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<document id="67C5CF8F18D125252685F6995E61C000" ID-CLB-Dataset="9693" ID-DOI="10.5852/ejt.2020.721.1127" ID-GBIF-Dataset="0cd61b23-1089-4c1b-bfc0-487887abad84" ID-Zenodo-Dep="4122118" ID-ZooBank="89E62EF8-2E45-4C59-94B7-6A5603E8939B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1603474447689" checkinUser="valdenar" docAuthor="Telnov, Dmitry" docDate="2020" docId="1A3787F69D0E9248FDE4FEB6CBCE2B94" docLanguage="en" docName="ejt-721_telnov.pdf" docOrigin="European Journal of Taxonomy 721" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Maechidius aiyura Telnov 2020, sp. nov." docType="treatment" docUuid="568C06B4-A3C4-41A1-B924-4E1232EF8A3B" docUuidSource="ZooBank" docVersion="10" lastPageNumber="20" masterDocId="E60EFF8E9D1C925BFF93FFBEC8562862" masterDocTitle="A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line" masterLastPageNumber="210" masterPageNumber="1" pageNumber="19" updateTime="1698846101371" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="6E7A12463D50156E1DD229961449C340">
<mods:title id="001BFFED7D6AC998BEDD8B0CB1BD3CF4">A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line</mods:title>
</mods:titleInfo>
<mods:name id="DF1DE8BB38D3A8250CA2B7C7EFA1DB4E" type="personal">
<mods:role id="A5A1F5A8EDC0C9FBE4B75D8AC4205367">
<mods:roleTerm id="559F3AA74F13DB325E711926AF9D3EB4">Author</mods:roleTerm>
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<mods:namePart id="C1645C6CD6BA5C25F68CC20015C5B8F4">Telnov, Dmitry</mods:namePart>
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<mods:typeOfResource id="EFFA63852AC80C7C511E2A7F5B707FCC">text</mods:typeOfResource>
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<mods:title id="A67D33221FBF0FB3D404373693263750">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="3C19486BBA5B4236D5788765F1C31F41">
<mods:date id="F93BB16698A118F8886AC4773838676D">2020</mods:date>
<mods:detail id="76EE199923A035ED0DF0E40056A85C08" type="pubDate">
<mods:number id="034C9848643B04CA043FFB0D74580D73">2020-10-19</mods:number>
</mods:detail>
<mods:detail id="9DF3E1E2B09F76F5D201963169A20F4D" type="volume">
<mods:number id="DF333581A7DAB18C4A7FF05F86CAA17B">721</mods:number>
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<mods:classification id="BA4DF941D0652BFB0A06D1A3F525C5C6">journal article</mods:classification>
<mods:identifier id="C478727100099237602DF8B4BD92FBF2" type="CLB-Dataset">9693</mods:identifier>
<mods:identifier id="C4BFC9759AA2B514A87E58F2051E1FC8" type="DOI">10.5852/ejt.2020.721.1127</mods:identifier>
<mods:identifier id="EA8F41B9EB0B0C54F106FBFEEDD04A0B" type="GBIF-Dataset">0cd61b23-1089-4c1b-bfc0-487887abad84</mods:identifier>
<mods:identifier id="0DD0B9BB0B2E64A006367DFBE07093AB" type="Zenodo-Dep">4122118</mods:identifier>
<mods:identifier id="B6CF2B9E5F209EF84BC8DB963AEC4BC1" type="ZooBank">89E62EF8-2E45-4C59-94B7-6A5603E8939B</mods:identifier>
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<treatment id="1A3787F69D0E9248FDE4FEB6CBCE2B94" ID-DOI="http://doi.org/10.5281/zenodo.4344333" ID-GBIF-Taxon="168648181" ID-Zenodo-Dep="4344333" LSID="urn:lsid:zoobank.org:act:568C06B4-A3C4-41A1-B924-4E1232EF8A3B" httpUri="http://treatment.plazi.org/id/1A3787F69D0E9248FDE4FEB6CBCE2B94" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
<subSubSection id="DA84656B9D0E9249FDE4FEB6CBEB2941" box="[631,957,264,291]" pageId="18" pageNumber="19" type="nomenclature">
<paragraph id="922136E09D0E9249FDE4FEB6CBEB2941" blockId="18.[367,1221,264,360]" box="[631,957,264,291]" pageId="18" pageNumber="19">
<heading id="C969818C9D0E9249FDE4FEB6CBEB2941" box="[631,957,264,291]" centered="true" fontSize="11" level="2" pageId="18" pageNumber="19" reason="2">
<taxonomicName id="559E4D639D0E9249FDE4FEB6CB0C2941" ID-CoL="7X3W3" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[631,858,264,291]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="aiyura" status="sp. nov.">
<emphasis id="A0EAEAF29D0E9249FDE4FEB6CB0C2941" bold="true" box="[631,858,264,291]" italics="true" pageId="18" pageNumber="19">Maechidius aiyura</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D0E9249FCF1FEB7CBEB2941" box="[866,957,265,291]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D0E9249FEFCFE95CC7F290A" pageId="18" pageNumber="19" type="description">
<paragraph id="922136E09D0E9249FEFCFE95CC932924" blockId="18.[367,1221,264,360]" box="[367,1221,299,326]" pageId="18" pageNumber="19">
<uri id="E60F3AE29D0E9249FEFCFE95CC932924" box="[367,1221,299,326]" pageId="18" pageNumber="19">
urn:lsid:zoobank.org:act:
<uuid id="E6380C359D0E9249FD07FE95CC932924" box="[660,1221,299,326]" pageId="18" pageNumber="19">568C06B4-A3C4-41A1-B924-4E1232EF8A3B</uuid>
</uri>
</paragraph>
<paragraph id="922136E09D0E9249FD98FEF0CC7F290A" blockId="18.[367,1221,264,360]" box="[523,1065,333,360]" pageId="18" pageNumber="19">
<figureCitation id="0AA52A659D0E9249FD98FEF0CA32290A" box="[523,612,334,360]" captionStart="Figs 18–21" captionStartId="17.[188,240,1928,1954]" captionTargetBox="[353,1214,268,1904]" captionTargetId="figure-12@17.[346,1240,250,1910]" captionTargetPageId="17" captionText="Figs 18–21. Papuan Maechidius Macleay, 1819, habitus, dorsal and lateral views. 18. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 19. M. aiyura sp. nov., holotype, ♂ (BMNH). 20–21. M. alesbezdeki sp. nov. ♂ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122130" httpUri="https://zenodo.org/record/4122130/files/figure.png" pageId="18" pageNumber="19">Figs 19</figureCitation>
,
<figureCitation id="0AA52A659D0E9249FDE2FEF0CAC7290A" box="[625,657,334,360]" captionStart="Figs 97–108" captionStartId="137.[189,241,1786,1812]" captionTargetBox="[351,1228,270,1756]" captionTargetId="figure-127@137.[351,1237,265,1772]" captionTargetPageId="137" captionText="Figs 97–108. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal view. 97. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 98. M. aiyura sp. nov., holotype, ♂ (BMNH). 99. M. alesbezdeki sp. nov., holotype, ♂ (NME). 100. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 101. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 102. M. bintang sp. nov., holotype, ♂ (NME). 103. M. awu sp. nov., holotype, ♂ (IECA). 104. Same, paratype, ♀ (IECA). 105. M. boessnecki sp. nov., holotype, ♂ (SNSD). 106. M. babyrousa sp. nov., holotype, ♂ (BMNH). 107. Same, paratype, ♀ (BMNH). 108. M. caperatus sp. nov., holotype, ♂ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122173" httpUri="https://zenodo.org/record/4122173/files/figure.png" pageId="18" pageNumber="19">98</figureCitation>
,
<figureCitation id="0AA52A659D0E9249FD0EFEF0CA9D290A" box="[669,715,334,360]" captionStart="Figs 193–207" captionStartId="145.[189,241,1716,1742]" captionTargetBox="[385,1203,267,1681]" captionTargetId="figure-159@145.[385,1203,265,1694]" captionTargetPageId="145" captionText="Figs 193–207. Papuan and Wallacean Maechidius Macleay, 1819, head (193–196) and pronotum (197– 207), dorsal view. 193. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 194. Same, ♂ (SNSD). 195. M. yamdena sp. nov., paratype, ♂ (BMNH). 196. Same, paratype, ♀ (BMNH). 197. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 198. M. aiyura sp. nov., holotype, ♂ (BMNH). 199. M. alesbezdeki sp. nov., holotype, ♂ (NME). 200. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 201. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 202. M. awu sp. nov., holotype, ♂ (IECA). 203. M. babyrousa sp. nov., holotype, ♂ (BMNH). 204. M. bintang sp. nov., holotype, ♂ (NME). 205. M. boessnecki sp. nov., holotype, ♂ (SNSD). 206. M. brocki sp. nov., holotype, ♂ (BMNH). 207. M. crypticus sp. nov., paratype, ♀ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122189" httpUri="https://zenodo.org/record/4122189/files/figure.png" pageId="18" pageNumber="19">198</figureCitation>
,
<figureCitation id="0AA52A659D0E9249FD44FEF0CB50290A" box="[727,774,334,360]" captionStart="Figs 286–301" captionStartId="152.[189,241,1716,1742]" captionTargetBox="[364,1224,266,1689]" captionTargetId="figure-172@152.[364,1224,266,1695]" captionTargetPageId="152" captionText="Figs 286–301. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 286. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 287. M. aiyura sp. nov., holotype, ♂ (BMNH). 288. M. alesbezdeki sp. nov., holotype, ♂ (NME). 289. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 290. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 291. M. paupianus Heller, 1910, ♂ (BMNH). 292. M. awu sp. nov., holotype, ♂ (IECA). 293. M. babyrousa sp. nov., holotype, ♂ (BMNH). 294. M. bintang sp. nov., holotype, ♂ (NME). 295. M. boessnecki sp. nov., holotype, ♂ (SNSD). 296. M. brocki sp. nov., holotype, ♂ (BMNH). 297. M. caperatus sp. nov., holotype, ♂ (NME). 298. M. ciliatus sp. nov., holotype, ♂ (IECA). 299. M. crypticus sp. nov., paratype, ♀ (BMNH). 300. M. dani sp. nov., holotype, ♂ (IECA). 301. M. deltouri sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122206" httpUri="https://zenodo.org/record/4122206/files/figure.png" pageId="18" pageNumber="19">287</figureCitation>
, 360, 452, 535, 570–572
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D0E9249FF2EFE30CD202A02" pageId="18" pageNumber="19" type="diagnosis">
<paragraph id="922136E09D0E9249FF2EFE30C99F29CA" blockId="18.[189,457,398,424]" box="[189,457,398,424]" pageId="18" pageNumber="19">
<heading id="C969818C9D0E9249FF2EFE30C99F29CA" bold="true" box="[189,457,398,424]" fontSize="11" level="3" pageId="18" pageNumber="19" reason="3">
<emphasis id="A0EAEAF29D0E9249FF2EFE30C99F29CA" bold="true" box="[189,457,398,424]" pageId="18" pageNumber="19">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D0E9249FF2EFE02CD202A02" blockId="18.[189,1399,444,608]" pageId="18" pageNumber="19">
Undoubtedly related to
<taxonomicName id="559E4D639D0E9249FE46FE03CBC029B4" authority="Heller, 1914" authorityName="Heller" authorityYear="1914" box="[469,918,444,470]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="interruptocarinulatus">
<emphasis id="A0EAEAF29D0E9249FE46FE03CAAC29B4" box="[469,762,444,470]" italics="true" pageId="18" pageNumber="19">M. interruptocarinulatus</emphasis>
<bibRefCitation id="F60F4B119D0E9249FC92FE02CBC029B4" author="Heller K. M." box="[769,918,444,470]" pageId="18" pageNumber="19" pagination="615 - 666" refId="ref98213" refString="Heller K. M. 1914. Coleoptera. Nova Guinea. Resultats de l'Expedition scientifique neerlandaise a la Nouvelle-Guinee en 1907 et 1909 sous les auspices du Dr. H. A. Lorentz. Zoologie 9: 615 - 666." type="journal article" year="1914">Heller, 1914</bibRefCitation>
</taxonomicName>
, but clearly different in lateral outline of pronotum in dorsal view (more regularly rounded in
<taxonomicName id="559E4D639D0E9249FCB8FE61CC19299A" authorityName="Heller" authorityYear="1914" box="[811,1103,478,504]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="interruptocarinulatus">
<emphasis id="A0EAEAF29D0E9249FCB8FE61CC19299A" box="[811,1103,478,504]" italics="true" pageId="18" pageNumber="19">M. interruptocarinulatus</emphasis>
</taxonomicName>
, constricted anteriad and posteriad in
<taxonomicName id="559E4D639D0E9249FEDCFDBFC9922A78" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[335,452,513,538]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="aiyura" status="sp. nov.">
<emphasis id="A0EAEAF29D0E9249FEDCFDBFC9922A78" box="[335,452,513,538]" italics="true" pageId="18" pageNumber="19">M. aiyura</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D0E9249FE58FDBFCA7F2A79" box="[459,553,513,539]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
), stronger sinuous lateral margins of labroclypeus, glabrous intervening spaces of elytra (covered with microscopical velvety pubescence in
<taxonomicName id="559E4D639D0E9249FC71FD9ACD502A5F" authorityName="Heller" authorityYear="1914" box="[994,1286,547,573]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="interruptocarinulatus">
<emphasis id="A0EAEAF29D0E9249FC71FD9ACD502A5F" box="[994,1286,547,573]" italics="true" pageId="18" pageNumber="19">M. interruptocarinulatus</emphasis>
</taxonomicName>
) and less distinct (narrower, more flat)elytral longitudinal carinae.Males are unknown for
<taxonomicName id="559E4D639D0E9249FBDCFDF8CD262A3D" authorityName="Heller" authorityYear="1914" box="[1103,1392,581,607]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="interruptocarinulatus">
<emphasis id="A0EAEAF29D0E9249FBDCFDF8CD262A3D" box="[1103,1392,581,607]" italics="true" pageId="18" pageNumber="19">M. interruptocarinulatus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D0E9249FF2EFD38CD212AAF" box="[189,1399,646,718]" pageId="18" pageNumber="19" type="etymology">
<paragraph id="922136E09D0E9249FF2EFD38C9122AC2" blockId="18.[189,324,646,672]" box="[189,324,646,672]" pageId="18" pageNumber="19">
<heading id="C969818C9D0E9249FF2EFD38C9122AC2" bold="true" box="[189,324,646,672]" fontSize="11" level="3" pageId="18" pageNumber="19" reason="3">
<emphasis id="A0EAEAF29D0E9249FF2EFD38C9122AC2" bold="true" box="[189,324,646,672]" pageId="18" pageNumber="19">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D0E9249FF2EFD0DCD212AAF" blockId="18.[189,1399,691,718]" box="[189,1399,691,718]" pageId="18" pageNumber="19">
Toponymic.The name derives from the Aiyura Valley, the
<typeStatus id="4D2588429D0E9249FCC7FD0ACBD12AAC" box="[852,903,692,718]" pageId="18" pageNumber="19">type</typeStatus>
locality of this species. Noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D0E9249FF2EFD4AC9242BA7" pageId="18" pageNumber="19" type="materials_examined">
<paragraph id="922136E09D0E9249FF2EFD4AC93A2B6C" blockId="18.[189,364,756,782]" box="[189,364,756,782]" pageId="18" pageNumber="19">
<heading id="C969818C9D0E9249FF2EFD4AC93A2B6C" bold="true" box="[189,364,756,782]" fontSize="11" level="3" pageId="18" pageNumber="19" reason="3">
<emphasis id="A0EAEAF29D0E9249FF2EFD4AC93A2B6C" bold="true" box="[189,364,756,782]" pageId="18" pageNumber="19">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D0E9249FF79FC9CC90A2B5E" blockId="18.[189,1399,802,965]" box="[234,348,802,828]" pageId="18" pageNumber="19">
<emphasis id="A0EAEAF29D0E9249FF79FC9CC90A2B5E" bold="true" box="[234,348,802,828]" pageId="18" pageNumber="19">
<typeStatus id="4D2588429D0E9249FF79FC9CC90A2B5E" box="[234,348,802,828]" pageId="18" pageNumber="19" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="922136E09D0E9249FF2EFCFAC9242BA7" blockId="18.[189,1399,802,965]" pageId="18" pageNumber="19">
<materialsCitation id="22F63CBD9D0E9249FF2EFCFAC9242BA7" ID-GBIF-Occurrence="2898878485" collectionCode="HAES, COLL, BMNH" collectorName="E. S. C. Smith Attr." country="Papua New Guinea" determinerName="R. B. Madge" location="Aiyura" municipality="Papua New Guinea East. High. Prov." pageId="18" pageNumber="19" specimenCount="1562" specimenCount-male="1561" typeStatus="holotype">
<collectingCountry id="EA8976709D0E9249FF2EFCFAC9B52B3C" box="[189,483,836,862]" name="Papua New Guinea" pageId="18" pageNumber="19">PAPUA NEW GUINEA</collectingCountry>
•
<specimenCount id="8498FD699D0E9249FD95FCFACA482B3C" box="[518,542,836,862]" pageId="18" pageNumber="19" type="male">♂</specimenCount>
; “
<collectingMunicipality id="7245AC9A9D0E9249FDA8FCFACBA52B3C" box="[571,1011,836,862]" country="Papua New Guinea" name="Papua New Guinea East. High. Prov." pageId="18" pageNumber="19">Papua New Guinea East.High.Prov.</collectingMunicipality>
, [p]
<collectionCode id="F48FAE259D0E9249FBA1FCFACC292B3C" box="[1074,1151,836,862]" pageId="18" pageNumber="19">HAES</collectionCode>
<location id="9741603B9D0E9249FB19FCFACC8D2B3C" LSID="urn:lsid:plazi:treatment:1A3787F69D0E9248FDE4FEB6CBCE2B94:9741603B9D0E9249FB19FCFACC8D2B3C" box="[1162,1243,836,862]" country="Papua New Guinea" municipality="Papua New Guinea East. High. Prov." name="Aiyura" pageId="18" pageNumber="19">Aiyura</location>
<specimenCount id="8498FD699D0E9249FB7BFCFACD6B2B3D" box="[1256,1341,836,863]" pageId="18" pageNumber="19" type="male">1560M</specimenCount>
30// NOV//1976 [h]
<collectorName id="3F6B53369D0E9249FEECFCD8CA192BE2" box="[383,591,870,896]" pageId="18" pageNumber="19">E.S.C.Smith Attr.</collectorName>
to U.V. light [p] //A327 [p] // 26961 [p]// C.I.E.
<collectionCode id="F48FAE259D0E9249FB18FCD8CC8D2BE2" box="[1163,1243,870,896]" pageId="18" pageNumber="19">COLL</collectionCode>
. A. [p] 9461 [h] [red label] //
<location id="9741603B9D0E9249FEE9FC37CA832BC1" LSID="urn:lsid:plazi:treatment:1A3787F69D0E9248FDE4FEB6CBCE2B94:9741603B9D0E9249FEE9FC37CA832BC1" box="[378,725,904,931]" country="Papua New Guinea" municipality="Papua New Guinea East. High. Prov." name="Pres by Comm Inst Ent B. M." pageId="18" pageNumber="19">Pres by Comm Inst Ent B.M.</location>
1977–1 [p] //
<taxonomicName id="559E4D639D0E9249FCEBFC36CC572BC0" authorityName="Macleay" authorityYear="1819" box="[888,1025,904,930]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="A0EAEAF29D0E9249FCEBFC36CC572BC0" box="[888,1025,904,930]" italics="true" pageId="18" pageNumber="19">
<location id="9741603B9D0E9249FCEBFC36CC572BC0" LSID="urn:lsid:plazi:treatment:1A3787F69D0E9248FDE4FEB6CBCE2B94:9741603B9D0E9249FCEBFC36CC572BC0" box="[888,1025,904,930]" country="Papua New Guinea" municipality="Papua New Guinea East. High. Prov." name="Maechidius" pageId="18" pageNumber="19">Maechidius</location>
</emphasis>
</taxonomicName>
<specimenCount id="8498FD699D0E9249FB9AFC37CC7C2BC1" box="[1033,1066,905,931]" pageId="18" pageNumber="19" type="generic">sp.</specimenCount>
[h] det.
<determinerName id="67FDDA249D0E9249FB02FC36CD4D2BC0" box="[1169,1307,904,930]" pageId="18" pageNumber="19">R.B.Madge</determinerName>
, 197[p] 7 [h]”;
<collectionCode id="F48FAE259D0E9249FE81FC15C9382BA7" box="[274,366,939,965]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="18" pageNumber="19">BMNH</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D0E9248FF2EFC55C9FD2B7B" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19" type="description">
<paragraph id="922136E09D0E9249FF2EFC55C9182C67" blockId="18.[189,334,1003,1029]" box="[189,334,1003,1029]" pageId="18" pageNumber="19">
<heading id="C969818C9D0E9249FF2EFC55C9182C67" bold="true" box="[189,334,1003,1029]" fontSize="11" level="3" pageId="18" pageNumber="19" reason="3">
<emphasis id="A0EAEAF29D0E9249FF2EFC55C9182C67" bold="true" box="[189,334,1003,1029]" pageId="18" pageNumber="19">Description</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D0E9249FF2EFBA4CD212C34" blockId="18.[189,1399,1049,1110]" pageId="18" pageNumber="19">
MEASUREMENTS. Total body length
<quantity id="55669B059D0E9249FDC2FBA7CAEA2C56" box="[593,700,1049,1076]" metricMagnitude="-3" metricUnit="m" metricValue="9.15" pageId="18" pageNumber="19" unit="mm" value="9.15">9.15 mm</quantity>
. Head
<quantity id="55669B059D0E9249FC9EFBA7CB232C56" box="[781,885,1049,1076]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="18" pageNumber="19" unit="mm" value="1.6">1.60 mm</quantity>
long, across eyes
<quantity id="55669B059D0E9249FBD8FBA7CCE22C56" box="[1099,1204,1049,1076]" metricMagnitude="-3" metricUnit="m" metricValue="2.15" pageId="18" pageNumber="19" unit="mm" value="2.15">2.15 mm</quantity>
wide. Pronotum 2.00 mm long, maximum width
<quantity id="55669B059D0E9249FDA7FB85CAC92C34" box="[564,671,1083,1110]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="18" pageNumber="19" unit="mm" value="3.2">3.20 mm</quantity>
. Elytral length
<quantity id="55669B059D0E9249FCC2FB85CBED2C34" box="[849,955,1083,1110]" metricMagnitude="-3" metricUnit="m" metricValue="5.55" pageId="18" pageNumber="19" unit="mm" value="5.55">5.55 mm</quantity>
, maximum combined width
<quantity id="55669B059D0E9249FA9BFB85CD252C34" box="[1288,1395,1083,1110]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="18" pageNumber="19" unit="mm" value="4.1">4.10 mm</quantity>
.
</paragraph>
<paragraph id="922136E09D0E9248FF2EFBC0CC6A2AC9" blockId="18.[189,1399,1149,2025]" lastBlockId="19.[189,1399,265,683]" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
Dorsum uniformly black-brown, venter, legs and labroclypeus castaneous brown. Head transverse, glossy dorsally and ventrally, flattened dorsally. Compound eye large, occupying about half side of head. Male labroclypeus broadly emarginate on anterior margin (
<figureCitation id="0AA52A659D0E9249FCEBFB7ACB992CBC" box="[888,975,1220,1246]" captionStart="Figs 97–108" captionStartId="137.[189,241,1786,1812]" captionTargetBox="[351,1228,270,1756]" captionTargetId="figure-127@137.[351,1237,265,1772]" captionTargetPageId="137" captionText="Figs 97–108. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal view. 97. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 98. M. aiyura sp. nov., holotype, ♂ (BMNH). 99. M. alesbezdeki sp. nov., holotype, ♂ (NME). 100. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 101. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 102. M. bintang sp. nov., holotype, ♂ (NME). 103. M. awu sp. nov., holotype, ♂ (IECA). 104. Same, paratype, ♀ (IECA). 105. M. boessnecki sp. nov., holotype, ♂ (SNSD). 106. M. babyrousa sp. nov., holotype, ♂ (BMNH). 107. Same, paratype, ♀ (BMNH). 108. M. caperatus sp. nov., holotype, ♂ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122173" httpUri="https://zenodo.org/record/4122173/files/figure.png" pageId="18" pageNumber="19">Fig. 98</figureCitation>
), its lateral margins sinuous in both dorsal and lateral views. Anterolateral angles of male labroclypeus slightly protruding anteriad, obtuse in dorsal view, bent up at ~90° to axis of frons in lateral view. Upper- and underside of labroclypeus with moderately long setae along anterior and lateral margins. Canthus nearly straight in dorsal view. Anterior and lateral margins of labroclypeus smooth. Head dorsal punctures circular to ovoid, deep and dense. Inner margin of some punctures with delicate membrane covered with microscopical velvety pubescence. Intervening spaces glossy, variably large. Canthus covered with microscopical velvety pubescence. Dirty-yellow inconspicuous suberect seta rises from anterior margin of each puncture, generally surpassing length of corresponding puncture. Antenna 9-segmented, club 3-lamellate. Scape large, with bulbous predistal projection on dorsal side, provided with two very long setae near distal margin. Pronotum strongly transverse, glossy dorsally and laterally, its anterior margin sinuous with median part (slightly) and anterolateral margins (stronger) protruding anteriad. Basal margin of pronotum broadly rounded. Lateral margin of pronotum rounded, more strongly constricted towards base than anteriad, crenulate all along (
<figureCitation id="0AA52A659D0E9249FD13F92ECAB02EC8" box="[640,742,1680,1706]" captionStart="Figs 193–207" captionStartId="145.[189,241,1716,1742]" captionTargetBox="[385,1203,267,1681]" captionTargetId="figure-159@145.[385,1203,265,1694]" captionTargetPageId="145" captionText="Figs 193–207. Papuan and Wallacean Maechidius Macleay, 1819, head (193–196) and pronotum (197– 207), dorsal view. 193. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 194. Same, ♂ (SNSD). 195. M. yamdena sp. nov., paratype, ♂ (BMNH). 196. Same, paratype, ♀ (BMNH). 197. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 198. M. aiyura sp. nov., holotype, ♂ (BMNH). 199. M. alesbezdeki sp. nov., holotype, ♂ (NME). 200. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 201. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 202. M. awu sp. nov., holotype, ♂ (IECA). 203. M. babyrousa sp. nov., holotype, ♂ (BMNH). 204. M. bintang sp. nov., holotype, ♂ (NME). 205. M. boessnecki sp. nov., holotype, ♂ (SNSD). 206. M. brocki sp. nov., holotype, ♂ (BMNH). 207. M. crypticus sp. nov., paratype, ♀ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122189" httpUri="https://zenodo.org/record/4122189/files/figure.png" pageId="18" pageNumber="19">Fig. 198</figureCitation>
). Inconspicuous suberect scale-like seta present between every two crenulae. Lateral margin of pronotum slightly sinuous in lateral view. Areas at antero- and posterolateral angles of pronotum slightly dorsally impressed. Pronotal punctures ovoid, very deep and dense, filled with microscopical velvety pubescence (
<figureCitation id="0AA52A659D0E9249FCA1F944CBDB2F76" box="[818,909,1786,1812]" captionStart="Figs 18–21" captionStartId="17.[188,240,1928,1954]" captionTargetBox="[353,1214,268,1904]" captionTargetId="figure-12@17.[346,1240,250,1910]" captionTargetPageId="17" captionText="Figs 18–21. Papuan Maechidius Macleay, 1819, habitus, dorsal and lateral views. 18. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 19. M. aiyura sp. nov., holotype, ♂ (BMNH). 20–21. M. alesbezdeki sp. nov. ♂ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122130" httpUri="https://zenodo.org/record/4122130/files/figure.png" pageId="18" pageNumber="19">Figs 19</figureCitation>
,
<figureCitation id="0AA52A659D0E9249FC08F944CB9D2F76" box="[923,971,1786,1812]" captionStart="Figs 193–207" captionStartId="145.[189,241,1716,1742]" captionTargetBox="[385,1203,267,1681]" captionTargetId="figure-159@145.[385,1203,265,1694]" captionTargetPageId="145" captionText="Figs 193–207. Papuan and Wallacean Maechidius Macleay, 1819, head (193–196) and pronotum (197– 207), dorsal view. 193. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 194. Same, ♂ (SNSD). 195. M. yamdena sp. nov., paratype, ♂ (BMNH). 196. Same, paratype, ♀ (BMNH). 197. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 198. M. aiyura sp. nov., holotype, ♂ (BMNH). 199. M. alesbezdeki sp. nov., holotype, ♂ (NME). 200. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 201. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 202. M. awu sp. nov., holotype, ♂ (IECA). 203. M. babyrousa sp. nov., holotype, ♂ (BMNH). 204. M. bintang sp. nov., holotype, ♂ (NME). 205. M. boessnecki sp. nov., holotype, ♂ (SNSD). 206. M. brocki sp. nov., holotype, ♂ (BMNH). 207. M. crypticus sp. nov., paratype, ♀ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122189" httpUri="https://zenodo.org/record/4122189/files/figure.png" pageId="18" pageNumber="19">198</figureCitation>
). Intervening spaces glossy, as large as to smaller than punctures. Pronotal setae inconspicuous, appressed to suberect; each seta rises from anterior margin of each puncture, not surpassing length of corresponding puncture. Basal and lateral margins of pronotum covered with microscopical velvety pubescence. Posterolateral angles densely covered with long scale-like setae. Hypomeron sinuous, very long setose on anterior margin opposite to compound eye, flange-like. Antennal pocket deep. Scutellar shield covered with microscopical velvety pubescence, narrowly rounded apically. Elytron glossy dorsally, maximum width across midlength, with obsolete humerus and distinct omoplate. Tracks of three flat glossy longitudinal carinae on each elytron; sutural stria not indicated.Elytral punctures delicate, ovoid, moderately deep, generally smaller than those on forebody, arranged in irregular longitudinal rows (
<figureCitation id="0AA52A659D0F9248FCA1FEEECBCC2908" box="[818,922,336,362]" captionStart="Figs 286–301" captionStartId="152.[189,241,1716,1742]" captionTargetBox="[364,1224,266,1689]" captionTargetId="figure-172@152.[364,1224,266,1695]" captionTargetPageId="152" captionText="Figs 286–301. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 286. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 287. M. aiyura sp. nov., holotype, ♂ (BMNH). 288. M. alesbezdeki sp. nov., holotype, ♂ (NME). 289. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 290. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 291. M. paupianus Heller, 1910, ♂ (BMNH). 292. M. awu sp. nov., holotype, ♂ (IECA). 293. M. babyrousa sp. nov., holotype, ♂ (BMNH). 294. M. bintang sp. nov., holotype, ♂ (NME). 295. M. boessnecki sp. nov., holotype, ♂ (SNSD). 296. M. brocki sp. nov., holotype, ♂ (BMNH). 297. M. caperatus sp. nov., holotype, ♂ (NME). 298. M. ciliatus sp. nov., holotype, ♂ (IECA). 299. M. crypticus sp. nov., paratype, ♀ (BMNH). 300. M. dani sp. nov., holotype, ♂ (IECA). 301. M. deltouri sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122206" httpUri="https://zenodo.org/record/4122206/files/figure.png" pageId="19" pageNumber="20">Fig. 287</figureCitation>
). Inner margin of each puncture covered with microscopical velvety pubescence (
<figureCitation id="0AA52A659D0F9248FD32FECACB5D29EC" box="[673,779,372,398]" captionStart="Figs 286–301" captionStartId="152.[189,241,1716,1742]" captionTargetBox="[364,1224,266,1689]" captionTargetId="figure-172@152.[364,1224,266,1695]" captionTargetPageId="152" captionText="Figs 286–301. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 286. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 287. M. aiyura sp. nov., holotype, ♂ (BMNH). 288. M. alesbezdeki sp. nov., holotype, ♂ (NME). 289. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 290. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 291. M. paupianus Heller, 1910, ♂ (BMNH). 292. M. awu sp. nov., holotype, ♂ (IECA). 293. M. babyrousa sp. nov., holotype, ♂ (BMNH). 294. M. bintang sp. nov., holotype, ♂ (NME). 295. M. boessnecki sp. nov., holotype, ♂ (SNSD). 296. M. brocki sp. nov., holotype, ♂ (BMNH). 297. M. caperatus sp. nov., holotype, ♂ (NME). 298. M. ciliatus sp. nov., holotype, ♂ (IECA). 299. M. crypticus sp. nov., paratype, ♀ (BMNH). 300. M. dani sp. nov., holotype, ♂ (IECA). 301. M. deltouri sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122206" httpUri="https://zenodo.org/record/4122206/files/figure.png" pageId="19" pageNumber="20">Fig. 287</figureCitation>
). Intervening spaces glossy, variably large. Setae of elytra inconspicuous, appressed to suberect, rising from anterior margin of corresponding punctures, not surpassing their length. Slightly longer suberect, sparse setae arranged in irregular longitudinal rows along remnants of elytral carinae. Male pygidium dorsally slightly convex and medially longitudinally impressed, with dense shallow ovoid punctures (
<figureCitation id="0AA52A659D0F9248FC91FDBCCB3A2A7F" box="[770,876,514,541]" captionStart="Figs 451–468" captionStartId="163.[189,241,1681,1707]" captionTargetBox="[456,1130,262,1650]" captionTargetId="figure-178@163.[456,1131,262,1657]" captionTargetPageId="163" captionText="Figs 451–468. Papuan and Wallacean Maechidius Macleay, 1819, pygidium, dorsal view. 451. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 452. M. aiyura sp. nov., holotype, ♂ (BMNH). 453. Same, different aspect (BMNH). 454. M. alesbezdeki sp. nov., holotype, ♂ (NME). 455. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 456. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 457. M. awu sp. nov., holotype, ♂ (IECA). 458. M. babyrousa sp. nov., holotype, ♂ (BMNH). 459. Same, paratype, ♀ (BMNH). 460. M. bintang sp. nov., holotype, ♂ (NME). 461. M. boessnecki sp. nov., holotype, ♂ (SNSD). 462. M. brocki sp. nov., holotype, ♂ (BMNH). 463. M. caperatus sp. nov., holotype, ♂ (NME). 464. M. ciliatus sp. nov., holotype, ♂ (IECA). 465. M. crypticus sp. nov., paratype, ♂ (BMNH). 466. Same, paratype, ♀ (BMNH). 467. M. dani sp. nov., holotype, ♂ (IECA). 468. M. deltouri sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122222" httpUri="https://zenodo.org/record/4122222/files/figure.png" pageId="19" pageNumber="20">Fig. 452</figureCitation>
). Intervening spaces in part microreticulate, generally smaller than punctures. Setae of pygidium inconspicuous, sparse, suberect. Male protibia with two moderately large distal teeth on external margin (Fig. 360). Male protibial terminal spur large, curved. Male lower meso- and metatibial terminal spur curved. Tarsal claws with pulvilli (in males only?). Spiculum gastrale as in Fig. 535. Male aedeagus as in
<figureCitation id="0AA52A659D0F9248FC07FD2ECC6E2AC9" box="[916,1080,656,683]" captionStart="Figs 570–585" captionStartId="170.[189,241,1892,1918]" captionTargetBox="[425,1155,265,1858]" captionTargetId="figure-74@170.[416,1172,265,1864]" captionTargetPageId="170" captionText="Figs 570–585. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 570– 572. M. aiyura sp. nov., holotype, ♂ (BMNH). 573–575. M. alesbezdeki sp. nov., holotype, ♂ (NME). 576–578. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 579–581. M. awu sp. nov., holotype, ♂ (IECA). 582–585. M. babyrousa sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122232" httpUri="https://zenodo.org/record/4122232/files/figure.png" pageId="19" pageNumber="20">Figs 570–572</figureCitation>
.
</paragraph>
<paragraph id="922136E09D0F9248FF2EFD6FC9E62A89" blockId="19.[189,432,721,747]" box="[189,432,721,747]" pageId="19" pageNumber="20">
<heading id="C969818C9D0F9248FF2EFD6FC9E62A89" bold="true" box="[189,432,721,747]" fontSize="11" level="3" pageId="19" pageNumber="20" reason="3">
<emphasis id="A0EAEAF29D0F9248FF2EFD6FC9E62A89" bold="true" box="[189,432,721,747]" pageId="19" pageNumber="20">Sexual dimorphism</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D0F9248FF2EFD41C9FD2B7B" blockId="19.[189,427,767,793]" box="[189,427,767,793]" pageId="19" pageNumber="20">Female is unknown.</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D0F9248FF2EFCFECA262BE5" box="[189,624,832,904]" pageId="19" pageNumber="20" type="biology_ecology">
<paragraph id="922136E09D0F9248FF2EFCFEC9762B38" blockId="19.[189,288,832,858]" box="[189,288,832,858]" pageId="19" pageNumber="20">
<heading id="C969818C9D0F9248FF2EFCFEC9762B38" bold="true" box="[189,288,832,858]" fontSize="11" level="3" pageId="19" pageNumber="20" reason="3">
<emphasis id="A0EAEAF29D0F9248FF2EFCFEC9762B38" bold="true" box="[189,288,832,858]" pageId="19" pageNumber="20">Ecology</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D0F9248FF2EFCD3CA262BE5" blockId="19.[189,624,877,904]" box="[189,624,877,904]" pageId="19" pageNumber="20">Occurs in lower montane rainforests.</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D0F9248FF2EFC10CBCE2B94" box="[189,920,942,1014]" pageId="19" pageNumber="20" type="distribution">
<paragraph id="922136E09D0F9248FF2EFC10C9002BAA" blockId="19.[189,342,942,968]" box="[189,342,942,968]" pageId="19" pageNumber="20">
<heading id="C969818C9D0F9248FF2EFC10C9002BAA" bold="true" box="[189,342,942,968]" fontSize="11" level="3" pageId="19" pageNumber="20" reason="3">
<emphasis id="A0EAEAF29D0F9248FF2EFC10C9002BAA" bold="true" box="[189,342,942,968]" pageId="19" pageNumber="20">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D0F9248FF2EFC62CBCE2B94" blockId="19.[189,920,988,1014]" box="[189,920,988,1014]" pageId="19" pageNumber="20">
Aiyura Valley, Eastern Highlands Province, East New
<collectingCountry id="EA8976709D0F9248FCAFFC62CBC32B94" box="[828,917,988,1014]" name="Guinea-Bissau" pageId="19" pageNumber="20">Guinea</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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