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101 lines
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<document id="1783C68E2DF544C2AD0D165D36A3F4FF" ID-CLB-Dataset="59555" ID-DOI="10.1007/s12542-023-00657-7" ID-GBIF-Dataset="6450bd33-f5f8-479c-9263-b450e4cc3030" ID-Zenodo-Dep="8206331" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="julia" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="julia" IM.tables_approvedBy="julia" IM.taxonomicNames_approvedBy="julia" IM.treatments_approvedBy="julia" checkinTime="1690914803121" checkinUser="julia" docAuthor="Dunlop 1, Jason A." docDate="2023" docId="14720107FF9FFFD5FF46F9BF651EDF12" docLanguage="en" docName="PalZ.2023.1-8.pdf" docOrigin="PalZ 2023" docTitle="Arthrolycosa Harger 1874" docType="treatment" docVersion="2" lastPageNumber="3" masterDocId="E84B797FFF9CFFD6FFCEFFCA612BD93B" masterDocTitle="The first Palaeozoic spider (Arachnida: Araneae) from Germany" masterLastPageNumber="8" masterPageNumber="1" pageNumber="3" updateTime="1690916557290" updateUser="ExternalLinkService">
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<mods:mods id="62A9F37F9FD1E369DB70B464F58AADCF" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="7E0EC1AF37D1C46523F9A37C3E796DC2">
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<mods:title id="F4A16CE1CF159CBD87FDA5BC6EF32F23">The first Palaeozoic spider (Arachnida: Araneae) from Germany</mods:title>
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</mods:titleInfo>
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<mods:name id="AF55B113EFF36902D9A37B4A9FA505F2" type="personal">
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<mods:role id="E90DBD40C5DFD95BA48F4269FFFF439E">
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<mods:roleTerm id="5AC1AE4FD1AC55751C5B2BED20733000">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="75B71537F602682C3935F537E2CA1298">Dunlop 1, Jason A.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="B0DAA989511FCB030E53DB197D49F32F">text</mods:typeOfResource>
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<mods:titleInfo id="369E4A8FAA562E613BA4AC3625F52E94">
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<mods:title id="A6FE1E9790B3677FCE5CCD843F3E3AC5">PalZ</mods:title>
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</mods:titleInfo>
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<mods:part id="4D7C7A88D622A000FC4835612A11DF9B">
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<mods:date id="5397EC63C962B3A232F89979C58152F0">2023</mods:date>
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<mods:detail id="05130753C1DB809D8706846041765514" type="pubDate">
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<mods:number id="81156D839E5BB89D46FA829152E8D3DF">2023-07-16</mods:number>
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</mods:detail>
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<mods:detail id="0B9E118126E67B16DA5774681DC777BC" type="volume">
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<mods:number id="35C0489A7A02C34AFDBD74B6E872382A">2023</mods:number>
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</mods:detail>
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<mods:extent id="D20F35FC9F18EC85E03755D5720F9152" unit="page">
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<mods:start id="12188CA230BA8F74413C5158F22DB8B7">1</mods:start>
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<mods:end id="38E9A446D2EFC39971F128C15D733B2A">8</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="05F063E0D2E4603407AE9FD68853C26D">journal article</mods:classification>
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<mods:identifier id="5AC7926BDE93303820DBEB7E25F85D49" type="CLB-Dataset">59555</mods:identifier>
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<mods:identifier id="A5A6B5013C362D1258A06A486EF588F2" type="DOI">10.1007/s12542-023-00657-7</mods:identifier>
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<mods:identifier id="4995BF864AB9BC64E907BDF4705C6701" type="GBIF-Dataset">6450bd33-f5f8-479c-9263-b450e4cc3030</mods:identifier>
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<mods:identifier id="949D835E5D5D9CF0067F27B3A145FF67" type="Zenodo-Dep">8206331</mods:identifier>
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</mods:mods>
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<treatment id="14720107FF9FFFD5FF46F9BF651EDF12" ID-DOI="http://doi.org/10.5281/zenodo.8206324" ID-Zenodo-Dep="8206324" LSID="urn:lsid:plazi:treatment:14720107FF9FFFD5FF46F9BF651EDF12" httpUri="http://treatment.plazi.org/id/14720107FF9FFFD5FF46F9BF651EDF12" lastPageNumber="3" pageId="3" pageNumber="3">
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<subSubSection id="D4C1E39AFF9FFFD5FF46F9BF60DDDFB6" box="[136,502,1653,1678]" pageId="3" pageNumber="3" type="nomenclature">
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<paragraph id="9C64B011FF9FFFD5FF46F9BF60DDDFB6" blockId="3.[136,502,1653,1678]" box="[136,502,1653,1678]" pageId="3" pageNumber="3">
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Genus
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<taxonomicName id="5BDBCB92FF9FFFD5FF1DF9BF60DDDFB6" authority="Harger, 1874" authorityName="Harger" authorityYear="1874" box="[211,502,1653,1678]" class="Arachnida" family="Arthrolycosidae" genus="Arthrolycosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="3" phylum="Arthropoda" rank="genus">
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<emphasis id="AEAF6C03FF9FFFD5FF1DF9BF6048DFB6" bold="true" box="[211,355,1653,1677]" italics="true" pageId="3" pageNumber="3">Arthrolycosa</emphasis>
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<bibRefCitation id="F84ACDE0FF9FFFD5FEA7F9BC60DDDFB6" author="Harger, O." box="[361,502,1653,1678]" journalOrPublisher="American Journal of Science" pageId="3" pageNumber="3" pagination="219 - 223" part="7" refId="ref4940" refString="Harger, O. 1874. Notice of a new spider from the Coal Measures of Illinois. American Journal of Science 7: 219 - 223." title="Notice of a new spider from the Coal Measures of Illinois" type="journal article" year="1874">Harger, 1874</bibRefCitation>
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="D4C1E39AFF9FFFD5FF46F97363BFDFEB" box="[136,660,1720,1744]" pageId="3" pageNumber="3" type="type_taxon">
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<paragraph id="9C64B011FF9FFFD5FF46F97363BFDFEB" blockId="3.[136,660,1720,1744]" box="[136,660,1720,1744]" pageId="3" pageNumber="3">
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<emphasis id="AEAF6C03FF9FFFD5FF46F973632ADFEB" box="[136,513,1720,1744]" italics="true" pageId="3" pageNumber="3">
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Type species.
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<taxonomicName id="5BDBCB92FF9FFFD5FED3F97263BFDFEB" authority="Harger, 1874" authorityName="Harger" authorityYear="1874" box="[285,660,1720,1744]" class="Arachnida" family="Arthrolycosidae" genus="Arthrolycosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="3" phylum="Arthropoda" rank="species" species="antiqua">
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Arthrolycosa antiqua
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<bibRefCitation id="F84ACDE0FF9FFFD5FDC6F97263BFDFEB" author="Harger, O." box="[520,660,1720,1744]" journalOrPublisher="American Journal of Science" pageId="3" pageNumber="3" pagination="219 - 223" part="7" refId="ref4940" refString="Harger, O. 1874. Notice of a new spider from the Coal Measures of Illinois. American Journal of Science 7: 219 - 223." title="Notice of a new spider from the Coal Measures of Illinois" type="journal article" year="1874">Harger, 1874</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="D4C1E39AFF9FFFD5FF46F9316316DE29" box="[136,573,1787,1811]" pageId="3" pageNumber="3" type="reference_group">
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<paragraph id="9C64B011FF9FFFD5FF46F9316316DE29" blockId="3.[136,573,1787,1811]" box="[136,573,1787,1811]" pageId="3" pageNumber="3">
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<emphasis id="AEAF6C03FF9FFFD5FF46F9316010DE29" box="[136,315,1787,1811]" italics="true" pageId="3" pageNumber="3">Included species</emphasis>
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.
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<taxonomicName id="5BDBCB92FF9FFFD5FE86F93160C8DE28" authority="Dunlop1, 2023" authorityName="Dunlop1" authorityYear="2023" box="[328,483,1787,1811]" class="Arachnida" family="Arthrolycosidae" genus="Arthrolycosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="3" phylum="Arthropoda" rank="species" species="wolterbeeki" status="sp. nov.">
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<emphasis id="AEAF6C03FF9FFFD5FE86F93160C8DE28" box="[328,483,1787,1811]" italics="true" pageId="3" pageNumber="3">A. wolterbeeki</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="B59CD178FF9FFFD5FE24F9316316DE29" box="[490,573,1787,1810]" pageId="3" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="D4C1E39AFF9FFFD5FF46F8F4651EDF12" pageId="3" pageNumber="3" type="discussion">
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<paragraph id="9C64B011FF9FFFD5FF46F8F4651EDF12" blockId="3.[136,772,1853,1911]" lastBlockId="3.[816,1452,853,1577]" pageId="3" pageNumber="3">
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<emphasis id="AEAF6C03FF9FFFD5FF46F8F46048DE6E" box="[136,355,1853,1877]" italics="true" pageId="3" pageNumber="3">Taxonomic remarks</emphasis>
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. Two arthrolycosid genera were recognised by
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<bibRefCitation id="F84ACDE0FF9FFFD5FEC8F8956089DE4C" author="Selden, P. A." box="[262,418,1887,1911]" editor="S. G. Lucas & W. A. DiMichele & B. D. Allen" journalOrPublisher="New Mexico Museum of Natural History and Science Bulletin" pageId="3" pageNumber="3" pagination="317 - 358" part="84" refId="ref6063" refString="Selden, P. A. 2021. New spiders (Araneae: Mesothelae), from the Carboniferous of New Mexico and England and a review of Paleozoic Araneae. In S. G. Lucas, W. A. DiMichele & B. D. Allen (eds.), Kinney Brick Quarry Lagerstatte. New Mexico Museum of Natural History and Science Bulletin 84: 317 - 358." title="New spiders (Araneae: Mesothelae), from the Carboniferous of New Mexico and England and a review of Paleozoic Araneae" type="journal article" volumeTitle="Kinney Brick Quarry Lagerstatte" year="2021">Selden (2021)</bibRefCitation>
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.
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<taxonomicName id="5BDBCB92FF9FFFD5FE61F89563F9DE4C" authority="Harger, 1874" authorityName="Harger" authorityYear="1874" box="[431,722,1887,1911]" class="Arachnida" family="Arthrolycosidae" genus="Arthrolycosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="3" phylum="Arthropoda" rank="genus">
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<emphasis id="AEAF6C03FF9FFFD5FE61F8956317DE4C" box="[431,572,1887,1911]" italics="true" pageId="3" pageNumber="3">Arthrolycosa</emphasis>
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<bibRefCitation id="F84ACDE0FF9FFFD5FD8AF89563F9DE4C" author="Harger, O." box="[580,722,1887,1911]" journalOrPublisher="American Journal of Science" pageId="3" pageNumber="3" pagination="219 - 223" part="7" refId="ref4940" refString="Harger, O. 1874. Notice of a new spider from the Coal Measures of Illinois. American Journal of Science 7: 219 - 223." title="Notice of a new spider from the Coal Measures of Illinois" type="journal article" year="1874">Harger, 1874</bibRefCitation>
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</taxonomicName>
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was redefined as having a carapace about as long as wide, with a distinct ridge behind the fovea (the depression in the centre of the carapace) and legs I and IV being longer than legs II and III.
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<taxonomicName id="5BDBCB92FF9FFFD5FC69FC7365FADAEA" authority="Roemer, 1865" authorityName="Roemer" authorityYear="1865" box="[935,1233,953,977]" class="Arachnida" family="Arthromygalidae" genus="Protolycosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="3" phylum="Arthropoda" rank="genus">
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<emphasis id="AEAF6C03FF9FFFD5FC69FC736500DAEA" box="[935,1067,953,977]" italics="true" pageId="3" pageNumber="3">Protolycosa</emphasis>
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Roemer, 1865
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</taxonomicName>
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was redefined on a carapace longer than wide and somewhat pointed anteriorly with a recurved posterior margin. The fovea may or may not be a deep pit, and leg I is characteristically shorter than the other legs. According to
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<bibRefCitation id="F84ACDE0FF9FFFD5FB41FBF5640EDD6C" author="Selden, P. A." box="[1167,1317,1087,1111]" editor="S. G. Lucas & W. A. DiMichele & B. D. Allen" journalOrPublisher="New Mexico Museum of Natural History and Science Bulletin" pageId="3" pageNumber="3" pagination="317 - 358" part="84" refId="ref6063" refString="Selden, P. A. 2021. New spiders (Araneae: Mesothelae), from the Carboniferous of New Mexico and England and a review of Paleozoic Araneae. In S. G. Lucas, W. A. DiMichele & B. D. Allen (eds.), Kinney Brick Quarry Lagerstatte. New Mexico Museum of Natural History and Science Bulletin 84: 317 - 358." title="New spiders (Araneae: Mesothelae), from the Carboniferous of New Mexico and England and a review of Paleozoic Araneae" type="journal article" volumeTitle="Kinney Brick Quarry Lagerstatte" year="2021">Selden (2021)</bibRefCitation>
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, both genera express rows of tubercles long the posterior margins of the opisthosoma (as does the Piesberg fossil), although earlier workers such as
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<bibRefCitation id="F84ACDE0FF9FFFD5FC12FB696592DD80" author="Petrunkevitch, A. I." box="[988,1209,1186,1211]" editor="R. C. Moore" journalOrPublisher="Boulder and Lawrence: Geological Society of America and University of Kansas Press" pageId="3" pageNumber="3" pagination="42 - 162" refId="ref5709" refString="Petrunkevitch, A. I. 1955. Arachnida. In Treatise on invertebrate paleontology, Part P, Arthropoda 2, ed. R. C. Moore, 42 - 162. Boulder and Lawrence: Geological Society of America and University of Kansas Press." title="Arachnida" type="book chapter" volumeTitle="Treatise on invertebrate paleontology, Part P, Arthropoda 2" year="1955">Petrunkevitch (1955)</bibRefCitation>
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cited this as diagnostic for
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<taxonomicName id="5BDBCB92FF9FFFD5FC99FB0E62F2DDE7" authorityName="Roemer" authorityYear="1865" box="[855,985,1220,1244]" class="Arachnida" family="Arthromygalidae" genus="Protolycosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="3" phylum="Arthropoda" rank="genus">
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<emphasis id="AEAF6C03FF9FFFD5FC99FB0E62F2DDE7" box="[855,985,1220,1244]" italics="true" pageId="3" pageNumber="3">Protolycosa</emphasis>
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</taxonomicName>
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only. Using these criteria, the full proportions of the carapace in the Piesberg fossil are uncertain, but the preserved curvature suggests that it was probably rather short and certainly not longer than wide. Also, the carapace posterior margin is not noticeably recurved. The preserved appendages suggest that leg I was not shorter than the succeeding legs, and a IV> I> II> III leg formula (from longest to shortest) seems likely. For these reasons, the new fossil is most consistent with
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<taxonomicName id="5BDBCB92FF9FFFD5FBBCFA0565D5DCDC" authorityName="Harger" authorityYear="1874" box="[1138,1278,1487,1511]" class="Arachnida" family="Arthrolycosidae" genus="Arthrolycosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="3" phylum="Arthropoda" rank="genus">
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<emphasis id="AEAF6C03FF9FFFD5FBBCFA0565D5DCDC" box="[1138,1278,1487,1511]" italics="true" pageId="3" pageNumber="3">Arthrolycosa</emphasis>
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</taxonomicName>
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, confirming the provisional report of the presence of this genus at Piesberg by
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<bibRefCitation id="F84ACDE0FF9FFFD5FC9FF9DB651BDF12" author="Leipner, A. & T. Fischer & P. Chellouche" box="[849,1072,1553,1577]" journalOrPublisher="Geoconservation Research" pageId="3" pageNumber="3" pagination="218 - 234" part="4" refId="ref5422" refString="Leipner, A., T. Fischer, and P. Chellouche. 2021. The Piesberg: A NW- German site of international importance for the Pennsylvanian (Late Carboniferous). Geoconservation Research 4: 218 - 234. https: // doi. org / 10.30486 / gcr. 2021.1913500. 1056." title="The Piesberg: A NW- German site of international importance for the Pennsylvanian (Late Carboniferous)" type="journal article" year="2021">Leipner et al. (2021)</bibRefCitation>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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</document>
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