|
|||
|
|
Patronymic, the species is dedicated to the memory of Otto Scheerpeltz (1888–1975), who marked the studied specimens as a new species but not described it.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="CD0E624BFFFEFFDB7DE6F877FABFCEDD" blockId="2.[151,1437,151,1976]" box="[199,1309,1951,1976]" pageId="2" pageNumber="199">
|
|||
|
|
<emphasis id="FFC5BE59FFFEFFDB7DE6F877FEE7CEDD" bold="true" box="[199,325,1951,1975]" inLineHeading="true" pageId="2" pageNumber="199" reason="1">Bionomics.</emphasis>
|
|||
|
|
Specimens were collected at elevation
|
|||
|
|
<quantity id="0A49CFAEFFFEFFDB7FC8F848FC98CEDD" box="[745,826,1952,1976]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="2" pageNumber="199" unit="m" value="4000.0">4000 m</quantity>
|
|||
|
|
a.s.l. Detailed bionomical data are unknown.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
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|
|
</document>
|