149 lines
16 KiB
XML
149 lines
16 KiB
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<document id="4DD5F33E1BF22AA2A0D70EA5C2EBADC5" ID-DOI="10.11646/zootaxa.3666.2.2" ID-GBIF-Dataset="930d392c-8720-4820-85eb-501985209cf3" ID-ISSN="1175-5326" ID-Zenodo-Dep="220466" ID-ZooBank="1F8D4E8A-2447-4556-9B98-D8229D858194" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460622323338" checkinUser="plazi" docAuthor="Prokin, Alexander A., Petrov, Pyotr N., Wang, Bo & Ponomarenko, Alexander G." docDate="2013" docId="03AB6D60FF9BD232FF4409E4B5984EB1" docLanguage="en" docName="zt03666p159.pdf" docOrigin="Zootaxa 3666 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Liadytes aspidytoides Prokin, Petrov, Wang & Ponomarenko, 2013, sp. n." docType="treatment" docVersion="8" lastPageNumber="141" masterDocId="FF921518FF9FD236FFD30973B6134A15" masterDocTitle="New fossil taxa and notes on the Mesozoic evolution of Liadytidae and Dytiscidae (Coleoptera)" masterLastPageNumber="159" masterPageNumber="137" pageNumber="141" updateTime="1698334052497" updateUser="plazi">
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<mods:titleInfo id="B075DAEDC70E4A7714F9C934B6DF9CA3">
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<mods:title id="5AFFCFBC15EB6CF1B758AC1F66458223">New fossil taxa and notes on the Mesozoic evolution of Liadytidae and Dytiscidae (Coleoptera)</mods:title>
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<mods:name id="BDA41709DE88E8984427F588FD61C282" type="personal">
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<mods:namePart id="CA84A32DA2043095A0C0BB364FE396D7">Prokin, Alexander A.</mods:namePart>
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</mods:name>
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<mods:namePart id="50947B838792D825E33140E42C4F8837">Petrov, Pyotr N.</mods:namePart>
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<mods:namePart id="B535452D7DE954D8A8D35EC8D265BBDB">Wang, Bo</mods:namePart>
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<mods:namePart id="5E1B58E63ABB85A0CD56AC1C8E584FE1">Ponomarenko, Alexander G.</mods:namePart>
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<mods:typeOfResource id="0C7CDCFE676DE6D6964FD354198374E5">text</mods:typeOfResource>
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<mods:title id="CD825A88FD89CF513065EFA4B9C0D32D">Zootaxa</mods:title>
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<mods:part id="DA44D0DCCACAD4BDD6711A94739A0BC6">
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<mods:date id="17F383FA6987033B1ACA750DA542E68B">2013</mods:date>
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<mods:detail id="C36A3D0252CE819F7FA7B9DBF0446775" type="volume">
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<mods:number id="960AE8DE180A6DC3CD22BAA220A446AD">3666</mods:number>
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<mods:number id="4DFF0F048727C245AF69E2233A9DB24F">2</mods:number>
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<mods:start id="438F9E76F397FDF016A564C6B382E798">137</mods:start>
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<mods:classification id="99ADBAE8E8BBA63662960563CA0EC061">journal article</mods:classification>
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<mods:identifier id="A05E74332EB632F69E7F166F84B3BE61" type="DOI">10.11646/zootaxa.3666.2.2</mods:identifier>
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<mods:identifier id="B902E3741391C11B196EE924065A42B3" type="GBIF-Dataset">930d392c-8720-4820-85eb-501985209cf3</mods:identifier>
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<mods:identifier id="281F0EB74221D382A6BD101E8A62EDB9" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="583E7193FDA0C871AEA67FBCF80CF655" type="Zenodo-Dep">220466</mods:identifier>
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<mods:identifier id="1752CF43125AA15F69BFAAC8E0080A76" type="ZooBank">1F8D4E8A-2447-4556-9B98-D8229D858194</mods:identifier>
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<treatment id="03AB6D60FF9BD232FF4409E4B5984EB1" ID-DOI="http://doi.org/10.5281/zenodo.5680598" ID-GBIF-Taxon="119569015" ID-Zenodo-Dep="5680598" LSID="urn:lsid:plazi:treatment:03AB6D60FF9BD232FF4409E4B5984EB1" httpUri="http://treatment.plazi.org/id/03AB6D60FF9BD232FF4409E4B5984EB1" lastPageNumber="141" pageId="4" pageNumber="141">
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<subSubSection id="C3188FFDFF9BD232FF4409E4B7CC4AA4" box="[151,479,151,177]" pageId="4" pageNumber="141" type="nomenclature">
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<paragraph id="8BBDDC76FF9BD232FF4409E4B7CC4AA4" blockId="4.[151,479,151,177]" box="[151,479,151,177]" pageId="4" pageNumber="141">
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<heading id="D0F56B1AFF9BD232FF4409E4B7CC4AA4" bold="true" box="[151,479,151,177]" fontSize="11" level="1" pageId="4" pageNumber="141" reason="1">
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<emphasis id="B9760064FF9BD232FF4409E4B7CC4AA4" bold="true" box="[151,479,151,177]" pageId="4" pageNumber="141">
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<taxonomicName id="4C02A7F5FF9BD232FF4409E4B7854AA4" box="[151,406,151,177]" class="Insecta" family="Liadytidae" genus="Liadytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="species" species="aspidytoides" status="sp. nov.">
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<emphasis id="B9760064FF9BD232FF4409E4B7854AA4" bold="true" box="[151,406,151,177]" italics="true" pageId="4" pageNumber="141">Liadytes aspidytoides</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A245BD1FFF9BD232FE4E09E4B7CC4AA4" box="[413,479,151,177]" pageId="4" pageNumber="141" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3188FFDFF9BD232FF4409ACB2514AE3" box="[151,1090,222,247]" pageId="4" pageNumber="141" type="etymology">
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<paragraph id="8BBDDC76FF9BD232FF4409ACB2514AE3" blockId="4.[151,1437,222,1189]" box="[151,1090,222,247]" pageId="4" pageNumber="141">
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<emphasis id="B9760064FF9BD232FF4409ACB70B4AE2" bold="true" box="[151,280,223,247]" pageId="4" pageNumber="141">Etymology</emphasis>
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: From the generic name
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<taxonomicName id="4C02A7F5FF9BD232FDE209ADB48F4AE3" box="[561,668,222,246]" class="Insecta" family="Aspidytidae" genus="Aspidytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="genus">
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<emphasis id="B9760064FF9BD232FDE209ADB48F4AE3" box="[561,668,222,246]" italics="true" pageId="4" pageNumber="141">Aspidytes</emphasis>
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</taxonomicName>
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; adjective in the nominative singular.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3188FFDFF9BD232FF140871B4654B93" pageId="4" pageNumber="141" type="materials_examined">
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<paragraph id="8BBDDC76FF9BD232FF140871B4654B93" blockId="4.[151,1437,222,1189]" pageId="4" pageNumber="141">
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<emphasis id="B9760064FF9BD232FF140871B73C4B0E" bold="true" box="[199,303,258,283]" pageId="4" pageNumber="141">Material</emphasis>
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:
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<typeStatus id="54B962D4FF9BD232FEEC0871B7B44B0F" box="[319,423,258,282]" pageId="4" pageNumber="141" type="holotype">Holotype</typeStatus>
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PIN 2470/942±, beetle without head, prothorax and legs; Shar Teg, Upper Jurassic, beds 441, layer 4;
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<typeStatus id="54B962D4FF9BD232FEFF0854B79E4B2A" box="[300,397,295,319]" pageId="4" pageNumber="141" type="paratype">paratype</typeStatus>
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2470/945, less complete impression of beetle from the same locality, beds 451/3; additional specimen (cannot be identified with certainty, therefore not included in the
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<typeStatus id="54B962D4FF9BD232FBD60838B2264B76" box="[1029,1077,331,355]" pageId="4" pageNumber="141">type</typeStatus>
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series) PIN 2470/938, isolated elytron from the same locality, beds 451/3.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3188FFDFF9BD232FF1408E1B5984EB1" pageId="4" pageNumber="141" type="description">
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<paragraph id="8BBDDC76FF9BD232FF1408E1B34248E4" blockId="4.[151,1437,222,1189]" pageId="4" pageNumber="141">
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<emphasis id="B9760064FF9BD232FF1408E1B7424BBE" bold="true" box="[199,337,402,427]" pageId="4" pageNumber="141">Description</emphasis>
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(
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<figureCitation id="1339C0F3FF9BD232FEB308E1B7DF4BBF" box="[352,460,402,426]" captionStart-0="FIGURES 1 – 4" captionStart-1="FIGURES 5 – 7" captionStartId-0="5.[151,264,1917,1940]" captionStartId-1="7.[151,264,1975,1998]" captionTargetBox-0="[152,1434,193,1895]" captionTargetBox-1="[175,1386,193,1949]" captionTargetId-0="figure@5.[152,1435,193,1896]" captionTargetId-1="figure@7.[175,1412,193,1950]" captionTargetPageId-0="5" captionTargetPageId-1="7" captionText-0="FIGURES 1 – 4. Liadytes aspidytoides sp. n.: (1) photo of holotype, (2) line drawing of holotype, (3) SEM photo of holotype, (4) SEM photo of details of metaventrite; arrows indicate metaventral suture." captionText-1="FIGURES 5 – 7. (5) Liadytes aspidytoides sp. n., photo of specimen PIN 2470 / 945; Liadyxianus kirejtshuki sp. n.: (6) photo of holotype, (7) line drawing of holotype." httpUri-0="https://zenodo.org/record/220467/files/figure.png" httpUri-1="https://zenodo.org/record/220468/files/figure.png" pageId="4" pageNumber="141">Figs. 1–5</figureCitation>
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): Beetle with dark elytra, which appear black due to carbonisation of organic substance in oxygen-free environment, and pale body. Maximum body width situated at level of elytral base. Mesocoxae only slightly separated, rounded. Maximum length of metaxocal plate (measured parallel to longitudinal body axis) 1.2 times as great as maximum length of lateral lobe of metaventrite (measured in same manner)3. Anterior margin of metacoxa weakly convex (metacoxal plate dilated anteriorly). Elevated median part of metacoxae with sides more or less straight, slightly diverging posteriad; metacoxal processes rounded posteriorly, with posterior margins meeting at an obtuse angle. Transverse suture of metaventrite present; median part of metacoxae elevated. Metepisternum 1.3 times as long as wide. Abdomen tapering from base of ventrite 4; ventrite 6 twice as long as ventrite 5. Elytron 3.4 times as long as wide. Epipleuron wide in its basal third, twice as wide as distally. Surface microsculpture of underside punctate, laterally on abdominal ventrites imbricate (according to Harris 1979).
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</paragraph>
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<paragraph id="8BBDDC76FF9BD232FF140B8EB3954900" blockId="4.[151,1437,222,1189]" box="[199,1414,765,789]" pageId="4" pageNumber="141">
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<emphasis id="B9760064FF9BD232FF140B8EB7654900" bold="true" box="[199,374,765,789]" pageId="4" pageNumber="141">Measurements</emphasis>
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: Body length, probably around
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<quantity id="4CFA7193FF9BD232FD0F0B8DB50C4900" box="[732,799,766,789]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="4" pageNumber="141" unit="mm" value="7.0">7 mm</quantity>
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; body width,
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<quantity id="4CFA7193FF9BD232FC640B8EB2514900" box="[951,1090,765,789]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" metricValueMax="3.7" metricValueMin="3.1" pageId="4" pageNumber="141" unit="mm" value="3.4" valueMax="3.7" valueMin="3.1">3.1–3.7 mm</quantity>
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; elytron length,
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<quantity id="4CFA7193FF9BD232FB240B8EB3924900" box="[1271,1409,765,789]" metricMagnitude="-3" metricUnit="m" metricValue="4.949999999999999" metricValueMax="5.3" metricValueMin="4.6" pageId="4" pageNumber="141" unit="mm" value="4.949999999999999" valueMax="5.3" valueMin="4.6">4.6–5.3 mm</quantity>
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.
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</paragraph>
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<paragraph id="8BBDDC76FF9BD232FF140A52B21749D9" blockId="4.[151,1437,222,1189]" pageId="4" pageNumber="141">
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<emphasis id="B9760064FF9BD232FF140A52B749492F" bold="true" box="[199,346,801,826]" pageId="4" pageNumber="141">Comparison</emphasis>
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: The new species is similar to
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<taxonomicName id="4C02A7F5FF9BD232FD6F0A51B209492C" authority="Ponomarenko, 1977" authorityName="Ponomarenko" authorityYear="1977" box="[700,1050,801,825]" class="Insecta" family="Liadytidae" genus="Liadytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="species" species="crassus">
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<emphasis id="B9760064FF9BD232FD6F0A51B53C492C" box="[700,815,802,825]" italics="true" pageId="4" pageNumber="141">L. crassus</emphasis>
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Ponomarenko,
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<quantity id="4CFA7193FF9BD232FC310A52B209492C" box="[994,1050,801,825]" metricMagnitude="1" metricUnit="m" metricValue="5.02158" pageId="4" pageNumber="141" unit="in" value="1977.0">1977</quantity>
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</taxonomicName>
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in the convex anterior margins of the metacoxae (dilated anteriorly), but differs from it, as well as from the other species of
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<taxonomicName id="4C02A7F5FF9BD232FB540A36B2F54948" box="[1159,1254,837,861]" class="Insecta" family="Liadytidae" genus="Liadytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="genus">
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<emphasis id="B9760064FF9BD232FB540A36B2F54948" box="[1159,1254,837,861]" italics="true" pageId="4" pageNumber="141">Liadytes</emphasis>
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</taxonomicName>
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, in the presence of the elevated median part of the metacoxae (forming plates that partly cover the metafemora, as in
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<taxonomicName id="4C02A7F5FF9BD232FAC70A1AB3884994" box="[1300,1435,873,897]" class="Insecta" family="Aspidytidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="family">Aspidytidae</taxonomicName>
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and
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<taxonomicName id="4C02A7F5FF9BD232FF1E0AE3B72C49BD" box="[205,319,912,936]" class="Insecta" family="Noteridae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="family">Noteridae</taxonomicName>
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; these structures are, apparently, derived from the broader femoral plates of the metacoxae4, homologous to those found in
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<taxonomicName id="4C02A7F5FF9BD232FE3F0AC7B47249D9" box="[492,609,948,972]" class="Insecta" family="Haliplidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="family">Haliplidae</taxonomicName>
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and many members of Geadephaga).
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</paragraph>
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<paragraph id="8BBDDC76FF9BD232FF140AABB5984EB1" blockId="4.[151,1437,222,1189]" pageId="4" pageNumber="141">
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<emphasis id="B9760064FF9BD232FF140AABB71949E5" bold="true" box="[199,266,984,1008]" pageId="4" pageNumber="141">Notes</emphasis>
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: The shape of femoral plates of the metacoxae and broad epipleuron of the new species are similar to those of
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<taxonomicName id="4C02A7F5FF9BD232FF2A0A8FB7774E01" box="[249,356,1020,1044]" class="Insecta" family="Aspidytidae" genus="Aspidytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="genus">
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<emphasis id="B9760064FF9BD232FF2A0A8FB7774E01" box="[249,356,1020,1044]" italics="true" pageId="4" pageNumber="141">Aspidytes</emphasis>
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</taxonomicName>
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, but the new species differs from it and from
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<taxonomicName id="4C02A7F5FF9BD232FCBD0A8FB5CF4E01" box="[878,988,1020,1044]" class="Insecta" family="Noteridae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="family">Noteridae</taxonomicName>
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in the mesocoxae located close to each other and in the absence of the anteromedial metaventral process (see above). The metacoxal plates dilated anteriorly are characteristic of
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<taxonomicName id="4C02A7F5FF9BD232FE260D37B46F4E49" box="[501,636,1092,1116]" class="Insecta" family="Aspidytidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="family">Aspidytidae</taxonomicName>
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and Liadytiscinae (
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<taxonomicName id="4C02A7F5FF9BD232FC8F0D37B5C44E49" box="[860,983,1092,1116]" class="Insecta" family="Dytiscidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="family">Dytiscidae</taxonomicName>
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) (see next section), but the new species differs from representatives of Liadytiscinae in the presence of the transverse suture on the metaventrite. Therefore, the new species is included here in the genus
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<taxonomicName id="4C02A7F5FF9BD232FD470DFFB5964EB1" box="[660,901,1164,1188]" class="Insecta" family="Liadytidae" genus="Liadytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="genus">
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<emphasis id="B9760064FF9BD232FD470DFFB4E04EB1" box="[660,755,1164,1188]" italics="true" pageId="4" pageNumber="141">Liadytes</emphasis>
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(Liadytidae)
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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</document>
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